427 lines
55 KiB
XML
427 lines
55 KiB
XML
<document id="B19FD5E5F96194241DBFC76A547E5895" ID-DOI="10.5281/zenodo.201469" ID-GBIF-Dataset="7f5320be-c542-478b-8c8b-a0834490a7ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="201469" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226411041" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo & Noriega, Jorge Ari" docDate="2011" docId="C46F87B0CF21A57BFF13F99BFAA718B3" docLanguage="en" docName="zt02747p052.pdf" docOrigin="Zootaxa 2747" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ananteris palmari Botero-Trujillo & Noriega, 2011, sp. nov." docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="3856FFC8CF28A575FF84FF9FFFA91B4B" masterDocTitle="On the identity of Microananteris, with a discussion on pectinal morphology, and description of a new Ananteris from Brazil (Scorpiones, Buthidae)" masterLastPageNumber="52" masterPageNumber="37" pageNumber="46" updateTime="1698238440825" updateUser="plazi">
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<mods:titleInfo id="A1C41C3F994BD218672389CFE27F7C37">
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<mods:title id="912D26C425F078245E13A59816B80D0D">On the identity of Microananteris, with a discussion on pectinal morphology, and description of a new Ananteris from Brazil (Scorpiones, Buthidae)</mods:title>
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<mods:namePart id="24898E6B1A59BDE1B0D9FAF6948305F8">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:namePart id="780B2C363F509DBC848D12D5C8FAEB2B">Noriega, Jorge Ari</mods:namePart>
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<mods:title id="0B78C5593547BACFFF1601D9146FE98E">Zootaxa</mods:title>
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<mods:part id="15DC7DB565782B2BC23881696DEBC99F">
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<mods:date id="2EE03875A6386AE6CD2A03335E13D0F0">2011</mods:date>
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<mods:start id="C37E362CFBA036FA388B66AFC01BF79F">37</mods:start>
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<mods:identifier id="3F9F42167CF9A79A25D3E4BADE4788F2" type="DOI">10.5281/zenodo.201469</mods:identifier>
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<mods:identifier id="2386287C91372769401A6CE058630A70" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8AB59321D171EFC57ABC08C65B3310C3" type="Zenodo-Dep">201469</mods:identifier>
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<treatment id="C46F87B0CF21A57BFF13F99BFAA718B3" ID-DOI="http://doi.org/10.5281/zenodo.6182461" ID-GBIF-Taxon="119386437" ID-Zenodo-Dep="6182461" LSID="urn:lsid:plazi:treatment:C46F87B0CF21A57BFF13F99BFAA718B3" httpUri="http://treatment.plazi.org/id/C46F87B0CF21A57BFF13F99BFAA718B3" lastPageId="14" lastPageNumber="50" pageId="9" pageNumber="46">
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<subSubSection id="04DC652DCF21A57CFF13F99BFE441D0A" pageId="9" pageNumber="46" type="nomenclature">
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<paragraph id="4C7936A6CF21A57CFF13F99BFE631D56" blockId="9.[151,493,1540,1601]" box="[151,458,1540,1565]" pageId="9" pageNumber="46">
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<heading id="173181CACF21A57CFF13F99BFE631D56" bold="true" box="[151,458,1540,1565]" fontSize="10" level="2" pageId="9" pageNumber="46" reason="2">
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<emphasis id="7EB2EAB4CF21A57CFF13F99BFE631D56" bold="true" box="[151,458,1540,1565]" pageId="9" pageNumber="46">
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<taxonomicName id="8BC64D25CF21A57CFF13F99BFEC01D56" ID-CoL="66LBR" box="[151,361,1540,1565]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
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<emphasis id="7EB2EAB4CF21A57CFF13F99BFEC01D56" bold="true" box="[151,361,1540,1565]" italics="true" pageId="9" pageNumber="46">Ananteris palmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="658157CFCF21A57CFEF5F99AFE631D56" box="[369,458,1541,1565]" pageId="9" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4C7936A6CF21A57CFF13F9B6FE441D0A" blockId="9.[151,493,1540,1601]" box="[151,493,1577,1601]" pageId="9" pageNumber="46">
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<figureCitation id="D4FD2A23CF21A57CFF13F9B6FE931D0A" box="[151,314,1577,1601]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1411,1434]" captionTargetBox="[153,1436,197,1389]" captionTargetId="figure@7.[151,1436,193,1390]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Left pectine of Ananteris palmari sp. nov., female holotype, under SEM. 19. Pectine. 20. Teeth. 21. Detail of teeth ornamentation. 22. Zoom of peg sensillae." httpUri="https://zenodo.org/record/201475/files/figure.png" pageId="9" pageNumber="46">Figures 19–22</figureCitation>
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,
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<figureCitation id="D4FD2A23CF21A57CFEC2F9B6FE391D0A" box="[326,400,1577,1601]" captionStart-0="FIGURES 31 – 36" captionStart-1="FIGURES 37 – 38" captionStart-2="FIGURE 39" captionStart-3="FIGURE 40" captionStartId-0="10.[151,264,1951,1974]" captionStartId-1="11.[151,264,949,972]" captionStartId-2="12.[151,250,2009,2032]" captionStartId-3="13.[151,250,846,869]" captionTargetBox-0="[168,1415,462,1902]" captionTargetBox-1="[155,1431,193,927]" captionTargetBox-2="[156,1431,1025,1984]" captionTargetBox-3="[176,1414,196,822]" captionTargetId-0="figure@10.[160,1420,445,1914]" captionTargetId-1="figure@11.[155,1432,193,928]" captionTargetId-2="figure@12.[153,1433,1021,1989]" captionTargetId-3="figure@13.[169,1418,193,825]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURES 31 – 36. Ananteris palmari sp. nov., female holotype. 31 – 35. Distribution of the trichobothria (illustrated from right pedipalp). 31. Femur, dorsal aspect. 32. Patella, dorsal aspect. 33. Patella, external aspect. 34. Chela, dorsoexternal aspect. 35. Chela, ventral aspect. 36. Right chelicera, dorsal aspect. Scale bars equal 0.5 mm." captionText-1="FIGURES 37 – 38. Female holotype of Ananteris palmari sp. nov., dorsal and ventral aspects. Scale bar equals 3 mm." captionText-2="FIGURE 39. Habitat of Ananteris palmari sp. nov. in the Natural Reserve Palmari, Brazil." captionText-3="FIGURE 40. Location of the type locality of Ananteris palmari sp. nov. (star)." httpUri-0="https://zenodo.org/record/201478/files/figure.png" httpUri-1="https://zenodo.org/record/201479/files/figure.png" httpUri-2="https://zenodo.org/record/201480/files/figure.png" httpUri-3="https://zenodo.org/record/201481/files/figure.png" pageId="9" pageNumber="46">31–40</figureCitation>
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;
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<tableCitation id="0144031DCF21A57CFE1EF9B6FE441D0A" box="[410,493,1577,1601]" captionStart="TABLE 1" captionStartId="14.[159,243,160,181]" captionText="TABLE 1. (continued)" pageId="9" pageNumber="46">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="04DC652DCF21A57CFF13F9EEFD621D9A" pageId="9" pageNumber="46" type="materials_examined">
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<paragraph id="4C7936A6CF21A57CFF13F9EEFD621D9A" blockId="9.[151,1436,1648,2033]" pageId="9" pageNumber="46">
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<emphasis id="7EB2EAB4CF21A57CFF13F9EEFEEC1DC2" bold="true" box="[151,325,1648,1673]" pageId="9" pageNumber="46">
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<typeStatus id="937D8804CF21A57CFF13F9EEFF7B1DC2" box="[151,210,1649,1673]" pageId="9" pageNumber="46">Type</typeStatus>
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material.
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</emphasis>
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<emphasis id="7EB2EAB4CF21A57CFEC8F9EEFE151DC2" box="[332,444,1649,1673]" italics="true" pageId="9" pageNumber="46">
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<typeStatus id="937D8804CF21A57CFEC8F9EEFE1F1DC2" box="[332,438,1649,1673]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="34D17636CF21A57CFE40F9EEFD871DC2" box="[452,558,1649,1673]" name="Brazil" pageId="9" pageNumber="46">BRAZIL</collectingCountry>
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:
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<emphasis id="7EB2EAB4CF21A57CFDBCF9EDFCB91DC2" box="[568,784,1649,1673]" italics="true" pageId="9" pageNumber="46">State of Amazonas:</emphasis>
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adult female from Atalaia do Norte, Natural Reserve
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<taxonomicName id="8BC64D25CF21A57CFAEBF9EEFF641DE6" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">Palmari</taxonomicName>
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,
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<quantity id="8B3E9B43CF21A57CFF5FF90AFE8D1DE6" box="[219,292,1685,1709]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="46" unit="km" value="31.0">31 km</quantity>
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west of Benjamin Constant,
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<quantity id="8B3E9B43CF21A57CFDF2F909FD1B1DE6" box="[630,690,1686,1709]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="9" pageNumber="46" unit="m" value="77.0">77 m</quantity>
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elev., 04˚17’17”S 70˚17’33”W, UV light at night,
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<date id="38781066CF21A57CFB7AF90AFF7A1D9A" pageId="9" pageNumber="46" value="2009-09-23">23 September 2009</date>
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; preserved in 70% ethanol. MPUJ-SCO-397.
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</paragraph>
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</subSubSection>
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<subSubSection id="04DC652DCF21A57CFF41F942FBC81DBE" box="[197,1121,1757,1782]" pageId="9" pageNumber="46" type="etymology">
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<paragraph id="4C7936A6CF21A57CFF41F942FBC81DBE" blockId="9.[151,1436,1648,2033]" box="[197,1121,1757,1782]" pageId="9" pageNumber="46">
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<emphasis id="7EB2EAB4CF21A57CFF41F942FEE51DBE" bold="true" box="[197,332,1757,1781]" pageId="9" pageNumber="46">Etymology.</emphasis>
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The species name is a noun in apposition taken from the
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<typeStatus id="937D8804CF21A57CFC4BF941FC561DBD" box="[975,1023,1758,1782]" pageId="9" pageNumber="46">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="04DC652DCF21A57CFF43F89FFC391C52" box="[199,912,1792,1818]" pageId="9" pageNumber="46" type="distribution">
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<paragraph id="4C7936A6CF21A57CFF43F89FFC391C52" blockId="9.[151,1436,1648,2033]" box="[199,912,1792,1818]" pageId="9" pageNumber="46">
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<emphasis id="7EB2EAB4CF21A57CFF43F89FFEC91C52" bold="true" box="[199,352,1792,1817]" pageId="9" pageNumber="46">Distribution.</emphasis>
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Known only from the
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<typeStatus id="937D8804CF21A57CFDE4F89DFD391C51" box="[608,656,1794,1818]" pageId="9" pageNumber="46">type</typeStatus>
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locality (
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<figureCitation id="D4FD2A23CF21A57CFD7EF89EFC2D1C52" box="[762,900,1793,1817]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStartId-0="12.[151,250,2009,2032]" captionStartId-1="13.[151,250,846,869]" captionTargetBox-0="[156,1431,1025,1984]" captionTargetBox-1="[176,1414,196,822]" captionTargetId-0="figure@12.[153,1433,1021,1989]" captionTargetId-1="figure@13.[169,1418,193,825]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 39. Habitat of Ananteris palmari sp. nov. in the Natural Reserve Palmari, Brazil." captionText-1="FIGURE 40. Location of the type locality of Ananteris palmari sp. nov. (star)." httpUri-0="https://zenodo.org/record/201480/files/figure.png" httpUri-1="https://zenodo.org/record/201481/files/figure.png" pageId="9" pageNumber="46">Figs. 39–40</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="04DC652DCF21A57FFF41F8BBFE3A1ACC" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46" type="diagnosis">
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<paragraph id="4C7936A6CF21A57FFF41F8BBFE3A1ACC" blockId="9.[151,1436,1648,2033]" lastBlockId="10.[151,1436,151,392]" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
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<emphasis id="7EB2EAB4CF21A57CFF41F8BBFD891C76" bold="true" box="[197,544,1828,1853]" pageId="9" pageNumber="46">Diagnosis (based on female).</emphasis>
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In having a small body size and low teeth count,
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<taxonomicName id="8BC64D25CF21A57CFBE3F8BAFA9F1C76" box="[1127,1334,1829,1853]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
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<emphasis id="7EB2EAB4CF21A57CFBE3F8BAFA9F1C76" box="[1127,1334,1829,1853]" italics="true" pageId="9" pageNumber="46">Ananteris palmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="658157CFCF21A57CFAC4F8BAFA351C76" box="[1344,1436,1829,1853]" pageId="9" pageNumber="46" rank="species">
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<emphasis id="7EB2EAB4CF21A57CFAC4F8BAFA351C76" bold="true" box="[1344,1436,1829,1853]" pageId="9" pageNumber="46">sp. nov.</emphasis>
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</taxonomicNameLabel>
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resembles
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<taxonomicName id="8BC64D25CF21A57CFE8BF8D6FED91C2A" box="[271,368,1865,1889]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="festae">
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<emphasis id="7EB2EAB4CF21A57CFE8BF8D6FED91C2A" box="[271,368,1865,1889]" italics="true" pageId="9" pageNumber="46">A. festae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8BC64D25CF21A57CFE2CF8D6FCCA1C2A" authority="Lourenco, 1999" authorityName="Lourenco" authorityYear="1999" box="[424,867,1865,1889]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="mariaelenae">
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<emphasis id="7EB2EAB4CF21A57CFE2CF8D6FD001C2A" box="[424,681,1865,1889]" italics="true" pageId="9" pageNumber="46">Ananteris mariaelenae</emphasis>
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<bibRefCitation id="28574B57CF21A57CFD35F8D5FCCA1C2A" author="Lourenco" box="[689,867,1865,1889]" pageId="9" pageNumber="51" refString="Lourenco W. R. (1999) Some remarks about Ananteris festae Borelli, 1899 and description of a new species of Ananteris Thorell from Ecuador (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 13 (160), 95 - 100." type="journal article" year="1999">Lourenço, 1999</bibRefCitation>
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</taxonomicName>
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from
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<collectingCountry id="34D17636CF21A57CFC2DF8D6FBA11C2A" box="[937,1032,1865,1889]" name="Ecuador" pageId="9" pageNumber="46">Ecuador</collectingCountry>
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, and
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<taxonomicName id="8BC64D25CF21A57CFBC1F8D6FB011C2A" box="[1093,1192,1865,1889]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="minor">
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<emphasis id="7EB2EAB4CF21A57CFBC1F8D6FB011C2A" box="[1093,1192,1865,1889]" italics="true" pageId="9" pageNumber="46">A. minor</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="34D17636CF21A57CFB6AF8D6FA311C2A" box="[1262,1432,1865,1889]" name="French Guiana" pageId="9" pageNumber="46">French Guiana</collectingCountry>
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. From the former one,
|
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<taxonomicName id="8BC64D25CF21A57CFE0AF8F2FDAE1CCE" box="[398,519,1901,1925]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF21A57CFE0AF8F2FDAE1CCE" box="[398,519,1901,1925]" italics="true" pageId="9" pageNumber="46">A. palmari</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF21A57CFD89F8F2FDCE1CCE" box="[525,615,1901,1925]" pageId="9" pageNumber="46" rank="species">
|
||
<emphasis id="7EB2EAB4CF21A57CFD89F8F2FDCE1CCE" bold="true" box="[525,615,1901,1925]" pageId="9" pageNumber="46">sp. nov.</emphasis>
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</taxonomicNameLabel>
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differs in having a complete reticular pattern on dorsal surface of chelicerae (
|
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<figureCitation id="D4FD2A23CF21A57CFF50F80EFE8E1CE2" box="[212,295,1937,1961]" captionStart="FIGURES 31 – 36" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[168,1415,462,1902]" captionTargetId="figure@10.[160,1420,445,1914]" captionTargetPageId="10" captionText="FIGURES 31 – 36. Ananteris palmari sp. nov., female holotype. 31 – 35. Distribution of the trichobothria (illustrated from right pedipalp). 31. Femur, dorsal aspect. 32. Patella, dorsal aspect. 33. Patella, external aspect. 34. Chela, dorsoexternal aspect. 35. Chela, ventral aspect. 36. Right chelicera, dorsal aspect. Scale bars equal 0.5 mm." httpUri="https://zenodo.org/record/201478/files/figure.png" pageId="9" pageNumber="46">Fig. 36</figureCitation>
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),
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<emphasis id="7EB2EAB4CF21A57CFEBFF80EFEC91CE2" box="[315,352,1937,1961]" italics="true" pageId="9" pageNumber="46">esb</emphasis>
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trichobothrium basal to
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<emphasis id="7EB2EAB4CF21A57CFDF7F80EFD261CE2" box="[627,655,1937,1961]" italics="true" pageId="9" pageNumber="46">db</emphasis>
|
||
(
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<emphasis id="7EB2EAB4CF21A57CFD1AF80EFD101CE2" box="[670,697,1937,1961]" italics="true" pageId="9" pageNumber="46">eb</emphasis>
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||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFD45F80EFD4F1CE2" box="[705,742,1937,1961]" italics="true" pageId="9" pageNumber="46">esb</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFD6AF80EFCA31CE2" box="[750,778,1937,1961]" italics="true" pageId="9" pageNumber="46">db</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFC96F80DFC981CE2" box="[786,817,1938,1961]" italics="true" pageId="9" pageNumber="46">est</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFCBCF80DFCE51CE2" box="[824,844,1938,1961]" italics="true" pageId="9" pageNumber="46">et</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFCD0F80EFCC01CE2" box="[852,873,1937,1961]" italics="true" pageId="9" pageNumber="46">dt</emphasis>
|
||
) (
|
||
<figureCitation id="D4FD2A23CF21A57CFC07F80EFC7C1CE2" box="[899,981,1937,1961]" captionStart="FIGURES 31 – 36" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[168,1415,462,1902]" captionTargetId="figure@10.[160,1420,445,1914]" captionTargetPageId="10" captionText="FIGURES 31 – 36. Ananteris palmari sp. nov., female holotype. 31 – 35. Distribution of the trichobothria (illustrated from right pedipalp). 31. Femur, dorsal aspect. 32. Patella, dorsal aspect. 33. Patella, external aspect. 34. Chela, dorsoexternal aspect. 35. Chela, ventral aspect. 36. Right chelicera, dorsal aspect. Scale bars equal 0.5 mm." httpUri="https://zenodo.org/record/201478/files/figure.png" pageId="9" pageNumber="46">Fig. 34</figureCitation>
|
||
), and ten pectinal teeth (
|
||
<figureCitation id="D4FD2A23CF21A57CFB62F80EFA901CE2" box="[1254,1337,1937,1961]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1411,1434]" captionTargetBox="[153,1436,197,1389]" captionTargetId="figure@7.[151,1436,193,1390]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Left pectine of Ananteris palmari sp. nov., female holotype, under SEM. 19. Pectine. 20. Teeth. 21. Detail of teeth ornamentation. 22. Zoom of peg sensillae." httpUri="https://zenodo.org/record/201475/files/figure.png" pageId="9" pageNumber="46">Fig. 19</figureCitation>
|
||
); in contrast,
|
||
<taxonomicName id="8BC64D25CF21A57CFF51F82AFE9E1C86" box="[213,311,1973,1997]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="festae">
|
||
<emphasis id="7EB2EAB4CF21A57CFF51F82AFE9E1C86" box="[213,311,1973,1997]" italics="true" pageId="9" pageNumber="46">A. festae</emphasis>
|
||
</taxonomicName>
|
||
has immaculate chelicerae,
|
||
<emphasis id="7EB2EAB4CF21A57CFDF1F82AFD381C86" box="[629,657,1973,1997]" italics="true" pageId="9" pageNumber="46">db</emphasis>
|
||
basal to
|
||
<emphasis id="7EB2EAB4CF21A57CFD73F82AFCB51C86" box="[759,796,1973,1997]" italics="true" pageId="9" pageNumber="46">esb</emphasis>
|
||
(
|
||
<emphasis id="7EB2EAB4CF21A57CFCA9F82AFCE11C86" box="[813,840,1973,1997]" italics="true" pageId="9" pageNumber="46">eb</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFCCBF82AFCC21C86" box="[847,875,1973,1997]" italics="true" pageId="9" pageNumber="46">db</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFCF7F82AFC311C86" box="[883,920,1973,1997]" italics="true" pageId="9" pageNumber="46">esb</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFC24F829FC161C86" box="[928,959,1974,1997]" italics="true" pageId="9" pageNumber="46">est</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFC43F829FC721C86" box="[967,987,1974,1997]" italics="true" pageId="9" pageNumber="46">et</emphasis>
|
||
:
|
||
<emphasis id="7EB2EAB4CF21A57CFC67F82AFC501C86" box="[995,1017,1973,1997]" italics="true" pageId="9" pageNumber="46">dt</emphasis>
|
||
), and 11–14 teeth (
|
||
<quantity id="8B3E9B43CF21A57CFB57F82AFA9F1C86" box="[1235,1334,1973,1997]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="9" pageNumber="46" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
|
||
females,
|
||
<quantity id="8B3E9B43CF21A57CFF13F846FF791CBA" box="[151,208,2009,2033]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="46" unit="in" value="14.0">14 in</quantity>
|
||
males). The new species differs from
|
||
<taxonomicName id="8BC64D25CF21A57CFD07F846FC871CBA" box="[643,814,2009,2033]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="mariaelenae">
|
||
<emphasis id="7EB2EAB4CF21A57CFD07F846FC871CBA" box="[643,814,2009,2033]" italics="true" pageId="9" pageNumber="46">A. mariaelenae</emphasis>
|
||
</taxonomicName>
|
||
in having ten pectinal teeth, pedipalp hand and fingers brownish (
|
||
<figureCitation id="D4FD2A23CF22A57FFE95FF08FE351BE4" box="[273,412,151,175]" captionStart="FIGURES 37 – 38" captionStartId="11.[151,264,949,972]" captionTargetBox="[155,1431,193,927]" captionTargetId="figure@11.[155,1432,193,928]" captionTargetPageId="11" captionText="FIGURES 37 – 38. Female holotype of Ananteris palmari sp. nov., dorsal and ventral aspects. Scale bar equals 3 mm." httpUri="https://zenodo.org/record/201479/files/figure.png" pageId="10" pageNumber="47">Figs. 37–38</figureCitation>
|
||
), and smaller morphometric values (
|
||
<tableCitation id="0144031DCF22A57FFCBBFF08FC3D1BE4" box="[831,916,151,175]" captionStart="TABLE 1" captionStartId="14.[159,243,160,181]" captionText="TABLE 1. (continued)" pageId="10" pageNumber="47">Table 1</tableCitation>
|
||
); instead,
|
||
<taxonomicName id="8BC64D25CF22A57FFB8BFF07FB151BFB" box="[1039,1212,152,176]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="mariaelenae">
|
||
<emphasis id="7EB2EAB4CF22A57FFB8BFF07FB151BFB" box="[1039,1212,152,176]" italics="true" pageId="10" pageNumber="47">A. mariaelenae</emphasis>
|
||
</taxonomicName>
|
||
has 12–13 pectinal teeth, pedipalp chela with yellowish hand and brownish fingers, and larger morphometric values (female; male unknown). Finally,
|
||
<taxonomicName id="8BC64D25CF22A57FFEF7FF7FFE421BB3" box="[371,491,224,248]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF22A57FFEF7FF7FFE421BB3" box="[371,491,224,248]" italics="true" pageId="10" pageNumber="47">A. palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF22A57FFE76FF7FFDE51BB3" box="[498,588,224,248]" pageId="10" pageNumber="47" rank="species">
|
||
<emphasis id="7EB2EAB4CF22A57FFE76FF7FFDE51BB3" bold="true" box="[498,588,224,248]" pageId="10" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="8BC64D25CF22A57FFD65FF7FFCEA1BB3" box="[737,835,224,248]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="7EB2EAB4CF22A57FFD65FF7FFCEA1BB3" box="[737,835,224,248]" italics="true" pageId="10" pageNumber="47">A. minor</emphasis>
|
||
</taxonomicName>
|
||
because its pectinal peg sensillae are conical not surrounded by seta-like structures (
|
||
<figureCitation id="D4FD2A23CF22A57FFD85FE9BFD211A57" box="[513,648,260,284]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1411,1434]" captionTargetBox="[153,1436,197,1389]" captionTargetId="figure@7.[151,1436,193,1390]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Left pectine of Ananteris palmari sp. nov., female holotype, under SEM. 19. Pectine. 20. Teeth. 21. Detail of teeth ornamentation. 22. Zoom of peg sensillae." httpUri="https://zenodo.org/record/201475/files/figure.png" pageId="10" pageNumber="47">Figs. 21–22</figureCitation>
|
||
) and the coxosternal region has abundant brown mottling throughout, whereas in
|
||
<taxonomicName id="8BC64D25CF22A57FFE9EFEB7FED71A0B" box="[282,382,296,320]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="7EB2EAB4CF22A57FFE9EFEB7FED71A0B" box="[282,382,296,320]" italics="true" pageId="10" pageNumber="47">A. minor</emphasis>
|
||
</taxonomicName>
|
||
the pectinal peg sensillae are bottle-like surrounded by seta-like structures and the venter is pale-yellow (female; male unknown). See
|
||
<bibRefCitation id="28574B57CF22A57FFDF5FED2FC8E1A2F" author="Lourenco" box="[625,807,332,356]" pageId="10" pageNumber="51" refString="Lourenco, W. R. (1982) Revision du genre Ananteris Thorell, 1891 (Scorpiones, Buthidae) et description de six especes nouvelles. Bulletin du Museum National d'Histoire Naturelle, 4 (1 / 2), 119 - 151." type="journal article" year="1982">Lourenço (1982</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="28574B57CF22A57FFCB6FED3FCC41A2F" author="Lourenco" box="[818,877,332,356]" pageId="10" pageNumber="51" refString="Lourenco, W. R. (1988) Ananteris festae Borelli, espece de scorpion caracteristique du centre d'endemisme " Chimborazo " en Equateur. Journal of Arachnology, 16, 266 - 267." type="journal article" year="1988">1988</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="28574B57CF22A57FFCFCFED3FC1A1A2F" author="Lourenco" box="[888,947,332,356]" pageId="10" pageNumber="51" refString="Lourenco W. R. (1999) Some remarks about Ananteris festae Borelli, 1899 and description of a new species of Ananteris Thorell from Ecuador (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 13 (160), 95 - 100." type="journal article" year="1999">1999</bibRefCitation>
|
||
, 2003) for information on
|
||
<taxonomicName id="8BC64D25CF22A57FFB5EFED3FA901A2F" box="[1242,1337,332,356]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="festae">
|
||
<emphasis id="7EB2EAB4CF22A57FFB5EFED3FA901A2F" box="[1242,1337,332,356]" italics="true" pageId="10" pageNumber="47">A. festae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8BC64D25CF22A57FFAC3FED3FF5F1AC3" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="mariaelenae">
|
||
<emphasis id="7EB2EAB4CF22A57FFAC3FED3FF5F1AC3" italics="true" pageId="10" pageNumber="47">A. mariaelenae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8BC64D25CF22A57FFEA8FEEFFE251AC3" box="[300,396,368,392]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="7EB2EAB4CF22A57FFEA8FEEFFE251AC3" box="[300,396,368,392]" italics="true" pageId="10" pageNumber="47">A. minor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="18B9662ECF22A57FFF13F800FC7A1CB9" httpUri="https://zenodo.org/record/201478/files/figure.png" pageId="10" pageNumber="47" targetBox="[168,1415,462,1902]" targetPageId="10">
|
||
<paragraph id="4C7936A6CF22A57FFF13F800FC7A1CB9" blockId="10.[151,1436,1951,2034]" pageId="10" pageNumber="47">
|
||
<emphasis id="7EB2EAB4CF22A57FFF13F800FEFD1CFE" bold="true" box="[151,340,1951,1974]" pageId="10" pageNumber="47">FIGURES 31–36.</emphasis>
|
||
<taxonomicName id="8BC64D25CF22A57FFEDEF83FFDBA1CFE" box="[346,531,1951,1973]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF22A57FFEDEF83FFDBA1CFE" box="[346,531,1951,1973]" italics="true" pageId="10" pageNumber="47">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF22A57FFD9EF83FFDC31CFE" box="[538,618,1952,1973]" pageId="10" pageNumber="47" rank="species">
|
||
<emphasis id="7EB2EAB4CF22A57FFD9EF83FFDC31CFE" bold="true" box="[538,618,1952,1973]" pageId="10" pageNumber="47">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female holotype.
|
||
<emphasis id="7EB2EAB4CF22A57FFCADF83FFCC71CFE" bold="true" box="[809,878,1952,1973]" pageId="10" pageNumber="47">31–35.</emphasis>
|
||
Distribution of the trichobothria (illustrated from right pedipalp).
|
||
<emphasis id="7EB2EAB4CF22A57FFE80F821FE8A1C98" bold="true" box="[260,291,1982,2003]" pageId="10" pageNumber="47">31.</emphasis>
|
||
Femur, dorsal aspect.
|
||
<emphasis id="7EB2EAB4CF22A57FFD8DF821FD831C98" bold="true" box="[521,554,1982,2003]" pageId="10" pageNumber="47">32.</emphasis>
|
||
Patella, dorsal aspect.
|
||
<emphasis id="7EB2EAB4CF22A57FFC97F821FC9A1C98" bold="true" box="[787,819,1982,2003]" pageId="10" pageNumber="47">33.</emphasis>
|
||
Patella, external aspect.
|
||
<emphasis id="7EB2EAB4CF22A57FFBB4F821FBF91C98" bold="true" box="[1072,1104,1982,2003]" pageId="10" pageNumber="47">34.</emphasis>
|
||
Chela, dorsoexternal aspect.
|
||
<emphasis id="7EB2EAB4CF22A57FFAF8F821FA351C98" bold="true" box="[1404,1436,1982,2003]" pageId="10" pageNumber="47">35.</emphasis>
|
||
Chela, ventral aspect.
|
||
<emphasis id="7EB2EAB4CF22A57FFEF3F842FE3E1CB9" bold="true" box="[375,407,2013,2034]" pageId="10" pageNumber="47">36.</emphasis>
|
||
Right chelicera, dorsal aspect. Scale bars equal 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="18B9662ECF23A57EFF13FC2AFA961887" box="[151,1343,949,972]" httpUri="https://zenodo.org/record/201479/files/figure.png" pageId="11" pageNumber="48" targetBox="[155,1431,193,927]" targetPageId="11">
|
||
<paragraph id="4C7936A6CF23A57EFF13FC2AFA961887" blockId="11.[151,1343,949,972]" box="[151,1343,949,972]" pageId="11" pageNumber="48">
|
||
<emphasis id="7EB2EAB4CF23A57EFF13FC2AFEFD1880" bold="true" box="[151,340,949,972]" pageId="11" pageNumber="48">FIGURES 37–38.</emphasis>
|
||
Female holotype of
|
||
<taxonomicName id="8BC64D25CF23A57EFDA1FC28FD741887" box="[549,733,950,972]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF23A57EFDA1FC28FD741887" box="[549,733,950,972]" italics="true" pageId="11" pageNumber="48">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF23A57EFD60FC29FC9D1880" box="[740,820,950,971]" pageId="11" pageNumber="48" rank="species">
|
||
<emphasis id="7EB2EAB4CF23A57EFD60FC29FC9D1880" bold="true" box="[740,820,950,971]" pageId="11" pageNumber="48">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, dorsal and ventral aspects. Scale bar equals 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="04DC652DCF23A57BFF41FC6CFAA718B3" lastPageId="14" lastPageNumber="51" pageId="11" pageNumber="48" type="description">
|
||
<paragraph id="4C7936A6CF23A57EFF41FC6CFDC61D48" blockId="11.[151,1437,1011,2007]" pageId="11" pageNumber="48">
|
||
<emphasis id="7EB2EAB4CF23A57EFF41FC6CFD061F47" bold="true" box="[197,687,1011,1036]" pageId="11" pageNumber="48">
|
||
Description of the
|
||
<typeStatus id="937D8804CF23A57EFE2AFC6BFDBE1F47" box="[430,535,1012,1036]" pageId="11" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
. Coloration.
|
||
</emphasis>
|
||
General coloration brownish with variegated pigmentation over almost the entire body (except ventrally) and appendages (
|
||
<figureCitation id="D4FD2A23CF23A57EFCA9FB88FC111F64" box="[813,952,1047,1071]" captionStart="FIGURES 37 – 38" captionStartId="11.[151,264,949,972]" captionTargetBox="[155,1431,193,927]" captionTargetId="figure@11.[155,1432,193,928]" captionTargetPageId="11" captionText="FIGURES 37 – 38. Female holotype of Ananteris palmari sp. nov., dorsal and ventral aspects. Scale bar equals 3 mm." httpUri="https://zenodo.org/record/201479/files/figure.png" pageId="11" pageNumber="48">Figs. 37–38</figureCitation>
|
||
). Carapace predominantly brown with yellow bands and spots; median and lateral ocular tubercles black. Chelicerae with coxa yellow; hand with complete reticular pattern on dorsal surface (
|
||
<figureCitation id="D4FD2A23CF23A57EFD99FBC0FDD81F3C" box="[541,625,1119,1143]" captionStart="FIGURES 31 – 36" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[168,1415,462,1902]" captionTargetId="figure@10.[160,1420,445,1914]" captionTargetPageId="10" captionText="FIGURES 31 – 36. Ananteris palmari sp. nov., female holotype. 31 – 35. Distribution of the trichobothria (illustrated from right pedipalp). 31. Femur, dorsal aspect. 32. Patella, dorsal aspect. 33. Patella, external aspect. 34. Chela, dorsoexternal aspect. 35. Chela, ventral aspect. 36. Right chelicera, dorsal aspect. Scale bars equal 0.5 mm." httpUri="https://zenodo.org/record/201478/files/figure.png" pageId="11" pageNumber="48">Fig. 36</figureCitation>
|
||
); movable and fixed fingers almost completely brown but yellowish distally; teeth reddish-yellow. Coxosternal region with abundant brown mottling throughout, except on sternum which yellow. Genital operculum and pectines whitish and spotless; pectinal basal piece light-yellow with two small inconspicuous brownish areas; sternites III–V completely yellow, without marks; VI–VII with some brown regions in the anterior margin, larger and more conspicuous in sternite VII; sternite III with a rather slit-like posterior median hyaline area. Tergites I–VI with two (small anterior, larger posterior) yellow areas at each side of the midline; each side of the tergites with a yellow arrow-like design pointing to the midline; tergite VII predominantly yellow with abundant brown spots and mottling; lateral margins of the tergites predominantly yellow, brown anteriorly. Metasomal segments I–IV predominantly yellow, segment V light reddish, all with brown mottling. Telson yellowish, with inconspicuous light brown region ventrolaterally and aculeus red. Pedipalps brown throughout, with lighter brown mottling in all surfaces of each segment; trichobothrial insertions yellow. Legs with variegated pigmentation, telotarsi completely yellow.
|
||
</paragraph>
|
||
<paragraph id="4C7936A6CF23A579FF41F990FBE4188C" blockId="11.[151,1437,1011,2007]" lastBlockId="12.[151,1436,151,967]" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48">
|
||
<emphasis id="7EB2EAB4CF23A57EFF41F990FEF71D6C" bold="true" box="[197,350,1551,1575]" pageId="11" pageNumber="48">Morphology.</emphasis>
|
||
Measurements in
|
||
<tableCitation id="0144031DCF23A57EFDAAF990FD2B1D6C" box="[558,642,1551,1575]" captionStart="TABLE 1" captionStartId="14.[159,243,160,181]" captionText="TABLE 1. (continued)" pageId="11" pageNumber="48">Table 1</tableCitation>
|
||
.
|
||
<emphasis id="7EB2EAB4CF23A57EFD14F98FFCAE1D6C" box="[656,775,1552,1575]" italics="true" pageId="11" pageNumber="48">Carapace:</emphasis>
|
||
brown regions densely covered with conspicuous rounded granulation; lateral margins not parallel, converging anteriorly; anterior margin moderately, evenly concave (not biconcave); median ocular carinae evident, others inconspicuous; median depression anterior to the ocular tubercle and complete median longitudinal furrow evident; median ocular tubercle low, located on posterior portion of anterior third of carapace; lateral ocular tubercles each with three linearly-arranged ocelli (plus a fourth ocelli-like structure located behind the median lateral ocelli in the left tubercle).
|
||
<emphasis id="7EB2EAB4CF23A57EFC2AF95CFB9B1D90" box="[942,1074,1731,1755]" italics="true" pageId="11" pageNumber="48">Chelicerae:</emphasis>
|
||
with abundant setae on internal and ventral surfaces; cheliceral dentition characteristic of
|
||
<taxonomicName id="8BC64D25CF23A57EFC9BF978FB9A1DB4" authority="Vachon 1963" authorityName="Vachon" authorityYear="1963" box="[799,1075,1767,1791]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="48" phylum="Arthropoda" rank="family">
|
||
Buthidae (
|
||
<bibRefCitation id="28574B57CF23A57EFC16F978FB821DB4" author="Vachon" box="[914,1067,1767,1791]" pageId="11" pageNumber="52" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle Paris (2 e ser.), 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Movable finger externally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth; internally with two well-developed teeth, one basal and one median, and one distal tooth. Fixed finger externally with one basal and one median tooth mounted onto a bicuspid, one subdistal, and one distal tooth; internally with only one tooth.
|
||
<emphasis id="7EB2EAB4CF23A57EFEC8F8E8FD871CC4" box="[332,558,1911,1935]" italics="true" pageId="11" pageNumber="48">Coxosternal region:</emphasis>
|
||
predominantly smooth, with few setae except for coxapophyses densely hirsute; sternum subpentagonal, with deep median depression and two anterolateral furrows.
|
||
<emphasis id="7EB2EAB4CF23A57EFB13F804FF521C9C" italics="true" pageId="11" pageNumber="48">Genital operculum and pectines:</emphasis>
|
||
genital operculum divided longitudinally, formed by subtriangular pieces; pectinal basal piece slightly wider than long, deeply notched anteriorly and convex posteriorly; pectines very small, not reaching posterior end of coxae IV; segments count on the pectines: marginal lamellae 3:3, middle lamellae 7:7, teeth 10:10, fulcra absent; pectinal proximal teeth absent (likely due to sexual dimorphism), distal tooth rounded, basal piece of the middle lamellae markedly enlarged (
|
||
<figureCitation id="D4FD2A23CF24A579FE65FE9BFD9C1A57" box="[481,565,260,284]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1411,1434]" captionTargetBox="[153,1436,197,1389]" captionTargetId="figure@7.[151,1436,193,1390]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Left pectine of Ananteris palmari sp. nov., female holotype, under SEM. 19. Pectine. 20. Teeth. 21. Detail of teeth ornamentation. 22. Zoom of peg sensillae." httpUri="https://zenodo.org/record/201475/files/figure.png" pageId="12" pageNumber="49">Fig. 19</figureCitation>
|
||
); pectinal peg sensillae conical and not surrounded by seta-like structures as revealed by scanning electron microscopy (
|
||
<figureCitation id="D4FD2A23CF24A579FD05FEB8FCA51A74" box="[641,780,295,319]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1411,1434]" captionTargetBox="[153,1436,197,1389]" captionTargetId="figure@7.[151,1436,193,1390]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Left pectine of Ananteris palmari sp. nov., female holotype, under SEM. 19. Pectine. 20. Teeth. 21. Detail of teeth ornamentation. 22. Zoom of peg sensillae." httpUri="https://zenodo.org/record/201475/files/figure.png" pageId="12" pageNumber="49">Figs. 21–22</figureCitation>
|
||
).
|
||
<emphasis id="7EB2EAB4CF24A579FCA4FEB7FC231A74" box="[800,906,296,319]" italics="true" pageId="12" pageNumber="49">Sternites:</emphasis>
|
||
with few fine translucent setae; III–V smooth; VI–VII with few weak lateral granulation; VII with very fine and almost undistinguishable granulation posteromedially, suggesting vestigial and markedly incomplete submedian carinae; spiracles semi-oval to oval elongate.
|
||
<emphasis id="7EB2EAB4CF24A579FAE5FEEEFF6E1AE7" italics="true" pageId="12" pageNumber="49">Tergites:</emphasis>
|
||
with similar granulation to that of carapace; tergites I–VI with axial carina evident posteriorly, dorsolateral and lateral carinae absent; VII tetracarinate (paired dorsolateral and lateral carinae, incomplete), with low and granulose median elevation in the position of axial carina.
|
||
<emphasis id="7EB2EAB4CF24A579FD77FE42FCD81ABF" box="[755,881,477,500]" italics="true" pageId="12" pageNumber="49">Metasoma:</emphasis>
|
||
with abundant setae, mostly in ventral and lateral surfaces; segment I with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); II–III with ten (median lateral carinae only evident on anterior two thirds); IV with six (ventrosubmedian and median lateral carinae absent); segment V with three (vestigial ventromedian and paired dorsolateral carinae); all carinae serrulose, except for ventromedian on segment V which made up of separated granules; intercarinal spaces with conspicuous granulation, primarily on the brown regions. Telson smooth, with very low elevations ventrally suggesting the presence of three vestigial carinae; aculeus shorter than vesicle; subaculear tubercle strong, rhomboidal to spinoid.
|
||
<emphasis id="7EB2EAB4CF24A579FD27FD47FCB519BB" box="[675,796,728,752]" italics="true" pageId="12" pageNumber="49">Pedipalps:</emphasis>
|
||
femur with only three evident longitudinal carinae (dorsoexternal, dorsointernal and ventroexternal), weak and scarcely granulose, and intercarinal surfaces smooth; patella smooth and acarinate, except for vestigial dorsointernal carina present on distal half; chela smooth and acarinate; fixed and movable fingers each with six and seven almost linear rows of granules (including short apical row in movable finger), respectively, and basal row longer than others. Pedipalps orthobothriotaxic
|
||
<typeStatus id="937D8804CF24A579FB42FCF7FB5F18CB" box="[1222,1270,872,896]" pageId="12" pageNumber="49">type</typeStatus>
|
||
A, femur with β configuration (
|
||
<bibRefCitation id="28574B57CF24A579FED4FC13FE4E18EF" author="Vachon" box="[336,487,908,932]" pageId="12" pageNumber="52" refString="Vachon, M. (1974) Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national de Histoire naturelle, Paris (3 e ser.), 104, 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="28574B57CF24A579FE75FC13FD8318EF" author="Vachon" box="[497,554,908,932]" pageId="12" pageNumber="52" refString="Vachon, M. (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des seances de l'Academie des Sciences, Paris (ser. D) 281, 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
|
||
) (
|
||
<figureCitation id="D4FD2A23CF24A579FDC6FC13FD6318EF" box="[578,714,908,932]" captionStart="FIGURES 31 – 36" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[168,1415,462,1902]" captionTargetId="figure@10.[160,1420,445,1914]" captionTargetPageId="10" captionText="FIGURES 31 – 36. Ananteris palmari sp. nov., female holotype. 31 – 35. Distribution of the trichobothria (illustrated from right pedipalp). 31. Femur, dorsal aspect. 32. Patella, dorsal aspect. 33. Patella, external aspect. 34. Chela, dorsoexternal aspect. 35. Chela, ventral aspect. 36. Right chelicera, dorsal aspect. Scale bars equal 0.5 mm." httpUri="https://zenodo.org/record/201478/files/figure.png" pageId="12" pageNumber="49">Figs. 31–35</figureCitation>
|
||
).
|
||
<emphasis id="7EB2EAB4CF24A579FD59FC12FCB518EF" box="[733,796,909,932]" italics="true" pageId="12" pageNumber="49">Legs:</emphasis>
|
||
tibia, basitarsus and telotarsus with abundant setae; tibial spur present on legs III–IV; prolateral and retrolateral pedal spurs present on all legs.
|
||
</paragraph>
|
||
<caption id="18B9662ECF24A579FF13F846FB9B1CBB" box="[151,1074,2009,2032]" httpUri="https://zenodo.org/record/201480/files/figure.png" pageId="12" pageNumber="49" targetBox="[156,1431,1025,1984]" targetPageId="12">
|
||
<paragraph id="4C7936A6CF24A579FF13F846FB9B1CBB" blockId="12.[151,1074,2009,2032]" box="[151,1074,2009,2032]" pageId="12" pageNumber="49">
|
||
<emphasis id="7EB2EAB4CF24A579FF13F846FEB61CA4" bold="true" box="[151,287,2009,2032]" pageId="12" pageNumber="49">FIGURE 39.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="8BC64D25CF24A579FE16F844FDE31CBB" box="[402,586,2010,2032]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF24A579FE16F844FDE31CBB" box="[402,586,2010,2032]" italics="true" pageId="12" pageNumber="49">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF24A579FDD5F845FD081CA4" box="[593,673,2010,2031]" pageId="12" pageNumber="49" rank="species">
|
||
<emphasis id="7EB2EAB4CF24A579FDD5F845FD081CA4" bold="true" box="[593,673,2010,2031]" pageId="12" pageNumber="49">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the Natural Reserve
|
||
<taxonomicName id="8BC64D25CF24A579FC16F845FC4D1CBB" box="[914,996,2010,2032]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="50" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">Palmari</taxonomicName>
|
||
, Brazil.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="18B9662ECF25A578FF13FCD1FC10182E" box="[151,953,846,869]" httpUri="https://zenodo.org/record/201481/files/figure.png" pageId="13" pageNumber="50" targetBox="[176,1414,196,822]" targetPageId="13">
|
||
<paragraph id="4C7936A6CF25A578FF13FCD1FC10182E" blockId="13.[151,953,846,869]" box="[151,953,846,869]" pageId="13" pageNumber="50">
|
||
<emphasis id="7EB2EAB4CF25A578FF13FCD1FEB6182F" bold="true" box="[151,287,846,869]" pageId="13" pageNumber="50">FIGURE 40.</emphasis>
|
||
Location of the type locality of
|
||
<taxonomicName id="8BC64D25CF25A578FDE2FCCFFCB6182E" box="[614,799,847,869]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF25A578FDE2FCCFFCB6182E" box="[614,799,847,869]" italics="true" pageId="13" pageNumber="50">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF25A578FCA1FCD0FCDC182F" box="[805,885,847,868]" pageId="13" pageNumber="50" rank="species">
|
||
<emphasis id="7EB2EAB4CF25A578FCA1FCD0FCDC182F" bold="true" box="[805,885,847,868]" pageId="13" pageNumber="50">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(star).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4C7936A6CF25A578FF41FC0EFEDC18BA" blockId="13.[151,1436,913,1009]" pageId="13" pageNumber="50">
|
||
<emphasis id="7EB2EAB4CF25A578FF41FC0EFE0B18E1" bold="true" box="[197,418,913,938]" pageId="13" pageNumber="50">Sympatric species.</emphasis>
|
||
<taxonomicName id="8BC64D25CF25A578FE2DFC0DFDDC18E1" box="[425,629,914,938]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF25A578FE2DFC0DFDDC18E1" box="[425,629,914,938]" italics="true" pageId="13" pageNumber="50">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF25A578FDF8FC0DFD7C18E1" box="[636,725,914,938]" pageId="13" pageNumber="50" rank="species">
|
||
<emphasis id="7EB2EAB4CF25A578FDF8FC0DFD7C18E1" bold="true" box="[636,725,914,938]" pageId="13" pageNumber="50">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found in sympatry with other four scorpion species:
|
||
<taxonomicName id="8BC64D25CF25A578FADDFC0CFE371886" authority="Pocock, 1897" authorityName="Pocock" authorityYear="1897" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="silvestris">
|
||
<emphasis id="7EB2EAB4CF25A578FADDFC0CFF531885" italics="true" pageId="13" pageNumber="50">Tityus silvestris</emphasis>
|
||
Pocock, 1897
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8BC64D25CF25A578FE2EFC28FCA81886" authority="Lourenco, 1984" authorityName="Lourenco" authorityYear="1984" box="[426,769,949,974]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="bastosi">
|
||
<emphasis id="7EB2EAB4CF25A578FE2EFC28FDED1885" box="[426,580,950,974]" italics="true" pageId="13" pageNumber="50">Tityus bastosi</emphasis>
|
||
Lourenço, 1984
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8BC64D25CF25A578FC89FC28FC181885" authority="Atreus" authorityName="Atreus" box="[781,945,950,974]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7EB2EAB4CF25A578FC89FC28FC181885" box="[781,945,950,974]" italics="true" pageId="13" pageNumber="50">Tityus (Atreus)</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName id="8BC64D25CF25A578FB96FC29FB271885" box="[1042,1166,950,974]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7EB2EAB4CF25A578FB96FC29FB271885" box="[1042,1166,950,974]" italics="true" pageId="13" pageNumber="50">Chactopsis</emphasis>
|
||
</taxonomicName>
|
||
sp. (identifications performed by R.B.-T.).
|
||
</paragraph>
|
||
<paragraph id="4C7936A6CF25A578FF13FBBAFE8B1FD7" blockId="13.[151,1436,1057,1314]" pageId="13" pageNumber="50">
|
||
<emphasis id="7EB2EAB4CF25A578FF13FBBAFEAD1F71" bold="true" box="[151,260,1061,1082]" pageId="13" pageNumber="50">
|
||
<tableCitation id="0144031DCF25A578FF13FBBAFF571F71" box="[151,254,1061,1082]" captionStart="TABLE 1" captionStartId="14.[159,243,160,181]" captionText="TABLE 1. (continued)" pageId="13" pageNumber="50">TABLE 1</tableCitation>
|
||
.
|
||
</emphasis>
|
||
Selected morphometric measurements (in millimeters) of
|
||
<taxonomicName id="8BC64D25CF25A578FCD6FBB9FBA21F70" box="[850,1035,1061,1083]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="palmari" status="sp. nov.">
|
||
<emphasis id="7EB2EAB4CF25A578FCD6FBB9FBA21F70" box="[850,1035,1061,1083]" italics="true" pageId="13" pageNumber="50">Ananteris palmari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="658157CFCF25A578FB95FBBAFBC81F71" box="[1041,1121,1061,1082]" pageId="13" pageNumber="50" rank="species">
|
||
<emphasis id="7EB2EAB4CF25A578FB95FBBAFBC81F71" bold="true" box="[1041,1121,1061,1082]" pageId="13" pageNumber="50">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female
|
||
<typeStatus id="937D8804CF25A578FB3DFBBAFABC1F70" box="[1209,1301,1061,1083]" pageId="13" pageNumber="50" type="holotype">holotype</typeStatus>
|
||
. 1Sum of carapace, mesosoma and metasoma. 2Sum of tergites
|
||
<date id="38781066CF25A578FD1EFBD8FD4F1F04" box="[666,742,1091,1116]" pageId="13" pageNumber="50">I–VII. 3</date>
|
||
Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella.
|
||
</paragraph>
|
||
<paragraph id="4C7936A6CF25A578FF1BFB25FAA91CF7" blockId="13.[151,1436,1057,1314]" lastBlockId="13.[159,1431,1504,2022]" pageId="13" pageNumber="50">Total body length:1 14.75 Carapace: length: 1.84 anterior width: 1.13 width: 1.05 depth: 0.87 Metasomal segment II: length: 0.89 width: 0.97 depth: 0.87 Metasomal segment III: length: 1.01 width: 0.95 depth: 0.87 Metasomal segment IV: length: 1.36 width: 0.91 depth: 0.89 Metasomal segment V: length: 2.39</paragraph>
|
||
<paragraph id="4C7936A6CF25A578FD3AFAA9FAA91E87" pageId="13" pageNumber="50">
|
||
<table id="3EC6C406CF255A8AFF13FAA9FA351E86" box="[151,1436,1334,1485]" gridcols="3" gridrows="4" pageId="13" pageNumber="50">
|
||
<tr id="F2F634E4CF255A8AFF13FAA9FA351E07" box="[151,1436,1334,1356]" gridrow="0" pageId="13" pageNumber="50" rowspan-0="1">
|
||
<th id="B1275D98CF255A8AFD3AFAA9FCC91E07" box="[702,864,1334,1356]" gridcol="1" gridrow="0" pageId="13" pageNumber="50">posterior width:</th>
|
||
<th id="B1275D98CF255A8AFB50FAA9FA351E07" box="[1236,1436,1334,1356]" gridcol="2" gridrow="0" pageId="13" pageNumber="50">1.75</th>
|
||
</tr>
|
||
<tr id="F2F634E4CF255A8AFF13FAC0FA351E33" box="[151,1436,1375,1400]" gridrow="1" pageId="13" pageNumber="50">
|
||
<th id="B1275D98CF255A8AFF13FAC0FE291E33" box="[151,384,1375,1400]" gridcol="0" gridrow="1" pageId="13" pageNumber="50">Mesosoma:</th>
|
||
<td id="B1275D98CF255A8AFD3AFAC0FCC91E33" box="[702,864,1375,1400]" gridcol="1" gridrow="1" pageId="13" pageNumber="50">total length:2</td>
|
||
<td id="B1275D98CF255A8AFB50FAC0FA351E33" box="[1236,1436,1375,1400]" gridcol="2" gridrow="1" pageId="13" pageNumber="50">4.12</td>
|
||
</tr>
|
||
<tr id="F2F634E4CF255A8AFF13FA14FA351EE8" box="[151,1436,1419,1443]" gridrow="2" pageId="13" pageNumber="50">
|
||
<th id="B1275D98CF255A8AFF13FA14FE291EE8" box="[151,384,1419,1443]" gridcol="0" gridrow="2" pageId="13" pageNumber="50">Metasoma:</th>
|
||
<td id="B1275D98CF255A8AFD3AFA14FCC91EE8" box="[702,864,1419,1443]" gridcol="1" gridrow="2" pageId="13" pageNumber="50">total length:3</td>
|
||
<td id="B1275D98CF255A8AFB50FA14FA351EE8" box="[1236,1436,1419,1443]" gridcol="2" gridrow="2" pageId="13" pageNumber="50">8.79</td>
|
||
</tr>
|
||
<tr id="F2F634E4CF255A8AFF13FA28FA351E86" box="[151,1436,1463,1485]" gridrow="3" pageId="13" pageNumber="50">
|
||
<th id="B1275D98CF255A8AFF13FA28FE291E86" box="[151,384,1463,1485]" gridcol="0" gridrow="3" pageId="13" pageNumber="50">Metasomal segment I:</th>
|
||
<td id="B1275D98CF255A8AFD3AFA28FCC91E86" box="[702,864,1463,1485]" gridcol="1" gridrow="3" pageId="13" pageNumber="50">length:</td>
|
||
<td id="B1275D98CF255A8AFB50FA28FA351E86" box="[1236,1436,1463,1485]" gridcol="2" gridrow="3" pageId="13" pageNumber="50">0.83</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="4C7936A6CF25A57BFB4FF84FFAA718B3" blockId="13.[159,1431,1504,2022]" lastBlockId="14.[159,1294,160,1016]" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50">continued next page width: 0.97 depth: 0.91 Telson: length: 2.31 vesicle width: 0.74 vesicle depth: 0.64 Pedipalp: total length:4 6.21 Pedipalp femur: length: 1.63 width: 0.49 depth: 0.41 Pedipalp patella: length: 2.08 width: 0.60 depth: 0.52 Pedipalp chela: length: 2.50 width: 0.41 depth: 0.43 fixed finger length:5 1.59 movable finger length: 1.83 palm length: 0.64 Pectines: length:6 1.13 teeth count (left/right): 10/10</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |