133 lines
12 KiB
XML
133 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="C4C3BE6BC0A0D972128B66F7651E5178" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Geotrochus kitteli Vermeulen, Liew & Schilthuizen, sp. n." docType="treatment" docUuid="AB073D02-62A6-4793-8420-BE7E44C87661" docUuidSource="ZooBank" docVersion="10" lastPageNumber="109" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="108" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>531</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>139</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
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<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144057526" LSID="urn:lsid:zoobank.org:act:AB073D02-62A6-4793-8420-BE7E44C87661" httpUri="http://treatment.plazi.org/id/C4C3BE6BC0A0D972128B66F7651E5178" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
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<subSubSection pageId="107" pageNumber="108" type="multiple">
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<paragraph pageId="107" pageNumber="108">Taxon classification Animalia Stylommatophora Trochomorphidae</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="nomenclature">
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<paragraph pageId="107" pageNumber="108">
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<taxonomicName LSID="http://zoobank.org/AB073D02-62A6-4793-8420-BE7E44C87661" authority="Vermeulen, Liew & Schilthuizen" class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus kitteli" order="Stylommatophora" pageId="107" pageNumber="108" phylum="Mollusca" rank="species" species="kitteli">Geotrochus kitteli Vermeulen, Liew & Schilthuizen</taxonomicName>
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<taxonomicNameLabel pageId="107" pageNumber="108">sp. n.</taxonomicNameLabel>
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Figure 89
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="holotype">
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<paragraph pageId="107" pageNumber="108">
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<materialsCitation collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectionCode="RMNH" collectorName="K. Kittel" country="Malaysia" httpUri="http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003930" location="West Coast Province" municipality="Sabah" specimenCode="RMNH.5003930" specimenCount="1" typeStatus="holotype">
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<typeStatus>Holotype</typeStatus>
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.
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<collectingCountry>Malaysia</collectingCountry>
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,
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<collectingMunicipality>Sabah</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:C4C3BE6BC0A0D972128B66F7651E5178:5D833EE4C694F058433EFBE6ECDB1A08" country="Malaysia" municipality="Sabah" name="West Coast Province">West Coast Province</location>
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,
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<location LSID="urn:lsid:plazi:treatment:C4C3BE6BC0A0D972128B66F7651E5178:41CB2A624E92ED3AE37712A5F418EA5D" country="Malaysia" municipality="Sabah" name="Kinabalu N. P.">Kinabalu N.P.</location>
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,
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<location LSID="urn:lsid:plazi:treatment:C4C3BE6BC0A0D972128B66F7651E5178:B909060DF400A7E038EC5D773873BD3E" country="Malaysia" municipality="Sabah" name="Headquarters area">Headquarters area</location>
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,
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<location LSID="urn:lsid:plazi:treatment:C4C3BE6BC0A0D972128B66F7651E5178:F6067452F2DFEF410B03A6046753AF3B" country="Malaysia" municipality="Sabah" name="Silau Silau">Silau Silau</location>
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(leg.
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<collectorName>K. Kittel</collectorName>
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,
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<specimenCode>RMNH.5003930</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="materials_examined">
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<paragraph pageId="107" pageNumber="108">Examined material from Sabah.</paragraph>
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<paragraph pageId="107" pageNumber="108">West Coast Province, Kinabalu N.P., Sayap-Nunuhon trail at 1776 m (leg. T.S. Liew, Dominik, J. Lapidin & Jasilin, BOR/MOL 4109).</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="description">
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<paragraph pageId="107" pageNumber="108">Description.</paragraph>
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<paragraph pageId="107" pageNumber="108">Shell rather small, rather thin, about opaque, yellowish brown, whorls slightly darker brown towards the periphery, moderately low-conical with approx. flat sides; apex not protruding. Surface dull or slightly shiny above the periphery, shiny below. Whorls: Apical whorls moderately convex, outer slightly convex, slightly shouldered; suture somewhat impressed, coinciding with the periphery; last whorl acutely angular, slightly compressed at the periphery, rounded below the periphery. Protoconch c. 2 1/8 whorls, smooth (see remark, below). Teleoconch. Radial sculpture: above the periphery rather distinct growth lines, raised at irregular intervals and wrinkling the shell surface, most conspicuously so below the suture, and giving the shell surface a somewhat rough appearance; below the periphery indistinct growth lines only. Spiral sculpture: last whorl with a sharp, somewhat pinched peripheral keel; start of fifth whorl with c. 15 thin, moderately spaced spiral threads: 1-2 slightly more distinct close to the periphery, as well as 13-14 inconspicuous above these; no spiral sculpture below the periphery. Umbilicus closed. Peristome thickened and reflexed. Dimensions: Height c. 5.5 mm; width c. 10 mm; h/w c. 0.55; diameters of the first 4 whorls c. 1.2 mm, c. 2.2 mm, c. 3.3 mm, c. 4.8 mm respectively; number of whorls c. 6 3/8, height aperture c. 3.5 mm; width aperture c. 5 mm.</paragraph>
