409 lines
76 KiB
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409 lines
76 KiB
XML
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<mods:title id="55E8E71AD3BCE8A764E670B938D0DCD4">Description of a new species of freshwater crab of the genus Arcithelphusa Pati & Sudha Devi, 2015 (Decapoda: Brachyura: Gecarcinucidae) from the Western Ghats, Kerala, India</mods:title>
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<mods:namePart id="CA2AC72FCDEC593066E2BFA4037B6A7D">Sujila, P. S.</mods:namePart>
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<mods:namePart id="9D4BA1B18A0C5684F04D8A1C90D0E8B8">Devi, A. R. Sudha</mods:namePart>
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<taxonomicName id="8AF64D24E03C7556FF3FF99CFDE4CA7E" ID-CoL="7NQY2" authority="Pati & Sudha Devi, 2015" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[151,637,1629,1656]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="205" phylum="Arthropoda" rank="genus">
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Pati & Sudha Devi, 2015
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(
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<figureCitation id="D5CD2A22E03C7556FF37F940FF75CA92" box="[159,236,1665,1691]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="2" pageNumber="205">Figs. 2</figureCitation>
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A–F, 3B, C, D, E–K, N–P, 4A–F)
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<emphasis id="7F82EAB5E03C7556FF3FF904FEBDCAD4" box="[151,292,1733,1757]" italics="true" pageId="2" pageNumber="205">Arcithelphusa</emphasis>
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<treatmentCitation id="CC5710B6E03C7556FE83F905FDCFCAD4" author="Pati, S. K. & Sudha Devi, A. R." box="[299,598,1732,1757]" page="35" pageId="2" pageNumber="205" year="2015">
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<bibRefCitation id="29674B56E03C7556FE83F905FDCFCAD4" author="Pati, S. K. & Sudha Devi, A. R." box="[299,598,1732,1757]" pageId="2" pageNumber="205" pagination="35" refId="ref8106" refString="Pati, S. K. & Sudha Devi, A. R. (2015 a) Description of a new genus and new species of freshwater crab (Brachyura: Gecarcinucidae) from the Western Ghats, Kerala India. Zoological Studies, 54, 35. https: // doi. org / 10.1186 / s 40555 - 015 - 0112 - 0" type="journal article" year="2015">Pati & Sudha Devi 2015a: 35</bibRefCitation>
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;
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<treatmentCitation id="CC5710B6E03C7556FDC9F905FCA3CAD4" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[609,826,1732,1757]" page="1297" pageId="2" pageNumber="205" year="2017">
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<bibRefCitation id="29674B56E03C7556FDC9F905FCA3CAD4" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[609,826,1732,1757]" pageId="2" pageNumber="205" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
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Pati
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<emphasis id="7F82EAB5E03C7556FD27F907FD22CAD4" box="[655,699,1733,1757]" italics="true" pageId="2" pageNumber="205">et al</emphasis>
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. 2017: 1297
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</bibRefCitation>
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</treatmentCitation>
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(list);
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<treatmentCitation id="CC5710B6E03C7556FCD5F905FBFECAD4" author="Rajesh, L. & Raj, S. & Pati, S. K. & Biju Kumar, A." box="[893,1127,1732,1757]" page="134" pageId="2" pageNumber="205" year="2017">
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<bibRefCitation id="29674B56E03C7556FCD5F905FBFECAD4" author="Rajesh, L. & Raj, S. & Pati, S. K. & Biju Kumar, A." box="[893,1127,1732,1757]" pageId="2" pageNumber="205" pagination="132 - 153" refId="ref8642" refString="Rajesh, L., Raj, S., Pati, S. K. & Biju Kumar, A. (2017) The freshwater crabs (Decapoda: Brachyura) of Kerala, India. Journal of Aquatic Biology & Fisheries, 5, 132 - 153." type="journal article" year="2017">
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Rajesh
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<emphasis id="7F82EAB5E03C7556FC60F907FC6CCAD4" box="[968,1013,1733,1757]" italics="true" pageId="2" pageNumber="205">et al</emphasis>
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. 2017: 134
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</bibRefCitation>
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</treatmentCitation>
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(list);
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<treatmentCitation id="CC5710B6E03C7556FB02F905FF79CAF5" author="Pati, S. K. & Thackeray, T." page="10" pageId="2" pageNumber="205" year="2018">
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<bibRefCitation id="29674B56E03C7556FB02F905FF79CAF5" author="Pati, S. K. & Thackeray, T." pageId="2" pageNumber="205" pagination="1 - 73" refId="ref8233" refString="Pati, S. K. & Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati & Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati & Thackeray 2018: 10</bibRefCitation>
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</treatmentCitation>
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(list).
