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<document id="69A0E0ED309028E4C7C45374268EDB8F" ID-DOI="10.11646/zootaxa.3722.2.9" ID-GBIF-Dataset="ca2d5b4e-4a87-4c98-ab92-d94875621d5f" ID-ISSN="1175-5326" ID-Zenodo-Dep="216561" ID-ZooBank="9D347B8C-0BCE-47A7-99DF-DBA9A38A4F44" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460691777826" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Paulay, Gustav" docDate="2013" docId="C445540AFFF410637AEBFC877D36A46A" docLanguage="en" docName="zt03722p300.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patagurus Anker &amp; Paulay, 2013, gen. nov." docType="treatment" docVersion="8" lastPageNumber="291" masterDocId="387C2C72FFF5106B7A7CFFE57F27A60B" masterDocTitle="A remarkable new crab-like hermit crab (Decapoda: Paguridae) from French Polynesia, with comments on carcinization in the Anomura" masterLastPageNumber="300" masterPageNumber="283" pageNumber="284" updateTime="1698351854519" updateUser="plazi">
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<mods:title id="47C5ED82178C9D5C6CC863CDEFB1FAFF">A remarkable new crab-like hermit crab (Decapoda: Paguridae) from French Polynesia, with comments on carcinization in the Anomura</mods:title>
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<mods:namePart id="C11F8229A3AF79BB884258B4DF03A5AC">Anker, Arthur</mods:namePart>
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<mods:namePart id="6BC10BE2D3666AB3DC8A0EDB9FBC955E">Paulay, Gustav</mods:namePart>
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<treatment id="C445540AFFF410637AEBFC877D36A46A" ID-DOI="http://doi.org/10.5281/zenodo.6154629" ID-GBIF-Taxon="119580089" ID-Zenodo-Dep="6154629" LSID="urn:lsid:plazi:treatment:C445540AFFF410637AEBFC877D36A46A" httpUri="http://treatment.plazi.org/id/C445540AFFF410637AEBFC877D36A46A" lastPageId="8" lastPageNumber="291" pageId="1" pageNumber="284">
<subSubSection id="04F6B697FFF4106A7AEBFC877EAEA577" box="[151,393,866,892]" pageId="1" pageNumber="284" type="nomenclature">
<paragraph id="4C53E51CFFF4106A7AEBFC877EAEA577" blockId="1.[151,393,866,892]" box="[151,393,866,892]" pageId="1" pageNumber="284">
<heading id="171B5270FFF4106A7AEBFC877EAEA577" bold="true" box="[151,393,866,892]" fontSize="11" level="1" pageId="1" pageNumber="284" reason="1">
<emphasis id="7E98390EFFF4106A7AEBFC877EAEA577" bold="true" box="[151,393,866,892]" pageId="1" pageNumber="284">
<taxonomicName id="8BEC9E9FFFF4106A7AEBFC877E33A577" ID-CoL="99382305-a477-4fee-bdd1-55bfa55f56a0" box="[151,276,866,892]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7E98390EFFF4106A7AEBFC877E33A577" bold="true" box="[151,276,866,892]" italics="true" pageId="1" pageNumber="284">Patagurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65AB8475FFF4106A7B67FC877EAEA577" box="[283,393,866,892]" pageId="1" pageNumber="284" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF4106A7AEBFC4C7C90A379" pageId="1" pageNumber="284" type="diagnosis">
<paragraph id="4C53E51CFFF4106A7AEBFC4C7C90A379" blockId="1.[151,1437,937,2006]" pageId="1" pageNumber="284">
<emphasis id="7E98390EFFF4106A7AEBFC4C7E36A5C9" bold="true" box="[151,273,937,962]" pageId="1" pageNumber="284">Diagnosis.</emphasis>
Anterior carapace vaulted, well calcified, gastric region slightly elevated; rostrum subtriangular; lateral margins each developed into two relatively small, subacute lobes; cervical groove clearly delineated; posterior carapace lobes very prominent, laterally projecting; posterior carapace calcified except for a narrow, posterior, membranous band tucked under calcified portion; branchiostegite with two large and two small calcified plates. Ocular acicles vestigial. Third maxillipeds with well-developed crista dentata and one accessory tooth. Ambulatory (second and third) pereopods similar, slender, with elongate dactyli. Fourth pereopod with rasp comprised of a single row of long, slender spines. Fifth pereopod non-chelate, with no discernible rasp and very small dactylus. Thoracic sternites rigidly articulated through sternite VII. Males with elongate right sexual tube directed mesially, left sexual tube shorter and stouter than right tube, both calcified and with long tuft of setae distally. Pleon narrow, elongate, segmentation of anterior pleomeres not discernable (tergal plates not delineated), segmentation of fifth pleomere faintly discernible, sixth tergite clearly defined, but feebly calcified; male pleopods 2–5 absent. Uropods well developed, symmetrical. Telson subquadrate, entire, folded firmly against ventral surface of sixth pleomere; position of anus not determined. Female characters presently unknown.
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF4106A7ABBFA987C5EA39D" box="[199,889,1405,1430]" pageId="1" pageNumber="284" type="materials_examined">
<paragraph id="4C53E51CFFF4106A7ABBFA987C5EA39D" blockId="1.[151,1437,937,2006]" box="[199,889,1405,1430]" pageId="1" pageNumber="284">
<emphasis id="7E98390EFFF4106A7ABBFA987E45A39D" bold="true" box="[199,354,1405,1430]" pageId="1" pageNumber="284">
<typeStatus id="93575BBEFFF4106A7ABBFA987E25A39D" box="[199,258,1405,1430]" pageId="1" pageNumber="284">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8BEC9E9FFFF4106A7B15FA9B7D21A39E" box="[361,518,1406,1429]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="7E98390EFFF4106A7B15FA9B7D21A39E" box="[361,518,1406,1429]" italics="true" pageId="1" pageNumber="284">Patagurus rex</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF4106A7872FA987D4FA39D" bold="true" box="[526,616,1405,1430]" pageId="1" pageNumber="284">
<taxonomicNameLabel id="65AB8475FFF4106A7872FA987D4FA39D" box="[526,616,1405,1430]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF4106A7ABBFA447FF1A02D" pageId="1" pageNumber="284" type="etymology">
<paragraph id="4C53E51CFFF4106A7ABBFA447FF1A02D" blockId="1.[151,1437,937,2006]" pageId="1" pageNumber="284">
<emphasis id="7E98390EFFF4106A7ABBFA447E69A3B1" bold="true" box="[199,334,1441,1466]" pageId="1" pageNumber="284">Etymology.</emphasis>
The new genus is named after one of the world’s most eminent carcinologists, the late Dr. Patsy (Pat) A. McLaughlin (
<date id="3852C3DCFFF4106A7BE8FA237D3FA3D5" box="[404,536,1478,1502]" pageId="1" pageNumber="284" value="1932" valueMax="2011">1932–2011</date>
). The generic name is a combination of McLaughlin’s preferred first name and the Greek word
<emphasis id="7E98390EFFF4106A7B30FA0F7E91A00A" box="[332,438,1514,1537]" italics="true" pageId="1" pageNumber="284">pagourus</emphasis>
(a kind of crab), which was the origin for the nominal pagurid genus,
<taxonomicName id="8BEC9E9FFFF4106A7EBBFA0F7FF5A02D" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7EBBFA0F7A01A00A" box="[1223,1318,1514,1537]" italics="true" pageId="1" pageNumber="284">Pagurus</emphasis>
Fabricius, 1775
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF410687ABBF9D47E67A55E" lastPageId="3" lastPageNumber="286" pageId="1" pageNumber="284" type="discussion">
<paragraph id="4C53E51CFFF410697ABBF9D47E34A76F" blockId="1.[151,1437,937,2006]" lastBlockId="2.[151,1436,151,356]" lastPageId="2" lastPageNumber="285" pageId="1" pageNumber="284">
<emphasis id="7E98390EFFF4106A7ABBF9D47E1CA041" bold="true" box="[199,315,1585,1610]" pageId="1" pageNumber="284">Remarks.</emphasis>
The number of gills, one of the differentiating characters between
<taxonomicName id="8BEC9E9FFFF4106A7E58F9D47BDCA041" box="[1060,1275,1585,1610]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7E58F9D47BDCA041" box="[1060,1275,1585,1610]" italics="true" pageId="1" pageNumber="284">Porcellanopagurus</emphasis>
</taxonomicName>
(11 pairs) and
<taxonomicName id="8BEC9E9FFFF4106A7AEBF9B07E72A065" box="[151,341,1621,1646]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7AEBF9B07E72A065" box="[151,341,1621,1646]" italics="true" pageId="1" pageNumber="284">Solitariopagurus</emphasis>
</taxonomicName>
(10 pairs), could not be determined in
<emphasis id="7E98390EFFF4106A7935F9B37CFDA066" box="[841,986,1622,1645]" italics="true" pageId="1" pageNumber="284">Patapagurus</emphasis>
, because penetration of the calcified branchiostegites would have resulted in serious damage to the branchial region of the single
<typeStatus id="93575BBEFFF4106A7EBCF99C7A05A099" box="[1216,1314,1657,1682]" pageId="1" pageNumber="284" type="holotype">holotype</typeStatus>
specimen. The dorsal surface of the carapace of
<taxonomicName id="8BEC9E9FFFF4106A7840F97B7D96A0BE" box="[572,689,1694,1717]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7840F97B7D96A0BE" box="[572,689,1694,1717]" italics="true" pageId="1" pageNumber="284">Patagurus</emphasis>
</taxonomicName>
(
<figureCitation id="D4D7F999FFF4106A78BCF9787DD9A0BD" box="[704,766,1693,1718]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="1" pageNumber="284">Fig. 1</figureCitation>
C) is most similar to those of species of
<taxonomicName id="8BEC9E9FFFF4106A7EB9F9787ABBA0BD" box="[1221,1436,1693,1718]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7EB9F9787ABBA0BD" box="[1221,1436,1693,1718]" italics="true" pageId="1" pageNumber="284">Porcellanopagurus</emphasis>
</taxonomicName>
(
<figureCitation id="D4D7F999FFF4106A7AE3F9247FC3A0D1" box="[159,228,1729,1754]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="1" pageNumber="284">Fig. 1</figureCitation>
A): they both generally have a pair of small lateral projections, two pairs of relatively small anterior carapace lobes, and a pair of more prominent posterior carapace lobes, which are posterior to, and clearly separated by, the cervical groove. However, the sclerotization of the posterior carapace is much more extensive in
<taxonomicName id="8BEC9E9FFFF4106A7F5CF8EF7AB3A12A" box="[1312,1428,1802,1825]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7F5CF8EF7AB3A12A" box="[1312,1428,1802,1825]" italics="true" pageId="1" pageNumber="284">Patagurus</emphasis>
</taxonomicName>
, leaving only a narrow, posterior, membranous band. In contrast,
<taxonomicName id="8BEC9E9FFFF4106A79FDF8C87B18A14D" box="[897,1087,1837,1862]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A79FDF8C87B18A14D" box="[897,1087,1837,1862]" italics="true" pageId="1" pageNumber="284">Solitariopagurus</emphasis>
</taxonomicName>
(
<figureCitation id="D4D7F999FFF4106A7E2CF8C87BB5A14D" box="[1104,1170,1837,1862]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="1" pageNumber="284">Fig. 1</figureCitation>
B) has a pair of strong lateral projections, three pairs of anterior carapace lobes, and a pair of small posterior carapace lobes. The dorsal surface of the shield is unarmed in all species of
<taxonomicName id="8BEC9E9FFFF4106A78A8F8907C8CA185" box="[724,939,1909,1934]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A78A8F8907C8CA185" box="[724,939,1909,1934]" italics="true" pageId="1" pageNumber="284">Porcellanopagurus</emphasis>
</taxonomicName>
(e.g., Borradaile 1916; Martin
<emphasis id="7E98390EFFF4106A7F6BF8937A75A185" box="[1303,1362,1909,1934]" italics="true" pageId="1" pageNumber="284">et al.</emphasis>
2009; McLaughlin 2000), as well as in the
<typeStatus id="93575BBEFFF4106A784EF87F7D45A1B9" box="[562,610,1946,1970]" pageId="1" pageNumber="284">type</typeStatus>
species of
<taxonomicName id="8BEC9E9FFFF4106A789CF87C7CB9A1B9" box="[736,926,1945,1970]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A789CF87C7CB9A1B9" box="[736,926,1945,1970]" italics="true" pageId="1" pageNumber="284">Solitariopagurus</emphasis>
</taxonomicName>
,
<taxonomicName id="8BEC9E9FFFF4106A79D0F87F7BFFA1B9" authority="Turkay, 1986" authorityName="Turkay" authorityYear="1986" box="[940,1240,1945,1970]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="profundus">
<emphasis id="7E98390EFFF4106A79D0F87F7B1EA1B9" box="[940,1081,1945,1970]" italics="true" pageId="1" pageNumber="284">S. profundus</emphasis>
Türkay, 1986
</taxonomicName>
. In the remaining three species of
<taxonomicName id="8BEC9E9FFFF4106A7B31F8587D2CA1DD" box="[333,523,1981,2006]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF4106A7B31F8587D2CA1DD" box="[333,523,1981,2006]" italics="true" pageId="1" pageNumber="284">Solitariopagurus</emphasis>
</taxonomicName>
, the shield is armed with a row of large tubercles behind the anterior margin and occasionally also by a few smaller tubercles elsewhere (Poupin &amp; McLaughlin 1996; McLaughlin 1997; McLaughlin 2000; Martin
<emphasis id="7E98390EFFF710697BB7FF587D21A6DE" box="[459,518,188,213]" italics="true" pageId="2" pageNumber="285">et al.</emphasis>
2009).
<taxonomicName id="8BEC9E9FFFF71069781EFF587DF0A6DF" box="[610,727,189,212]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF71069781EFF587DF0A6DF" box="[610,727,189,212]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
(
<figureCitation id="D4D7F999FFF710697895FF597C0DA6DF" box="[745,810,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="2" pageNumber="285">Fig. 1</figureCitation>
C) not only has the surface of the shield, but also the surfaces of the well-developed posterior carapace lobes covered with an abundance of tubercles of various sizes. Although the posterior carapace is well developed in all species of
<taxonomicName id="8BEC9E9FFFF7106979F9FEE17B7BA716" box="[901,1116,260,285]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF7106979F9FEE17B7BA716" box="[901,1116,260,285]" italics="true" pageId="2" pageNumber="285">Porcellanopagurus</emphasis>
</taxonomicName>
, it is reduced in members of
<taxonomicName id="8BEC9E9FFFF710697AEBFEC27E72A74B" box="[151,341,295,320]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697AEBFEC27E72A74B" box="[151,341,295,320]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
and presumably partly reduced and partly fused to the posterior lobes in the
<typeStatus id="93575BBEFFF710697E91FECD7A3AA74B" box="[1261,1309,296,320]" pageId="2" pageNumber="285">type</typeStatus>
species of
<taxonomicName id="8BEC9E9FFFF710697AEBFEA87E2BA76F" box="[151,268,333,356]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697AEBFEA87E2BA76F" box="[151,268,333,356]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="1893B594FFF710697AEBF9627B49A0D1" httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="2" pageNumber="285" targetBox="[164,1420,430,1584]" targetPageId="2">
<paragraph id="4C53E51CFFF710697AEBF9627B49A0D1" blockId="2.[151,1436,1671,1754]" pageId="2" pageNumber="285">
<emphasis id="7E98390EFFF710697AEBF9627E29A097" bold="true" box="[151,270,1671,1693]" pageId="2" pageNumber="285">FIGURE 1</emphasis>
. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A,
<taxonomicName id="8BEC9E9FFFF710697EAEF9627AB2A096" box="[1234,1429,1671,1693]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697EAEF9627AB2A096" box="[1234,1429,1671,1693]" italics="true" pageId="2" pageNumber="285">Porcellanopagurus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="8BEC9E9FFFF710697AC9F9407E47A0B0" box="[181,352,1701,1723]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697AC9F9407E47A0B0" box="[181,352,1701,1723]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
; C,
<taxonomicName id="8BEC9E9FFFF710697BF1F9427ED0A0B7" box="[397,503,1703,1724]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7E98390EFFF710697BF1F9427ED0A0B7" box="[397,503,1703,1724]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF710697B81F9437D7BA0B0" bold="true" box="[509,604,1702,1723]" pageId="2" pageNumber="285">
<taxonomicNameLabel id="65AB8475FFF710697B81F9437D7BA0B0" box="[509,604,1702,1723]" pageId="2" pageNumber="285" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
Figures not to scale. Abbreviations: ac1, ac2, ac3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum.
</paragraph>
</caption>
<paragraph id="4C53E51CFFF710687ABBF8E47D8CA6DE" blockId="2.[151,1436,1793,2034]" lastBlockId="3.[151,1437,151,717]" lastPageId="3" lastPageNumber="286" pageId="2" pageNumber="285">
The thoracic sternites of
<taxonomicName id="8BEC9E9FFFF710697BA3F8E67D74A111" box="[479,595,1795,1818]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697BA3F8E67D74A111" box="[479,595,1795,1818]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
are quite similar to those of
<taxonomicName id="8BEC9E9FFFF7106979E5F8E47B70A111" box="[921,1111,1793,1818]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF7106979E5F8E47B70A111" box="[921,1111,1793,1818]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
and both differ considerably from the sternites of
<taxonomicName id="8BEC9E9FFFF710697B01F8C07D73A135" box="[381,596,1829,1854]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697B01F8C07D73A135" box="[381,596,1829,1854]" italics="true" pageId="2" pageNumber="285">Porcellanopagurus</emphasis>
</taxonomicName>
. In
<taxonomicName id="8BEC9E9FFFF710697803F8C37DD3A136" box="[639,756,1830,1853]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697803F8C37DD3A136" box="[639,756,1830,1853]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="8BEC9E9FFFF710697955F8C07CC0A135" box="[809,999,1829,1854]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697955F8C07CC0A135" box="[809,999,1829,1854]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
, sternites III through VII are broad and rigidly articulated, with sternite VII flexed dorsally at a right angle relative to sternite VI. Similarly, asymmetrically paired male sexual tubes are seen in
<taxonomicName id="8BEC9E9FFFF710697841F88B7D96A18E" box="[573,689,1902,1925]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697841F88B7D96A18E" box="[573,689,1902,1925]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
and in all four species of
<taxonomicName id="8BEC9E9FFFF71069799DF8887BB8A18D" box="[993,1183,1901,1926]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF71069799DF8887BB8A18D" box="[993,1183,1901,1926]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
, but are lacking in all species of
<taxonomicName id="8BEC9E9FFFF710697B66F8747ED6A1A1" box="[282,497,1937,1962]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697B66F8747ED6A1A1" box="[282,497,1937,1962]" italics="true" pageId="2" pageNumber="285">Porcellanopagurus</emphasis>
</taxonomicName>
. Although nothing is known about the female secondary sexual characters of
<taxonomicName id="8BEC9E9FFFF710697AEBF8527E2BA1C5" box="[151,268,1975,1998]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697AEBF8527E2BA1C5" box="[151,268,1975,1998]" italics="true" pageId="2" pageNumber="285">Patagurus</emphasis>
</taxonomicName>
, females of
<taxonomicName id="8BEC9E9FFFF710697BE9F8507D4BA1C5" box="[405,620,1973,1998]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697BE9F8507D4BA1C5" box="[405,620,1973,1998]" italics="true" pageId="2" pageNumber="285">Porcellanopagurus</emphasis>
</taxonomicName>
all have paired gonopores, whilst females of
<taxonomicName id="8BEC9E9FFFF710697E16F8507A0FA1C5" box="[1130,1320,1973,1998]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697E16F8507A0FA1C5" box="[1130,1320,1973,1998]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
have only a single left gonopore. Propodal rasps are present on the fifth pereopod in
<taxonomicName id="8BEC9E9FFFF710697E5CF83C7BF9A1F9" box="[1056,1246,2009,2034]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF710697E5CF83C7BF9A1F9" box="[1056,1246,2009,2034]" italics="true" pageId="2" pageNumber="285">Solitariopagurus</emphasis>
</taxonomicName>
, but are lost in
<taxonomicName id="8BEC9E9FFFF610687AEBFF7C7E2BA6BB" box="[151,268,153,176]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687AEBFF7C7E2BA6BB" box="[151,268,153,176]" italics="true" pageId="3" pageNumber="286">Patagurus</emphasis>
</taxonomicName>
. Rasps on the fourth pereopod are modified to a single row of sharp, corneous spines in both genera, rather than well-developed as in most pagurids.
</paragraph>
<paragraph id="4C53E51CFFF610687ABBFF057D62A734" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="286">
In species of
<taxonomicName id="8BEC9E9FFFF610687B23FF3A7D11A6F3" box="[351,566,223,248]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687B23FF3A7D11A6F3" box="[351,566,223,248]" italics="true" pageId="3" pageNumber="286">Porcellanopagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="8BEC9E9FFFF61068780CFF3A7C09A6F3" box="[624,814,223,248]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF61068780CFF3A7C09A6F3" box="[624,814,223,248]" italics="true" pageId="3" pageNumber="286">Solitariopagurus</emphasis>
</taxonomicName>
, even in the largest specimens examined (McLaughlin 2000; de Saint Laurent &amp; McLaughlin 2000), the right cheliped was not shorter than the left, as is the case in the male of the
<typeStatus id="93575BBEFFF610687B60FECD7E6BA74B" box="[284,332,296,320]" pageId="3" pageNumber="286">type</typeStatus>
species of
<taxonomicName id="8BEC9E9FFFF610687BB5FECC7D19A74B" box="[457,574,297,320]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687BB5FECC7D19A74B" box="[457,574,297,320]" italics="true" pageId="3" pageNumber="286">Patagurus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4C53E51CFFF610687ABBFEA97B0FA4C6" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="286">
The number and
<typeStatus id="93575BBEFFF610687BE5FEA87EEEA76E" box="[409,457,333,357]" pageId="3" pageNumber="286">type</typeStatus>
of gills and several important female characters remain unknown for
<taxonomicName id="8BEC9E9FFFF610687F5CFEA87AB3A76F" box="[1312,1428,333,356]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687F5CFEA87AB3A76F" box="[1312,1428,333,356]" italics="true" pageId="3" pageNumber="286">Patagurus</emphasis>
</taxonomicName>
. Therefore, and in the absence of a more complete phylogenetic data for the family
<taxonomicName id="8BEC9E9FFFF610687E34FE8A7B9CA783" box="[1096,1211,367,392]" class="Malacostraca" family="Paguridae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
, speculations of the phylogenetic relationship of
<taxonomicName id="8BEC9E9FFFF610687B8EFE707D41A7A7" box="[498,614,405,428]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687B8EFE707D41A7A7" box="[498,614,405,428]" italics="true" pageId="3" pageNumber="286">Patagurus</emphasis>
</taxonomicName>
with
<taxonomicName id="8BEC9E9FFFF6106878CBFE717CA9A7A6" box="[695,910,404,429]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF6106878CBFE717CA9A7A6" box="[695,910,404,429]" italics="true" pageId="3" pageNumber="286">Porcellanopagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="8BEC9E9FFFF6106879AAFE717BB3A7A6" box="[982,1172,404,429]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF6106879AAFE717BB3A7A6" box="[982,1172,404,429]" italics="true" pageId="3" pageNumber="286">Solitariopagurus</emphasis>
</taxonomicName>
would be premature.
<taxonomicName id="8BEC9E9FFFF610687AEBFE527E49A7DB" box="[151,366,439,464]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687AEBFE527E49A7DB" box="[151,366,439,464]" italics="true" pageId="3" pageNumber="286">Porcellanopagurus</emphasis>
</taxonomicName>
,
<taxonomicName id="8BEC9E9FFFF610687BF8FE527D65A7DB" box="[388,578,439,464]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687BF8FE527D65A7DB" box="[388,578,439,464]" italics="true" pageId="3" pageNumber="286">Solitariopagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="8BEC9E9FFFF6106878FBFE5C7DDCA7DB" box="[647,763,441,464]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF6106878FBFE5C7DDCA7DB" box="[647,763,441,464]" italics="true" pageId="3" pageNumber="286">Patagurus</emphasis>
</taxonomicName>
share numerous synapomorphies and appear to form a morphologically distinctive clade. However, the phylogenetic position of this clade within the large and heterogeneous family
<taxonomicName id="8BEC9E9FFFF610687BEBFE1A7D20A413" box="[407,519,511,536]" class="Malacostraca" family="Paguridae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
remains poorly understood. There is at least an indication of some affinity of
<taxonomicName id="8BEC9E9FFFF610687AEBFDC17E49A436" box="[151,366,548,573]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687AEBFDC17E49A436" box="[151,366,548,573]" italics="true" pageId="3" pageNumber="286">Porcellanopagurus</emphasis>
</taxonomicName>
and
<taxonomicName id="8BEC9E9FFFF610687BDAFDC17D43A436" box="[422,612,548,573]" class="Malacostraca" family="Paguridae" genus="Solitariopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687BDAFDC17D43A436" box="[422,612,548,573]" italics="true" pageId="3" pageNumber="286">Solitariopagurus</emphasis>
</taxonomicName>
to several other pagurid genera characterized by a more or less calcified posterior carapace, reduced abdomen and absence of pleopods in males, especially
<taxonomicName id="8BEC9E9FFFF610687E37FDAD7E61A48F" authority="McLaughlin &amp; Lemaitre 1997" authorityName="McLaughlin &amp; Lemaitre" authorityYear="1997" class="Malacostraca" family="Paguridae" genus="Ostraconotus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687E37FDAD7BC5A454" box="[1099,1250,584,607]" italics="true" pageId="3" pageNumber="286">Ostraconotus</emphasis>
(McLaughlin &amp; Lemaitre 1997)
</taxonomicName>
(see Discussion).
<taxonomicName id="8BEC9E9FFFF610687865FD897DD7A48E" box="[537,752,620,645]" class="Malacostraca" family="Paguridae" genus="Porcellanopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="7E98390EFFF610687865FD897DD7A48E" box="[537,752,620,645]" italics="true" pageId="3" pageNumber="286">Porcellanopagurus</emphasis>
</taxonomicName>
was included in recent phylogenetic analysis of anomurans (Schnabel
<emphasis id="7E98390EFFF610687B6DFD747E6BA4A3" box="[273,332,655,680]" italics="true" pageId="3" pageNumber="286">et al.</emphasis>
2011, Bracken-Grissom
<emphasis id="7E98390EFFF610687812FD747D8DA4A3" box="[622,682,655,680]" italics="true" pageId="3" pageNumber="286">et al.</emphasis>
2013) and recovered well within the
<taxonomicName id="8BEC9E9FFFF610687E18FD6A7BF0A4A3" box="[1124,1239,655,680]" class="Malacostraca" family="Paguridae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
, sister to a clade comprised of several northern temperate
<taxonomicName id="8BEC9E9FFFF61068781EFD517DF5A4C6" box="[610,722,692,717]" class="Malacostraca" family="Paguridae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
and representative king crabs.
</paragraph>
<paragraph id="4C53E51CFFF610687AEBFCFD7E67A55E" blockId="3.[151,411,791,853]" pageId="3" pageNumber="286">
<emphasis id="7E98390EFFF610687AEBFCFD7EBCA53B" bold="true" box="[151,411,791,816]" pageId="3" pageNumber="286">
<taxonomicName id="8BEC9E9FFFF610687AEBFCFD7E1DA524" box="[151,314,792,815]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="7E98390EFFF610687AEBFCFD7E1DA524" bold="true" box="[151,314,792,815]" italics="true" pageId="3" pageNumber="286">Patagurus rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65AB8475FFF610687B3EFCF27EBCA53B" box="[322,411,791,816]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D4D7F999FFF610687AE3FCD97FCEA55F" box="[159,233,828,853]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="3" pageNumber="286">Figs. 1</figureCitation>
C, 2–6)
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF610687AEBFC617E7DA5EF" pageId="3" pageNumber="286" type="materials_examined">
<paragraph id="4C53E51CFFF610687AEBFC617E7DA5EF" blockId="3.[151,1436,900,2005]" pageId="3" pageNumber="286">
<emphasis id="7E98390EFFF610687AEBFC617E61A596" bold="true" box="[151,326,900,925]" pageId="3" pageNumber="286">
<typeStatus id="93575BBEFFF610687AEBFC617FF5A596" box="[151,210,900,925]" pageId="3" pageNumber="286">Type</typeStatus>
material.
</emphasis>
<typeStatus id="93575BBEFFF610687B33FC617E90A596" box="[335,439,900,925]" pageId="3" pageNumber="286" type="holotype">Holotype</typeStatus>
male (sl
<quantity id="8B1448F9FFF61068785FFC617D58A597" box="[547,639,900,925]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="286" unit="mm" value="5.1">5.1 mm</quantity>
), UF 23548,
<collectingCountry id="34FBA58CFFF610687969FC617CFBA596" box="[789,988,900,925]" name="French Polynesia" pageId="3" pageNumber="286">French Polynesia</collectingCountry>
, Society Islands, Moorea, R/V
<emphasis id="7E98390EFFF610687F3EFC617A49A596" box="[1346,1390,900,925]" italics="true" pageId="3" pageNumber="286">Alis</emphasis>
sta. DW 3471,
<geoCoordinate id="29D883DBFFF610687B6FFC427E59A5CB" box="[275,382,935,960]" direction="west" orientation="longitude" pageId="3" pageNumber="286" precision="5" value="-149.9167">149.9167</geoCoordinate>
°W,
<geoCoordinate id="29D883DBFFF610687BD1FC4D7D02A5CB" box="[429,549,936,960]" direction="south" orientation="latitude" pageId="3" pageNumber="286" precision="5" value="-17.4833">17.4833°S</geoCoordinate>
, Warén dredge, depth
<quantity id="8B1448F9FFF61068795DFC4D7C4DA5B4" box="[801,874,936,960]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="286" unit="m" value="400.0">400 m</quantity>
, mud and rocks, some shells, leg. G. Paulay
<emphasis id="7E98390EFFF610687F20FC4C7AB2A5CB" box="[1372,1429,935,960]" italics="true" pageId="3" pageNumber="286">et al.</emphasis>
,
<date id="3852C3DCFFF610687AEBFC297E70A5EF" box="[151,343,972,997]" pageId="3" pageNumber="286" value="2009-10-21">21 October 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFF610637ABBFC0A7BD3A791" lastPageId="8" lastPageNumber="291" pageId="3" pageNumber="286" type="description">
<paragraph id="4C53E51CFFF610687ABBFC0A7AA8A347" blockId="3.[151,1436,900,2005]" pageId="3" pageNumber="286">
<emphasis id="7E98390EFFF610687ABBFC0A7E7EA203" bold="true" box="[199,345,1007,1032]" pageId="3" pageNumber="286">Description.</emphasis>
Shield (
<figureCitation id="D4D7F999FFF610687BC9FC0A7ED8A203" box="[437,511,1007,1032]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[164,1420,430,1584]" captionTargetId="figure@2.[151,1436,409,1649]" captionTargetPageId="2" captionText="FIGURE 1. Diagrammatic dorsal view of cephalothorax in three similarly carcinized pagurid genera: A, Porcellanopagurus; B, Solitariopagurus; C, Patagurus gen. nov. Figures not to scale. Abbreviations: ac 1, ac 2, ac 3, first, second and third anterior carapace lobes; cg, cervical groove; lp, lateral projection; pcl, posterior carapace lobe; r, rostrum." httpUri="https://zenodo.org/record/216562/files/figure.png" pageId="3" pageNumber="286">Figs. 1</figureCitation>
C, 2A) considerably wider than long; anterior margins between rostrum and lateral projections broad, each with slight median protrusion; lateral projections (
<figureCitation id="D4D7F999FFF61068799AFBF17B0CA227" box="[998,1067,1044,1069]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Fig. 2</figureCitation>
B) widely separated, moderately developed, subacute. Rostrum subtriangular, prominently drawn out into appreciably narrower tip with a middorsal ridge. Anterolateral margins of shield developed as subrectangular protrusions with relatively small first and second anterior carapace lobes (
<figureCitation id="D4D7F999FFF610687B81FB9A7D66A293" box="[509,577,1151,1176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Fig. 2</figureCitation>
A). Posterior carapace lobes extremely developed, sclerotized, subtriangular, each with dentate anterior margin; a narrow membranous portion, completely concealed in dorsal view, remaining near posterior margin of carapace. Dorsal surface of carapace with abundant tubercles and ridges of various sizes and shapes as illustrated (
<figureCitation id="D4D7F999FFF610687BC9FB097D25A30F" box="[437,514,1260,1285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Figs. 2</figureCitation>
A, B, 4A). Branchiostegite with two large and two small calcified plates (
<figureCitation id="D4D7F999FFF610687F44FB097A5BA30F" box="[1336,1404,1260,1285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="3" pageNumber="286">Fig. 4</figureCitation>
E). Ocular peduncles about 0.2 shield length, each distinctly swollen in proximal half; corneas approximately half peduncular length. Ocular acicles very small, hidden in dorsal view by proximally broadened margins of rostrum.
</paragraph>
<paragraph id="4C53E51CFFF610687ABBFAB27C57A043" blockId="3.[151,1436,900,2005]" pageId="3" pageNumber="286">
Antennular peduncle (
<figureCitation id="D4D7F999FFF610687BBBFAB27D3FA37B" box="[455,536,1367,1392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Figs. 2</figureCitation>
B, C) considerably longer than antennal peduncle, overreaching distal corneal margin by more than full length of ultimate segment, unarmed, but with several long setae on distodorsal margin of ultimate segment (omitted from
<figureCitation id="D4D7F999FFF610687B83FA7A7D66A3B3" box="[511,577,1439,1464]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Fig. 2</figureCitation>
C). Antennal peduncle (
<figureCitation id="D4D7F999FFF610687936FA7A7CBFA3B3" box="[842,920,1439,1464]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Figs. 2</figureCitation>
B, D) overreaching distal corneal margin by at least full length of ultimate segment, but considerably shorter than antennular peduncle; epistomial plate and first peduncular segment unarmed and lacking setae; second segment with dorsolateral distal angle produced, unarmed and terminally subacute; third through fifth segments each with few short setae; antennal acicle reaching to distal margin of fourth peduncular segment, unarmed, terminally blunt.
</paragraph>
<paragraph id="4C53E51CFFF610687ABBF9B17A42A09B" blockId="3.[151,1436,900,2005]" pageId="3" pageNumber="286">
Mouthparts not removed. Third maxilliped with basis armed with subdistal tooth on mesial surface; ischium with well-developed crista dentata and one accessory tooth (
<figureCitation id="D4D7F999FFF61068788EF9927C13A09B" box="[754,820,1655,1680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="3" pageNumber="286">Fig. 2</figureCitation>
E); merus, carpus, propodus and dactylus unarmed.
</paragraph>
<paragraph id="4C53E51CFFF610687ABBF9797C94A14F" blockId="3.[151,1436,900,2005]" pageId="3" pageNumber="286">
Thoracic sternite III prominently produced anteriorly into two triangular, subacute lobes, each with distinctly elevated, oblique ridge (
<figureCitation id="D4D7F999FFF610687BCAF95A7ED8A0D3" box="[438,511,1727,1752]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="3" pageNumber="286">Fig. 4</figureCitation>
C, D). Sternite IV roundly subrectangular, unequally divided into anterior and posterior portions by broad suture; posterior portion rigidly articulated with sternite V (
<figureCitation id="D4D7F999FFF610687ED8F9017BCAA0F7" box="[1188,1261,1764,1789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="3" pageNumber="286">Fig. 4</figureCitation>
D). Sternite VI narrow, rigidly articulated with sternite VII at ~90° angle, latter extending dorsad. Sternite VIII narrow, anterodorsal to sternite VII, separated from latter by membranous area.
</paragraph>
<paragraph id="4C53E51CFFF6106E7ABBF8AA7C49A1DA" blockId="3.[151,1436,900,2005]" lastBlockId="5.[151,1436,1724,2001]" lastPageId="5" lastPageNumber="288" pageId="3" pageNumber="286">
Right cheliped much larger and more robust than left cheliped, but slightly shorter in overall length; propodocarpal articulation perpendicular to main axis of cheliped, without pronounced rotation (
<figureCitation id="D4D7F999FFF610687E03F8917BE3A187" box="[1151,1220,1908,1933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="3" pageNumber="286">Fig. 4</figureCitation>
B, 5B). Chela (
<figureCitation id="D4D7F999FFF610687F12F8917F82A1BB" captionStart="FIGURE 3" captionStartId="5.[151,250,1571,1593]" captionTargetBox="[179,1418,205,1524]" captionTargetId="figure@5.[151,1436,193,1549]" captionTargetPageId="5" captionText="FIGURE 3. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view)." httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="3" pageNumber="286">Fig. 3</figureCitation>
C, 4B) approximately as long as carpus; dorsal surface convex, smooth, drawn out mesially and laterally into thin margins; ventral surface of palm and dactylus with scattered fine setae; dactylus approximately 0.6 length of palm, unarmed, with rounded margins, some scattered setae present; cutting edges of dactylus and fixed finger calcareous, each with two low teeth, proximal larger than distal; fingers proximally gaping when closed; fingertips with small corneous point. Carpus (
<figureCitation id="D4D7F999FFF0106E785BF8E17D4EA117" box="[551,617,1796,1821]" captionStart="FIGURE 3" captionStartId="5.[151,250,1571,1593]" captionTargetBox="[179,1418,205,1524]" captionTargetId="figure@5.[151,1436,193,1549]" captionTargetPageId="5" captionText="FIGURE 3. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view)." httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="5" pageNumber="288">Fig. 3</figureCitation>
A, B) approximately as long as merus, trapezoidal, with dorsomesial and dorsolateral margins each elevated into low, unarmed crest, dorsal surface elevated in midline and with median row of tubercles, forming a distinctly rounded ridge distally; ventral surface with scattered, moderately long setae. Merus roundly triangular; lateral surface minutely granular; ventromesial and ventrolateral margins each with row of spinulose tubercles, ventral surface with some larger tubercles, smaller granules and fine setae. Ischium with faint rugosities or granules mesially, remaining surface unarmed.
</paragraph>
<caption id="1893B594FFF1106F7AEBF8FF7A55A1A1" httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="4" pageNumber="287" targetBox="[176,1385,210,1764]" targetPageId="4">
<paragraph id="4C53E51CFFF1106F7AEBF8FF7A55A1A1" blockId="4.[151,1436,1818,1962]" pageId="4" pageNumber="287">
<emphasis id="7E98390EFFF1106F7AEBF8FF7E28A124" bold="true" box="[151,271,1818,1840]" pageId="4" pageNumber="287">FIGURE 2</emphasis>
.
<taxonomicName id="8BEC9E9FFFF1106F7B63F8FE7E96A13B" box="[287,433,1819,1840]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="7E98390EFFF1106F7B63F8FE7E96A13B" box="[287,433,1819,1840]" italics="true" pageId="4" pageNumber="287">Patagurus rex</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF1106F7BC7F8FF7D44A124" bold="true" box="[443,611,1818,1840]" pageId="4" pageNumber="287">gen. &amp; sp. nov.</emphasis>
, holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view).
</paragraph>
</caption>
<caption id="1893B594FFF0106E7AEBF9C67C8BA09F" httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="5" pageNumber="288" targetBox="[179,1418,205,1524]" targetPageId="5">
<paragraph id="4C53E51CFFF0106E7AEBF9C67C8BA09F" blockId="5.[151,1436,1571,1684]" pageId="5" pageNumber="288">
<emphasis id="7E98390EFFF0106E7AEBF9C67E29A033" bold="true" box="[151,270,1571,1593]" pageId="5" pageNumber="288">FIGURE 3</emphasis>
.
<taxonomicName id="8BEC9E9FFFF0106E7B61F9C17E89A032" box="[285,430,1572,1593]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="7E98390EFFF0106E7B61F9C17E89A032" box="[285,430,1572,1593]" italics="true" pageId="5" pageNumber="288">Patagurus rex</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF0106E7BCBF9C67D7AA033" bold="true" box="[439,605,1571,1593]" pageId="5" pageNumber="288">gen. &amp; sp. nov.</emphasis>
, holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view).
</paragraph>
</caption>
<caption id="1893B594FFF3106D7AEBF89E7EDCA1E6" httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="6" pageNumber="289" targetBox="[226,1355,193,1878]" targetPageId="6">
<paragraph id="4C53E51CFFF3106D7AEBF89E7EDCA1E6" blockId="6.[151,1436,1915,2029]" pageId="6" pageNumber="289">
<emphasis id="7E98390EFFF3106D7AEBF89E7E28A19B" bold="true" box="[151,271,1915,1937]" pageId="6" pageNumber="289">FIGURE 4</emphasis>
.
<taxonomicName id="8BEC9E9FFFF3106D7B63F8997E96A19A" box="[287,433,1916,1937]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="7E98390EFFF3106D7B63F8997E96A19A" box="[287,433,1916,1937]" italics="true" pageId="6" pageNumber="289">Patagurus rex</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF3106D7BC6F89E7D45A19B" bold="true" box="[442,610,1915,1937]" pageId="6" pageNumber="289">gen. &amp; sp. nov.</emphasis>
, holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped
<emphasis id="7E98390EFFF3106D797AF87F7C62A1A4" box="[774,837,1946,1967]" italics="true" pageId="6" pageNumber="289">in situ</emphasis>
(dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell
<taxonomicName id="8BEC9E9FFFF3106D7B98F85D7D7CA1C5" box="[484,603,1976,1998]" class="Bivalvia" family="Mytilidae" genus="Gregariella" kingdom="Animalia" order="Mytiloida" pageId="6" pageNumber="289" phylum="Mollusca" rank="genus">
<emphasis id="7E98390EFFF3106D7B98F85D7D7CA1C5" box="[484,603,1976,1998]" italics="true" pageId="6" pageNumber="289">Gregariella</emphasis>
</taxonomicName>
sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view.
</paragraph>
</caption>
<caption id="1893B594FFF2106C7AEBFB767E8FA2CC" httpUri="https://zenodo.org/record/216566/files/figure.png" pageId="7" pageNumber="290" targetBox="[151,1436,178,1134]" targetPageId="7">
<paragraph id="4C53E51CFFF2106C7AEBFB767E8FA2CC" blockId="7.[151,1437,1171,1223]" pageId="7" pageNumber="290">
<emphasis id="7E98390EFFF2106C7AEBFB767E2AA2A3" bold="true" box="[151,269,1171,1193]" pageId="7" pageNumber="290">FIGURE 5</emphasis>
.
<taxonomicName id="8BEC9E9FFFF2106C7B65FB717E8FA2A2" box="[281,424,1172,1193]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="7E98390EFFF2106C7B65FB717E8FA2A2" box="[281,424,1172,1193]" italics="true" pageId="7" pageNumber="290">Patagurus rex</emphasis>
</taxonomicName>
<emphasis id="7E98390EFFF2106C7BD3FB767D69A2A3" bold="true" box="[431,590,1171,1193]" pageId="7" pageNumber="290">gen. &amp; sp. nov.</emphasis>
, holotype male (UF 23548), habitus and coloration in dorsal view (shell detached). Photograph: Arthur Anker.
</paragraph>
</caption>
<paragraph id="4C53E51CFFF2106C7ABBFB0C7E7BA025" blockId="7.[151,1437,1257,2016]" pageId="7" pageNumber="290">
Left cheliped elongate, slender; dactylus and fixed finger distinctly arched ventrally. Chela shorter than carpus; dorsal surface unarmed, slightly convex, with rounded margins; ventral surface with numerous tufts of moderately long setae on palm and especially on fixed finger; cutting edge of fixed finger with row of small corneous teeth; dactylus more than twice length of palm, unarmed, with tufts of moderately long setae; margins rounded; cutting edge with row of minute corneous teeth (
<figureCitation id="D4D7F999FFF2106C7842FA8A7DA5A38C" box="[574,642,1391,1416]" captionStart="FIGURE 3" captionStartId="5.[151,250,1571,1593]" captionTargetBox="[179,1418,205,1524]" captionTargetId="figure@5.[151,1436,193,1549]" captionTargetPageId="5" captionText="FIGURE 3. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view)." httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="7" pageNumber="290">Fig. 3</figureCitation>
D–F). Carpus approximately equal to merus length; dorsomesial and dorsolateral margins each elevated into unarmed crest; dorsal surface elevated in midline and armed with row of subacute spiniform tubercles; lateral face minutely granular; ventral surface with scattered small tubercles and moderately long setae. Merus roundly triangular; lateral surface minutely granular; ventral surface with irregular row of spinulose tubercles adjacent to lateral margin, some tubercles and fine setae present on ventral surface and mesially. Ischium unarmed.
</paragraph>
<paragraph id="4C53E51CFFF2106C7ABBF9D27B32A0FD" blockId="7.[151,1437,1257,2016]" pageId="7" pageNumber="290">
Ambulatory legs (second and third pereopods) similar in shape and length (
<figureCitation id="D4D7F999FFF2106C7E73F9D27B7CA044" box="[1039,1115,1591,1616]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">Figs. 2</figureCitation>
F, 5). Dactyli slightly shorter than propodi, each with sparse, moderately long setae dorsally and one corneous spinule adjacent to terminal corneous unguis; mesial and lateral faces with few shorter, scattered setae; ventral margins each with row of 11 or 12 corneous spines (
<figureCitation id="D4D7F999FFF2106C7A97F97E7E08A0B8" box="[235,303,1691,1716]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">Fig. 2</figureCitation>
G). Propodi approximately 1.5 length of carpi, unarmed, with scattered setae. Carpi short, slightly less than 0.5 lengths of meri; unarmed, with few scattered setae. Meri unarmed except for minute protuberances on ventral margin. Ischia unarmed; lengths of ischium of second and third pereopods similar.
</paragraph>
<paragraph id="4C53E51CFFF2106C7ABBF91A7BC8A196" blockId="7.[151,1437,1257,2016]" pageId="7" pageNumber="290">
Fourth pereopod (
<figureCitation id="D4D7F999FFF2106C7BF7F91A7EEBA11C" box="[395,460,1791,1816]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">Fig. 2</figureCitation>
H) with dactylus and propodus bearing scattered setae; dactylus somewhat sickle-shaped, about as long as propodus, covered with minute tubercles (
<figureCitation id="D4D7F999FFF2106C796FF8C57C73A132" box="[787,852,1824,1849]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">
Fig.
<date id="3852C3DCFFF2106C7939F8C47C73A132" box="[837,852,1825,1849]" pageId="7" pageNumber="290">2</date>
</figureCitation>
I); propodal rasp consisting of single row of five, long corneous spines; carpus and merus unarmed, each with few setae. Fifth pereopod (
<figureCitation id="D4D7F999FFF2106C7E51F8A47B57A151" box="[1069,1136,1857,1882]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">Fig. 2</figureCitation>
J) with very small dactylus obscured by terminal tufts of long setae originating from propodus, without rasp. Coxae of male fifth pereopods each with well-developed sexual tube, right longer and more slender, both densely setose distally (
<figureCitation id="D4D7F999FFF2106C7EF7F8617BEAA196" box="[1163,1229,1924,1949]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[176,1385,210,1764]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURE 2. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, cephalothorax and eyestalks (dorsal view); B, anterior portion of carapace and cephalic appendages (dorsal view); C, ultimate segment and flagellum of left antennular peduncle (lateral view, dorsolateral distal tuft of setae on ultimate segment omitted); D, left antennal peduncle (lateral view); E, right third maxilliped (ventrolateral view); F, right first ambulatory leg (lateral view); G, same, dactylus; H, left fourth pereopod (lateral view); I, same, tip of dactylus (mesial view); J, right fifth pereopod (lateral view); K, sexual tubes (posterior view)." httpUri="https://zenodo.org/record/216563/files/figure.png" pageId="7" pageNumber="290">Fig. 2</figureCitation>
K).
</paragraph>
<paragraph id="4C53E51CFFF210637ABBF8407E9AA6DA" blockId="7.[151,1437,1257,2016]" lastBlockId="8.[151,1436,151,610]" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290">
Male pleon markedly reduced (
<figureCitation id="D4D7F999FFF2106C7853F8407DA7A1B5" box="[559,640,1957,1982]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="7" pageNumber="290">Figs. 4</figureCitation>
D, 5C, E), lacking pleopods 2–5. Uropods symmetrical; endopod and exopod each with ovate rasp of corneous scales, exopodal rasps larger than endopodal rasps (
<figureCitation id="D4D7F999FFF2106C7EA1F8227A04A1D4" box="[1245,1315,1991,2016]" captionStart="FIGURE 3" captionStartId="5.[151,250,1571,1593]" captionTargetBox="[179,1418,205,1524]" captionTargetId="figure@5.[151,1436,193,1549]" captionTargetPageId="5" captionText="FIGURE 3. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view)." httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="7" pageNumber="290">Fig. 3</figureCitation>
G). Telson subquadrate, with trace of lateral indentation; dorsal surface proximally with median depression; posterior margin entire, unarmed (
<figureCitation id="D4D7F999FFFD10637B2AFF5C7EBFA6DA" box="[342,408,185,210]" captionStart="FIGURE 3" captionStartId="5.[151,250,1571,1593]" captionTargetBox="[179,1418,205,1524]" captionTargetId="figure@5.[151,1436,193,1549]" captionTargetPageId="5" captionText="FIGURE 3. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, carpus of right cheliped (dorsal view); B, same (lateral view); C, chela of right cheliped (dorsal view); D, chela of left cheliped (dorsal view); E, same, distal portion of fingers (ventral view); F, same, detail of dentition of cutting edges (ventral view); G, distal portion of pleon with sixth somite and uropods (dorsal view, telson not visible); H, telson and uropods (dorsal view)." httpUri="https://zenodo.org/record/216564/files/figure.png" pageId="8" pageNumber="291">Fig. 3</figureCitation>
H).
</paragraph>
<paragraph id="4C53E51CFFFD10637ABBFF3F7EB7A6F8" blockId="8.[151,1436,151,610]" box="[199,400,218,243]" pageId="8" pageNumber="291">Female unknown.</paragraph>
<paragraph id="4C53E51CFFFD10637ABBFF197BD3A791" blockId="8.[151,1436,151,610]" pageId="8" pageNumber="291">
<emphasis id="7E98390EFFFD10637ABBFF197E7BA71E" bold="true" box="[199,348,252,277]" pageId="8" pageNumber="291">Color in life</emphasis>
. Calcified portions of carapace hyaline-whitish with scattered tints of orange and pink; tip of rostrum and first and second anterior carapace lobes darker orange; ocular peduncles white; antennular and antennal peduncles and flagella orange; ventral cephalothorax pale orangish-white; chelipeds bright orange dorsally, lighter orange ventrally; ambulatory legs with orange dactyli, propodi and carpi each reddish-orange with white band at distal margin, meri reddish-orange, each with irregular splotches of white (
<figureCitation id="D4D7F999FFFD10637E07FE647BEDA791" box="[1147,1226,385,410]" captionStart="FIGURE 5" captionStartId="7.[151,250,1171,1193]" captionTargetBox="[151,1436,178,1134]" captionTargetId="figure@7.[151,1436,177,1136]" captionTargetPageId="7" captionText="FIGURE 5. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548), habitus and coloration in dorsal view (shell detached). Photograph: Arthur Anker." httpUri="https://zenodo.org/record/216566/files/figure.png" pageId="8" pageNumber="291">Figs. 5</figureCitation>
,
<figureCitation id="D4D7F999FFFD10637EAAFE647BC1A791" box="[1238,1254,385,410]" captionStart="FIGURE 6" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[151,1436,649,1939]" captionTargetId="figure@8.[151,1436,649,1939]" captionTargetPageId="8" captionText="FIGURE 6. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548), habitus and coloration: A, anterodorsal view, carrying bivalve shell Gregariella sp., immediately after capture; B, frontal view, shell detached; C, anterolateral view, pleon not visible; D, ventral view, pleon exposed. Photographs: Gustav Paulay (A), Arthur Anker (B – D)." httpUri="https://zenodo.org/record/216567/files/figure.png" pageId="8" pageNumber="291">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFFD10637ABBFE477EE3A7D6" pageId="8" pageNumber="291" type="etymology">
<paragraph id="4C53E51CFFFD10637ABBFE477EE3A7D6" blockId="8.[151,1436,151,610]" pageId="8" pageNumber="291">
<emphasis id="7E98390EFFFD10637ABBFE477E61A7B0" bold="true" box="[199,326,418,443]" pageId="8" pageNumber="291">Etymology</emphasis>
. Referring to the extraordinary albeit superficial resemblance of this new species to some king crabs (
<taxonomicName id="8BEC9E9FFFFD10637A9EFE207E23A7D7" box="[226,260,453,476]" class="Malacostraca" family="Paguridae" genus="Patagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="rex">
<emphasis id="7E98390EFFFD10637A9EFE207E23A7D7" box="[226,260,453,476]" italics="true" pageId="8" pageNumber="291">rex</emphasis>
</taxonomicName>
= king in Latin).
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFFD10637ABBFE007C31A415" pageId="8" pageNumber="291" type="distribution">
<paragraph id="4C53E51CFFFD10637ABBFE007A4EA7F5" blockId="8.[151,1436,151,610]" box="[199,1385,485,510]" pageId="8" pageNumber="291">
<emphasis id="7E98390EFFFD10637ABBFE007E47A7F5" bold="true" box="[199,352,485,510]" pageId="8" pageNumber="291">Distribution.</emphasis>
Presently known only from the
<typeStatus id="93575BBEFFFD106378B5FE037DDEA7F5" box="[713,761,486,510]" pageId="8" pageNumber="291">type</typeStatus>
locality off Moorea, Society Islands,
<collectingCountry id="34FBA58CFFFD10637EE3FE007A42A7F5" box="[1183,1381,485,510]" name="French Polynesia" pageId="8" pageNumber="291">French Polynesia</collectingCountry>
.
</paragraph>
<paragraph id="4C53E51CFFFD10637ABBFDE37C31A415" blockId="8.[151,1436,151,610]" box="[199,790,518,543]" pageId="8" pageNumber="291">
<emphasis id="7E98390EFFFD10637ABBFDE37E0DA414" bold="true" box="[199,298,518,543]" pageId="8" pageNumber="291">Habitat.</emphasis>
Mud and rocks bottom at a depth of
<specimenCount id="5AEA2E95FFFD106378B4FDE27C31A415" box="[712,790,519,543]" pageId="8" pageNumber="291" type="generic">
<quantity id="8B1448F9FFFD106378B4FDE27C35A415" box="[712,786,519,543]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="291" unit="m" value="400.0">400 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="04F6B697FFFD10637ABBFDC27D36A46A" pageId="8" pageNumber="291" type="description">
<paragraph id="4C53E51CFFFD10637ABBFDC27D36A46A" blockId="8.[151,1436,151,610]" pageId="8" pageNumber="291">
<emphasis id="7E98390EFFFD10637ABBFDC27EABA44B" bold="true" box="[199,396,551,576]" pageId="8" pageNumber="291">Biological notes.</emphasis>
The hermit crab was covering its miniature abdomen with a valve of the mytilid bivalve,
<taxonomicName id="8BEC9E9FFFFD10637AEBFDAC7E3DA469" box="[151,282,585,610]" class="Bivalvia" family="Mytilidae" genus="Gregariella" kingdom="Animalia" order="Mytiloida" pageId="8" pageNumber="291" phylum="Mollusca" rank="genus">
<emphasis id="7E98390EFFFD10637AEBFDAC7E3DA469" box="[151,282,585,610]" italics="true" pageId="8" pageNumber="291">Gregariella</emphasis>
</taxonomicName>
sp. (
<figureCitation id="D4D7F999FFFD10637B2CFDAC7EB9A46A" box="[336,414,585,610]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[226,1355,193,1878]" captionTargetId="figure@6.[226,1357,193,1878]" captionTargetPageId="6" captionText="FIGURE 4. Patagurus rex gen. &amp; sp. nov., holotype male (UF 23548): A, anterodorsal view of cephalothorax with eyes, cephalic appendages, and part of chelipeds; B, right cheliped in situ (dorsal view); C, ventral view of cephalothorax and pleon, latter concealed by bivalve shell Gregariella sp.; D, sternum and pleon without bivalve shell (ventral view); E, branchiostegial area of cephalothorax, lateral view." httpUri="https://zenodo.org/record/216565/files/figure.png" pageId="8" pageNumber="291">Figs. 4</figureCitation>
C, 6A, B).
</paragraph>
</subSubSection>
</treatment>
</document>
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