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<caption pageId="107" pageNumber="108">
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<paragraph pageId="107" pageNumber="108">
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Figure 89-92. 89
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus kitteli" order="Stylommatophora" pageId="107" pageNumber="108" phylum="Mollusca" rank="species" species="kitteli">Geotrochus kitteli</taxonomicName>
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sp. n. Frontal view, shell 5.5 mm high (Malaysia, Sabah, West Coast Province, Kinabalu N.P., Headquarters area, Silau Silau (RMNH.5003930, holotype) 90
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus whiteheadi" order="Stylommatophora" pageId="107" pageNumber="108" phylum="Mollusca" rank="species" species="whiteheadi">Geotrochus whiteheadi</taxonomicName>
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(E.A. Smith, 1895) 90A Frontal view, shell 5.5 mm high 90B Umbilical view (Malaysia, Sabah, West Coast Province, Kinabalu N.P.,
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<normalizedToken originalValue="Carson’s" pageId="107" pageNumber="108">Carson's</normalizedToken>
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Falls, V 12710) 91
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus subscalaris" order="Stylommatophora" pageId="107" pageNumber="108" phylum="Mollusca" rank="species" species="subscalaris">Geotrochus subscalaris</taxonomicName>
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sp. n. Frontal view, shell 7.0 mm high (Malaysia, Sabah, Sandakan Province, Kinabatangan valley, Batu Pangi, RMNH.5003931, holotype) 92
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus meristotrochus" order="Stylommatophora" pageId="107" pageNumber="108" phylum="Mollusca" rank="species" species="meristotrochus">Geotrochus meristotrochus</taxonomicName>
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sp. n. 92A Frontal view, shell 7.0 mm high 92B Umbilical view (Malaysia, Sabah, Interior Province, Sepulut valley, Batu Punggul, RMNH.5003932, holotype).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="habitat in sabah and distribution">
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<paragraph pageId="107" pageNumber="108">Habitat in Sabah and distribution.</paragraph>
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<paragraph pageId="107" pageNumber="108">Mixed montane forest, 1500-1800 m alt. Sabah: Mount Kinabalu only. Endemic to Sabah.</paragraph>
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</subSubSection>
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<subSubSection pageId="108" pageNumber="109" type="cross diagnosis">
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<paragraph pageId="108" pageNumber="109">
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<pageBreakToken pageId="108" pageNumber="109" start="start">Cross</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph pageId="108" pageNumber="109">
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In size and general shape most similar to
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrochus whiteheadi" order="Stylommatophora" pageId="108" pageNumber="109" phylum="Mollusca" rank="species" species="whiteheadi">Geotrochus whiteheadi</taxonomicName>
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, it differs by the higher ratio h/w, and by the more distinct growth lines above the periphery, which give the shell surface a rough appearance. It differs from
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<taxonomicName class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochomorpha rhysa" order="Stylommatophora" pageId="108" pageNumber="109" phylum="Mollusca" rank="species" species="rhysa">Trochomorpha rhysa</taxonomicName>
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by its somewhat pinched peripheral keel and the not-nodular spiral sculpture above the periphery.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="108" pageNumber="109" type="remarks">
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<paragraph pageId="108" pageNumber="109">Remarks.</paragraph>
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<paragraph pageId="108" pageNumber="109">The single shell available is somewhat worn in the apical region. Traces of what may be radial riblets are present in the suture of the protoconch, close to the transition to the teleoconch.</paragraph>
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</subSubSection>
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<subSubSection pageId="108" pageNumber="109" type="etymology">
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<paragraph pageId="108" pageNumber="109">Etymology.</paragraph>
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<paragraph pageId="108" pageNumber="109">Named after Mr. Klaus Kittel, Germany, who collected the type specimen.</paragraph>
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</subSubSection>
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</treatment>
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</document> |