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</paragraph>
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<paragraph id="4D4936A7E03C7556FF3FF8EBFA12CB4D" blockId="2.[151,1436,1833,2041]" box="[151,1419,1833,1860]" pageId="2" pageNumber="205">
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<emphasis id="7F82EAB5E03C7556FF3FF8EBFEABCB4D" bold="true" box="[151,306,1833,1860]" pageId="2" pageNumber="205">
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<typeStatus id="924D8805E03C7556FF3FF8EBFF4BCB4D" box="[151,210,1834,1860]" pageId="2" pageNumber="205">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="8AF64D24E03C7556FE91F8EBFC29CB4D" ID-CoL="6828H" authority="Pati & Sudha Devi, 2015" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[313,944,1833,1860]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="cochleariformis">
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<emphasis id="7F82EAB5E03C7556FE91F8EBFD14CB4D" box="[313,653,1834,1860]" italics="true" pageId="2" pageNumber="205">Arcithelphusa cochleariformis</emphasis>
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Pati & Sudha Devi, 2015
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, by original designation; gender feminine.
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</paragraph>
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<subSubSection id="05EC652CE03C7557FF6FF88CFB7CCE8E" lastPageId="3" lastPageNumber="206" pageId="2" pageNumber="205" type="diagnosis">
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<paragraph id="4D4936A7E03C7557FF6FF88CFB7CCE8E" blockId="2.[151,1436,1833,2041]" lastBlockId="3.[151,1437,153,647]" lastPageId="3" lastPageNumber="206" pageId="2" pageNumber="205">
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<emphasis id="7F82EAB5E03C7556FF6FF88CFEA3CB61" bold="true" box="[199,314,1869,1896]" pageId="2" pageNumber="205">Diagnosis</emphasis>
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[emended from
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<bibRefCitation id="29674B56E03C7556FE50F88FFCA9CB61" author="Pati, S. K. & Sudha Devi, A. R." box="[504,816,1869,1896]" pageId="2" pageNumber="205" pagination="35" refId="ref8106" refString="Pati, S. K. & Sudha Devi, A. R. (2015 a) Description of a new genus and new species of freshwater crab (Brachyura: Gecarcinucidae) from the Western Ghats, Kerala India. Zoological Studies, 54, 35. https: // doi. org / 10.1186 / s 40555 - 015 - 0112 - 0" type="journal article" year="2015">Pati & Sudha Devi (2015a)</bibRefCitation>
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]. Carapace ovate, broader than long (cw/cl = 1.2–1.4), relatively deep (ch/cw = 0.6–0.7); dorsal surface arched, almost smooth; anterolateral margin lacking distinct serrations; front strongly deflexed, with narrow frontal margin (fw/cw = 0.25); epigastric cristae only visible as 2 low, broad, oval protuberances; postorbital cristae indistinct; external orbital angle indistinct; epibranchial tooth indistinct; branchial regions highly inflated; cervical grooves relatively shallow, not reaching beyond level of postorbital cristae; frontal median triangle incomplete, lateral margins indiscernible; epistome posterior margin with triangular median lobe and sinuous lateral lobes (
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<figureCitation id="D5CD2A22E03D7557FDE4FF7CFC8BCCDE" box="[588,786,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2A, B, D, E</figureCitation>
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,
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||
<figureCitation id="D5CD2A22E03D7557FCB6FF7CFC3ECCDE" box="[798,935,189,215]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="3" pageNumber="206">4A, B, D, E</figureCitation>
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). Third maxilliped exopod longer than ischium, lacking flagellum or with relatively short flagellum, ca. 0.5× width of merus (
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<figureCitation id="D5CD2A22E03D7557FB85FF20FB31CCF2" box="[1069,1192,225,251]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">Fig. 3B, N</figureCitation>
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). Chelipeds relatively smooth (
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<figureCitation id="D5CD2A22E03D7557FF51FEC4FEDCCD16" box="[249,325,261,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2</figureCitation>
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A–F, 4A–F). Ambulatory legs slender, long (
|
||
<figureCitation id="D5CD2A22E03D7557FC90FEC4FC1DCD16" box="[824,900,261,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2</figureCitation>
|
||
A–F, 4A–F). Suture between male thoracic sternites s2/s3 shallow or deep, broad, not reaching edge of sternum; suture between male thoracic sternites s3/s4 deep, broad, reaching edge of sternum (
|
||
<figureCitation id="D5CD2A22E03D7557FDB3FE8CFD04CD6E" box="[539,669,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FD03FE8CFD54CD6E" box="[683,717,333,359]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">3C</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FD74FE8CFC99CD6E" box="[732,768,333,359]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="3" pageNumber="206">4C</figureCitation>
|
||
). Pleonal locking mechanism with low, anteriorly directed tubercles on each antero-submedial part of sternite 5 (
|
||
<figureCitation id="D5CD2A22E03D7557FD5CFEB0FCDACD82" box="[756,835,369,395]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">
|
||
Fig.
|
||
<date id="39481067E03D7557FC81FEB0FCDACD82" box="[809,835,369,395]" pageId="3" pageNumber="206">3I</date>
|
||
</figureCitation>
|
||
). Male sternopleonal cavity long, reaching imaginary line joining medial part of cheliped coxae (
|
||
<figureCitation id="D5CD2A22E03D7557FD2AFE54FC9BCDA6" box="[642,770,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FCA7FE54FCAACDA6" box="[783,819,405,431]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="3" pageNumber="206">4C</figureCitation>
|
||
). Male pleon narrowly triangular, with concave lateral margins; pleonal somite 6 trapezoidal, broader than long, with gently convex lateral margins (
|
||
<figureCitation id="D5CD2A22E03D7557FB1AFE78FAA9CDDA" box="[1202,1328,441,467]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FA93FE78FAC6CDDA" box="[1339,1375,441,467]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">3D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FAC3FE78FA09CDDA" box="[1387,1424,441,467]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="3" pageNumber="206">4C</figureCitation>
|
||
). Male telson narrow, with strongly concave lateral margins; apex rounded (
|
||
<figureCitation id="D5CD2A22E03D7557FC7DFE1CFBCACDFE" box="[981,1107,477,503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206">Figs. 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FBF6FE1CFB1BCDFE" box="[1118,1154,477,503]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">3D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03D7557FB26FE1CFB2BCDFE" box="[1166,1202,477,503]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="3" pageNumber="206">4C</figureCitation>
|
||
). G1 relatively stout, with large flexible zone, tip not reaching pleonal locking structure; terminal segment relatively short, ca. 0.3–0.4× length of subterminal segment, medial portion distinctly curved outwards, dorsal flap absent or low, broad; subterminal segment relatively stout, with strongly convex outer margin at basal half (
|
||
<figureCitation id="D5CD2A22E03D7557FB8EFD88FBFECE6A" box="[1062,1127,585,611]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">Fig. 3</figureCitation>
|
||
E–G, I, J, O). G2 short, ca. 0.6–0.7× length of G1; distal segment very short, ca. 0.2× length of basal segment (
|
||
<figureCitation id="D5CD2A22E03D7557FB96FDACFB41CE8E" box="[1086,1240,621,647]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="3" pageNumber="206">Fig. 3H, K, P</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1989662FE03D7557FF3FF888FB21CB8C" ID-DOI="http://doi.org/10.5281/zenodo.3458929" ID-Zenodo-Dep="3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="3" pageNumber="206" startId="3.[151,250,1865,1890]" targetBox="[151,1435,661,1846]" targetPageId="3">
|
||
<paragraph id="4D4936A7E03D7557FF3FF888FB21CB8C" blockId="3.[151,1436,1865,1926]" pageId="3" pageNumber="206">
|
||
<emphasis id="7F82EAB5E03D7557FF3FF888FE8CCB6B" bold="true" box="[151,277,1865,1890]" pageId="3" pageNumber="206">FIGURE 2.</emphasis>
|
||
<taxonomicName id="8AF64D24E03D7557FEB5F88BFD8FCB6B" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[285,534,1866,1890]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E03D7557FEB5F88BFD8FCB6B" box="[285,534,1866,1890]" italics="true" pageId="3" pageNumber="206">Arcithelphusa tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E03D7557FDB6F88BFDE8CB6B" bold="true" box="[542,625,1866,1890]" pageId="3" pageNumber="206">
|
||
<taxonomicNameLabel id="64B157CEE03D7557FDB6F88BFDE8CB6B" box="[542,625,1866,1890]" pageId="3" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (ZSI-WRC C.1816) (A–C);
|
||
<taxonomicName id="8AF64D24E03D7557FB95F88BFB62CB6B" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[1085,1275,1866,1890]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E03D7557FB95F88BFB62CB6B" box="[1085,1275,1866,1890]" italics="true" pageId="3" pageNumber="206">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="05EC652CE03A7550FF6FFF58FAC4CAA0" pageId="4" pageNumber="207" type="discussion">
|
||
<paragraph id="4D4936A7E03A7550FF6FFF58FAD9CE88" blockId="4.[151,1437,153,1776]" pageId="4" pageNumber="207">
|
||
<emphasis id="7F82EAB5E03A7550FF6FFF58FEA2CCBA" bold="true" box="[199,315,153,179]" pageId="4" pageNumber="207">Remarks.</emphasis>
|
||
Among the Indian genera of gecarcinucid crabs,
|
||
<taxonomicName id="8AF64D24E03A7550FCCDFF58FB98CCBA" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[869,1025,153,179]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FCCDFF58FB98CCBA" box="[869,1025,153,179]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
is most close to
|
||
<taxonomicName id="8AF64D24E03A7550FB16FF58FAE6CCBA" authorityName="Alcock" authorityYear="1909" box="[1214,1407,153,179]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FB16FF58FAE6CCBA" box="[1214,1407,153,179]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
in carapace morphology and gonopod structures (see
|
||
<bibRefCitation id="29674B56E03A7550FD78FF7DFC6BCCDE" author="Pati, S. K. & Sudha Devi, A. R." box="[720,1010,188,215]" pageId="4" pageNumber="207" pagination="35" refId="ref8106" refString="Pati, S. K. & Sudha Devi, A. R. (2015 a) Description of a new genus and new species of freshwater crab (Brachyura: Gecarcinucidae) from the Western Ghats, Kerala India. Zoological Studies, 54, 35. https: // doi. org / 10.1186 / s 40555 - 015 - 0112 - 0" type="journal article" year="2015">Pati & Sudha Devi 2015a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="29674B56E03A7550FC55FF7DFB32CCDE" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[1021,1195,188,215]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FB87FF7CFBF9CCDF" box="[1071,1120,188,214]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
). Both the genera can be distinguished from other Indian gecarcinucid genera mainly by the presence of a relatively deep carapace (ch/cw = 0.5–0.7), a narrow frontal margin (ca. 0.25 times the carapace width), an incomplete frontal median triangle, and a short G2 distal segment (ca. 0.2–0.3 times the length of the basal segment) (
|
||
<figureCitation id="D5CD2A22E03A7550FC43FEE6FBF5CD48" box="[1003,1132,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="4" pageNumber="207">Figs. 2B, E</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03A7550FBD0FEE6FB40CD48" box="[1144,1241,295,321]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">3H, K, P</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03A7550FB4DFEE6FABECD48" box="[1253,1319,295,321]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="4" pageNumber="207">4B, E</figureCitation>
|
||
; see
|
||
<bibRefCitation id="29674B56E03A7550FAF6FEE6FEBDCD6D" author="Bahir, M. M. & Yeo, D. C. J." pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: figs. 4D, 5B;
|
||
<bibRefCitation id="29674B56E03A7550FE63FE8BFDE5CD6D" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[459,636,330,357]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FE56FE8AFDA8CD6D" box="[510,561,330,356]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 11b, 12d, 13b, 14d). Other major characters common to both are their strongly convex dorsal surface, the highly inflated branchial regions, the prominent and triangular median lobe on the epistome posterior margin, a narrowly triangular male pleon, and a short G2 (
|
||
<figureCitation id="D5CD2A22E03A7550FBE1FE50FB0CCDA2" box="[1097,1173,401,427]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="4" pageNumber="207">Figs. 2</figureCitation>
|
||
A–F, 3D, E, H, J, K, O, P, 4A–E; see
|
||
<bibRefCitation id="29674B56E03A7550FE84FE74FE65CDC6" author="Bahir, M. M. & Yeo, D. C. J." box="[300,508,437,463]" pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: figs. 4A, D, 5A–C;
|
||
<bibRefCitation id="29674B56E03A7550FD4AFE74FC09CDC6" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[738,912,437,463]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FCBDFE77FCDFCDC6" box="[789,838,437,463]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 11a–c, h, 12a, d, 13a–c, h, 14a, d). While they share many features of the carapace and gonopods, the flagellum on the exopod of the third maxilliped shows considerable variations. The flagellum on the third maxilliped exopod is altogether absent in
|
||
<taxonomicName id="8AF64D24E03A7550FB61FE3DFA05CE1F" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[1225,1436,508,534]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E03A7550FB61FE3DFA05CE1F" box="[1225,1436,508,534]" italics="true" pageId="4" pageNumber="207">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="D5CD2A22E03A7550FF08FDDEFF6ECE30" box="[160,247,543,569]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3N</figureCitation>
|
||
) whereas it is distinct but short in
|
||
<taxonomicName id="8AF64D24E03A7550FDDAFDE1FD67CE30" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[626,766,543,570]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E03A7550FDDAFDE1FD67CE30" box="[626,766,543,570]" italics="true" pageId="4" pageNumber="207">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E03A7550FCACFDDEFCC4CE30" bold="true" box="[772,861,543,569]" pageId="4" pageNumber="207">
|
||
<taxonomicNameLabel id="64B157CEE03A7550FCACFDDEFCC4CE30" box="[772,861,543,569]" pageId="4" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, reaching nearly half the merus width (
|
||
<figureCitation id="D5CD2A22E03A7550FAA7FDDEFAFFCE30" box="[1295,1382,543,569]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3B</figureCitation>
|
||
). On the other hand, all the species of
|
||
<taxonomicName id="8AF64D24E03A7550FDAEFD82FD5ECE54" authorityName="Alcock" authorityYear="1909" box="[518,711,579,605]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FDAEFD82FD5ECE54" box="[518,711,579,605]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
have a well-developed and relatively long flagellum on the third maxilliped exopod, which reaches up to 0.6–0.7 times the merus width (see
|
||
<bibRefCitation id="29674B56E03A7550FC43FDA7FB01CE89" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[1003,1176,614,641]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FBB5FDA6FBD7CE89" box="[1053,1102,614,640]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs.
|
||
<date id="39481067E03A7550FB73FDA7FA99CE89" box="[1243,1280,614,640]" pageId="4" pageNumber="207" value="2011-01">11i</date>
|
||
,
|
||
<date id="39481067E03A7550FAA4FDA7FAAACE88" box="[1292,1331,614,641]" pageId="4" pageNumber="207" value="2013-01">13i</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03A7550FF6FFD48FC68C844" blockId="4.[151,1437,153,1776]" pageId="4" pageNumber="207">
|
||
The variations in the length of the flagellum on the third maxilliped exopod could be associated with the ecological requirements of each species of
|
||
<taxonomicName id="8AF64D24E03A7550FDCDFD6CFC98CECE" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[613,769,685,711]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FDCDFD6CFC98CECE" box="[613,769,685,711]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AF64D24E03A7550FC95FD6CFC67CECE" authorityName="Alcock" authorityYear="1909" box="[829,1022,685,711]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FC95FD6CFC67CECE" box="[829,1022,685,711]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
. According to
|
||
<bibRefCitation id="29674B56E03A7550FB06FD6CFA0ECECE" author="Cumberlidge, N." box="[1198,1431,685,711]" pageId="4" pageNumber="207" refId="ref7512" refString="Cumberlidge, N. (1999) The freshwater crabs of West Africa. Family Potamonautidae. Faune et Flore Tropicales 35. Institut de Recherche pour le Developpement, Paris, 382 pp." type="book" year="1999">Cumberlidge (1999)</bibRefCitation>
|
||
, the presence of a long flagellum on the third maxilliped exopod of a crab is a plesiomorphic state, and the absence of the said flagellum is an apomorphic state, which is common in air-breathing crabs and mostly associated with the adaptations of the respiratory system by the relatively terrestrial species of freshwater crabs (see
|
||
<bibRefCitation id="29674B56E03A7550FAA8FCD9FF4ACF5C" author="Ng, P. K. L. & Shokita, S." pageId="4" pageNumber="207" pagination="1 - 7" refId="ref7982" refString="Ng, P. K. L. & Shokita, S. (1995) Ryukyum, a new genus of terrestrial crab from the Ryukyu Islands (Brachyura: Potamidae). Crustacean Research, 24, 1 - 7. https: // doi. org / 10.18353 / crustacea. 24.0 _ 1" type="journal article" year="1995">Ng & Shokita 1995</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="29674B56E03A7550FF40FCFAFE55CF5C" author="Cumberlidge, N." box="[232,460,827,853]" pageId="4" pageNumber="207" refId="ref7512" refString="Cumberlidge, N. (1999) The freshwater crabs of West Africa. Family Potamonautidae. Faune et Flore Tropicales 35. Institut de Recherche pour le Developpement, Paris, 382 pp." type="book" year="1999">Cumberlidge (1999)</bibRefCitation>
|
||
also noted that this character is useful to distinguish some genera of the West African crabs. The presence or absence of a flagellum on the third maxilliped exopod, however, is not a reliable character to exclusively define
|
||
<taxonomicName id="8AF64D24E03A7550FEC4FC43FD91CF95" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[364,520,898,924]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FEC4FC43FD91CF95" box="[364,520,898,924]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AF64D24E03A7550FDE9FC43FC9BCF95" authorityName="Alcock" authorityYear="1909" box="[577,770,898,924]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FDE9FC43FC9BCF95" box="[577,770,898,924]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
because the condition of the flagellum varies considerably among their species depending upon the habitat preferences.
|
||
<bibRefCitation id="29674B56E03A7550FCE7FC67FC74CFB6" author="Huang, C." box="[847,1005,933,960]" pageId="4" pageNumber="207" pagination="173 - 179" refId="ref7665" refString="Huang, C. (2018) Revision of Yarepotamon Dai & Turkay, 1997 (Brachyura: Potamidae), freshwater crabs endemic to southern China, with descriptions of two new genera and four new species. Journal of Crustacean Biology, 38 (2), 173 - 179. https: // doi. org / 10.1093 / jcbiol / rux 120" type="journal article" year="2018">Huang (2018)</bibRefCitation>
|
||
recently found that some crab genera are strikingly similar in external morphology, and the use of the G1 characters alone could be problematic to define them. In these situations, he suggested using a combination of characters that are apparently under less environmental selective pressure, such as male pleon, G1, and female vulvae. Furthermore, a molecular study has now become necessary to support morphological data (see
|
||
<bibRefCitation id="29674B56E03A7550FD3FFBF2FCD5C844" author="Shih, H. - T. & Huang, C. & Ng, P. K. L." box="[663,844,1075,1102]" pageId="4" pageNumber="207" pagination="309 - 331" refId="ref8693" refString="Shih, H. - T., Huang, C. & Ng, P. K. L. (2016) A re-appraisal of the widely-distributed freshwater crab genus Sinopotamon Bott, 1967, from China, with establishment of a new genus (Crustacea: Decapoda: Potamidae). Zootaxa, 4138 (2), 309 - 331. https: // doi. org / 10.11646 / zootaxa. 4138.2.5" type="journal article" year="2016">
|
||
Shih
|
||
<emphasis id="7F82EAB5E03A7550FD79FBF5FC9BC844" box="[721,770,1075,1101]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="29674B56E03A7550FCFFFBF5FC7FC844" author="Huang, C." box="[855,998,1075,1102]" pageId="4" pageNumber="207" pagination="173 - 179" refId="ref7665" refString="Huang, C. (2018) Revision of Yarepotamon Dai & Turkay, 1997 (Brachyura: Potamidae), freshwater crabs endemic to southern China, with descriptions of two new genera and four new species. Journal of Crustacean Biology, 38 (2), 173 - 179. https: // doi. org / 10.1093 / jcbiol / rux 120" type="journal article" year="2018">Huang 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03A7550FF6FFB97FC1DC8F6" blockId="4.[151,1437,153,1776]" pageId="4" pageNumber="207">
|
||
Currently, no molecular data is available for
|
||
<taxonomicName id="8AF64D24E03A7550FD69FB96FCC4C878" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[705,861,1111,1137]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FD69FB96FCC4C878" box="[705,861,1111,1137]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AF64D24E03A7550FC3DFB96FBCFC878" authorityName="Alcock" authorityYear="1909" box="[917,1110,1111,1137]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FC3DFB96FBCFC878" box="[917,1110,1111,1137]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
, and the female vulvae were not described for all the species. Their male pleon is quite similar in morphology. On the contrary, the G1 structure is a reliable character to set them apart. While the generic treatment of
|
||
<taxonomicName id="8AF64D24E03A7550FC1EFB5CFBCBC8BE" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[950,1106,1181,1207]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FC1EFB5CFBCBC8BE" box="[950,1106,1181,1207]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
needs investigation based on morphological and molecular data, we still continue to recognize the genus and provide here an emended diagnosis to accommodate both
|
||
<taxonomicName id="8AF64D24E03A7550FE27FB24FDC6C8F7" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[399,607,1252,1279]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="cochleariformis">
|
||
<emphasis id="7F82EAB5E03A7550FE27FB24FDC6C8F7" box="[399,607,1252,1279]" italics="true" pageId="4" pageNumber="207">A. cochleariformis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8AF64D24E03A7550FD3EFB24FCBAC8F7" authority="Pati & Sujila & Devi, 2019" authorityName="Pati & Sujila & Devi" authorityYear="2019" box="[662,803,1252,1279]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="tumpikkai" status="sp. nov.">
|
||
<emphasis id="7F82EAB5E03A7550FD3EFB24FCBAC8F7" box="[662,803,1252,1279]" italics="true" pageId="4" pageNumber="207">A. tumpikkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7F82EAB5E03A7550FC83FB24FC1DC8F6" bold="true" box="[811,900,1253,1279]" pageId="4" pageNumber="207">
|
||
<taxonomicNameLabel id="64B157CEE03A7550FC83FB24FC1DC8F6" box="[811,900,1253,1279]" pageId="4" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4D4936A7E03A7550FF6FFAC9FAC4CAA0" blockId="4.[151,1437,153,1776]" pageId="4" pageNumber="207">
|
||
<taxonomicName id="8AF64D24E03A7550FF6FFAC9FEFAC92B" authorityName="Pati & Sudha Devi" authorityYear="2015" box="[199,355,1288,1314]" class="Malacostraca" family="Gecarcinucidae" genus="Arcithelphusa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FF6FFAC9FEFAC92B" box="[199,355,1288,1314]" italics="true" pageId="4" pageNumber="207">Arcithelphusa</emphasis>
|
||
</taxonomicName>
|
||
can be now distinguished from
|
||
<taxonomicName id="8AF64D24E03A7550FD6EFAC9FC1EC92B" authorityName="Alcock" authorityYear="1909" box="[710,903,1288,1314]" class="Malacostraca" family="Gecarcinucidae" genus="Cylindrotelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F82EAB5E03A7550FD6EFAC9FC1EC92B" box="[710,903,1288,1314]" italics="true" pageId="4" pageNumber="207">Cylindrotelphusa</emphasis>
|
||
</taxonomicName>
|
||
by the following suite of characters: the anterolateral margins of the carapace lack distinct serrations (
|
||
<figureCitation id="D5CD2A22E03A7550FD52FAEAFCE6C94F" box="[762,895,1323,1350]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="4" pageNumber="207">Figs. 2A, D</figureCitation>
|
||
,
|
||
<figureCitation id="D5CD2A22E03A7550FC22FAEAFC57C94F" box="[906,974,1323,1350]" captionStart="FIGURE 4" captionStartId="8.[151,250,1420,1445]" captionTargetBox="[151,1435,382,1367]" captionTargetId="figure@8.[151,1435,382,1367]" captionTargetPageId="8" captionText="FIGURE 4. Arcithelphusa tumpikkai sp. nov., paratype, male (ZSI-WRC C.1818) (A–C), paratype, female (ZSI-WRC C.1818) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458935" httpUri="https://zenodo.org/record/3458935/files/figure.png" pageId="4" pageNumber="207">4A, D</figureCitation>
|
||
) (vs. with distinct serrations; see
|
||
<bibRefCitation id="29674B56E03A7550FAEAFAEAFE9CC960" author="Bahir, M. M. & Yeo, D. C. J." pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: fig. 5A;
|
||
<bibRefCitation id="29674B56E03A7550FEC4FA8EFD81C960" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[364,536,1359,1385]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FE36FA91FE56C960" box="[414,463,1359,1385]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 11a, 13a); the third maxilliped exopod lacks a flagellum (
|
||
<figureCitation id="D5CD2A22E03A7550FB7DFA8EFAB5C960" box="[1237,1324,1359,1385]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3N</figureCitation>
|
||
) or with a short flagellum that reaches half the merus width (
|
||
<figureCitation id="D5CD2A22E03A7550FD69FAB3FC8FC985" box="[705,790,1394,1420]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3B</figureCitation>
|
||
) (vs. with a well-developed flagellum that reaches beyond half the merus width; see
|
||
<bibRefCitation id="29674B56E03A7550FE1BFA57FDC7C9B9" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[435,606,1430,1456]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FE4DFA56FD8CC9B9" box="[485,533,1430,1456]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs.
|
||
<date id="39481067E03A7550FD37FA57FD5DC9B9" box="[671,708,1430,1456]" pageId="4" pageNumber="207" value="2011-01">11i</date>
|
||
,
|
||
<date id="39481067E03A7550FD67FA57FD6FC9B8" box="[719,758,1430,1457]" pageId="4" pageNumber="207" value="2013-01">13i</date>
|
||
); the chelipeds are relatively smooth (
|
||
<figureCitation id="D5CD2A22E03A7550FB33FA57FB7FC9B9" box="[1179,1254,1430,1456]" captionStart="FIGURE 2" captionStartId="3.[151,250,1865,1890]" captionTargetBox="[151,1435,661,1846]" captionTargetId="figure@3.[151,1435,661,1846]" captionTargetPageId="3" captionText="FIGURE 2. Arcithelphusa tumpikkai sp. nov., holotype male (ZSI-WRC C.1816) (A–C); A. cochleariformis, holotype male (ZSI-WRC C.1188) (D–F). A, D, dorsal view; B, E, frontal view; C, F, ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3458929" httpUri="https://zenodo.org/record/3458929/files/figure.png" pageId="4" pageNumber="207">Figs. 2</figureCitation>
|
||
A–F, 4A–F) (vs. rugose; see
|
||
<bibRefCitation id="29674B56E03A7550FEB2FA78FE70C9DA" author="Bahir, M. M. & Yeo, D. C. J." box="[282,489,1465,1492]" pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: fig. 5A–C;
|
||
<bibRefCitation id="29674B56E03A7550FDD9FA78FC84C9DA" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[625,797,1465,1492]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FD0BFA7BFD4DC9DA" box="[675,724,1465,1491]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 11a, c, d, 13a, c, d); the G1 is relatively stout with a large flexible zone (
|
||
<figureCitation id="D5CD2A22E03A7550FEDDFA1CFE2DC9FE" box="[373,436,1501,1527]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3</figureCitation>
|
||
E–G, J, O) (vs. relatively slender with a reduced flexible zone; see
|
||
<bibRefCitation id="29674B56E03A7550FB36FA1CFAF5C9FE" author="Bahir, M. M. & Yeo, D. C. J." box="[1182,1388,1501,1527]" pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: fig. 4A–C;
|
||
<bibRefCitation id="29674B56E03A7550FF40F9C1FE0CCA13" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[232,405,1536,1563]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FEB2F9C0FED2CA13" box="[282,331,1536,1562]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 12a–c, 14a–c); the G1 terminal segment is relatively short, ca. 0.3–0.4 times the length of the subterminal segment (
|
||
<figureCitation id="D5CD2A22E03A7550FE70F9E5FDF0CA37" box="[472,617,1572,1598]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3E, J, O</figureCitation>
|
||
) (vs. relatively long, ca. 0.4–0.6 times the length of the subterminal segment; see
|
||
<bibRefCitation id="29674B56E03A7550FEAFF986FE4ACA68" author="Bahir, M. M. & Yeo, D. C. J." box="[263,467,1607,1634]" pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: fig. 4A;
|
||
<bibRefCitation id="29674B56E03A7550FD91F986FD7ACA68" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[569,739,1607,1634]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FDC2F989FD02CA68" box="[618,667,1607,1633]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 12a, 14a); the G1 subterminal segment is relatively stout, with a strongly convex outer margin at the basal half (
|
||
<figureCitation id="D5CD2A22E03A7550FD40F9ABFC17CA8C" box="[744,910,1642,1669]" captionStart="FIGURE 3" captionStartId="6.[151,250,1823,1848]" captionTargetBox="[255,1346,170,1792]" captionTargetId="figure@6.[237,1350,166,1803]" captionTargetPageId="6" captionText="FIGURE 3. Arcithelphusa tumpikkai sp. nov., holotype, male (ZSI-WRC C.1816) (A–I), paratype, male (ZSI-WRC C.1818) (J, K), paratype, female (ZSI-WRC C.1818) (L); A. cochleariformis, holotype, male (ZSI-WRC C.1188) (M–P). A, M, posterior margin of carapace; B, N, left third maxilliped; C, anterior thoracic sternites (s1–s4); D, pleon; E, J, dorsal view of left G1; F, ventral view of left G1; G, mesial view of left G1; H, K, left G2; I, ventral view showing sternopleonal cavity with G1 in situ; L, ventral view showing sternopleonal cavity with vulvae; O, dorsal view of right G1; P, right G2. Scale bars = 5 mm (A, M), 2 mm (B–D, I, L, N), 1 mm (E–H, J, K, O, P)." figureDoi="http://doi.org/10.5281/zenodo.3458933" httpUri="https://zenodo.org/record/3458933/files/figure.png" pageId="4" pageNumber="207">Fig. 3E, F, J, O</figureCitation>
|
||
) (vs. relatively slender, with a straight to gently convex outer margin at the basal half; see
|
||
<bibRefCitation id="29674B56E03A7550FDC7F94FFCA6CAA1" author="Bahir, M. M. & Yeo, D. C. J." box="[623,831,1678,1705]" pageId="4" pageNumber="207" pagination="309 - 354" refId="ref7465" refString="Bahir, M. M. & Yeo, D. C. J. (2007) The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 309 - 354." type="journal article" year="2007">Bahir & Yeo 2007</bibRefCitation>
|
||
: fig. 4A, B;
|
||
<bibRefCitation id="29674B56E03A7550FC61F94FFBEFCAA1" author="Pati, S. K. & Rajesh, L. & Raj, S. & Sheeja, V. U. & Kumar, A. B. & Sureshan, P. M." box="[969,1142,1678,1705]" pageId="4" pageNumber="207" pagination="1295 - 1330" refId="ref8399" refString="Pati, S. K., Rajesh, L., Raj, S., Sheeja, V. U., Kumar, A. B. & Sureshan, P. M. (2017) Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India. Journal of Natural History, 51 (23 - 24), 1295 - 1330. https: // doi. org / 10.1080 / 00222933.2017.1324054" type="journal article" year="2017">
|
||
Pati
|
||
<emphasis id="7F82EAB5E03A7550FC53F94EFBB4CAA1" box="[1019,1069,1678,1704]" italics="true" pageId="4" pageNumber="207">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
: figs. 12a, c, 14a, c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EC652CE03A7550FF6FF970FEDDCAF9" pageId="4" pageNumber="207" type="distribution">
|
||
<paragraph id="4D4936A7E03A7550FF6FF970FEDDCAF9" blockId="4.[151,1437,153,1776]" pageId="4" pageNumber="207">
|
||
<emphasis id="7F82EAB5E03A7550FF6FF970FEF8CAC5" bold="true" box="[199,353,1713,1740]" pageId="4" pageNumber="207">Distribution.</emphasis>
|
||
The genus is endemic to the Western Ghats of
|
||
<collectingCountry id="35E17637E03A7550FCD7F970FC20CAC2" box="[895,953,1713,1739]" name="India" pageId="4" pageNumber="207">India</collectingCountry>
|
||
and known only from Wayanad district of
|
||
<collectingRegion id="8F32F845E03A7550FF3FF914FF7BCAE6" box="[151,226,1749,1775]" country="India" name="Kerala" pageId="4" pageNumber="207">Kerala</collectingRegion>
|
||
(
|
||
<figureCitation id="D5CD2A22E03A7550FF59F914FEABCAF9" box="[241,306,1749,1776]" captionStart="FIGURE 1" captionStartId="1.[151,250,1897,1922]" captionTargetBox="[176,1411,166,1877]" captionTargetId="figure@1.[176,1411,166,1877]" captionTargetPageId="1" captionText="FIGURE 1. Map showing distribution of Arcithelphusa tumpikkai sp. nov. and A. cochleariformis. Collection/observation sites (1–17): 1, Makkimala (type locality); 2, Kambamala; 3, Puthiyidam; 4, Kappikalam; 5, Varayal; 6, Thirunelly; 7, Bavali; 8, 9, Ondayangadi; 10, Pakkom; 11, Mananthavady; 12, Chettapalam; 13, Ellumannam; 14, Panamaram; 15, Padinjarathara; 16, Kambalakkad; 17, Muthanga." figureDoi="http://doi.org/10.5281/zenodo.3458927" httpUri="https://zenodo.org/record/3458927/files/figure.png" pageId="4" pageNumber="207">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |