treatments-xml/data/C5/3C/DE/C53CDE032678FF88AB85FADEBFC98FC4.xml

<document id="7B194C6B03BCB789FB2019BB17A10AD0" ID-CLB-Dataset="3915" ID-DOI="10.11646/zootaxa.3881.1.6" ID-GBIF-Dataset="6bb77067-0860-413d-a018-eab590cae615" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949744" ID-ZooBank="F1D2B596-8630-49E2-B704-76CBBDF5353E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623706258562" checkinUser="felipe" docAuthor="Matsui, Masafumi, Belabut, Daicus M. &amp; Ahmad, Norhayati" docDate="2014" docId="C53CDE032678FF88AB85FADEBFC98FC4" docLanguage="en" docName="zt03881p093.pdf" docOrigin="Zootaxa 3881 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnonectes utara Matsui &amp; Belabut &amp; Ahmad 2014, sp. nov." docType="treatment" docUuid="19AC8174-73A9-475E-A965-A60911F42CA7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="86" masterDocId="3905A67B267DFF83AB12FF85BA598D34" masterDocTitle="Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae)" masterLastPageNumber="93" masterPageNumber="75" pageNumber="80" updateTime="1699015699284" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F158AA2C18B53C801EFA729E104801DD">Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae)</mods:title>
</mods:titleInfo>
<mods:name id="4B274427B3F768BBD602738CD0A1AFF6" type="personal">
<mods:role id="C141489CF2CB5478470C8A9573A83E76">
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<mods:namePart id="D439A6CFC69C8D26DBA86F640C4229D7">Matsui, Masafumi</mods:namePart>
<mods:affiliation id="5CD92560206F09C3DA3744E373B0C1E7">Graduate School of Human and Environmental Studies, Kyoto University, Kyoto 606 - 8501, Japan</mods:affiliation>
<mods:nameIdentifier id="DA043BCA431DB573961A0CC573CE3AEE" type="email">fumi@zoo.zool.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="B772C063794C5D9A4634F89B80FF121D">Belabut, Daicus M.</mods:namePart>
<mods:affiliation id="7793995DA15568164A1644EE2AD9D340">Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603 Kuala Lumpur, Malaysia &amp; Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia</mods:affiliation>
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<mods:namePart id="6EC6D0D2F4C656BA9179C49368BE2241">Ahmad, Norhayati</mods:namePart>
<mods:affiliation id="664B4FB43E337C36769F7BA57B2C9C21">Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia</mods:affiliation>
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<mods:title id="529702BD9A835AC0CC83A2CC5077C6D7">Zootaxa</mods:title>
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<mods:date id="0B4177E680CF90D5C83FCCAEC2C834EB">2014</mods:date>
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<mods:number id="77A85C1ADD8A04CE267004586897FF31">2014-11-03</mods:number>
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<mods:number id="F9991A680EDA812E0132F5AB8BE1026D">3881</mods:number>
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<treatment id="C53CDE032678FF88AB85FADEBFC98FC4" ID-DOI="http://doi.org/10.5281/zenodo.5593083" ID-GBIF-Taxon="189161837" ID-Zenodo-Dep="5593083" LSID="urn:lsid:zoobank.org:act:19AC8174-73A9-475E-A965-A60911F42CA7" httpUri="http://treatment.plazi.org/id/C53CDE032678FF88AB85FADEBFC98FC4" lastPageId="11" lastPageNumber="86" pageId="5" pageNumber="80">
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<heading id="1662D8792678FF86AB85FADEBB878842" bold="true" box="[151,478,1371,1398]" fontSize="11" level="1" pageId="5" pageNumber="80" reason="1">
<emphasis id="7FE1B3072678FF86AB85FADEBB878842" bold="true" box="[151,478,1371,1398]" pageId="5" pageNumber="80">
<taxonomicName id="8A9514962678FF86AB85FADEBB238841" ID-CoL="6QDC6" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[151,378,1371,1397]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072678FF86AB85FADEBB238841" bold="true" box="[151,378,1371,1397]" italics="true" pageId="5" pageNumber="80">Limnonectes utara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64D20E7C2678FF86AA92FAD9BB878842" box="[384,478,1372,1398]" pageId="5" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E2678FF86AB84FA27B9E488E3" pageId="5" pageNumber="80" type="reference_group">
<paragraph id="4D2A6F152678FF86AB84FA27B9E488E3" blockId="5.[150,1436,1442,1495]" pageId="5" pageNumber="80">
<emphasis id="7FE1B3072678FF86AB84FA27BB48888C" bold="true" box="[150,273,1442,1464]" pageId="5" pageNumber="80">Synonymy:</emphasis>
<taxonomicName id="8A9514962678FF86AA08FA26B8C7888C" authority=": Boulenger, 1912, p. 229" authorityName=": Boulenger" authorityPageNumber="229" authorityYear="1912" box="[282,670,1442,1464]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072678FF86AA08FA26BBCA888C" box="[282,403,1442,1464]" italics="true" pageId="5" pageNumber="80">Rana kuhlii</emphasis>
: 
<bibRefCitation id="290412E42678FF86AAB0FA27B816888C" author="Boulenger, G. A." box="[418,591,1442,1464]" pageId="5" pageNumber="80" refId="ref12043" refString="Boulenger, G. A. (1912) A Vertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including Adjacent Islands. Taylor and Francis, London, 294 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10813" type="book" year="1912">Boulenger, 1912</bibRefCitation>
, p. 229
</taxonomicName>
(part); 
<bibRefCitation id="290412E42678FF86A9FDFA27B92D888C" author="Taylor, E. H." box="[751,884,1442,1464]" pageId="5" pageNumber="80" pagination="265 - 599" refId="ref13631" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. The University of Kansas Science Bulletin, 43, 265 - 599." type="journal article" year="1962">Taylor, 1962</bibRefCitation>
, p. 408 (part); 
<bibRefCitation id="290412E42678FF86AF06FA26BEC8888C" author="Berry, P. Y." box="[1044,1169,1442,1464]" pageId="5" pageNumber="80" refId="ref12019" refString="Berry, P. Y. (1975) The Amphibian Fauna of Peninsular Malaysia. Tropical Press, Kuala Lumpur, 130 pp" type="book" year="1975">Berry, 1975</bibRefCitation>
, p. 71 (part). 
<taxonomicName id="8A9514962678FF86AE36FA26BBDF88E3" authority=": Inger, 1966, p. 196" authorityName=": Inger" authorityPageNumber="196" authorityYear="1966" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="kuhli">
<emphasis id="7FE1B3072678FF86AE36FA26BFCC888C" box="[1316,1429,1442,1464]" italics="true" pageId="5" pageNumber="80">Rana kuhli</emphasis>
: 
<bibRefCitation id="290412E42678FF86ABD5FA44BB6488E3" author="Inger, R. F." box="[199,317,1473,1495]" pageId="5" pageNumber="80" pagination="1 - 402" refId="ref12569" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana, Zoology, 52, 1 - 402." type="journal article" year="1966">Inger, 1966</bibRefCitation>
, p. 196
</taxonomicName>
(part). 
<taxonomicName id="8A9514962678FF86AAC0FA47B9E088E3" ID-CoL="72DKQ" authority=": Grismer et al., 2010, p. 152" authorityName="Grismer" authorityPageNumber="152" authorityYear="2010" box="[466,953,1473,1495]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="80" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072678FF86AAC0FA47B8CB88E3" box="[466,658,1473,1495]" italics="true" pageId="5" pageNumber="80">Limnonectes kuhlii</emphasis>
: 
<bibRefCitation id="290412E42678FF86A98DFA44B93588E3" author="Grismer, L. L. &amp; Chan, K. O. &amp; Grismer, J. L. &amp; Wood, P. L. Jr. &amp; Norhayati, A." box="[671,876,1473,1495]" pageId="5" pageNumber="80" pagination="147 - 160" refId="ref12508" refString="Grismer, L. L., Chan, K. O., Grismer, J. L., Wood, P. L. Jr. &amp; Norhayati, A. (2010) A checklist of the herpetofauna of the Banjaran Bintang, Peninsular Malaysia. Russian Journal of Herpetology, 17, 147 - 160." type="journal article" year="2010">
Grismer 
<emphasis id="7FE1B3072678FF86A9EAFA47B97C88E3" box="[760,805,1473,1495]" italics="true" pageId="5" pageNumber="80">et al</emphasis>
., 2010
</bibRefCitation>
, p. 152
</taxonomicName>
.
</paragraph>
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<materialsCitation id="FDFD65482678FF86AB85FA7BBAB58B56" ID-GBIF-Occurrence="3391168304" collectingDate="2011-02-28" collectionCode="UKMHC, KUHE" country="Malaysia" elevation="1160" latitude="4.85" location="Bukit Larut" longLatPrecision="1306" longitude="100.8" municipality="State" pageId="5" pageNumber="80" specimenCode="UKMHC 528, KUHE 54064" specimenCount="1" stateProvince="Perak" typeStatus="holotype">
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<typeStatus id="922ED1B72678FF86AB85FA7BBB5F8B23" box="[151,262,1534,1559]" pageId="5" pageNumber="80" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="1D33C76E2678FF86AA01FA7BBBF78B23" box="[275,430,1534,1559]" collectionCode="UKMHC" pageId="5" pageNumber="80">UKMHC 528</specimenCode>
(former 
<specimenCode id="1D33C76E2678FF86A906FA7AB8ED8B23" box="[532,692,1534,1559]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 54064</specimenCode>
), an adult male from the upper part of 
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(= Larut Hill), 
<collectingMunicipality id="AD4EF56F2678FF86AB85F9A0BA968B0A" box="[151,207,1573,1598]" pageId="5" pageNumber="80">State</collectingMunicipality>
of 
<collectingRegion id="8F51A1F72678FF86ABE4F9A0BB608B0A" box="[246,313,1573,1598]" country="Malaysia" name="Perak" pageId="5" pageNumber="80">Perak</collectingRegion>
, 
<collectingCountry id="35822F852678FF86AA57F9A0BBF58B0A" box="[325,428,1573,1598]" name="Malaysia" pageId="5" pageNumber="80">Malaysia</collectingCountry>
(
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4 
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51' N
</geoCoordinate>
, 
<geoCoordinate id="28A109D22678FF86A90DF9A0B8D78B09" box="[543,654,1570,1598]" degrees="100" direction="east" minutes="48" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="100.8">
100 
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48' E
</geoCoordinate>
, 
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<quantity id="8A6DC2F02678FF86A988F9A0B8B68B09" box="[666,751,1573,1598]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="80" unit="m" value="1160.0">1160 m</quantity>
a.s.l.
</elevation>
: 
<figureCitation id="D5AE73902678FF86A82EF9A0B9D78B0A" box="[828,910,1573,1598]" captionStart="FIGURE 10" captionStartId="16.[151,250,1380,1402]" captionTargetBox="[313,1209,424,1343]" captionTargetId="figure-364@16.[294,1229,415,1354]" captionTargetPageId="16" captionText="FIGURE 10. Schematic map of the Malay Peninsula, illustrating the five mountain ranges (from west to east: A, Banjaran Bintang; B, Banjaran Titiwangsa; C, Banjaran Benom; D, Banjaran Gunung Tahan; E, Banjaran Timur), the type localities (filled triangles) of Limnonectes utara sp. nov. (Larut Hill) and L. selatan sp. nov. (Genting Highlands), and known localities of either of the two or unidentified species (dark circles: 1, Hulu Terengganu; 2, Ulu Gombak; 3, Gunung Bubu; 4, Gunung Lawit; 5, Endau Rompin). Hatching indicates areas 300 m a.s.l. and above in elevation." figureDoi="http://doi.org/10.5281/zenodo.4949769" httpUri="https://zenodo.org/record/4949769/files/figure.png" pageId="5" pageNumber="80">Fig. 10</figureCitation>
), collected on 
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<collectingDate id="296FB03D2678FF86AF24F9A0BEA78B0A" box="[1078,1278,1573,1598]" pageId="5" pageNumber="80" value="2011-02-28">28 February 2011</collectingDate>
</date>
by Masafumi Matsui.
</materialsCitation>
</paragraph>
<paragraph id="4D2A6F152678FF86ABD5F9E8B89D8A22" blockId="5.[151,1436,1534,1994]" pageId="5" pageNumber="80">
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<typeStatus id="922ED1B72678FF86ABD5F9E8BB188BB2" box="[199,321,1645,1670]" pageId="5" pageNumber="80" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="1D33C76E2678FF86AA5FF9EBBBB28BB2" box="[333,491,1645,1670]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 15447</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86AAE4F9E8B8668BB2" box="[502,575,1645,1670]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">15463</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86A958F9E8B8CA8BB2" box="[586,659,1645,1670]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">15465</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86A98CF9E8B8BD8BB2" box="[670,740,1645,1670]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">15514</specimenCode>
(
<specimenCount id="5B93A49C2678FF86A9E1F9E8B93B8BB2" box="[755,866,1645,1670]" pageId="5" pageNumber="80" type="adult">four adult</specimenCount>
females)
</materialsCitation>
, 
<materialsCitation id="FDFD65482678FF86A8CAF9EBBF1A8BB2" ID-GBIF-Occurrence="3391168311" box="[984,1347,1645,1670]" collectionCode="KUHE" pageId="5" pageNumber="80" specimenCode="KUHE 15442" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86A8CAF9EBBE2A8BB2" box="[984,1139,1645,1670]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 15442</specimenCode>
(one young male)
</materialsCitation>
, 
<materialsCitation id="FDFD65482678FF86AE5CF9EBBBA08B9E" ID-GBIF-Occurrence="3391168306" collectionCode="KUHE" pageId="5" pageNumber="80" specimenCode="KUHE 15441" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86AE5CF9EBBA848B9E" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 15441</specimenCode>
(one young female), and
</materialsCitation>
<materialsCitation id="FDFD65482678FF86A910F917B8968BFA" ID-GBIF-Occurrence="3391168312" collectingDate="1993-01-03" collectionCode="KUHE" elevation="680" pageId="5" pageNumber="80" specimenCode="KUHE 15461" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86A910F917B8C48B9E" box="[514,669,1681,1706]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 15461</specimenCode>
(one unsexed young) from lower part of Bukit Larut (
<elevation id="C6B888262678FF86AE14F914BFD58B9E" box="[1286,1420,1681,1706]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="80" unit="m" value="680.0">
<quantity id="8A6DC2F02678FF86AE14F914BF098B9D" box="[1286,1360,1681,1706]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="80" unit="m" value="680.0">680 m</quantity>
a.s.l.
</elevation>
), collected on 
<date id="392B49D52678FF86AA3BF930BB8E8BFA" box="[297,471,1717,1742]" pageId="5" pageNumber="80" value="1993-01-03">
<collectingDate id="296FB03D2678FF86AA3BF930BB8E8BFA" box="[297,471,1717,1742]" pageId="5" pageNumber="80" value="1993-01-03">3 January 1993</collectingDate>
</date>
by Masafumi Matsui
</materialsCitation>
. 
<materialsCitation id="FDFD65482678FF86A9C9F933BEC68BFA" ID-GBIF-Occurrence="3391168319" box="[731,1183,1717,1742]" collectionCode="KUHE" pageId="5" pageNumber="80" specimenCode="KUHE 54065, 54089" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86A9C9F933B9228BFA" box="[731,891,1717,1742]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 54065</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86A895F930B9948BFA" box="[903,973,1717,1742]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">54089</specimenCode>
(
<specimenCount id="5B93A49C2678FF86A8CFF930BE108BFA" box="[989,1097,1717,1742]" pageId="5" pageNumber="80" type="adult">two adult</specimenCount>
males)
</materialsCitation>
, 
<materialsCitation id="FDFD65482678FF86AFBEF933BBDA8BC6" ID-GBIF-Occurrence="3391168317" collectionCode="KUHE" pageId="5" pageNumber="80" specimenCode="KUHE 54086-54088" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86AFBEF933BFC58BFA" box="[1196,1436,1717,1742]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 54086–54088</specimenCode>
(
<specimenCount id="5B93A49C2678FF86AB8CF95CBB418BC6" box="[158,280,1753,1778]" pageId="5" pageNumber="80" type="adult">three adult</specimenCount>
females)
</materialsCitation>
, 
<materialsCitation id="FDFD65482678FF86AA9DF95FB8998A22" ID-GBIF-Occurrence="3391168302" collectingDate="2011-02-27" collectingDateMax="2011-02-28" collectingDateMin="2011-02-27" collectionCode="KUHE" pageId="5" pageNumber="80" specimenCode="KUHE 54090" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D33C76E2678FF86AA9DF95FB8738BC6" box="[399,554,1753,1778]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 54090</specimenCode>
(one unsexed young) from the type locality, collected from 
<date id="392B49D52678FF86AFC0F95CBA978A22" pageId="5" pageNumber="80" value="2011-02-27" valueMax="2011-02-28" valueMin="2011-02-27">
<collectingDate id="296FB03D2678FF86AFC0F95CBA978A22" pageId="5" pageNumber="80" value="2011-02-27" valueMax="2011-02-28" valueMin="2011-02-27">27 to 28 February 2011</collectingDate>
</date>
by Masafumi Matsui and Norihiro Kuraishi
</materialsCitation>
.
</paragraph>
<paragraph id="4D2A6F152678FF86ABD5F8A4BF478AB6" blockId="5.[151,1436,1534,1994]" pageId="5" pageNumber="80">
<emphasis id="7FE1B3072678FF86ABD5F8A4BBE28A0E" bold="true" box="[199,443,1825,1850]" inLineHeading="true" pageId="5" pageNumber="80" reason="1">Referred specimens.</emphasis>
<materialsCitation id="FDFD65482678FF86AAD7F8A4BEF48A6A" ID-GBIF-Occurrence="3391168315" collectionCode="UKMHC" collectorName="Daicus M. Belabut" country="Malaysia" county="State" elevation="169" latitude="5.4333334" location="Sungai Tembat" longLatPrecision="1305" longitude="102.96667" municipality="Hulu" pageId="5" pageNumber="80" specimenCode="UKMHC 700, 703, 704" specimenCount="1" stateProvince="Terengganu">
<specimenCode id="1D33C76E2678FF86AAD7F8A4B83A8A0E" box="[453,611,1825,1850]" collectionCode="UKMHC" pageId="5" pageNumber="80">UKMHC 700</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86A963F8A4B8C78A0E" box="[625,670,1825,1850]" collectionCode="UKMHC" pageId="5" pageNumber="80">703</specimenCode>
, 
<specimenCode id="1D33C76E2678FF86A9B9F8A4B88C8A0E" box="[683,725,1825,1850]" collectionCode="UKMHC" pageId="5" pageNumber="80">704</specimenCode>
from 
<location id="484A39CE2678FF86A80DF8A4B9948A0E" LSID="urn:lsid:plazi:treatment:C53CDE032678FF88AB85FADEBFC98FC4:484A39CE2678FF86A80DF8A4B9948A0E" box="[799,973,1825,1850]" country="Malaysia" county="State" latitude="5.4333334" longLatPrecision="1305" longitude="102.96667" municipality="Hulu" name="Sungai Tembat" pageId="5" pageNumber="80" stateProvince="Terengganu">Sungai Tembat</location>
, 
<collectingMunicipality id="AD4EF56F2678FF86A8C8F8A4BE4B8A0E" box="[986,1042,1825,1850]" pageId="5" pageNumber="80">Hulu</collectingMunicipality>
<collectingRegion id="8F51A1F72678FF86AF09F8A7BEFB8A0E" box="[1051,1186,1826,1850]" country="Malaysia" name="Terengganu" pageId="5" pageNumber="80">Terengganu</collectingRegion>
, 
<collectingCounty id="A44B17992678FF86AFA2F8A4BEB18A0E" box="[1200,1256,1825,1850]" pageId="5" pageNumber="80">State</collectingCounty>
of 
<collectingRegion id="8F51A1F72678FF86AE03F8A7BFC18A0E" box="[1297,1432,1826,1850]" country="Malaysia" name="Terengganu" pageId="5" pageNumber="80">Terengganu</collectingRegion>
, 
<collectingCountry id="35822F852678FF86AB85F8C0BAA78A6A" box="[151,254,1861,1886]" name="Malaysia" pageId="5" pageNumber="80">Malaysia</collectingCountry>
(ca. 
<geoCoordinate id="28A109D22678FF86AA24F8C0BBD78A69" box="[310,398,1861,1886]" degrees="5" direction="north" minutes="26" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="5.4333334">5°26' N</geoCoordinate>
; 
<geoCoordinate id="28A109D22678FF86AA88F8C0B8558A69" box="[410,524,1861,1886]" degrees="102" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="102.96667">102°58' E</geoCoordinate>
, 
<elevation id="C6B888262678FF86A905F8C0B88A8A6A" box="[535,723,1861,1886]" metricMagnitude="2" metricUnit="m" metricValue="1.69" metricValueMax="1.7" metricValueMin="1.68" pageId="5" pageNumber="80" unit="m" value="169.0" valueMax="170.0" valueMin="168.0">
<quantity id="8A6DC2F02678FF86A905F8C0B8CE8A69" box="[535,663,1861,1886]" metricMagnitude="2" metricUnit="m" metricValue="1.69" metricValueMax="1.7" metricValueMin="1.68" pageId="5" pageNumber="80" unit="m" value="169.0" valueMax="170.0" valueMin="168.0">168–170 m</quantity>
a.s.l.
</elevation>
), collected in 2009 by 
<collectorName id="E0600AC32678FF86A8C4F8C0BEF48A6A" box="[982,1197,1861,1886]" pageId="5" pageNumber="80">Daicus M. Belabut</collectorName>
</materialsCitation>
; 
<materialsCitation id="FDFD65482678FF86AFABF8C3BF438AB6" ID-GBIF-Occurrence="3391168301" collectingDate="1993-01-03" collectionCode="KUHE" collectorName="Masafumi Matsui" country="Malaysia" elevation="600" location="Bukit Larut" pageId="5" pageNumber="80" specimenCode="KUHE 49514" specimenCount="1" specimenCount-larva="1" stateProvince="Terengganu">
<specimenCode id="1D33C76E2678FF86AFABF8C3BF0A8A6A" box="[1209,1363,1861,1886]" collectionCode="KUHE" country="Japan" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="80">KUHE 49514</specimenCode>
(
<specimenCount id="5B93A49C2678FF86AE70F8C0BA848AB6" pageId="5" pageNumber="80" type="larva">three larvae</specimenCount>
) from lower part of 
<location id="484A39CE2678FF86AAD2F8ECB8188AB5" LSID="urn:lsid:plazi:treatment:C53CDE032678FF88AB85FADEBFC98FC4:484A39CE2678FF86AAD2F8ECB8188AB5" box="[448,577,1897,1922]" country="Malaysia" name="Bukit Larut" pageId="5" pageNumber="80" stateProvince="Terengganu">Bukit Larut</location>
(
<elevation id="C6B888262678FF86A943F8ECB88C8AB6" box="[593,725,1897,1922]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="80" unit="m" value="600.0">
<quantity id="8A6DC2F02678FF86A943F8ECB8C38AB5" box="[593,666,1897,1922]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="80" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
), collected on 
<date id="392B49D52678FF86A869F8ECBE7F8AB6" box="[891,1062,1897,1922]" pageId="5" pageNumber="80" value="1993-01-03">
<collectingDate id="296FB03D2678FF86A869F8ECBE7F8AB6" box="[891,1062,1897,1922]" pageId="5" pageNumber="80" value="1993-01-03">3 January 1993</collectingDate>
</date>
by 
<collectorName id="E0600AC32678FF86AF42F8ECBF438AB6" box="[1104,1306,1897,1922]" pageId="5" pageNumber="80">Masafumi Matsui</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E2678FF86ABD5F808B90E8AFE" pageId="5" pageNumber="80" type="etymology">
<paragraph id="4D2A6F152678FF86ABD5F808B90E8AFE" blockId="5.[151,1436,1534,1994]" pageId="5" pageNumber="80">
<emphasis id="7FE1B3072678FF86ABD5F808BB178A92" bold="true" box="[199,334,1933,1958]" inLineHeading="true" pageId="5" pageNumber="80" reason="1">Etymology.</emphasis>
The specific epithet utara is a Malay word denoting north, alluding to the northerly-restricted distribution of the new species within the Peninsular 
<collectingCountry id="35822F852678FF86A9FBF834B90A8AFE" box="[745,851,1969,1994]" name="Malaysia" pageId="5" pageNumber="80">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="19EA3F9D267BFF85AB85FD47B95C8FC3" ID-DOI="http://doi.org/10.5281/zenodo.4949753" ID-Zenodo-Dep="4949753" httpUri="https://zenodo.org/record/4949753/files/figure.png" pageId="6" pageNumber="81" startId="6.[151,250,706,728]" targetBox="[158,1436,193,672]" targetPageId="6">
<paragraph id="4D2A6F15267BFF85AB85FD47B95C8FC3" blockId="6.[151,1437,706,759]" pageId="6" pageNumber="81">
<emphasis id="7FE1B307267BFF85AB85FD47BB578FEC" bold="true" box="[151,270,706,728]" pageId="6" pageNumber="81">FIGURE 3</emphasis>
. Dorsolateral views of male holotypes of 
<taxonomicName id="8A951496267BFF85A9D0FD46B9D88FEC" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[706,897,707,728]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B307267BFF85A9D0FD46B9D88FEC" box="[706,897,707,728]" italics="true" pageId="6" pageNumber="81">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B307267BFF85A89BFD47B9828FEC" bold="true" box="[905,987,706,728]" pageId="6" pageNumber="81">
<taxonomicNameLabel id="64D20E7C267BFF85A89BFD47B9828FEC" box="[905,987,706,728]" pageId="6" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 528) from Larut Hill (A) and 
<taxonomicName id="8A951496267BFF85AE95FD46BAB98FC3" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B307267BFF85AE95FD46BAB98FC3" italics="true" pageId="6" pageNumber="81">L. selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B307267BFF85ABF4FD64BB6E8FC3" bold="true" box="[230,311,737,759]" pageId="6" pageNumber="81">
<taxonomicNameLabel id="64D20E7C267BFF85ABF4FD64BB6E8FC3" box="[230,311,737,759]" pageId="6" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 529) from Genting Highlands (B).
</paragraph>
</caption>
<subSubSection id="058F3C9E267BFF85ABD5FCA1BB6B89FC" pageId="6" pageNumber="81" type="diagnosis">
<paragraph id="4D2A6F15267BFF85ABD5FCA1BB6B89FC" blockId="6.[151,1437,804,2017]" pageId="6" pageNumber="81">
<emphasis id="7FE1B307267BFF85ABD5FCA1BB188E09" bold="true" box="[199,321,804,829]" inLineHeading="true" pageId="6" pageNumber="81" reason="1">Diagnosis.</emphasis>
The new species is assigned to the genus 
<taxonomicName id="8A951496267BFF85A83BFCA0B9EF8E08" authorityName="Fitzinger" authorityYear="1843" box="[809,950,805,828]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis id="7FE1B307267BFF85A83BFCA0B9EF8E08" box="[809,950,805,828]" italics="true" pageId="6" pageNumber="81">Limnonectes</emphasis>
</taxonomicName>
only by molecular phylogenetic evidence, because morphological diagnostic characters of the genus previously proposed (e.g. 
<bibRefCitation id="290412E4267BFF85AF47FCCDBFCE8E54" author="Emerson, S. B. &amp; Berrigan, D." box="[1109,1431,839,864]" pageId="6" pageNumber="81" pagination="22 - 31" refId="ref12295" refString="Emerson, S. B. &amp; Berrigan, D. (1993) Systematics of Southeast Asian ranids: Multiple origins of voicelessness in the subgenus Limnonectes (Fitzinger). Herpetologica, 49, 22 - 31." type="journal article" year="1993">Emerson and Berrigan 1993</bibRefCitation>
; 
<bibRefCitation id="290412E4267BFF85AB85FCE9BB198EB1" author="Fei, L. &amp; Ye, C. - Y. &amp; Huang, Y. - Z. &amp; Jiang, J. - P. &amp; Xie, F." box="[151,320,876,901]" pageId="6" pageNumber="81" refId="ref12335" refString="Fei, L., Ye, C. - Y., Huang, Y. - Z., Jiang, J. - P. &amp; Xie, F. (Eds.) (2005) An Illustrated Key to Chinese Amphibians. Sichuan Publishing House of Science and Technology, Chongqing, 340 pp." type="book" year="2005">
Fei 
<emphasis id="7FE1B307267BFF85ABD1FCE8BAA48EB1" box="[195,253,876,901]" italics="true" pageId="6" pageNumber="81">et al.</emphasis>
2005
</bibRefCitation>
) are mostly osteological ones, which we did not examine. Moreover, osteological synapomorphic characters proposed by previous authors suffer from limited taxon sampling and require reexamination. Morphological assignment of the new species to 
<taxonomicName id="8A951496267BFF85A9D4FC30B90A8EF8" authorityName="Fitzinger" authorityYear="1843" box="[710,851,949,972]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis id="7FE1B307267BFF85A9D4FC30B90A8EF8" box="[710,851,949,972]" italics="true" pageId="6" pageNumber="81">Limnonectes</emphasis>
</taxonomicName>
was based only on the single characteristic usually found in the genus, enlarged “fangs” (tooth-like projections [odontoid processes]) on the lower jaw. A mediumsized species morphologically similar to 
<taxonomicName id="8A951496267BFF85A962FC78B8888921" box="[624,721,1020,1045]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B307267BFF85A962FC78B8888921" box="[624,721,1020,1045]" italics="true" pageId="6" pageNumber="81">L. kuhlii</emphasis>
</taxonomicName>
, with adult SVL 
<quantity id="8A6DC2F0267BFF85A88DFC79BE428927" box="[927,1051,1020,1045]" metricMagnitude="-2" metricUnit="m" metricValue="7.45" metricValueMax="7.9" metricValueMin="7.0" pageId="6" pageNumber="81" unit="mm" value="74.5" valueMax="79.0" valueMin="70.0">70–79 mm</quantity>
in males, 
<quantity id="8A6DC2F0267BFF85AF8AFC79BF4D8927" box="[1176,1300,1020,1045]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="7.9" metricValueMin="5.9" pageId="6" pageNumber="81" unit="mm" value="69.0" valueMax="79.0" valueMin="59.0">59–79 mm</quantity>
in females; males with relatively longer head than females; tibia dorsally densely covered by warts; toe webbing full, with very shallow excision between toes; first finger usually slightly longer than second, and nuptial pad present on first finger of males; morphologically differs from 
<taxonomicName id="8A951496267BFF85A9B9FBECB94589B4" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[683,796,1127,1152]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="81" phylum="Chordata" rank="species" species="selatan">
<emphasis id="7FE1B307267BFF85A9B9FBECB94589B4" box="[683,796,1127,1152]" italics="true" pageId="6" pageNumber="81">L. selatan</emphasis>
</taxonomicName>
, described below, in smoother dorsal skin, less densely arranged circum-cloacal warts, more confluent dorsal dark markings, and lack of large dark blotches on rear of thigh (
<figureCitation id="D5AE7390267BFF85ABF2FB2ABB7D89FC" box="[224,292,1199,1224]" captionStart="FIGURE 6" captionStartId="8.[151,250,1772,1794]" captionTargetBox="[151,1436,1412,1749]" captionTargetId="figure-421@8.[151,1436,1412,1749]" captionTargetPageId="8" captionText="FIGURE 6. Posterior views of left thigh of preserved male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B), showing circum-cloacal warts and dark blotches along dorso-ventral border. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4949761" httpUri="https://zenodo.org/record/4949761/files/figure.png" pageId="6" pageNumber="81">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E267BFF88ABD5FB51BED58DC3" lastPageId="11" lastPageNumber="86" pageId="6" pageNumber="81" type="description">
<paragraph id="4D2A6F15267BFF85ABD5FB51BED18B04" blockId="6.[151,1437,804,2017]" pageId="6" pageNumber="81">
<emphasis id="7FE1B307267BFF85ABD5FB51B8A689D9" bold="true" box="[199,767,1236,1261]" inLineHeading="true" pageId="6" pageNumber="81" reason="1">
Description of 
<typeStatus id="922ED1B7267BFF85AA65FB51BB8489D9" box="[375,477,1236,1261]" pageId="6" pageNumber="81" type="holotype">holotype</typeStatus>
(measurements in mm).
</emphasis>
SVL 79.1; habitus stocky (
<figureCitation id="D5AE7390267BFF85AF26FB51BEC189D9" box="[1076,1176,1236,1261]" captionStart="FIGURE 3" captionStartId="6.[151,250,706,728]" captionTargetBox="[158,1436,193,672]" captionTargetId="figure-607@6.[158,1443,193,672]" captionTargetPageId="6" captionText="FIGURE 3. Dorsolateral views of male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B)." figureDoi="http://doi.org/10.5281/zenodo.4949753" httpUri="https://zenodo.org/record/4949753/files/figure.png" pageId="6" pageNumber="81">Figs. 3A</figureCitation>
, 
<figureCitation id="D5AE7390267BFF85AFB7FB51BEBC89D9" box="[1189,1253,1236,1261]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[151,1436,337,1622]" captionTargetId="figure-277@7.[151,1436,337,1622]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A, B) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="6" pageNumber="81">4A,B</figureCitation>
); head enlarged, longer (HL 36.0, 45.5%SVL) than broad (HW 33.8, 42.7%SVL); snout obtusely pointed, obtuse in profile, projecting beyond lower jaw; eye length (EL 11.0,13.9%SVL) shorter than snout length (SL 13.0, 16.4%SVL); canthus rounded; lore sloping, concave; nostril dorsolateral on canthus, closer to tip of snout than to eye; internarial distance (IND 5.8, 7.3%SVL) equal to upper eyelid (UEW 5.8, 7.3%SVL) and wider than interorbital distance (IOD 5.4, 6.8%SVL); pineal spot visible; tympanic annulus barely visible through skin; vomerine teeth in oblique groups, between and behind line connecting anterior rims of choanae, groups separated from one another by onefifth length of one group and from choana by about one-fourth length of one group; lower jaw with a pair of toothlike projections (odontoid processes) near symphysis, more than twice depth of mandible at base of projections; tongue oval, deeply notched posteriorly, without papillae; vocal sac and vocal slits absent.
</paragraph>
<paragraph id="4D2A6F15267BFF85ABD5F9B9BFC48A64" blockId="6.[151,1437,804,2017]" pageId="6" pageNumber="81">
Forelimb heavy, relatively short (FLL 41.9, 53.0% SVL); fingers moderately slender; finger length formula: II &lt;I &lt;IV &lt;III (
<figureCitation id="D5AE7390267BFF85AA56F9DABBC68B4C" box="[324,415,1631,1656]" captionStart="FIGURE 5" captionStartId="8.[151,250,1279,1301]" captionTargetBox="[151,1436,661,1258]" captionTargetId="figure-412@8.[151,1436,661,1258]" captionTargetPageId="8" captionText="FIGURE 5. Ventral views of hand (A) and foot (B) of male holotype of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill, and preaxial view of hand (C) and ventral view of foot (D) of male holotype of L. selatan sp. nov. (UKMHC 529) from Genting Highlands. In anesthetized state, scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949757" httpUri="https://zenodo.org/record/4949757/files/figure.png" pageId="6" pageNumber="81">Fig. 5A</figureCitation>
), first finger slightly longer than second, length of first (11.0, 13.9%SVL, measured from distal edge of inner palmar tubercle) equal to length of eye; fourth finger much longer than second; tips of fingers bluntly rounded, forming small pads without circummarginal grooves; no webs between fingers; inner palmar tubercle moderate (4.1, 5.2% SVL), oval, not elevated; middle palmar tubercle oval, smaller than inner palmar tubercle, not contacting outer or inner palmar tubercles; outer palmar tubercle slightly smaller than middle tubercle; proximal subarticular tubercles round and elevated; distal subarticular tubercles low and indistinct; no supernumerary metacarpal tubercles; narrow, but distinct flaps of skin along both edges of second and third fingers.
</paragraph>
<paragraph id="4D2A6F15267BFF84ABD5F8D9BE488C29" blockId="6.[151,1437,804,2017]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="82" pageId="6" pageNumber="81">
Hindlimb heavy, relatively short (HLL 117.0,147.9% SVL) about 2.8 times length of forelimb; tibia short (TL 35.2, 44.5% SVL), heels not overlapping when limbs are held at right angles to body; tibiotarsal articulation of adpressed limb reaching to point posterior to eye; foot (FL 37.1, 46.9% SVL) slightly longer than tibia; toe length formula I &lt;II &lt;V &lt;III &lt;IV; tips of toes expanded into round, elevated pads lacking grooves (disk diameter of fourth toe, 4TDW 2.0, 2.5% SVL); all toes webbed to base of disks (
<figureCitation id="D5AE7390267AFF84A888FF12B9AB8D84" box="[922,1010,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1279,1301]" captionTargetBox="[151,1436,661,1258]" captionTargetId="figure-412@8.[151,1436,661,1258]" captionTargetPageId="8" captionText="FIGURE 5. Ventral views of hand (A) and foot (B) of male holotype of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill, and preaxial view of hand (C) and ventral view of foot (D) of male holotype of L. selatan sp. nov. (UKMHC 529) from Genting Highlands. In anesthetized state, scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949757" httpUri="https://zenodo.org/record/4949757/files/figure.png" pageId="7" pageNumber="82">Fig. 5B</figureCitation>
); webbing formula: I 0 – 0 II 0 – 0 III 0 – 0 IV 0 – 0 V; a distinct, movable flap of skin on outer edge of fifth toe and on inner edge of first toe; subarticular tubercles oval and distinct; an elongate inner metatarsal tubercle, length (IMTL 5.7, 7.2%SVL), about half length of first toe (1TOEL 11.2, 14.2% SVL); no outer metatarsal tubercle.
</paragraph>
<caption id="19EA3F9D267AFF84AB85F9E9BF178B94" ID-DOI="http://doi.org/10.5281/zenodo.4949755" ID-Zenodo-Dep="4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="7" pageNumber="82" startId="7.[151,250,1644,1666]" targetBox="[151,1436,337,1622]" targetPageId="7">
<paragraph id="4D2A6F15267AFF84AB85F9E9BF178B94" blockId="7.[151,1436,1644,1696]" pageId="7" pageNumber="82">
<emphasis id="7FE1B307267AFF84AB85F9E9BB578BB6" bold="true" box="[151,270,1644,1666]" pageId="7" pageNumber="82">FIGURE 4</emphasis>
. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of 
<taxonomicName id="8A951496267AFF84AF04F9E9BE8D8BB5" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[1046,1236,1644,1665]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="82" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B307267AFF84AF04F9E9BE8D8BB5" box="[1046,1236,1644,1665]" italics="true" pageId="7" pageNumber="82">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B307267AFF84AFCEF9E9BF748BB6" bold="true" box="[1244,1325,1644,1666]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="64D20E7C267AFF84AFCEF9E9BF748BB6" box="[1244,1325,1644,1666]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 528) from Larut Hill (A, B) and 
<taxonomicName id="8A951496267AFF84AAF2F90EB81D8B94" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[480,580,1674,1696]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="82" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B307267AFF84AAF2F90EB81D8B94" box="[480,580,1674,1696]" italics="true" pageId="7" pageNumber="82">L. selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B307267AFF84A958F90FB8C38B94" bold="true" box="[586,666,1674,1696]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="64D20E7C267AFF84A958F90FB8C38B94" box="[586,666,1674,1696]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="4D2A6F15267AFF8BABD5F948B85A8C74" blockId="7.[151,1436,1740,2018]" lastBlockId="8.[151,1437,151,608]" lastPageId="8" lastPageNumber="83" pageId="7" pageNumber="82">
Dorsum anteriorly very weakly rugose, without warts, posteriorly with very low ridges radiating from scattered, low warts (
<figureCitation id="D5AE7390267AFF84AA9EF974BBB08A3E" box="[396,489,1777,1802]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[151,1436,337,1622]" captionTargetId="figure-277@7.[151,1436,337,1622]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A, B) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="7" pageNumber="82">Fig. 4A</figureCitation>
); top of snout without tubercles, but eyelid covered with small warts and wrinkles with medial, longitudinal ridge formed by fused warts; weak transverse fold between posterior margins of eyes; a strong, supratympanic fold from eye to above axilla; posterior side of trunk scattered with tubercles; circum-cloacal warts small being sparsely and narrowly distributed anterior to cloaca, posteriorly more scattered but widely distributed, increasing size to end at dorso-ventral border of thigh (
<figureCitation id="D5AE7390267AFF84A8B3F804B9A78AAE" box="[929,1022,1921,1946]" captionStart="FIGURE 6" captionStartId="8.[151,250,1772,1794]" captionTargetBox="[151,1436,1412,1749]" captionTargetId="figure-421@8.[151,1436,1412,1749]" captionTargetPageId="8" captionText="FIGURE 6. Posterior views of left thigh of preserved male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B), showing circum-cloacal warts and dark blotches along dorso-ventral border. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4949761" httpUri="https://zenodo.org/record/4949761/files/figure.png" pageId="7" pageNumber="82">Fig. 6A</figureCitation>
); dorsal surface of thigh smooth on proximal two-thirds and scattered with small, low warts tipped with translucent spinules on distal one-third, continuing to tibia; tibia dorsally covered with numerous large and small round warts each tipped with large whitish cone surrounded by clusters of much smaller, whitish asperities; tarsus less densely covered by similar warts dorsolaterally; tarsus with a thick dermal ridge extending proximally from metatarsal tubercle; throat and chest weakly rugose, abdomen smooth (
<figureCitation id="D5AE73902675FF8BA94AFF5AB8E88DCC" box="[600,689,223,248]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[151,1436,337,1622]" captionTargetId="figure-277@7.[151,1436,337,1622]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A, B) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="8" pageNumber="83">Fig. 4B</figureCitation>
); a distinct creamy tinge, with minute asperities, forming a nuptial pad covering medial surface of first finger from its base to level of subarticular tubercle, sharply set off from remainder of skin on first finger.
</paragraph>
<paragraph id="4D2A6F152675FF8BABD5FEC9BF6C8F54" blockId="8.[151,1437,151,608]" pageId="8" pageNumber="83">
<emphasis id="7FE1B3072675FF8BABD5FEC9BB498C51" bold="true" box="[199,272,332,357]" inLineHeading="true" pageId="8" pageNumber="83" reason="1">Color.</emphasis>
In life, dorsum light brown with confluent dark brown markings (
<figureCitation id="D5AE73902675FF8BAF1FFEC9BE2A8C51" box="[1037,1139,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,706,728]" captionTargetBox="[158,1436,193,672]" captionTargetId="figure-607@6.[158,1443,193,672]" captionTargetPageId="6" captionText="FIGURE 3. Dorsolateral views of male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B)." figureDoi="http://doi.org/10.5281/zenodo.4949753" httpUri="https://zenodo.org/record/4949753/files/figure.png" pageId="8" pageNumber="83">Figs. 3A</figureCitation>
, 
<figureCitation id="D5AE73902675FF8BAF93FEC9BEF18C51" box="[1153,1192,332,357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[151,1436,337,1622]" captionTargetId="figure-277@7.[151,1436,337,1622]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A, B) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="8" pageNumber="83">4A</figureCitation>
); head with a narrow light band anterior to dark interorbital bar; blackish brown stripe on canthus rostralis; side of head pale brown with dark markings; oblique blackish brown temporal stripe on and along supratympanic fold beginning behind eye reaching inguinal area, and continuing on flank; lips barred with dark brown; dark brown stripe on anterior side of upper arm; limbs marked dorsally with dark-brown crossbars; dorsal and ventral border of posterior thigh scattered with small brown spots (
<figureCitation id="D5AE73902675FF8BAAA9FE7AB8408F2C" box="[443,537,511,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1772,1794]" captionTargetBox="[151,1436,1412,1749]" captionTargetId="figure-421@8.[151,1436,1412,1749]" captionTargetPageId="8" captionText="FIGURE 6. Posterior views of left thigh of preserved male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B), showing circum-cloacal warts and dark blotches along dorso-ventral border. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4949761" httpUri="https://zenodo.org/record/4949761/files/figure.png" pageId="8" pageNumber="83">Fig. 6A</figureCitation>
); throat weakly mottled with pale gray, spotted with dark brown posteriorly; abdomen immaculate cream (
<figureCitation id="D5AE73902675FF8BAAF0FDA1B8638F09" box="[482,570,548,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1644,1666]" captionTargetBox="[151,1436,337,1622]" captionTargetId="figure-277@7.[151,1436,337,1622]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal (A, C) and ventral (B, D) views of anesthetized male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A, B) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (C, D) Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949755" httpUri="https://zenodo.org/record/4949755/files/figure.png" pageId="8" pageNumber="83">Fig. 4B</figureCitation>
); ventral surfaces of hand and foot dark brown (
<figureCitation id="D5AE73902675FF8BAF46FDA1BE838F09" box="[1108,1242,548,573]" captionStart="FIGURE 5" captionStartId="8.[151,250,1279,1301]" captionTargetBox="[151,1436,661,1258]" captionTargetId="figure-412@8.[151,1436,661,1258]" captionTargetPageId="8" captionText="FIGURE 5. Ventral views of hand (A) and foot (B) of male holotype of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill, and preaxial view of hand (C) and ventral view of foot (D) of male holotype of L. selatan sp. nov. (UKMHC 529) from Genting Highlands. In anesthetized state, scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4949757" httpUri="https://zenodo.org/record/4949757/files/figure.png" pageId="8" pageNumber="83">Figs. 5 A,B</figureCitation>
). In preservative, the dorsal coloration has slightly faded, but otherwise no obvious change in color or pattern has occurred.
</paragraph>
<caption id="19EA3F9D2675FF8BAB85FB7AB8A68867" ID-DOI="http://doi.org/10.5281/zenodo.4949757" ID-Zenodo-Dep="4949757" httpUri="https://zenodo.org/record/4949757/files/figure.png" pageId="8" pageNumber="83" startId="8.[151,250,1279,1301]" targetBox="[151,1436,661,1258]" targetPageId="8">
<paragraph id="4D2A6F152675FF8BAB85FB7AB8A68867" blockId="8.[151,1436,1279,1363]" pageId="8" pageNumber="83">
<emphasis id="7FE1B3072675FF8BAB85FB7ABB558821" bold="true" box="[151,268,1279,1301]" pageId="8" pageNumber="83">FIGURE 5</emphasis>
. Ventral views of hand (A) and foot (B) of male holotype of 
<taxonomicName id="8A9514962675FF8BA861FA85BE698821" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[883,1072,1280,1301]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="83" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072675FF8BA861FA85BE698821" box="[883,1072,1280,1301]" italics="true" pageId="8" pageNumber="83">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072675FF8BAF27FB7ABEDF8821" bold="true" box="[1077,1158,1279,1301]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="64D20E7C2675FF8BAF27FB7ABEDF8821" box="[1077,1158,1279,1301]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 528) from Larut Hill, and preaxial view of hand (C) and ventral view of foot (D) of male holotype of 
<taxonomicName id="8A9514962675FF8BAF10FA9ABE3F8800" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[1026,1126,1310,1332]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="83" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B3072675FF8BAF10FA9ABE3F8800" box="[1026,1126,1310,1332]" italics="true" pageId="8" pageNumber="83">L. selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072675FF8BAF7CFA9BBEE68800" bold="true" box="[1134,1215,1310,1332]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="64D20E7C2675FF8BAF7CFA9BBEE68800" box="[1134,1215,1310,1332]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 529) from Genting Highlands. In anesthetized state, scale bar = 10 mm.
</paragraph>
</caption>
<caption id="19EA3F9D2675FF8BAB85F969B8698A0B" ID-DOI="http://doi.org/10.5281/zenodo.4949761" ID-Zenodo-Dep="4949761" httpUri="https://zenodo.org/record/4949761/files/figure.png" pageId="8" pageNumber="83" startId="8.[151,250,1772,1794]" targetBox="[151,1436,1412,1749]" targetPageId="8">
<paragraph id="4D2A6F152675FF8BAB85F969B8698A0B" blockId="8.[151,1436,1772,1855]" pageId="8" pageNumber="83">
<emphasis id="7FE1B3072675FF8BAB85F969BB558A36" bold="true" box="[151,268,1772,1794]" pageId="8" pageNumber="83">FIGURE 6</emphasis>
. Posterior views of left thigh of preserved male holotypes of 
<taxonomicName id="8A9514962675FF8BA866F969BE688A35" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[884,1073,1772,1793]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="83" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072675FF8BA866F969BE688A35" box="[884,1073,1772,1793]" italics="true" pageId="8" pageNumber="83">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072675FF8BAF25F969BEDE8A36" bold="true" box="[1079,1159,1772,1794]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="64D20E7C2675FF8BAF25F969BEDE8A36" box="[1079,1159,1772,1794]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 528) from Larut Hill (A) and 
<taxonomicName id="8A9514962675FF8BAA09F88EBB268A14" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[283,383,1802,1824]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="83" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B3072675FF8BAA09F88EBB268A14" box="[283,383,1802,1824]" italics="true" pageId="8" pageNumber="83">L. selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072675FF8BAA94F88FBB8E8A14" bold="true" box="[390,471,1802,1824]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="64D20E7C2675FF8BAA94F88FBB8E8A14" box="[390,471,1802,1824]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UKMHC 529) from Genting Highlands (B), showing circum-cloacal warts and dark blotches along dorso-ventral border. Not to scale.
</paragraph>
</caption>
<paragraph id="4D2A6F152675FF8AABD5F8E3BE678C51" blockId="8.[151,1437,1894,2027]" lastBlockId="9.[151,1436,151,357]" lastPageId="9" lastPageNumber="84" pageId="8" pageNumber="83">
<emphasis id="7FE1B3072675FF8BABD5F8E3BB188A4B" bold="true" box="[199,321,1894,1919]" inLineHeading="true" pageId="8" pageNumber="83" reason="1">Variation.</emphasis>
Individual variation in size and body proportions is given in 
<tableCitation id="00175AAE2675FF8BA8E5F8E3BE158A4B" box="[1015,1100,1894,1919]" captionStart="TABLE 3" captionStartId="9.[151,235,405,427]" captionTargetPageId="9" captionText="TABLE 3. Measurements in adults of two new Limnonectes species from Peninsular Malaysia. SVL (mean±1SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations." httpUri="http://table.plazi.org/id/19EA3F9D2674FF8AAB85FE10BF028CFB" pageId="8" pageNumber="83" tableUuid="19EA3F9D2674FF8AAB85FE10BF028CFB">Table 3</tableCitation>
. Males have relatively longer head than females. Well-developed warts are invariably present, but there are some variations in qualitative traits. The postorbital light-colored bar was absent (50% of adults examined) or only vaguely traced (42%), and a few (8%) had a thin bar. The majority (83%) had a wide temporal stripe, and at least narrow temporal stripe was present (17%). Distinct spots were found widely distributed on dorsum in many (83%) individuals, and were found on at least part of the dorsum in others (17%). Many (75%) had weak spots or dots on the chin, but some had dusty marking (17%) or lacked chin spots (8%). The dark stripe on the upper arm was clear and continuous (38%) or weak or disjunct (46%), but was absent in some individuals (15%). Dorsal warts on the body were present usually (75%) only partially, and some (25 %) had very few warts. The first finger was usually longer than the second (69%), but was subequal (15%), or shorter than the first (15%) in some individuals.
</paragraph>
<caption id="19EA3F9D2674FF8AAB85FE10BF028CFB" ID-Table-UUID="19EA3F9D2674FF8AAB85FE10BF028CFB" httpUri="http://table.plazi.org/id/19EA3F9D2674FF8AAB85FE10BF028CFB" pageId="9" pageNumber="84" startId="9.[151,235,405,427]" targetBox="[159,1320,491,2023]" targetIsTable="true" targetPageId="9">
<paragraph id="4D2A6F152674FF8AAB85FE10BF028CFB" blockId="9.[151,1435,405,463]" pageId="9" pageNumber="84">
<emphasis id="7FE1B3072674FF8AAB85FE10BB5D8C9F" bold="true" box="[151,260,405,427]" pageId="9" pageNumber="84">TABLE 3.</emphasis>
Measurements in adults of two new 
<taxonomicName id="8A9514962674FF8AA96FFE13B9148C9F" box="[637,845,405,427]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7FE1B3072674FF8AA96FFE13B8A48C9F" box="[637,765,406,427]" italics="true" pageId="9" pageNumber="84">Limnonectes</emphasis>
species
</taxonomicName>
from Peninsular Malaysia. SVL (mean±1SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations.
</paragraph>
</caption>
<paragraph id="4D2A6F152674FF8AAA43FE69BEAC8AD3" pageId="9" pageNumber="84">
<table id="3F959DB52674007CAB8DFE6EBF718AD3" box="[159,1320,491,2023]" gridcols="5" gridrows="38" pageId="9" pageNumber="84">
<tr id="F3A56D572674007CAB8DFE6EBF718F35" box="[159,1320,491,513]" gridrow="0" pageId="9" pageNumber="84" rowspan-0="1" rowspan-2="1" rowspan-4="1">
<th id="B074042B2674007CAA43FE6EBBAC8F35" box="[337,501,491,513]" gridcol="1" gridrow="0" pageId="9" pageNumber="84">
<taxonomicName id="8A9514962674FF8AAA43FE69BBC68F35" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[337,415,492,513]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="84" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072674FF8AAA43FE69BBC68F35" box="[337,415,492,513]" italics="true" pageId="9" pageNumber="84">L. utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072674FF8AAAB7FE6EBBAC8F35" bold="true" box="[421,501,491,513]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="64D20E7C2674FF8AAAB7FE6EBBAC8F35" box="[421,501,491,513]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="B074042B2674007CA89FFE6EBE1B8F35" box="[909,1090,491,513]" gridcol="3" gridrow="0" pageId="9" pageNumber="84">
<taxonomicName id="8A9514962674FF8AA89FFE69B9B28F35" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[909,1003,492,513]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="84" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B3072674FF8AA89FFE69B9B28F35" box="[909,1003,492,513]" italics="true" pageId="9" pageNumber="84">L. selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072674FF8AA8E0FE6EBE1B8F35" bold="true" box="[1010,1090,491,513]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="64D20E7C2674FF8AA8E0FE6EBE1B8F35" box="[1010,1090,491,513]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="F3A56D572674007CAB8DFD91BF718F1E" box="[159,1320,532,554]" gridrow="1" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FD91BBAC8F1E" box="[337,501,532,554]" gridcol="1" gridrow="1" pageId="9" pageNumber="84">4 males</td>
<td id="B074042B2674007CA94CFD91B8B48F1E" box="[606,749,532,554]" gridcol="2" gridrow="1" pageId="9" pageNumber="84">6 females</td>
<td id="B074042B2674007CA89FFD91BE1B8F1E" box="[909,1090,532,554]" gridcol="3" gridrow="1" pageId="9" pageNumber="84">3 males</td>
<td id="B074042B2674007CAF8BFD91BF718F1E" box="[1177,1320,532,554]" gridcol="4" gridrow="1" pageId="9" pageNumber="84">8 females</td>
</tr>
<tr id="F3A56D572674007CAB8DFDB8BF718F67" box="[159,1320,573,595]" gridrow="2" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFDB8BAA58F67" box="[159,252,573,595]" gridcol="0" gridrow="2" pageId="9" pageNumber="84">SVL</th>
<td id="B074042B2674007CAA43FDB8BBAC8F67" box="[337,501,573,595]" gridcol="1" gridrow="2" pageId="9" pageNumber="84">74.2±4.6</td>
<td id="B074042B2674007CA94CFDB8B8B48F67" box="[606,749,573,595]" gridcol="2" gridrow="2" pageId="9" pageNumber="84">69.5±6.8</td>
<td id="B074042B2674007CA89FFDB8BE1B8F67" box="[909,1090,573,595]" gridcol="3" gridrow="2" pageId="9" pageNumber="84">53.5±1.2</td>
<td id="B074042B2674007CAF8BFDB8BF718F67" box="[1177,1320,573,595]" gridcol="4" gridrow="2" pageId="9" pageNumber="84">70.0±3.8</td>
</tr>
<tr id="F3A56D572674007CAB8DFDE0BF718F4F" box="[159,1320,613,635]" gridrow="3" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FDE0BBAC8F4F" box="[337,501,613,635]" gridcol="1" gridrow="3" pageId="9" pageNumber="84">(70.0–79.1)</td>
<td id="B074042B2674007CA94CFDE0B8B48F4F" box="[606,749,613,635]" gridcol="2" gridrow="3" pageId="9" pageNumber="84">(59.2–78.7)</td>
<td id="B074042B2674007CA89FFDE0BE1B8F4F" box="[909,1090,613,635]" gridcol="3" gridrow="3" pageId="9" pageNumber="84">(52.2–54.4)</td>
<td id="B074042B2674007CAF8BFDE0BF718F4F" box="[1177,1320,613,635]" gridcol="4" gridrow="3" pageId="9" pageNumber="84">(61.6–73.0)</td>
</tr>
<tr id="F3A56D572674007CAB8DFD0BBF718F90" box="[159,1320,654,676]" gridrow="4" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFD0BBAA58F90" box="[159,252,654,676]" gridcol="0" gridrow="4" pageId="9" pageNumber="84">RHL</th>
<td id="B074042B2674007CAA43FD0BBBAC8F90" box="[337,501,654,676]" gridcol="1" gridrow="4" pageId="9" pageNumber="84">46.0</td>
<td id="B074042B2674007CA94CFD0BB8B48F90" box="[606,749,654,676]" gridcol="2" gridrow="4" pageId="9" pageNumber="84">42.4</td>
<td id="B074042B2674007CA89FFD0BBE1B8F90" box="[909,1090,654,676]" gridcol="3" gridrow="4" pageId="9" pageNumber="84">44.5</td>
<td id="B074042B2674007CAF8BFD0BBF718F90" box="[1177,1320,654,676]" gridcol="4" gridrow="4" pageId="9" pageNumber="84">42.4</td>
</tr>
<tr id="F3A56D572674007CAB8DFD32BF718FF9" box="[159,1320,695,717]" gridrow="5" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FD32BBAC8FF9" box="[337,501,695,717]" gridcol="1" gridrow="5" pageId="9" pageNumber="84">(44.5–47.6)</td>
<td id="B074042B2674007CA94CFD32B8B48FF9" box="[606,749,695,717]" gridcol="2" gridrow="5" pageId="9" pageNumber="84">(40.5–44.0)</td>
<td id="B074042B2674007CA89FFD32BE1B8FF9" box="[909,1090,695,717]" gridcol="3" gridrow="5" pageId="9" pageNumber="84">(42.5–46.0)</td>
<td id="B074042B2674007CAF8BFD32BF718FF9" box="[1177,1320,695,717]" gridcol="4" gridrow="5" pageId="9" pageNumber="84">(39.8–44.3)</td>
</tr>
<tr id="F3A56D572674007CAB8DFD65BF718FC2" box="[159,1320,736,758]" gridrow="6" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFD65BAA58FC2" box="[159,252,736,758]" gridcol="0" gridrow="6" pageId="9" pageNumber="84">RSL</th>
<td id="B074042B2674007CAA43FD65BBAC8FC2" box="[337,501,736,758]" gridcol="1" gridrow="6" pageId="9" pageNumber="84">16.6</td>
<td id="B074042B2674007CA94CFD65B8B48FC2" box="[606,749,736,758]" gridcol="2" gridrow="6" pageId="9" pageNumber="84">15.5</td>
<td id="B074042B2674007CA89FFD65BE1B8FC2" box="[909,1090,736,758]" gridcol="3" gridrow="6" pageId="9" pageNumber="84">16.0</td>
<td id="B074042B2674007CAF8BFD65BF718FC2" box="[1177,1320,736,758]" gridcol="4" gridrow="6" pageId="9" pageNumber="84">15.3</td>
</tr>
<tr id="F3A56D572674007CAB8DFC8CBF718E2B" box="[159,1320,777,799]" gridrow="7" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FC8CBBAC8E2B" box="[337,501,777,799]" gridcol="1" gridrow="7" pageId="9" pageNumber="84">(15.9–17.1)</td>
<td id="B074042B2674007CA94CFC8CB8B48E2B" box="[606,749,777,799]" gridcol="2" gridrow="7" pageId="9" pageNumber="84">(14.9–16.7)</td>
<td id="B074042B2674007CA89FFC8CBE1B8E2B" box="[909,1090,777,799]" gridcol="3" gridrow="7" pageId="9" pageNumber="84">(15.7–16.3)</td>
<td id="B074042B2674007CAF8BFC8CBF718E2B" box="[1177,1320,777,799]" gridcol="4" gridrow="7" pageId="9" pageNumber="84">(14.4–15.9)</td>
</tr>
<tr id="F3A56D572674007CAB8DFCB7BF718E7C" box="[159,1320,818,840]" gridrow="8" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFCB7BAA58E7C" box="[159,252,818,840]" gridcol="0" gridrow="8" pageId="9" pageNumber="84">REL</th>
<td id="B074042B2674007CAA43FCB7BBAC8E7C" box="[337,501,818,840]" gridcol="1" gridrow="8" pageId="9" pageNumber="84">14.4</td>
<td id="B074042B2674007CA94CFCB7B8B48E7C" box="[606,749,818,840]" gridcol="2" gridrow="8" pageId="9" pageNumber="84">15.3</td>
<td id="B074042B2674007CA89FFCB7BE1B8E7C" box="[909,1090,818,840]" gridcol="3" gridrow="8" pageId="9" pageNumber="84">14.5</td>
<td id="B074042B2674007CAF8BFCB7BF718E7C" box="[1177,1320,818,840]" gridcol="4" gridrow="8" pageId="9" pageNumber="84">13.6</td>
</tr>
<tr id="F3A56D572674007CAB8DFCDFBF718E44" box="[159,1320,858,880]" gridrow="9" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FCDFBBAC8E44" box="[337,501,858,880]" gridcol="1" gridrow="9" pageId="9" pageNumber="84">(13.7–14.7)</td>
<td id="B074042B2674007CA94CFCDFB8B48E44" box="[606,749,858,880]" gridcol="2" gridrow="9" pageId="9" pageNumber="84">(14.3–16.1)</td>
<td id="B074042B2674007CA89FFCDFBE1B8E44" box="[909,1090,858,880]" gridcol="3" gridrow="9" pageId="9" pageNumber="84">(14.0–14.8)</td>
<td id="B074042B2674007CAF8BFCDFBF718E44" box="[1177,1320,858,880]" gridcol="4" gridrow="9" pageId="9" pageNumber="84">(12.7–14.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DFC06BF718EAD" box="[159,1320,899,921]" gridrow="10" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFC06BAA58EAD" box="[159,252,899,921]" gridcol="0" gridrow="10" pageId="9" pageNumber="84">RED</th>
<td id="B074042B2674007CAA43FC06BBAC8EAD" box="[337,501,899,921]" gridcol="1" gridrow="10" pageId="9" pageNumber="84">11.3</td>
<td id="B074042B2674007CA94CFC06B8B48EAD" box="[606,749,899,921]" gridcol="2" gridrow="10" pageId="9" pageNumber="84">11.8</td>
<td id="B074042B2674007CA89FFC06BE1B8EAD" box="[909,1090,899,921]" gridcol="3" gridrow="10" pageId="9" pageNumber="84">11.1</td>
<td id="B074042B2674007CAF8BFC06BF718EAD" box="[1177,1320,899,921]" gridcol="4" gridrow="10" pageId="9" pageNumber="84">10.4</td>
</tr>
<tr id="F3A56D572674007CAB8DFC29BF718EF6" box="[159,1320,940,962]" gridrow="11" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FC29BBAC8EF6" box="[337,501,940,962]" gridcol="1" gridrow="11" pageId="9" pageNumber="84">(10.4–11.5)</td>
<td id="B074042B2674007CA94CFC29B8B48EF6" box="[606,749,940,962]" gridcol="2" gridrow="11" pageId="9" pageNumber="84">(10.9–13.2)</td>
<td id="B074042B2674007CA89FFC29BE1B8EF6" box="[909,1090,940,962]" gridcol="3" gridrow="11" pageId="9" pageNumber="84">(10.8–12.1)</td>
<td id="B074042B2674007CAF8BFC29BF718EF6" box="[1177,1320,940,962]" gridcol="4" gridrow="11" pageId="9" pageNumber="84">(9.9–11.0)</td>
</tr>
<tr id="F3A56D572674007CAB8DFC50BF718EDF" box="[159,1320,981,1003]" gridrow="12" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFC50BAA58EDF" box="[159,252,981,1003]" gridcol="0" gridrow="12" pageId="9" pageNumber="84">RHW</th>
<td id="B074042B2674007CAA43FC50BBAC8EDF" box="[337,501,981,1003]" gridcol="1" gridrow="12" pageId="9" pageNumber="84">42.8</td>
<td id="B074042B2674007CA94CFC50B8B48EDF" box="[606,749,981,1003]" gridcol="2" gridrow="12" pageId="9" pageNumber="84">40.3</td>
<td id="B074042B2674007CA89FFC50BE1B8EDF" box="[909,1090,981,1003]" gridcol="3" gridrow="12" pageId="9" pageNumber="84">42.9</td>
<td id="B074042B2674007CAF8BFC50BF718EDF" box="[1177,1320,981,1003]" gridcol="4" gridrow="12" pageId="9" pageNumber="84">41.7</td>
</tr>
<tr id="F3A56D572674007CAB8DFC78BF718927" box="[159,1320,1021,1043]" gridrow="13" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FC78BBAC8927" box="[337,501,1021,1043]" gridcol="1" gridrow="13" pageId="9" pageNumber="84">(41.9–43.4)</td>
<td id="B074042B2674007CA94CFC78B8B48927" box="[606,749,1021,1043]" gridcol="2" gridrow="13" pageId="9" pageNumber="84">(38.9–42.3)</td>
<td id="B074042B2674007CA89FFC78BE1B8927" box="[909,1090,1021,1043]" gridcol="3" gridrow="13" pageId="9" pageNumber="84">(41.7–42.9)</td>
<td id="B074042B2674007CAF8BFC78BF718927" box="[1177,1320,1021,1043]" gridcol="4" gridrow="13" pageId="9" pageNumber="84">(39.8–44.0)</td>
</tr>
<tr id="F3A56D572674007CAB8DFBA3BF718908" box="[159,1320,1062,1084]" gridrow="14" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFBA3BAA58908" box="[159,252,1062,1084]" gridcol="0" gridrow="14" pageId="9" pageNumber="84">RIND</th>
<td id="B074042B2674007CAA43FBA3BBAC8908" box="[337,501,1062,1084]" gridcol="1" gridrow="14" pageId="9" pageNumber="84">7.7</td>
<td id="B074042B2674007CA94CFBA3B8B48908" box="[606,749,1062,1084]" gridcol="2" gridrow="14" pageId="9" pageNumber="84">7.5</td>
<td id="B074042B2674007CA89FFBA3BE1B8908" box="[909,1090,1062,1084]" gridcol="3" gridrow="14" pageId="9" pageNumber="84">8.6</td>
<td id="B074042B2674007CAF8BFBA3BF718908" box="[1177,1320,1062,1084]" gridcol="4" gridrow="14" pageId="9" pageNumber="84">8.0</td>
</tr>
<tr id="F3A56D572674007CAB8DFBCABF718951" box="[159,1320,1103,1125]" gridrow="15" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FBCABBAC8951" box="[337,501,1103,1125]" gridcol="1" gridrow="15" pageId="9" pageNumber="84">(7.3–8.3)</td>
<td id="B074042B2674007CA94CFBCAB8B48951" box="[606,749,1103,1125]" gridcol="2" gridrow="15" pageId="9" pageNumber="84">(7.3–8.3)</td>
<td id="B074042B2674007CA89FFBCABE1B8951" box="[909,1090,1103,1125]" gridcol="3" gridrow="15" pageId="9" pageNumber="84">(8.3–9.5)</td>
<td id="B074042B2674007CAF8BFBCABF718951" box="[1177,1320,1103,1125]" gridcol="4" gridrow="15" pageId="9" pageNumber="84">(7.7–8.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DFBFDBF7189BA" box="[159,1320,1144,1166]" gridrow="16" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFBFDBAA589BA" box="[159,252,1144,1166]" gridcol="0" gridrow="16" pageId="9" pageNumber="84">RIOD</th>
<td id="B074042B2674007CAA43FBFDBBAC89BA" box="[337,501,1144,1166]" gridcol="1" gridrow="16" pageId="9" pageNumber="84">7.5</td>
<td id="B074042B2674007CA94CFBFDB8B489BA" box="[606,749,1144,1166]" gridcol="2" gridrow="16" pageId="9" pageNumber="84">7.4</td>
<td id="B074042B2674007CA89FFBFDBE1B89BA" box="[909,1090,1144,1166]" gridcol="3" gridrow="16" pageId="9" pageNumber="84">7.0</td>
<td id="B074042B2674007CAF8BFBFDBF7189BA" box="[1177,1320,1144,1166]" gridcol="4" gridrow="16" pageId="9" pageNumber="84">7.5</td>
</tr>
<tr id="F3A56D572674007CAB8DFB24BF718983" box="[159,1320,1185,1207]" gridrow="17" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FB24BBAC8983" box="[337,501,1185,1207]" gridcol="1" gridrow="17" pageId="9" pageNumber="84">(6.8–8.1)</td>
<td id="B074042B2674007CA94CFB24B8B48983" box="[606,749,1185,1207]" gridcol="2" gridrow="17" pageId="9" pageNumber="84">(6.4–7.7)</td>
<td id="B074042B2674007CA89FFB24BE1B8983" box="[909,1090,1185,1207]" gridcol="3" gridrow="17" pageId="9" pageNumber="84">(6.9–9.8)</td>
<td id="B074042B2674007CAF8BFB24BF718983" box="[1177,1320,1185,1207]" gridcol="4" gridrow="17" pageId="9" pageNumber="84">(6.3–8.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DFB4FBF7189D4" box="[159,1320,1226,1248]" gridrow="18" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFB4FBAA589D4" box="[159,252,1226,1248]" gridcol="0" gridrow="18" pageId="9" pageNumber="84">RUEW</th>
<td id="B074042B2674007CAA43FB4FBBAC89D4" box="[337,501,1226,1248]" gridcol="1" gridrow="18" pageId="9" pageNumber="84">7.7</td>
<td id="B074042B2674007CA94CFB4FB8B489D4" box="[606,749,1226,1248]" gridcol="2" gridrow="18" pageId="9" pageNumber="84">7.7</td>
<td id="B074042B2674007CA89FFB4FBE1B89D4" box="[909,1090,1226,1248]" gridcol="3" gridrow="18" pageId="9" pageNumber="84">8.1</td>
<td id="B074042B2674007CAF8BFB4FBF7189D4" box="[1177,1320,1226,1248]" gridcol="4" gridrow="18" pageId="9" pageNumber="84">7.2</td>
</tr>
<tr id="F3A56D572674007CAB8DFB77BF71883C" box="[159,1320,1266,1288]" gridrow="19" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FB77BBAC883C" box="[337,501,1266,1288]" gridcol="1" gridrow="19" pageId="9" pageNumber="84">(6.7–8.7)</td>
<td id="B074042B2674007CA94CFB77B8B4883C" box="[606,749,1266,1288]" gridcol="2" gridrow="19" pageId="9" pageNumber="84">(6.7–7.9)</td>
<td id="B074042B2674007CA89FFB77BE1B883C" box="[909,1090,1266,1288]" gridcol="3" gridrow="19" pageId="9" pageNumber="84">(8.0–8.2)</td>
<td id="B074042B2674007CAF8BFB77BF71883C" box="[1177,1320,1266,1288]" gridcol="4" gridrow="19" pageId="9" pageNumber="84">(6.5–7.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DFA9EBF718805" box="[159,1320,1307,1329]" gridrow="20" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFA9EBAA58805" box="[159,252,1307,1329]" gridcol="0" gridrow="20" pageId="9" pageNumber="84">RLAL</th>
<td id="B074042B2674007CAA43FA9EBBAC8805" box="[337,501,1307,1329]" gridcol="1" gridrow="20" pageId="9" pageNumber="84">43.9</td>
<td id="B074042B2674007CA94CFA9EB8B48805" box="[606,749,1307,1329]" gridcol="2" gridrow="20" pageId="9" pageNumber="84">42.9</td>
<td id="B074042B2674007CA89FFA9EBE1B8805" box="[909,1090,1307,1329]" gridcol="3" gridrow="20" pageId="9" pageNumber="84">44.1</td>
<td id="B074042B2674007CAF8BFA9EBF718805" box="[1177,1320,1307,1329]" gridcol="4" gridrow="20" pageId="9" pageNumber="84">44.1</td>
</tr>
<tr id="F3A56D572674007CAB8DFAC1BF71886E" box="[159,1320,1348,1370]" gridrow="21" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FAC1BBAC886E" box="[337,501,1348,1370]" gridcol="1" gridrow="21" pageId="9" pageNumber="84">(43.5-44.4)</td>
<td id="B074042B2674007CA94CFAC1B8B4886E" box="[606,749,1348,1370]" gridcol="2" gridrow="21" pageId="9" pageNumber="84">(42.1–44.0)</td>
<td id="B074042B2674007CA89FFAC1BE1B886E" box="[909,1090,1348,1370]" gridcol="3" gridrow="21" pageId="9" pageNumber="84">(43.4–44.3)</td>
<td id="B074042B2674007CAF8BFAC1BF71886E" box="[1177,1320,1348,1370]" gridcol="4" gridrow="21" pageId="9" pageNumber="84">(42.2–45.1)</td>
</tr>
<tr id="F3A56D572674007CAB8DFAE8BF7188B7" box="[159,1320,1389,1411]" gridrow="22" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFAE8BAA588B7" box="[159,252,1389,1411]" gridcol="0" gridrow="22" pageId="9" pageNumber="84">RFLL</th>
<td id="B074042B2674007CAA43FAE8BBAC88B7" box="[337,501,1389,1411]" gridcol="1" gridrow="22" pageId="9" pageNumber="84">55.0</td>
<td id="B074042B2674007CA94CFAE8B8B488B7" box="[606,749,1389,1411]" gridcol="2" gridrow="22" pageId="9" pageNumber="84">54.1</td>
<td id="B074042B2674007CA89FFAE8BE1B88B7" box="[909,1090,1389,1411]" gridcol="3" gridrow="22" pageId="9" pageNumber="84">52.0</td>
<td id="B074042B2674007CAF8BFAE8BF7188B7" box="[1177,1320,1389,1411]" gridcol="4" gridrow="22" pageId="9" pageNumber="84">53.6</td>
</tr>
<tr id="F3A56D572674007CAB8DFA10BF71889F" box="[159,1320,1429,1451]" gridrow="23" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FA10BBAC889F" box="[337,501,1429,1451]" gridcol="1" gridrow="23" pageId="9" pageNumber="84">(53.0–56.0)</td>
<td id="B074042B2674007CA94CFA10B8B4889F" box="[606,749,1429,1451]" gridcol="2" gridrow="23" pageId="9" pageNumber="84">(51.1–56.1)</td>
<td id="B074042B2674007CA89FFA10BE1B889F" box="[909,1090,1429,1451]" gridcol="3" gridrow="23" pageId="9" pageNumber="84">(51.7–52.9)</td>
<td id="B074042B2674007CAF8BFA10BF71889F" box="[1177,1320,1429,1451]" gridcol="4" gridrow="23" pageId="9" pageNumber="84">(51.0–56.1)</td>
</tr>
<tr id="F3A56D572674007CAB8DFA3BBF7188E0" box="[159,1320,1470,1492]" gridrow="24" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DFA3BBAA588E0" box="[159,252,1470,1492]" gridcol="0" gridrow="24" pageId="9" pageNumber="84">RHLL</th>
<td id="B074042B2674007CAA43FA3BBBAC88E0" box="[337,501,1470,1492]" gridcol="1" gridrow="24" pageId="9" pageNumber="84">148.3</td>
<td id="B074042B2674007CA94CFA3BB8B488E0" box="[606,749,1470,1492]" gridcol="2" gridrow="24" pageId="9" pageNumber="84">146.5</td>
<td id="B074042B2674007CA89FFA3BBE1B88E0" box="[909,1090,1470,1492]" gridcol="3" gridrow="24" pageId="9" pageNumber="84">151.1</td>
<td id="B074042B2674007CAF8BFA3BBF7188E0" box="[1177,1320,1470,1492]" gridcol="4" gridrow="24" pageId="9" pageNumber="84">150.4</td>
</tr>
<tr id="F3A56D572674007CAB8DFA62BF7188C9" box="[159,1320,1511,1533]" gridrow="25" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43FA62BBAC88C9" box="[337,501,1511,1533]" gridcol="1" gridrow="25" pageId="9" pageNumber="84">(144.9–149.4)</td>
<td id="B074042B2674007CA94CFA62B8B488C9" box="[606,749,1511,1533]" gridcol="2" gridrow="25" pageId="9" pageNumber="84">(143.6–150.3)</td>
<td id="B074042B2674007CA89FFA62BE1B88C9" box="[909,1090,1511,1533]" gridcol="3" gridrow="25" pageId="9" pageNumber="84">(149.4–152.2)</td>
<td id="B074042B2674007CAF8BFA62BF7188C9" box="[1177,1320,1511,1533]" gridcol="4" gridrow="25" pageId="9" pageNumber="84">(146.2–157.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DF995BF718B12" box="[159,1320,1552,1574]" gridrow="26" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF995BAA58B12" box="[159,252,1552,1574]" gridcol="0" gridrow="26" pageId="9" pageNumber="84">RTHGH</th>
<td id="B074042B2674007CAA43F995BBAC8B12" box="[337,501,1552,1574]" gridcol="1" gridrow="26" pageId="9" pageNumber="84">48.8</td>
<td id="B074042B2674007CA94CF995B8B48B12" box="[606,749,1552,1574]" gridcol="2" gridrow="26" pageId="9" pageNumber="84">49.1</td>
<td id="B074042B2674007CA89FF995BE1B8B12" box="[909,1090,1552,1574]" gridcol="3" gridrow="26" pageId="9" pageNumber="84">51.0</td>
<td id="B074042B2674007CAF8BF995BF718B12" box="[1177,1320,1552,1574]" gridcol="4" gridrow="26" pageId="9" pageNumber="84">48.9</td>
</tr>
<tr id="F3A56D572674007CAB8DF9BCBF718B7B" box="[159,1320,1593,1615]" gridrow="27" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F9BCBBAC8B7B" box="[337,501,1593,1615]" gridcol="1" gridrow="27" pageId="9" pageNumber="84">(47.6–49.4)</td>
<td id="B074042B2674007CA94CF9BCB8B48B7B" box="[606,749,1593,1615]" gridcol="2" gridrow="27" pageId="9" pageNumber="84">(45.6–52.6)</td>
<td id="B074042B2674007CA89FF9BCBE1B8B7B" box="[909,1090,1593,1615]" gridcol="3" gridrow="27" pageId="9" pageNumber="84">(47.9–51.1)</td>
<td id="B074042B2674007CAF8BF9BCBF718B7B" box="[1177,1320,1593,1615]" gridcol="4" gridrow="27" pageId="9" pageNumber="84">(48.3–50.7)</td>
</tr>
<tr id="F3A56D572674007CAB8DF9E7BF718B4C" box="[159,1320,1634,1656]" gridrow="28" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF9E7BAA58B4C" box="[159,252,1634,1656]" gridcol="0" gridrow="28" pageId="9" pageNumber="84">RTL</th>
<td id="B074042B2674007CAA43F9E7BBAC8B4C" box="[337,501,1634,1656]" gridcol="1" gridrow="28" pageId="9" pageNumber="84">44.9</td>
<td id="B074042B2674007CA94CF9E7B8B48B4C" box="[606,749,1634,1656]" gridcol="2" gridrow="28" pageId="9" pageNumber="84">45.8</td>
<td id="B074042B2674007CA89FF9E7BE1B8B4C" box="[909,1090,1634,1656]" gridcol="3" gridrow="28" pageId="9" pageNumber="84">46.6</td>
<td id="B074042B2674007CAF8BF9E7BF718B4C" box="[1177,1320,1634,1656]" gridcol="4" gridrow="28" pageId="9" pageNumber="84">46.5</td>
</tr>
<tr id="F3A56D572674007CAB8DF90FBF718B94" box="[159,1320,1674,1696]" gridrow="29" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F90FBBAC8B94" box="[337,501,1674,1696]" gridcol="1" gridrow="29" pageId="9" pageNumber="84">(43.2–45.7)</td>
<td id="B074042B2674007CA94CF90FB8B48B94" box="[606,749,1674,1696]" gridcol="2" gridrow="29" pageId="9" pageNumber="84">(43.8–46.1)</td>
<td id="B074042B2674007CA89FF90FBE1B8B94" box="[909,1090,1674,1696]" gridcol="3" gridrow="29" pageId="9" pageNumber="84">(46.1–46.6)</td>
<td id="B074042B2674007CAF8BF90FBF718B94" box="[1177,1320,1674,1696]" gridcol="4" gridrow="29" pageId="9" pageNumber="84">(44.8–48.4)</td>
</tr>
<tr id="F3A56D572674007CAB8DF936BF718BFD" box="[159,1320,1715,1737]" gridrow="30" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF936BAA58BFD" box="[159,252,1715,1737]" gridcol="0" gridrow="30" pageId="9" pageNumber="84">RFL</th>
<td id="B074042B2674007CAA43F936BBAC8BFD" box="[337,501,1715,1737]" gridcol="1" gridrow="30" pageId="9" pageNumber="84">47.4</td>
<td id="B074042B2674007CA94CF936B8B48BFD" box="[606,749,1715,1737]" gridcol="2" gridrow="30" pageId="9" pageNumber="84">47.5</td>
<td id="B074042B2674007CA89FF936BE1B8BFD" box="[909,1090,1715,1737]" gridcol="3" gridrow="30" pageId="9" pageNumber="84">49.1</td>
<td id="B074042B2674007CAF8BF936BF718BFD" box="[1177,1320,1715,1737]" gridcol="4" gridrow="30" pageId="9" pageNumber="84">48.8</td>
</tr>
<tr id="F3A56D572674007CAB8DF959BF718BC6" box="[159,1320,1756,1778]" gridrow="31" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F959BBAC8BC6" box="[337,501,1756,1778]" gridcol="1" gridrow="31" pageId="9" pageNumber="84">(46.9–48.7)</td>
<td id="B074042B2674007CA94CF959B8B48BC6" box="[606,749,1756,1778]" gridcol="2" gridrow="31" pageId="9" pageNumber="84">(46.3–50.5)</td>
<td id="B074042B2674007CA89FF959BE1B8BC6" box="[909,1090,1756,1778]" gridcol="3" gridrow="31" pageId="9" pageNumber="84">(46.6–49.2)</td>
<td id="B074042B2674007CAF8BF959BF718BC6" box="[1177,1320,1756,1778]" gridcol="4" gridrow="31" pageId="9" pageNumber="84">(46.9–51.8)</td>
</tr>
<tr id="F3A56D572674007CAB8DF880BF718A2F" box="[159,1320,1797,1819]" gridrow="32" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF880BAA58A2F" box="[159,252,1797,1819]" gridcol="0" gridrow="32" pageId="9" pageNumber="84">RIMTL</th>
<td id="B074042B2674007CAA43F880BBAC8A2F" box="[337,501,1797,1819]" gridcol="1" gridrow="32" pageId="9" pageNumber="84">7.7</td>
<td id="B074042B2674007CA94CF880B8B48A2F" box="[606,749,1797,1819]" gridcol="2" gridrow="32" pageId="9" pageNumber="84">7.6</td>
<td id="B074042B2674007CA89FF880BE1B8A2F" box="[909,1090,1797,1819]" gridcol="3" gridrow="32" pageId="9" pageNumber="84">6.5</td>
<td id="B074042B2674007CAF8BF880BF718A2F" box="[1177,1320,1797,1819]" gridcol="4" gridrow="32" pageId="9" pageNumber="84">6.7</td>
</tr>
<tr id="F3A56D572674007CAB8DF8A8BF718A77" box="[159,1320,1837,1859]" gridrow="33" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F8A8BBAC8A77" box="[337,501,1837,1859]" gridcol="1" gridrow="33" pageId="9" pageNumber="84">(7.2–8.1)</td>
<td id="B074042B2674007CA94CF8A8B8B48A77" box="[606,749,1837,1859]" gridcol="2" gridrow="33" pageId="9" pageNumber="84">(7.0–7.8)</td>
<td id="B074042B2674007CA89FF8A8BE1B8A77" box="[909,1090,1837,1859]" gridcol="3" gridrow="33" pageId="9" pageNumber="84">(6.5–6.6)</td>
<td id="B074042B2674007CAF8BF8A8BF718A77" box="[1177,1320,1837,1859]" gridcol="4" gridrow="33" pageId="9" pageNumber="84">(6.1–7.7)</td>
</tr>
<tr id="F3A56D572674007CAB8DF8D3BF718A58" box="[159,1320,1878,1900]" gridrow="34" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF8D3BAA58A58" box="[159,252,1878,1900]" gridcol="0" gridrow="34" pageId="9" pageNumber="84">R1TOEL</th>
<td id="B074042B2674007CAA43F8D3BBAC8A58" box="[337,501,1878,1900]" gridcol="1" gridrow="34" pageId="9" pageNumber="84">13.6</td>
<td id="B074042B2674007CA94CF8D3B8B48A58" box="[606,749,1878,1900]" gridcol="2" gridrow="34" pageId="9" pageNumber="84">13.2</td>
<td id="B074042B2674007CA89FF8D3BE1B8A58" box="[909,1090,1878,1900]" gridcol="3" gridrow="34" pageId="9" pageNumber="84">15.3</td>
<td id="B074042B2674007CAF8BF8D3BF718A58" box="[1177,1320,1878,1900]" gridcol="4" gridrow="34" pageId="9" pageNumber="84">15.1</td>
</tr>
<tr id="F3A56D572674007CAB8DF8FABF718AA1" box="[159,1320,1919,1941]" gridrow="35" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F8FABBAC8AA1" box="[337,501,1919,1941]" gridcol="1" gridrow="35" pageId="9" pageNumber="84">(12.0–14.2)</td>
<td id="B074042B2674007CA94CF8FAB8B48AA1" box="[606,749,1919,1941]" gridcol="2" gridrow="35" pageId="9" pageNumber="84">(12.8–14.5)</td>
<td id="B074042B2674007CA89FF8FABE1B8AA1" box="[909,1090,1919,1941]" gridcol="3" gridrow="35" pageId="9" pageNumber="84">(14.8–15.6)</td>
<td id="B074042B2674007CAF8BF8FABF718AA1" box="[1177,1320,1919,1941]" gridcol="4" gridrow="35" pageId="9" pageNumber="84">(14.0–16.6)</td>
</tr>
<tr id="F3A56D572674007CAB8DF82DBF718A8A" box="[159,1320,1960,1982]" gridrow="36" pageId="9" pageNumber="84">
<th id="B074042B2674007CAB8DF82DBAA58A8A" box="[159,252,1960,1982]" gridcol="0" gridrow="36" pageId="9" pageNumber="84">R4TDW</th>
<td id="B074042B2674007CAA43F82DBBAC8A8A" box="[337,501,1960,1982]" gridcol="1" gridrow="36" pageId="9" pageNumber="84">2.3</td>
<td id="B074042B2674007CA94CF82DB8B48A8A" box="[606,749,1960,1982]" gridcol="2" gridrow="36" pageId="9" pageNumber="84">2.0</td>
<td id="B074042B2674007CA89FF82DBE1B8A8A" box="[909,1090,1960,1982]" gridcol="3" gridrow="36" pageId="9" pageNumber="84">1.7</td>
<td id="B074042B2674007CAF8BF82DBF718A8A" box="[1177,1320,1960,1982]" gridcol="4" gridrow="36" pageId="9" pageNumber="84">2.1</td>
</tr>
<tr id="F3A56D572674007CAB8DF854BF718AD3" box="[159,1320,2001,2023]" gridrow="37" pageId="9" pageNumber="84" rowspan-0="1">
<td id="B074042B2674007CAA43F854BBAC8AD3" box="[337,501,2001,2023]" gridcol="1" gridrow="37" pageId="9" pageNumber="84">(2.1–2.5)</td>
<td id="B074042B2674007CA94CF854B8B48AD3" box="[606,749,2001,2023]" gridcol="2" gridrow="37" pageId="9" pageNumber="84">(1.8–2.7)</td>
<td id="B074042B2674007CA89FF854BE1B8AD3" box="[909,1090,2001,2023]" gridcol="3" gridrow="37" pageId="9" pageNumber="84">(1.5–2.0)</td>
<td id="B074042B2674007CAF8BF854BF718AD3" box="[1177,1320,2001,2023]" gridcol="4" gridrow="37" pageId="9" pageNumber="84">(1.7–2.3)</td>
</tr>
</table>
</paragraph>
<paragraph id="4D2A6F152677FF89ABD5FF12B9258C74" blockId="10.[151,1437,151,320]" pageId="10" pageNumber="85">
<emphasis id="7FE1B3072677FF89ABD5FF12BBFF8D84" bold="true" box="[199,422,151,176]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Eggs and tadpoles.</emphasis>
The diameter of nine ovulated eggs from a female (KUHE 15447) ranged from 2.59–3.09 (mean±1SD = 2.74±0.17) mm. The animal hemisphere of the egg is dark brown and the vegetal hemisphere is pale yellow in color. Three tadpoles putatively assigned to the new species and at stage (
<bibRefCitation id="290412E42677FF89AF76FF5ABEA68DCC" author="Gosner, K. L." box="[1124,1279,223,248]" pageId="10" pageNumber="85" pagination="183 - 190" refId="ref12477" refString="Gosner, K. L. (1960) A simplified table for staging anuran embryos and larvae with note on identification. Herpetologica, 16, 183 - 190." type="journal article" year="1960">Gosner, 1960</bibRefCitation>
) 35–36 (total length = 39.0–39.9 [39.6±0.5] mm, head-body length = 13.6–14.1 [13.8±0.3] mm) are nearly same as those of 
<taxonomicName id="8A9514962677FF89AE97FE80BAB18C74" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="selatan">
<emphasis id="7FE1B3072677FF89AE97FE80BAB18C74" italics="true" pageId="10" pageNumber="85">L. selatan</emphasis>
</taxonomicName>
described below in body shape and coloration (see 
<figureCitation id="D5AE73902677FF89A839FEA2B9368C74" box="[811,879,295,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1369,1391]" captionTargetBox="[151,1436,373,1348]" captionTargetId="figure-468@10.[151,1436,373,1348]" captionTargetPageId="10" captionText="FIGURE 7. Lateral (A, D), dorsal (B, E), and ventral (C, F) views, and oral disk (G, H) of larvae putatively assigned to Limnonectes utara sp. nov. from Larut Hill (A, B, C, G: KUHE 49514, Stage 36, total length = 39.9 mm) and confirmed as Limnonectes selatan sp. nov. from Genting Highlands (D, E, F, H: KUHE 49515, Stage 38, total length = 45.4 mm) after preservation. Scale bar for A–F= 10 mm. G, H=not to scale." figureDoi="http://doi.org/10.5281/zenodo.4949763" httpUri="https://zenodo.org/record/4949763/files/figure.png" pageId="10" pageNumber="85">Fig. 7</figureCitation>
).
</paragraph>
<caption id="19EA3F9D2677FF89AB85FADCB8AD88FF" ID-DOI="http://doi.org/10.5281/zenodo.4949763" ID-Zenodo-Dep="4949763" httpUri="https://zenodo.org/record/4949763/files/figure.png" pageId="10" pageNumber="85" startId="10.[151,250,1369,1391]" targetBox="[151,1436,373,1348]" targetPageId="10">
<paragraph id="4D2A6F152677FF89AB85FADCB8AD88FF" blockId="10.[151,1436,1369,1483]" pageId="10" pageNumber="85">
<emphasis id="7FE1B3072677FF89AB85FADCBB49885B" bold="true" box="[151,272,1369,1391]" pageId="10" pageNumber="85">FIGURE 7</emphasis>
. Lateral (A, D), dorsal (B, E), and ventral (C, F) views, and oral disk (G, H) of larvae putatively assigned to 
<taxonomicName id="8A9514962677FF89AB85FAFCBB0F88BA" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[151,342,1401,1422]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072677FF89AB85FAFCBB0F88BA" box="[151,342,1401,1422]" italics="true" pageId="10" pageNumber="85">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072677FF89AA4FFAFDBBE988BA" bold="true" box="[349,432,1400,1422]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="64D20E7C2677FF89AA4FFAFDBBE988BA" box="[349,432,1400,1422]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Larut Hill (A, B, C, G: KUHE 49514, Stage 36, total length = 39.9 mm) and confirmed as 
<taxonomicName id="8A9514962677FF89AB85FA12BB308898" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[151,361,1430,1452]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B3072677FF89AB85FA12BB308898" box="[151,361,1430,1452]" italics="true" pageId="10" pageNumber="85">Limnonectes selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072677FF89AA61FA13BB9F8898" bold="true" box="[371,454,1430,1452]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="64D20E7C2677FF89AA61FA13BB9F8898" box="[371,454,1430,1452]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Genting Highlands (D, E, F, H: KUHE 49515, Stage 38, total length = 45.4 mm) after preservation. Scale bar for A–F= 10 mm. G, H=not to scale.
</paragraph>
</caption>
<paragraph id="4D2A6F152677FF88ABD5FA77BED58DC3" blockId="10.[151,1437,1522,2015]" lastBlockId="11.[151,1436,151,752]" lastPageId="11" lastPageNumber="86" pageId="10" pageNumber="85">
<emphasis id="7FE1B3072677FF89ABD5FA77BB348B3F" bold="true" box="[199,365,1522,1547]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Comparisons.</emphasis>
<taxonomicName id="8A9514962677FF89AA61FA76B81D8B3E" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[371,580,1523,1546]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="utara">
<emphasis id="7FE1B3072677FF89AA61FA76B81D8B3E" box="[371,580,1523,1546]" italics="true" pageId="10" pageNumber="85">Limnonectes utara</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="8A9514962677FF89A802FA76B9278B3F" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[784,894,1522,1547]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="selatan" status="sp. nov.">
<emphasis id="7FE1B3072677FF89A802FA76B9468B3E" box="[784,799,1523,1546]" italics="true" pageId="10" pageNumber="85">L</emphasis>
. 
<emphasis id="7FE1B3072677FF89A83FFA77B9278B3F" box="[813,894,1522,1547]" italics="true" pageId="10" pageNumber="85">selatan</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072677FF89A896FA77B9848B3F" bold="true" box="[900,989,1522,1547]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="64D20E7C2677FF89A896FA77B9848B3F" box="[900,989,1522,1547]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but is significantly larger in male SVL, and has relatively smaller internarial space, tibia, hindlimb, and first toe, and larger inner metatarsal tubercle and fourth toe disk. The dorsal dark markings are clearer and more confluent, marking on rear of thigh is less clear, and the circum-cloacal warts are less developed than in 
<taxonomicName id="8A9514962677FF89A9F1F9DAB90A8B43" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[739,851,1630,1655]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="selatan">
<emphasis id="7FE1B3072677FF89A9F1F9DAB8AB8B42" box="[739,754,1631,1654]" italics="true" pageId="10" pageNumber="85">L</emphasis>
. 
<emphasis id="7FE1B3072677FF89A810F9DBB90A8B43" box="[770,851,1630,1655]" italics="true" pageId="10" pageNumber="85">selatan</emphasis>
</taxonomicName>
(
<figureCitation id="D5AE73902677FF89A871F9DBB9F08B43" box="[867,937,1630,1655]" captionStart="FIGURE 6" captionStartId="8.[151,250,1772,1794]" captionTargetBox="[151,1436,1412,1749]" captionTargetId="figure-421@8.[151,1436,1412,1749]" captionTargetPageId="8" captionText="FIGURE 6. Posterior views of left thigh of preserved male holotypes of Limnonectes utara sp. nov. (UKMHC 528) from Larut Hill (A) and L. selatan sp. nov. (UKMHC 529) from Genting Highlands (B), showing circum-cloacal warts and dark blotches along dorso-ventral border. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4949761" httpUri="https://zenodo.org/record/4949761/files/figure.png" pageId="10" pageNumber="85">Fig. 6</figureCitation>
). The new species, along with 
<taxonomicName id="8A9514962677FF89AE18F9DABF238B43" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[1290,1402,1630,1655]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="selatan">
<emphasis id="7FE1B3072677FF89AE18F9DABF408B42" box="[1290,1305,1631,1654]" italics="true" pageId="10" pageNumber="85">L</emphasis>
. 
<emphasis id="7FE1B3072677FF89AE3BF9DBBF238B43" box="[1321,1402,1630,1655]" italics="true" pageId="10" pageNumber="85">selatan</emphasis>
</taxonomicName>
, is differentiated from all the other named species except for some Bornean 
<taxonomicName id="8A9514962677FF89A8DAF906BE7F8BAF" box="[968,1062,1666,1691]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072677FF89A8DAF906B98E8BAE" box="[968,983,1667,1690]" italics="true" pageId="10" pageNumber="85">L</emphasis>
. 
<emphasis id="7FE1B3072677FF89A8F4F907BE7F8BAF" box="[998,1062,1666,1691]" italics="true" pageId="10" pageNumber="85">kuhlii</emphasis>
</taxonomicName>
-like frogs by their tibia, which is heavily covered by large, conical tubercles (vs. tibia at most with weak tubercles dorsally). In addition, it differs from 
<taxonomicName id="8A9514962677FF89ABC6F94EBB138BD6" baseAuthorityName="Stejneger" baseAuthorityYear="1901" box="[212,330,1739,1762]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="namiyei">
<emphasis id="7FE1B3072677FF89ABC6F94EBB138BD6" box="[212,330,1739,1762]" italics="true" pageId="10" pageNumber="85">L. namiyei</emphasis>
</taxonomicName>
by the lack of vocal openings, and from 
<taxonomicName id="8A9514962677FF89A81DF94EB9248BD7" box="[783,893,1738,1763]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="fragilis">
<emphasis id="7FE1B3072677FF89A81DF94EB9248BD7" box="[783,893,1738,1763]" italics="true" pageId="10" pageNumber="85">L. fragilis</emphasis>
</taxonomicName>
by smaller subarticular tubercles and possession of nuptial pads in males. From 
<taxonomicName id="8A9514962677FF89AAE0F96AB8168A33" box="[498,591,1774,1799]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072677FF89AAE0F96AB8168A33" box="[498,591,1774,1799]" italics="true" pageId="10" pageNumber="85">L. kuhlii</emphasis>
</taxonomicName>
, it differs in possession of nuptial pads in males and larger body size (mean SVL 
<quantity id="8A6DC2F02677FF89ABC6F897BB7D8A1E" box="[212,292,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="10" pageNumber="85" unit="mm" value="74.0">74 mm</quantity>
in males and 
<quantity id="8A6DC2F02677FF89AAD4F897B84F8A1E" box="[454,534,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="85" unit="mm" value="70.0">70 mm</quantity>
in females vs. 
<quantity id="8A6DC2F02677FF89A9D7F897B94D8A1E" box="[709,788,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="85" unit="mm" value="62.0">62 mm</quantity>
and 
<quantity id="8A6DC2F02677FF89A85DF897B9F88A1E" box="[847,929,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="85" unit="mm" value="59.0">59 mm</quantity>
, respectively, in 
<taxonomicName id="8A9514962677FF89AF73F896BE998A1F" box="[1121,1216,1810,1835]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072677FF89AF73F896BE998A1F" box="[1121,1216,1810,1835]" italics="true" pageId="10" pageNumber="85">L. kuhlii</emphasis>
</taxonomicName>
). The new species, with a back usually without distinct ridges, and nuptial pad only on the first finger, differs from 
<taxonomicName id="8A9514962677FF89AFE9F8B2BFCC8A7B" authorityName="Ye &amp; Fei" authorityYear="1994" box="[1275,1429,1846,1871]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="fujianensis">
<emphasis id="7FE1B3072677FF89AFE9F8B2BFCC8A7B" box="[1275,1429,1846,1871]" italics="true" pageId="10" pageNumber="85">L. fujianensis</emphasis>
</taxonomicName>
, which has a back with many ridges, including a dorsolateral ridge, and nuptial pads on the two inner fingers. From 
<taxonomicName id="8A9514962677FF89AB85F8FABB6C8AA3" authorityName="Ye, Fei, Xie &amp; Jiang" authorityYear="2007" box="[151,309,1918,1943]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="bannaensis">
<emphasis id="7FE1B3072677FF89AB85F8FABB6C8AA3" box="[151,309,1918,1943]" italics="true" pageId="10" pageNumber="85">L. bannaensis</emphasis>
</taxonomicName>
, it differs by lacking nuptial pad on the second finger. 
<taxonomicName id="8A9514962677FF89A8AAF8FABED58AA2" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[952,1164,1919,1942]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072677FF89A8AAF8FABED58AA2" box="[952,1164,1919,1942]" italics="true" pageId="10" pageNumber="85">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072677FF89AF87F8FBBEAB8AA3" bold="true" box="[1173,1266,1918,1943]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="64D20E7C2677FF89AF87F8FBBEAB8AA3" box="[1173,1266,1918,1943]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
invariably has confluent dark dorsal marking unlike 
<taxonomicName id="8A9514962677FF89A948F826B8888A8E" box="[602,721,1955,1978]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="jarujini">
<emphasis id="7FE1B3072677FF89A948F826B8888A8E" box="[602,721,1955,1978]" italics="true" pageId="10" pageNumber="85">L. jarujini</emphasis>
</taxonomicName>
and 
<taxonomicName id="8A9514962677FF89A806F826B9C78A8E" box="[788,926,1955,1978]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="isanensis">
<emphasis id="7FE1B3072677FF89A806F826B9C78A8E" box="[788,926,1955,1978]" italics="true" pageId="10" pageNumber="85">L. isanensis</emphasis>
</taxonomicName>
, which usually lack dark dorsal markings. 
<taxonomicName id="8A9514962677FF89AB85F84DBB308AEB" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[151,361,1992,2015]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072677FF89AB85F84DBB308AEB" box="[151,361,1992,2015]" italics="true" pageId="10" pageNumber="85">Limnonectes utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072677FF89AA62F843BB938AEB" bold="true" box="[368,458,1990,2015]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="64D20E7C2677FF89AA62F843BB938AEB" box="[368,458,1990,2015]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="8A9514962677FF89A94DF84DB8918AEB" box="[607,712,1990,2015]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="85" phylum="Chordata" rank="species" species="taylori">
<emphasis id="7FE1B3072677FF89A94DF84DB8918AEB" box="[607,712,1990,2015]" italics="true" pageId="10" pageNumber="85">L. taylori</emphasis>
</taxonomicName>
in having few dorsal warts, which are much more abundant and widely present in 
<taxonomicName id="8A9514962676FF88AA4FFF1CBB9C8D84" box="[349,453,151,176]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="taylori">
<emphasis id="7FE1B3072676FF88AA4FFF1CBB9C8D84" box="[349,453,151,176]" italics="true" pageId="11" pageNumber="86">L. taylori</emphasis>
</taxonomicName>
. From 
<taxonomicName id="8A9514962676FF88A907FF1CB89D8D9B" box="[533,708,152,176]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="megastomias">
<emphasis id="7FE1B3072676FF88A907FF1CB89D8D9B" box="[533,708,152,176]" italics="true" pageId="11" pageNumber="86">L. megastomias</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88A9C0FF1CB9758D84" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[722,812,153,176]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072676FF88A9C0FF1CB9758D84" box="[722,812,153,176]" italics="true" pageId="11" pageNumber="86">L. utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072676FF88A820FF12B9D28D84" bold="true" box="[818,907,151,176]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="64D20E7C2676FF88A820FF12B9D28D84" box="[818,907,151,176]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs in having a smaller body size, relatively shorter head, lacking a nuptial pad on the second finger, and not having a heavily pigmented venter. In addition, the second finger is not constantly longer than first in the new species, unlike 
<taxonomicName id="8A9514962676FF88A8C7FF64BEDC8DC3" box="[981,1157,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="megastomias">
<emphasis id="7FE1B3072676FF88A8C7FF64BEDC8DC3" box="[981,1157,224,248]" italics="true" pageId="11" pageNumber="86">L. megastomias</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E2676FF88ABD5FE81B9E88C99" pageId="11" pageNumber="86" type="distribution">
<paragraph id="4D2A6F152676FF88ABD5FE81B9E88C99" blockId="11.[151,1436,151,752]" pageId="11" pageNumber="86">
<emphasis id="7FE1B3072676FF88ABD5FE81BB438C29" bold="true" box="[199,282,260,285]" inLineHeading="true" pageId="11" pageNumber="86" reason="1">Range.</emphasis>
Peninsular 
<collectingCountry id="35822F852676FF88AAB4FE81B8488C29" box="[422,529,260,285]" name="Malaysia" pageId="11" pageNumber="86">Malaysia</collectingCountry>
: Bukit Larut (= Larut Hill), State of 
<collectingRegion id="8F51A1F72676FF88A8D4FE81BE508C29" box="[966,1033,260,285]" country="Malaysia" name="Perak" pageId="11" pageNumber="86">Perak</collectingRegion>
, (
<quantity id="8A6DC2F02676FF88AF32FE81BEFD8C2F" box="[1056,1188,260,285]" metricMagnitude="2" metricUnit="m" metricValue="6.4" metricValueMax="6.8" metricValueMin="6.0" pageId="11" pageNumber="86" unit="m" value="640.0" valueMax="680.0" valueMin="600.0">600–680 m</quantity>
a.s.l., 
<quantity id="8A6DC2F02676FF88AFE7FE81BF148C2F" box="[1269,1357,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="11" pageNumber="86" unit="m" value="1160.0">1160 m</quantity>
a.s.l.), Sungai Tembat, Hulu 
<collectingRegion id="8F51A1F72676FF88AA9EFEADB84A8C74" box="[396,531,296,320]" country="Malaysia" name="Terengganu" pageId="11" pageNumber="86">Terengganu</collectingRegion>
, State of 
<collectingRegion id="8F51A1F72676FF88A968FEADB9588C74" box="[634,769,296,320]" country="Malaysia" name="Terengganu" pageId="11" pageNumber="86">Terengganu</collectingRegion>
, (
<quantity id="8A6DC2F02676FF88A806FEA2B9CD8C0B" box="[788,916,295,320]" metricMagnitude="2" metricUnit="m" metricValue="1.69" metricValueMax="1.7" metricValueMin="1.68" pageId="11" pageNumber="86" unit="m" value="169.0" valueMax="170.0" valueMin="168.0">168–170 m</quantity>
a.s.l.). Records of 
<taxonomicName id="8A9514962676FF88AF75FEACBEFB8C74" box="[1127,1186,297,320]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis id="7FE1B3072676FF88AF75FEACBEFB8C74" box="[1127,1186,297,320]" italics="true" pageId="11" pageNumber="86">Rana</emphasis>
</taxonomicName>
or 
<taxonomicName id="8A9514962676FF88AFDAFEADBFC58C74" box="[1224,1436,295,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="7FE1B3072676FF88AFDAFEADBFC58C74" box="[1224,1436,295,320]" italics="true" pageId="11" pageNumber="86">Limnonectes kuhlii</emphasis>
</taxonomicName>
from Larut Hills at about 
<quantity id="8A6DC2F02676FF88AADBFEC9B8448C51" box="[457,541,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="11" pageNumber="86" unit="ft" value="4500.0">4500 ft</quantity>
(
<bibRefCitation id="290412E42676FF88A93DFEC9B8EF8C51" author="Berry, P. Y." box="[559,694,332,357]" pageId="11" pageNumber="86" refId="ref12019" refString="Berry, P. Y. (1975) The Amphibian Fauna of Peninsular Malaysia. Tropical Press, Kuala Lumpur, 130 pp" type="book" year="1975">Berry 1975</bibRefCitation>
), Ulu Kenas Recreational Forest, Gunung Bubu (
<bibRefCitation id="290412E42676FF88AFEAFEC9BA8A8CBC" author="Grismer, L. L. &amp; Chan, K. O. &amp; Grismer, J. L. &amp; Wood, P. L. Jr. &amp; Norhayati, A." pageId="11" pageNumber="86" pagination="147 - 160" refId="ref12508" refString="Grismer, L. L., Chan, K. O., Grismer, J. L., Wood, P. L. Jr. &amp; Norhayati, A. (2010) A checklist of the herpetofauna of the Banjaran Bintang, Peninsular Malaysia. Russian Journal of Herpetology, 17, 147 - 160." type="journal article" year="2010">
Grismer 
<emphasis id="7FE1B3072676FF88AE72FEC8BFCC8C51" box="[1376,1429,332,357]" italics="true" pageId="11" pageNumber="86">et al</emphasis>
. 2010
</bibRefCitation>
), and Gunung Lawit (
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), State of 
<collectingRegion id="8F51A1F72676FF88A988FEF5B9478CBC" box="[666,798,368,392]" country="Malaysia" name="Terengganu" pageId="11" pageNumber="86">Terengganu</collectingRegion>
(
<bibRefCitation id="290412E42676FF88A83DFEEAB9EC8CBC" author="Dring, J. C." box="[815,949,367,392]" pageId="11" pageNumber="86" pagination="181 - 241" refId="ref12249" refString="Dring, J. C. (1979) Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum Natural History (Zoology), 34, 181 - 241." type="journal article" year="1979">Dring 1979</bibRefCitation>
) are thought to represent this species (see Discussion). Known localities range in altitude from 
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a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E2676FF88ABD5FE32BB9C8FFF" pageId="11" pageNumber="86" type="biology_ecology">
<paragraph id="4D2A6F152676FF88ABD5FE32BB9C8FFF" blockId="11.[151,1436,151,752]" pageId="11" pageNumber="86">
<emphasis id="7FE1B3072676FF88ABD5FE32BBDD8CE4" bold="true" box="[199,388,439,464]" inLineHeading="true" pageId="11" pageNumber="86" reason="1">Natural history.</emphasis>
At the 
<typeStatus id="922ED1B72676FF88AAC4FE3DB85F8CE4" box="[470,518,440,464]" pageId="11" pageNumber="86">type</typeStatus>
locality (
<quantity id="8A6DC2F02676FF88A97DFE32B89D8CFB" box="[623,708,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="11" pageNumber="86" unit="m" value="1160.0">1160 m</quantity>
a.s.l.), the 
<typeStatus id="922ED1B72676FF88A82CFE3DB9378CE4" box="[830,878,440,464]" pageId="11" pageNumber="86">type</typeStatus>
series of 
<taxonomicName id="8A9514962676FF88A8CBFE3CBE6A8CE4" authority="Matsui &amp; Belabut &amp; Ahmad, 2014" authorityName="Matsui &amp; Belabut &amp; Ahmad" authorityYear="2014" box="[985,1075,441,464]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="utara" status="sp. nov.">
<emphasis id="7FE1B3072676FF88A8CBFE3CBE6A8CE4" box="[985,1075,441,464]" italics="true" pageId="11" pageNumber="86">L. utara</emphasis>
</taxonomicName>
<emphasis id="7FE1B3072676FF88AF2BFE32BECB8CE4" bold="true" box="[1081,1170,439,464]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="64D20E7C2676FF88AF2BFE32BECB8CE4" box="[1081,1170,439,464]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in and along a small stream in a roadside ditch (width &lt;
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) in a secondary forest, while at the lower elevation site (
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a.s.l.), frogs were found at the edge of the shaded main stream (width &lt;
<quantity id="8A6DC2F02676FF88A8EEFE7ABE748F2C" box="[1020,1069,511,536]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="86" unit="m" value="3.0">3 m</quantity>
) in dipterocarp forest. Females collected in early January and late February had fully developed ova in ovaries. Tadpoles in later stages of development were also found in early January. Therefore, the breeding period may be relatively extended. Other amphibian species at the 
<typeStatus id="922ED1B72676FF88AADCFDE9BBA78FB0" box="[462,510,620,644]" pageId="11" pageNumber="86">type</typeStatus>
locality were 
<taxonomicName id="8A9514962676FF88A9A1FDE9B9C78FB1" authorityName="Boulenger" authorityYear="1886" box="[691,926,620,645]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="longipes">
<emphasis id="7FE1B3072676FF88A9A1FDE9B9C78FB1" box="[691,926,620,645]" italics="true" pageId="11" pageNumber="86">Megophrys longipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88A8A6FDE9BEFE8FB1" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[948,1191,620,645]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="7FE1B3072676FF88A8A6FDE9BEFE8FB1" box="[948,1191,620,645]" italics="true" pageId="11" pageNumber="86">Leptolalax heteropus</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88AFAEFDE9BFCD8FB1" authorityName="Inger" authorityYear="1960" box="[1212,1428,620,645]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="malayana">
<emphasis id="7FE1B3072676FF88AFAEFDE9BFCD8FB1" box="[1212,1428,620,645]" italics="true" pageId="11" pageNumber="86">Ansonia malayana</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88AB85FD15BBC78F9C" baseAuthorityName="Anderson" baseAuthorityYear="1871" box="[151,414,655,680]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="khasianus">
<emphasis id="7FE1B3072676FF88AB85FD15BBC78F9C" box="[151,414,655,680]" italics="true" pageId="11" pageNumber="86">Limnonectes khasianus</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88AAA2FD14B8418F9C" baseAuthorityName="Boulenger" baseAuthorityYear="1920" box="[432,536,655,680]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="blythii">
<emphasis id="7FE1B3072676FF88AAA2FD14BBE68F9C" box="[432,447,657,680]" italics="true" pageId="11" pageNumber="86">L</emphasis>
. 
<emphasis id="7FE1B3072676FF88AAC3FD0AB8418F9C" box="[465,536,655,680]" italics="true" pageId="11" pageNumber="86">blythii</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88A938FD0AB95D8F9C" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[554,772,655,680]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="larutensis">
<emphasis id="7FE1B3072676FF88A938FD0AB95D8F9C" box="[554,772,655,680]" italics="true" pageId="11" pageNumber="86">Amolops larutensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88A807FD0AB99F8F9C" baseAuthorityName="Boulenger" baseAuthorityYear="1891" box="[789,966,655,680]" class="Amphibia" family="Ranidae" genus="Odorrana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="hosii">
<emphasis id="7FE1B3072676FF88A807FD0AB99F8F9C" box="[789,966,655,680]" italics="true" pageId="11" pageNumber="86">Odorrana hosii</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88A8C5FD0ABE988F9C" baseAuthorityName="Leong &amp; Lim" baseAuthorityYear="2003" box="[983,1217,655,680]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="banjarana">
<emphasis id="7FE1B3072676FF88A8C5FD0ABE988F9C" box="[983,1217,655,680]" italics="true" pageId="11" pageNumber="86">Hylarana banjarana</emphasis>
</taxonomicName>
, 
<taxonomicName id="8A9514962676FF88AFC1FD0ABFCC8F93" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[1235,1429,655,680]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="petersi">
<emphasis id="7FE1B3072676FF88AFC1FD0ABFCC8F93" box="[1235,1429,655,680]" italics="true" pageId="11" pageNumber="86">Philautus petersi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8A9514962676FF88ABD4FD31BBE78FF9" authorityName="Taylor" authorityYear="1960" box="[198,446,692,717]" class="Amphibia" family="Ichthyophiidae" genus="Ichthyophis" kingdom="Animalia" order="Gymnophiona" pageId="11" pageNumber="86" phylum="Chordata" rank="species" species="larutensis">
<emphasis id="7FE1B3072676FF88ABD4FD31BBE78FF9" box="[198,446,692,717]" italics="true" pageId="11" pageNumber="86">Ichthyophis larutensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="058F3C9E2676FF88ABD5FD52BFC98FC4" box="[199,1424,727,752]" pageId="11" pageNumber="86" type="description">
<paragraph id="4D2A6F152676FF88ABD5FD52BFC98FC4" blockId="11.[151,1436,151,752]" box="[199,1424,727,752]" pageId="11" pageNumber="86">
<emphasis id="7FE1B3072676FF88ABD5FD52BB178FC4" bold="true" box="[199,334,727,752]" inLineHeading="true" pageId="11" pageNumber="86" reason="1">Karyotype.</emphasis>
The diploid chromosome number is 22, with six large and five small pairs (Matsui, unpub. data).
</paragraph>
</subSubSection>
</treatment>
</document>