172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="54653D74B2FEC61604442DDCC4896FEB" ID-CLB-Dataset="46730" ID-DOI="10.5281/zenodo.201491" ID-GBIF-Dataset="30aff672-89b2-441c-af59-1a1b9eb6a7c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="201491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365005065" checkinUser="plazi" docAuthor="Leschen, Richard A. B." docDate="2011" docId="C563080BFFDF3A70FF216CF2F51EFA49" docLanguage="en" docName="zt02908p044.pdf" docOrigin="Zootaxa 2908" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laricobius kovalevi Nikitsky" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="395A7073FFD23A7EFFB66A19F02FFFF6" masterDocTitle="World review of Laricobius (Coleoptera: Derodontidae)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="14" updateTime="1698295152186" updateUser="ExternalLinkService">
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<mods:mods id="7A7BDC98694891B81C926B136DEE8BDE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CA251F8C297B55163C89FF97E656D891">
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<mods:title id="5C2A846F9D9FF0FB324D3D94C66C2359">World review of Laricobius (Coleoptera: Derodontidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="681A0CCE5528E68F77D4844A92199111" type="personal">
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<mods:roleTerm id="F4970022F8E648F60BFF66CAF3C22A57">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A82323E34216A47E92CB0CB918DC407E">Leschen, Richard A. B.</mods:namePart>
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<mods:typeOfResource id="04031BEBB108770430F31F539E77FB4E">text</mods:typeOfResource>
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<mods:title id="16B4907C1AE0C923EB54CB21FE79F14B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EC82329E622A1FF2F7B5F64175431583">
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<mods:date id="C563D0D1FB4F499FFDD8578E9647FA89">2011</mods:date>
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<mods:detail id="50654F5823991617F32B262DC73E47E1" type="volume">
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<mods:number id="7CD4FF5AF061F1C29624196E27338D06">2908</mods:number>
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<mods:start id="C43DB8279632947D590E3E704A23F03C">1</mods:start>
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<mods:end id="E70C2778615A0F77B9449ECA3B24CDB9">44</mods:end>
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<mods:classification id="B75A754325ECE80E27F83C59C4E54FFD">journal article</mods:classification>
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<mods:identifier id="4FDA12B2EACB12063022D6B8E4D1AE2A" type="CLB-Dataset">46730</mods:identifier>
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<mods:identifier id="0484FD489142C3693781C86EE01294D6" type="DOI">10.5281/zenodo.201491</mods:identifier>
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<mods:identifier id="0291036AB4288A4C5E71FB2DB1B4960B" type="GBIF-Dataset">30aff672-89b2-441c-af59-1a1b9eb6a7c7</mods:identifier>
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<mods:identifier id="DE1CE4BAB365B5FA7ACAC9EB359F5910" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B995F14E868AA9DD418F52F46BA1E47E" type="Zenodo-Dep">201491</mods:identifier>
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</mods:mods>
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<treatment id="C563080BFFDF3A70FF216CF2F51EFA49" ID-DOI="http://doi.org/10.5281/zenodo.5661226" ID-GBIF-Taxon="119410636" ID-Zenodo-Dep="5661226" LSID="urn:lsid:plazi:treatment:C563080BFFDF3A70FF216CF2F51EFA49" httpUri="http://treatment.plazi.org/id/C563080BFFDF3A70FF216CF2F51EFA49" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection id="05D0EA96FFDF3A73FF216CF2F378F8F3" box="[151,855,1771,1798]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="4D75B91DFFDF3A73FF216CF2F378F8F3" blockId="13.[151,855,1771,1830]" box="[151,855,1771,1798]" pageId="13" pageNumber="14">
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<heading id="163D0E71FFDF3A73FF216CF2F378F8F3" bold="true" box="[151,855,1771,1798]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="7FBE650FFFDF3A73FF216CF2F378F8F3" bold="true" box="[151,855,1771,1798]" pageId="13" pageNumber="14">
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9.
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<taxonomicName id="8ACAC29EFFDF3A73FF026CF5F23DF8F3" authority="Nikitsky" authorityName="Nikitsky" box="[180,530,1771,1798]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kovalevi">
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<emphasis id="7FBE650FFFDF3A73FF026CF5F18EF8F0" bold="true" box="[180,417,1772,1798]" italics="true" pageId="13" pageNumber="14">Laricobius kovalevi</emphasis>
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Nikitsky
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</taxonomicName>
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in
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<bibRefCitation id="295BC4ECFFDF3A73FD8F6CF2F378F8F3" author="Nikitsky" box="[569,855,1771,1797]" pageId="13" pageNumber="14" refString="Nikitsky, N. B. & Lafer, G. Sh. (1992) Fam. Derodontidae. In: Ler, P. A. (Ed), Keys to the identification of insects of the Far Eastern of the USSR, vol. 3, Coleoptera 2. Nauka, Peterburg, pp. 40 - 44 (in Russian)." type="book chapter" year="1992">Nikitsky & Lafer, 1992</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDF3A73FF216D17F10EF8D0" box="[151,289,1806,1830]" pageId="13" pageNumber="14" type="description">
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<paragraph id="4D75B91DFFDF3A73FF216D17F10EF8D0" blockId="13.[151,855,1771,1830]" box="[151,289,1806,1830]" pageId="13" pageNumber="14">(figs 10, 40)</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDF3A73FF216D4FF4DEF82C" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="4D75B91DFFDF3A73FF216D4FF4DEF82C" blockId="13.[151,1436,1878,2010]" pageId="13" pageNumber="14">
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<emphasis id="7FBE650FFFDF3A73FF216D4FF13EF899" bold="true" box="[151,273,1878,1903]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum dark; posterior tooth absent. Scutellum dark. Surface of elytra not incurvate; elytral punctures not confluent. Epipleura at base dark. Ventrites dark. Femora light. Tibiae light. Aedeagus without median carina on phallobase; medium lobe subacute; apices of parameres without an internal ridge.
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</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDC3A70FF736A8EF30CFC76" pageId="14" pageNumber="15" type="description">
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<paragraph id="4D75B91DFFDC3A70FF736A8EF30CFC76" blockId="14.[151,1436,151,1471]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736A8EF178FF46" bold="true" box="[197,343,151,176]" pageId="14" pageNumber="15">Description.</emphasis>
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Length
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<quantity id="8A3214F8FFDC3A70FE0C6A8EF20AFF59" box="[442,549,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="15" unit="mm" value="2.8">2.80 mm</quantity>
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. Body bicolored with head, prothorax, scutellum, elytra along sides to level above abdominal ventrite 2 (covering stria 10), epipleuron, and venter chocolate brown; elytra reddish brown to dark tan; antenna reddish brown to dark brown with AI and club darker, legs reddish brown to dark tan. Head with ocelli; u-shaped furrow present; interocular distance about
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<date id="39749FDDFFDC3A70FC826B1DF37FFEEA" box="[820,848,260,284]" pageId="14" pageNumber="15">4x</date>
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the width of the eye; macropunctures rounded, distinct to indistinct and not rounded, fusing sometimes to form irregular trenches and present at the center of the frons; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2/3 the width of the eye. Antenna with length of A3 about equal to A4, ratios 1.9:1.6:1.4:1.3:1.3:1.2:1.1:1.0:1.5:1.6:2.4; A11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides evenly rounded; about 0.73 x as long as wide (pronotal length/greatest pronotal width = 0.73); depth = 0.42; pronotum laterally broadly explanate, lateral carina with a distinct sharp edge; anterior angle variable (90º to about 60° and rounded), not forming a tooth; posterior angle indistinct, short tooth absent at the posterior of the lateral margin; foveae visible dorsally; macropunctures on disc distinct, well-impressed and well-separated; micropunctures coarse; setae erect, long with average length 2/3 or equal to width of eye. Elytra about 3.48 x as long as wide (elytral length/greatest elytral width = 3.48) and 3.37 x as long as pronotum (elytral length/pronotal length = 3.37); surface not incurvate at basal third; macropunctures not coalescing to form grooves, separated by an average of 1/2 puncture diameter; micropunctures fine; microsculpture visible; setae suberect to erect, moderately long averaging about 2/3the length of the eye. Aedeagus with phallobase quadrate, about 1/2 the length of the median lobe, median carina absent; median lobe not projecting beyond apices of parameres, narrow with its greatest width less than the width of the parameres, apex subacute and converging apically at about apical 1/3, median groove or carina absent, ostium subapical; parameres broad and subacute, short setae present on apices and along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDC3A70FF736992F2A0FB1E" pageId="14" pageNumber="15" type="discussion">
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<paragraph id="4D75B91DFFDC3A70FF736992F0FBFBE6" blockId="14.[151,1436,151,1471]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736992F162FC52" bold="true" box="[197,333,907,932]" pageId="14" pageNumber="15">Comments.</emphasis>
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Among the ocelli-bearing species with an explanate pronotal margin,
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<taxonomicName id="8ACAC29EFFDC3A70FBCF6994F4D9FC52" box="[1145,1270,908,932]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kovalevi">
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<emphasis id="7FBE650FFFDC3A70FBCF6994F4D9FC52" box="[1145,1270,908,932]" italics="true" pageId="14" pageNumber="15">L. kovalevi</emphasis>
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</taxonomicName>
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can be distinguished by the black head and pronotum and lighter elytra. The posterior pronotal tooth is absent in
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<taxonomicName id="8ACAC29EFFDC3A70FAAE69A8F5BAFC3E" box="[1304,1429,944,968]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kovalevi">
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<emphasis id="7FBE650FFFDC3A70FAAE69A8F5BAFC3E" box="[1304,1429,944,968]" italics="true" pageId="14" pageNumber="15">L. kovalevi</emphasis>
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</taxonomicName>
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, which is present in
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<taxonomicName id="8ACAC29EFFDC3A70FED869CCF1CEFC1A" box="[366,481,980,1004]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rubidus">
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<emphasis id="7FBE650FFFDC3A70FED869CCF1CEFC1A" box="[366,481,980,1004]" italics="true" pageId="14" pageNumber="15">L. rubidus</emphasis>
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</taxonomicName>
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, while the femora and tibiae are light which can distinguish
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<taxonomicName id="8ACAC29EFFDC3A70FBC969CCF4D6FC1A" box="[1151,1273,980,1004]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kovalevi">
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<emphasis id="7FBE650FFFDC3A70FBC969CCF4D6FC1A" box="[1151,1273,980,1004]" italics="true" pageId="14" pageNumber="15">L. kovalevi</emphasis>
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</taxonomicName>
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from
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<emphasis id="7FBE650FFFDC3A70FA8A69CCF0FBFBE6" italics="true" pageId="14" pageNumber="15">
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<taxonomicName id="8ACAC29EFFDC3A70FA8A69CCF0FEFBE6" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="erichsonii">L. erichsonii</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph id="4D75B91DFFDC3A70FF736E04F0D4FB32" blockId="14.[151,1436,151,1471]" pageId="14" pageNumber="15">
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I had two
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<typeStatus id="927107BFFFDC3A70FE8F6E04F18AFBC3" box="[313,421,1053,1077]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
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in relatively poor condition from ZMUM for study and only the male genitalia were dissected from one of these. A single measured specimen fell outside of the published size range of the species (
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<quantity id="8A3214F8FFDC3A70FADC6E59F0DEFB8A" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="14" pageNumber="15" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1– 2.5 mm</quantity>
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). At the time this manuscript was in review, additional specimens of this species from
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<collectingCountry id="35DDF98DFFDC3A70FB016E7CF4D8FB8A" box="[1207,1271,1125,1148]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
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were sent sent to N. Havill (USDA) and myself by S. Shiyake (OMNH), and based on dissection, this species is confirmed as a
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<taxonomicName id="8ACAC29EFFDC3A70FA336E90F0DBFB32" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kovalevi">
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<emphasis id="7FBE650FFFDC3A70FA336E90F0DBFB32" italics="true" pageId="14" pageNumber="15">L. kovalevi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="4D75B91DFFDC3A70FF736EC9F2A0FB1E" blockId="14.[151,1436,151,1471]" box="[197,655,1232,1256]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736EC9F120FB1E" bold="true" box="[197,271,1232,1256]" pageId="14" pageNumber="15">Hosts.</emphasis>
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Plant:
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<taxonomicName id="8ACAC29EFFDC3A70FEE86EC8F2A0FB1E" authority="(Pall.) Regel." authorityName="(Pall.) Regel." box="[350,655,1232,1256]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pumila">
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<emphasis id="7FBE650FFFDC3A70FEE86EC8F1DCFB1E" box="[350,499,1232,1256]" italics="true" pageId="14" pageNumber="15">Pinus pumila</emphasis>
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(Pall.) Regel.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDC3A70FF736EEAF2FAFAFA" box="[197,725,1267,1292]" pageId="14" pageNumber="15" type="distribution">
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<paragraph id="4D75B91DFFDC3A70FF736EEAF2FAFAFA" blockId="14.[151,1436,151,1471]" box="[197,725,1267,1292]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736EEAF171FAFA" bold="true" box="[197,350,1267,1292]" pageId="14" pageNumber="15">Distribution.</emphasis>
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<collectingCountry id="35DDF98DFFDC3A70FED36EEDF19BFAFA" box="[357,436,1268,1292]" name="Russia" pageId="14" pageNumber="15">Russia</collectingCountry>
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: Far East.
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<collectingCountry id="35DDF98DFFDC3A70FD9F6EECF242FAFA" box="[553,621,1269,1292]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
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: Honshū.
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</paragraph>
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</subSubSection>
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<subSubSection id="05D0EA96FFDC3A70FF736F01F51EFA49" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="4D75B91DFFDC3A70FF736F01F4AEFAA2" blockId="14.[151,1436,151,1471]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736F01F1C2FAC6" bold="true" box="[197,493,1303,1328]" pageId="14" pageNumber="15">
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<typeStatus id="927107BFFFDC3A70FF736F01F12FFAC6" box="[197,256,1304,1328]" pageId="14" pageNumber="15">Type</typeStatus>
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material examined.
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</emphasis>
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<typeStatus id="927107BFFFDC3A70FE426F0EF2A0FAD9" box="[500,655,1303,1327]" pageId="14" pageNumber="15" type="paratype">PARATYPES</typeStatus>
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(
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<collectionCode id="2BDB21D8FFDC3A70FD2B6F0EF2D4FAD9" box="[669,763,1303,1327]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="14" pageNumber="15">ZMUM</collectionCode>
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): 2,
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<collectionCode id="2BDB21D8FFDC3A70FC9D6F01F3A7FAC6" box="[811,904,1304,1328]" pageId="14" pageNumber="15">ИАГАД</collectionCode>
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. обп. ЛГ–19 Стоковое, 11 KM ЮВ КУЛУ, 14 0 6 1976, Л. Глумкова,
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<typeStatus id="927107BFFFDC3A70FE286F24F223FAA3" box="[414,524,1341,1365]" pageId="14" pageNumber="15" type="paratype">Paratypus</typeStatus>
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<taxonomicName id="8ACAC29EFFDC3A70FDA56F25F376FAA2" authority="Nikitsky" authorityName="Nikitsky" box="[531,857,1340,1364]" class="Insecta" family="Derodontidae" genus="Laricobius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kovalevi">Laricobius kovalevi Nikitsky</taxonomicName>
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(red label with name hw).
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</paragraph>
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<paragraph id="4D75B91DFFDC3A70FF736F46F51EFA49" blockId="14.[151,1436,151,1471]" pageId="14" pageNumber="15">
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<emphasis id="7FBE650FFFDC3A70FF736F46F21CFA8E" bold="true" box="[197,563,1375,1400]" pageId="14" pageNumber="15">Additional material examined.</emphasis>
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5, Tekari-dake, Senzu, Honkawane-cho, Haibara-gun, Shizuoka Pref.,
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<collectingCountry id="35DDF98DFFDC3A70FAE36F79F5B7FA81" box="[1365,1432,1376,1399]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
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,
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<date id="39749FDDFFDC3A70FF216F9DF114FA6A" box="[151,315,1412,1436]" pageId="14" pageNumber="15">24. September</date>
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, 2001, Y. Tahira leg., altitude
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<quantity id="8A3214F8FFDC3A70FD3C6F9DF2CEFA6A" box="[650,737,1412,1436]" metricMagnitude="3" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="m" value="2540.0">2,540m</quantity>
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, on
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<taxonomicName id="8ACAC29EFFDC3A70FCA66F9DF38CFA6A" box="[784,931,1412,1436]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pumila">Pinus pumila</taxonomicName>
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(2,
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<collectionCode id="2BDB21D8FFDC3A70FC796F9DF407FA6A" box="[975,1064,1412,1436]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="14" pageNumber="15">OMNH</collectionCode>
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; 2,
|
||
<collectionCode id="2BDB21D8FFDC3A70FBF96F9DF4B0FA6A" LSID="urn:lsid:biocol.org:col:34070" box="[1103,1183,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="14" pageNumber="15">NZAC</collectionCode>
|
||
; 1,
|
||
<collectionCode id="2BDB21D8FFDC3A70FB736F9DF528FA6A" LSID="urn:lsid:biocol.org:col:34880" box="[1221,1287,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="14" pageNumber="15">YPM</collectionCode>
|
||
); 1, Mt. Hakkoda, Aomori Pref.,
|
||
<collectingCountry id="35DDF98DFFDC3A70FECB6FB1F1EFFA49" box="[381,448,1448,1471]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
|
||
,
|
||
<date id="39749FDDFFDC3A70FE7D6FBEF253FA49" box="[459,636,1447,1471]" pageId="14" pageNumber="15" value="2007-10-04">4 October 2007</date>
|
||
, S. Shiyake leg., altitude
|
||
<quantity id="8A3214F8FFDC3A70FC2E6FBEF3D9FA49" box="[920,1014,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="14" pageNumber="15" unit="m" value="1520.0">1,520 m</quantity>
|
||
, on
|
||
<taxonomicName id="8ACAC29EFFDC3A70FB926FBEF497FA49" box="[1060,1208,1447,1471]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pumila">Pinus pumila</taxonomicName>
|
||
(1,
|
||
<collectionCode id="2BDB21D8FFDC3A70FB526FBEF509FA49" LSID="urn:lsid:biocol.org:col:34880" box="[1252,1318,1447,1471]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="14" pageNumber="15">YPM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |