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<document id="EA31E1E263F979947BE35FC5FAC55AAD" ID-DOI="10.1080/00222933.2020.1759721" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609053" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615923843413" checkinUser="carolina" docAuthor="Bevilaqua, Marcus &amp; Fonseca, Claudio Ruy Vasconcelos da" docDate="2020" docId="C4008788690A9E14FEF4FCA881436A53" docLanguage="en" docName="JNATHIST.54.5-6.321-350.pdf" docOrigin="Journal of Natural History 54 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222933.2020.1759721" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Passalus (Pertinax) cuneatus Hincks 1950" docType="treatment" docVersion="3" lastPageNumber="331" masterDocId="3839FFF0690D9E1FFFCAFF9A83526814" masterDocTitle="Redescription of the species of Passalus Fabricius, 1792 (Coleoptera: Passalidae) described by Walter Douglas Hincks (1906 - 1961) deposited in the Museum of the University of Manchester" masterLastPageNumber="350" masterPageNumber="321" pageNumber="327" updateTime="1698887187464" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="132ABF582AFDC5E84420137CCC590783">Redescription of the species of Passalus Fabricius, 1792 (Coleoptera: Passalidae) described by Walter Douglas Hincks (1906 - 1961) deposited in the Museum of the University of Manchester</mods:title>
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<mods:namePart id="D79510AC60A03BC3DC5DF34809F6D27F">Bevilaqua, Marcus</mods:namePart>
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<mods:namePart id="4B695A2748D98ACD2DF8FF8F8C8C7A87">Fonseca, Claudio Ruy Vasconcelos da</mods:namePart>
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<mods:title id="6DB0A819014D55A9678896922EAEF36F">Journal of Natural History</mods:title>
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<mods:date id="46FB98D71298A76759A2A6C0D4C09E43">2020</mods:date>
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<paragraph id="4C16369E690A9E18FEF4FCA880B76B5F" blockId="7.[318,997,818,843]" box="[318,997,818,843]" pageId="7" pageNumber="327">
<heading id="175E81F2690A9E18FEF4FCA880B76B5F" box="[318,997,818,843]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="327" reason="2">
<taxonomicName id="8BA94D1D690A9E18FEF4FCA880176B5F" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[318,837,818,843]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="327" phylum="Arthropoda" rank="species" species="cuneatus" subGenus="Pertinax">
<emphasis id="7EDDEA8C690A9E18FEF4FCA880176B5F" bold="true" box="[318,837,818,843]" pageId="7" pageNumber="327">
<emphasis id="7EDDEA8C690A9E18FEF4FCA882F56B5F" bold="true" box="[318,423,818,843]" italics="true" pageId="7" pageNumber="327">Passalus</emphasis>
(
<emphasis id="7EDDEA8C690A9E18FE7DFCA8814F6B5F" bold="true" box="[439,541,818,843]" italics="true" pageId="7" pageNumber="327">Pertinax</emphasis>
)
<emphasis id="7EDDEA8C690A9E18FDE4FCA881CE6B5F" bold="true" box="[558,668,818,843]" italics="true" pageId="7" pageNumber="327">cuneatus</emphasis>
<bibRefCitation id="28384B6F690A9E18FD6FFCA880176B5F" author="Hincks WD" box="[677,837,818,843]" pageId="7" pageNumber="327" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="D4922A1B690A9E18FC9FFCA8808F6B5F" box="[853,989,818,843]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="8.[160,225,1482,1505]" captionStartId-1="9.[160,225,1329,1352]" captionTargetBox-0="[163,1153,142,1451]" captionTargetBox-1="[163,1153,142,1297]" captionTargetId-0="figure-55@8.[163,1153,142,1451]" captionTargetId-1="figure-112@9.[163,1153,142,1298]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." captionText-1="Figure 4. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.4609065" figureDoi-1="http://doi.org/10.5281/zenodo.4609067" httpUri-0="https://zenodo.org/record/4609065/files/figure.png" httpUri-1="https://zenodo.org/record/4609067/files/figure.png" pageId="7" pageNumber="327">
Figures 3
<emphasis id="7EDDEA8C690A9E18FC0BFCA8809D6B5F" box="[961,975,818,843]" italics="true" pageId="7" pageNumber="327"></emphasis>
4
</figureCitation>
)
</heading>
</paragraph>
<paragraph id="4C16369E690A9E18FF0AFCFD87D16B6B" blockId="7.[160,1155,871,930]" box="[192,1155,871,895]" pageId="7" pageNumber="327">
<bibRefCitation id="28384B6F690A9E18FF0AFCFD82C36B6B" author="Hincks WD" box="[192,401,871,895]" pageId="7" pageNumber="327" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks 1950: 1043</bibRefCitation>
, fig. 2;
<bibRefCitation id="28384B6F690A9E18FE2FFCFD805C6B6B" author="Hincks WD &amp; Dibb JR" box="[485,782,871,895]" pageId="7" pageNumber="327" pagination="1 - 33" refId="ref13308" refString="Hincks WD, Dibb JR. 1958. Supplementa. Passalidae. Coleopterorum Catalogus. Berlin: W. Junk, J. Schenkling, es-Gravenhage; p. 1 - 33." type="book chapter" year="1958">Hincks and Dibb 1958: 16</bibRefCitation>
;
<bibRefCitation id="28384B6F690A9E18FCD1FCFD877C6B6B" author="Reyes-Castillo P" box="[795,1070,871,895]" pageId="7" pageNumber="327" pagination="1 - 240" refId="ref13746" refString="Reyes-Castillo P. 1970. Coleoptera, Passalidae: morfologia y division en grandes grupos; generos americanos. Folia Entomol Mex. 20 (22): 1 - 240." type="journal article" year="1970">Reyes-Castillo 1970: 204</bibRefCitation>
; Reyes-
</paragraph>
</subSubSection>
<subSubSection id="04B36515690A9E18FF6AFC1081AC6BB6" box="[160,766,905,930]" pageId="7" pageNumber="327" type="description">
<paragraph id="4C16369E690A9E18FF6AFC1081AC6BB6" blockId="7.[160,1155,871,930]" box="[160,766,905,930]" pageId="7" pageNumber="327">
Castillo and Castillo 1992: 357;
<bibRefCitation id="28384B6F690A9E18FE37FC1081FC6BB6" author="Miles C" box="[509,686,906,930]" pageId="7" pageNumber="327" pagination="109 - 126" refId="ref13705" refString="Miles C. 2017. The Passalidae collection of the Manchester Museum, UK, with a complete type catalogue. Russ Entomol J. 26 (2): 109 - 126. doi: 10.15298 / rusentj. 26.2.03" type="journal article" year="2017">Miles 2017: 119</bibRefCitation>
, fig 12.
</paragraph>
</subSubSection>
<subSubSection id="04B36515690A9E18FF6AFC5783896CE8" pageId="7" pageNumber="327" type="diagnosis">
<paragraph id="4C16369E690A9E18FF6AFC5783896CE8" blockId="7.[160,1156,973,1276]" pageId="7" pageNumber="327">
<emphasis id="7EDDEA8C690A9E18FF6AFC5782486BF3" bold="true" box="[160,282,973,999]" italics="true" pageId="7" pageNumber="327">Diagnosis.</emphasis>
Large size species; flat body; large and straight mandibles; straight anterior edge of the front; labrum with a notch in the
<emphasis id="7EDDEA8C690A9E18FD69FC6881FB6C1E" box="[675,681,1010,1034]" italics="true" pageId="7" pageNumber="327"></emphasis>
V
<emphasis id="7EDDEA8C690A9E18FD72FC6881EC6C1E" box="[696,702,1010,1034]" italics="true" pageId="7" pageNumber="327"></emphasis>
shaped; large and smooth mediofrontal area with a row of punctures close to the anterior edge of the front; laterofrontal + mediofrontal tubercles practically attached to inner tubercles; posterofrontal ridges larger and lower than the anterofrontal ridges; practically absent pronotal punctuations; deep mesosternal scars; poorly delimited metasternal disc; metasternum glabrous with a small group of punctures in the posterior region; aedeagus with parameres reaching almost the apex of the median lobe and tegmen with wide opening in the median region in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="04B36515690A9E15FF6AFAB282996EAB" lastPageId="10" lastPageNumber="330" pageId="7" pageNumber="327" type="description">
<paragraph id="4C16369E690A9E15FF6AFAB280D76CA2" blockId="7.[160,1156,1320,1727]" lastBlockId="10.[160,1156,141,1206]" lastPageId="10" lastPageNumber="330" pageId="7" pageNumber="327">
<emphasis id="7EDDEA8C690A9E18FF6AFAB282146D56" bold="true" box="[160,326,1320,1346]" italics="true" pageId="7" pageNumber="327">Redescription.</emphasis>
<emphasis id="7EDDEA8C690A9E18FE9CFAB082C16D56" bold="true" box="[342,403,1322,1346]" pageId="7" pageNumber="327">Body</emphasis>
(
<figureCitation id="D4922A1B690A9E18FE6BFAB082AE6D56" box="[417,508,1322,1346]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="7" pageNumber="327">Figure 3</figureCitation>
):
<emphasis id="7EDDEA8C690A9E18FDDAFAB381336D56" box="[528,609,1321,1346]" italics="true" pageId="7" pageNumber="327">habitus</emphasis>
: flat;
<emphasis id="7EDDEA8C690A9E18FD51FAB381906D56" box="[667,706,1321,1346]" italics="true" pageId="7" pageNumber="327">size</emphasis>
: big (&lt;
<quantity id="8B519B7B690A9E18FCC7FAB0800C6D56" box="[781,862,1322,1346]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="327" unit="mm" value="30.0">30 mm</quantity>
in length).
<emphasis id="7EDDEA8C690A9E18FC16FAB087486D56" bold="true" box="[988,1050,1322,1346]" pageId="7" pageNumber="327">Head</emphasis>
(
<figureCitation id="D4922A1B690A9E18FBE2FAB087D16D56" box="[1064,1155,1322,1346]" captionStart="Figure 4" captionStartId="9.[160,225,1329,1352]" captionTargetBox="[163,1153,142,1297]" captionTargetId="figure-112@9.[163,1153,142,1298]" captionTargetPageId="9" captionText="Figure 4. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4609067" httpUri="https://zenodo.org/record/4609067/files/figure.png" pageId="7" pageNumber="327">Figure 4</figureCitation>
(a,c)):
<emphasis id="7EDDEA8C690A9E18FF2FFAD6826B6D71" box="[229,313,1356,1381]" italics="true" pageId="7" pageNumber="327">Labrum</emphasis>
: anterior border concave.
<emphasis id="7EDDEA8C690A9E18FDA4FAD681936D71" box="[622,705,1356,1381]" italics="true" pageId="7" pageNumber="327">Clypeus</emphasis>
: hidden under the frons; with anterior angles under the lateral + mediofrontal tubercles.
<emphasis id="7EDDEA8C690A9E18FD3BFAF480896D93" box="[753,987,1390,1415]" italics="true" pageId="7" pageNumber="327">Anterior frontal edge</emphasis>
: straight, with a notch in the median region.
<emphasis id="7EDDEA8C690A9E18FDC9FA0B80216DBE" box="[515,883,1425,1450]" italics="true" pageId="7" pageNumber="327">Secondary mediofrontal tubercles</emphasis>
: absent.
<emphasis id="7EDDEA8C690A9E18FC17FA0B87386DBE" box="[989,1130,1425,1450]" italics="true" pageId="7" pageNumber="327">Laterofrontal</emphasis>
+
<emphasis id="7EDDEA8C690A9E18FF6AFA2E82CB6DD9" box="[160,409,1460,1485]" italics="true" pageId="7" pageNumber="327">mediofrontal tubercles</emphasis>
: conspicuous, acute, and projected forward.
<emphasis id="7EDDEA8C690A9E18FC7FFA2E872F6DD9" box="[949,1149,1460,1485]" italics="true" pageId="7" pageNumber="327">Mediofrontal area</emphasis>
: caliciform 2.2x wider than long, flat and deep; with coarse punctuations only in the anterior region, near the anterior frontal edge.
<emphasis id="7EDDEA8C690A9E18FD63FA6380046E06" box="[681,854,1529,1554]" italics="true" pageId="7" pageNumber="327">Cephalic nodule</emphasis>
: large but not well marked and rounded shape.
<emphasis id="7EDDEA8C690A9E18FE45F986817D6E21" box="[399,559,1564,1589]" italics="true" pageId="7" pageNumber="327">Inner tubercles</emphasis>
: large, conspicuous with acute apexes, little smaller than the laterofrontal + mediofrontal tubercles, of which are practically attached by the anterofrontal ridges.
<emphasis id="7EDDEA8C690A9E18FE5EF9FB81206E6E" box="[404,626,1633,1658]" italics="true" pageId="7" pageNumber="327">Anterofrontal ridges</emphasis>
: straight, smaller, and taller than the posterofrontal ridges.
<emphasis id="7EDDEA8C690A9E18FE9EF91E81646E89" box="[340,566,1668,1693]" italics="true" pageId="7" pageNumber="327">Posterofrontal ridges</emphasis>
: low, strong, and straight, beginning at the apex of the central tubercle.
<emphasis id="7EDDEA8C690A9E18FE53F93C813F6EAB" box="[409,621,1702,1727]" italics="true" pageId="7" pageNumber="327">Laterofrontal areas</emphasis>
: slightly convex and smooth surface.
<emphasis id="7EDDEA8C690A9E17FBFFF93C83A46E45" italics="true" lastPageId="8" lastPageNumber="328" pageId="7" pageNumber="327">Central tubercle</emphasis>
: conical, small, low, with non-free apex and dilated back.
<emphasis id="7EDDEA8C69059E17FCBEF9A2872F6E45" box="[884,1149,1592,1617]" italics="true" pageId="8" pageNumber="328">Lateroposterior tubercles</emphasis>
: inconspicuous, far from the central tubercle, with rounded shape.
<emphasis id="7EDDEA8C69059E17FC51F9C187196E60" box="[923,1099,1627,1652]" italics="true" pageId="8" pageNumber="328">Postfrontal area</emphasis>
: flat, smooth surface.
<emphasis id="7EDDEA8C69059E17FE93F9E481736E83" box="[345,545,1662,1687]" italics="true" pageId="8" pageNumber="328">Postfrontal groove</emphasis>
: well marked and with a slight notch located medially.
<emphasis id="7EDDEA8C69059E17FF6AF93A82F36EAD" box="[160,417,1696,1721]" italics="true" pageId="8" pageNumber="328">Lateropostfrontal areas</emphasis>
: shallow, smooth, and glabrous surface.
<emphasis id="7EDDEA8C69059E17FC48F93A87176EAD" box="[898,1093,1696,1721]" italics="true" pageId="8" pageNumber="328">Epicranial sutures</emphasis>
: well marked.
<emphasis id="7EDDEA8C69049E16FECFFA2E82F36DD9" box="[261,417,1460,1485]" italics="true" pageId="9" pageNumber="329">Epicranial pits</emphasis>
: deep and very visible.
<emphasis id="7EDDEA8C69049E16FD74FA2E80A46DD9" box="[702,1014,1460,1485]" italics="true" pageId="9" pageNumber="329">Anterior angles of the head</emphasis>
: small, with obtuse apices, smaller than the laterofrontal + mediofrontal tubercles.
<emphasis id="7EDDEA8C69049E16FC1CFA4C872C6DFB" box="[982,1150,1494,1519]" italics="true" pageId="9" pageNumber="329">Canthus ocular</emphasis>
: apexes rounded not reaching half eye.
<emphasis id="7EDDEA8C69049E16FDAEFA6381986E06" box="[612,714,1529,1554]" italics="true" pageId="9" pageNumber="329">Antennas</emphasis>
: trilamellate, with robust lamellae, the distal being wider than the other two.
<emphasis id="7EDDEA8C69049E16FD9CF98781B06E21" bold="true" box="[598,738,1565,1589]" pageId="9" pageNumber="329">Mouthparts</emphasis>
(
<figureCitation id="D4922A1B69049E16FD38F987800B6E21" box="[754,857,1565,1589]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="9" pageNumber="329">Figures 3</figureCitation>
(b), 4(b)):
<taxonomicName id="8BA94D1D69049E16FC0FF98687556E21" box="[965,1031,1564,1589]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="9" pageNumber="329" phylum="Platyhelminthes" rank="genus">
<emphasis id="7EDDEA8C69049E16FC0FF98687556E21" box="[965,1031,1564,1589]" italics="true" pageId="9" pageNumber="329">Ligula</emphasis>
</taxonomicName>
: tridentate with a larger middle tooth and narrower than the lateral teeth, which are inconspicuous; punctuations mainly concentrated in the median region.
<emphasis id="7EDDEA8C69049E16FC9EF9FB87666E6E" box="[852,1076,1633,1658]" italics="true" pageId="9" pageNumber="329">Hypostomal process</emphasis>
: wide, glabrous and near the mentum.
<emphasis id="7EDDEA8C69049E16FDE1F91E801C6E89" box="[555,846,1668,1693]" italics="true" pageId="9" pageNumber="329">Mentum mediobasal area</emphasis>
: not dilated, smooth and glabrous, with non-protruding anterior region, notched in the middle.
<emphasis id="7EDDEA8C69049E15FC12F93C838A68B2" italics="true" lastPageId="10" lastPageNumber="330" pageId="9" pageNumber="329">Mentum lateral lobes</emphasis>
: rounded apexes.
<emphasis id="7EDDEA8C69079E15FE6EFF1781D468B2" box="[420,646,141,166]" italics="true" pageId="10" pageNumber="330">Mentum lateral scars</emphasis>
: deep and oval-shape.
<emphasis id="7EDDEA8C69079E15FC4DFF1780A468B2" box="[903,1014,141,166]" italics="true" pageId="10" pageNumber="330">Mandibles</emphasis>
: incisor lobe with three well-formed teeth at apex; robust suprainternal teeth; infrabasal pits absent.
<emphasis id="7EDDEA8C69079E15FF6AFF4983BD68F8" box="[160,239,211,236]" italics="true" pageId="10" pageNumber="330">Maxilla</emphasis>
: lacinia bidentate at the apex.
<emphasis id="7EDDEA8C69079E15FDB9FF4E81B968F8" bold="true" box="[627,747,212,236]" pageId="10" pageNumber="330">Prothorax</emphasis>
. Pronotum (
<figureCitation id="D4922A1B69079E15FC4FFF4E80A068F8" box="[901,1010,212,236]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figures 3</figureCitation>
(a,c), 4(a
<emphasis id="7EDDEA8C69079E15FB9FFF49873068FF" box="[1109,1122,211,235]" italics="true" pageId="10" pageNumber="330"></emphasis>
c)):
<emphasis id="7EDDEA8C69079E15FF6AFF6F8261691A" box="[160,307,245,270]" italics="true" pageId="10" pageNumber="330">Anterior edge</emphasis>
: slightly sinuous.
<emphasis id="7EDDEA8C69079E15FE37FF6F81F1691A" box="[509,675,245,270]" italics="true" pageId="10" pageNumber="330">Anterior angles</emphasis>
: straight and slightly protruding.
<emphasis id="7EDDEA8C69079E15FBE8FF6F83B96925" italics="true" pageId="10" pageNumber="330">Marginal groove</emphasis>
: well-marked, deep in the lateral region, with coarse punctures throughout its extension, without dilation at the apex; reaching almost two-thirds of the pronotal width in the anterior margin.
<emphasis id="7EDDEA8C69079E15FE77FEC7810B6962" box="[445,601,349,374]" italics="true" pageId="10" pageNumber="330">Lateral fossae</emphasis>
: small, well marked and slighty deep; slightly rounded shape.
<emphasis id="7EDDEA8C69079E15FE91FE1A811D698D" box="[347,591,384,409]" italics="true" pageId="10" pageNumber="330">Pronotal punctuations</emphasis>
: practically absent, formed by a group of two to five punctures concentrated only in the lateral fossae. Prosternum (
<figureCitation id="D4922A1B69079E15FC78FE3E874F69A8" box="[946,1053,420,444]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figures 3</figureCitation>
(b), 4(d)):
<emphasis id="7EDDEA8C69079E15FF6AFE5F827D69CA" box="[160,303,453,478]" italics="true" pageId="10" pageNumber="330">Prepisternum</emphasis>
: not dilated in the anterior region; fine pubescence throughout the lateroposterior region, being almost inconspicuously present on the margins of the anterior region.
<emphasis id="7EDDEA8C69079E15FF30FD91822C6A30" box="[250,382,523,548]" italics="true" pageId="10" pageNumber="330">Prepimerum</emphasis>
: glabrous.
<emphasis id="7EDDEA8C69079E15FDCBFD9181C26A30" box="[513,656,523,548]" italics="true" pageId="10" pageNumber="330">Prosternelum</emphasis>
: rhomboid with broad and rounded apex.
<emphasis id="7EDDEA8C69079E15FF6AFDB482116A52" bold="true" box="[160,323,558,582]" pageId="10" pageNumber="330">Mesosternum</emphasis>
(
<figureCitation id="D4922A1B69079E15FE9EFDB482EC6A52" box="[340,446,558,582]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figures 3</figureCitation>
(b), 4(d)): smooth, shiny and glabrous.
<emphasis id="7EDDEA8C69079E15FCB1FDB7876E6A52" box="[891,1084,557,582]" italics="true" pageId="10" pageNumber="330">Mesosternal scars</emphasis>
: long, broad and deep; without punctuations or pubescence.
<emphasis id="7EDDEA8C69079E15FCC6FDCB80FE6A7D" bold="true" box="[780,940,593,617]" pageId="10" pageNumber="330">Metasternum</emphasis>
(
<figureCitation id="D4922A1B69079E15FC71FDCB87706A7D" box="[955,1058,593,617]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figures 3</figureCitation>
(b), 4(e)):
<emphasis id="7EDDEA8C69079E15FF6AFDE8821D6A9F" box="[160,335,626,651]" italics="true" pageId="10" pageNumber="330">Metasternal disc</emphasis>
: poorly delimited, but more dilated than the lateral region of the metasternum.
<emphasis id="7EDDEA8C69079E15FF26FD0F81566ABA" box="[236,516,661,686]" italics="true" pageId="10" pageNumber="330">Metasternal punctuations</emphasis>
: group of punctures only in the lateroposterior region.
<emphasis id="7EDDEA8C69079E15FF6AFD2282F46AC5" box="[160,422,696,721]" italics="true" pageId="10" pageNumber="330">Metasternal pubescence</emphasis>
: absent.
<emphasis id="7EDDEA8C69079E15FDC7FD22807C6AC5" box="[525,814,696,721]" italics="true" pageId="10" pageNumber="330">Metasternal lateral groove</emphasis>
: narrow, thinner than mesotibiae, deep, without punctures or bristles.
<emphasis id="7EDDEA8C69079E15FDBDFD4181EF6AE7" bold="true" box="[631,701,731,755]" pageId="10" pageNumber="330">Elytra</emphasis>
(
<figureCitation id="D4922A1B69079E15FD06FD41807A6AE7" box="[716,808,731,755]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figure 3</figureCitation>
(a)): Approximately 2.6x longer and 1.2x wider than pronotum.
<emphasis id="7EDDEA8C69079E15FDC0FD6781156B02" box="[522,583,765,790]" italics="true" pageId="10" pageNumber="330">Striae</emphasis>
: narrower than the interstriae, marked with round and inconspicuous punctuations on the dorsal striae and well defined in lateral striae.
<taxonomicName id="8BA94D1D69079E15FF6AFCD882576B4F" authorityName="Walker" authorityYear="1861" box="[160,261,834,859]" class="Insecta" family="Uraniidae" genus="Epipleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="7EDDEA8C69079E15FF6AFCD882576B4F" box="[160,261,834,859]" italics="true" pageId="10" pageNumber="330">Epipleura</emphasis>
</taxonomicName>
: glabrous.
<emphasis id="7EDDEA8C69079E15FEB5FCD8829C6B4F" box="[383,462,834,859]" italics="true" pageId="10" pageNumber="330">Humeri</emphasis>
: finely pubescent.
<emphasis id="7EDDEA8C69079E15FD6AFCD981856B4F" bold="true" box="[672,727,835,859]" pageId="10" pageNumber="330">Legs</emphasis>
(
<figureCitation id="D4922A1B69079E15FD2FFCD980196B4F" box="[741,843,835,859]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figures 3</figureCitation>
(a,b), 4(e)):
<emphasis id="7EDDEA8C69079E15FC0CFCD887786B4F" box="[966,1066,834,859]" italics="true" pageId="10" pageNumber="330">Profemur</emphasis>
: ventral anterior border with poorly marked groove, not reaching the apex of the profemur; ventral posterior border with few setae only near the apex.
<emphasis id="7EDDEA8C69079E15FC4DFC1280B56BB5" box="[903,999,904,929]" italics="true" pageId="10" pageNumber="330">Protibiae</emphasis>
: not dilated.
<emphasis id="7EDDEA8C69079E15FF6AFC3082476BD7" box="[160,277,938,963]" italics="true" pageId="10" pageNumber="330">Mesotibiae</emphasis>
: one or two small spines on the outer face.
<emphasis id="7EDDEA8C69079E15FCE8FC3080C46BD7" box="[802,918,938,963]" italics="true" pageId="10" pageNumber="330">Metatibiae</emphasis>
: one small spine on the outer face.
<emphasis id="7EDDEA8C69079E15FE9CFC5482996BF2" bold="true" box="[342,459,974,998]" pageId="10" pageNumber="330">Abdomen</emphasis>
(
<figureCitation id="D4922A1B69079E15FE15FC5481126BF2" box="[479,576,974,998]" captionStart="Figure 3" captionStartId="8.[160,225,1482,1505]" captionTargetBox="[163,1153,142,1451]" captionTargetId="figure-55@8.[163,1153,142,1451]" captionTargetPageId="8" captionText="Figure 3. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="10" pageNumber="330">Figure 3</figureCitation>
(b)): sternite VII with well-marked and incomplete groove; smooth sides.
<emphasis id="7EDDEA8C69079E15FE6BFC6B81456C1D" bold="true" box="[417,535,1009,1033]" pageId="10" pageNumber="330">Aedeagus</emphasis>
(
<figureCitation id="D4922A1B69079E15FDEDFC6B81D46C1D" box="[551,646,1009,1033]" captionStart="Figure 4" captionStartId="9.[160,225,1329,1352]" captionTargetBox="[163,1153,142,1297]" captionTargetId="figure-112@9.[163,1153,142,1298]" captionTargetPageId="9" captionText="Figure 4. Passalus (Pertinax) cuneatus Hincks, 1950 male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4609067" httpUri="https://zenodo.org/record/4609067/files/figure.png" pageId="10" pageNumber="330">Figure 4</figureCitation>
(f)): Median lobe with smaller width than the parameres and the basal piece together and almost as long as these. In ventral view with two sclerotised plaques on the lateral margin; basal piece and parameres fused forming a tegmen; parameres projected upwardly encapsulating the median lobe; anterior margin of parameres straight in the
<emphasis id="7EDDEA8C69079E15FE39FBE182AA6C87" box="[499,504,1147,1171]" italics="true" pageId="10" pageNumber="330"></emphasis>
V
<emphasis id="7EDDEA8C69079E15FDCDFBE1815E6C87" box="[519,524,1147,1171]" italics="true" pageId="10" pageNumber="330"></emphasis>
-shape. In lateral view, the apex of the parameres are slightly rounded. In dorsal view with widely separated parameres.
</paragraph>
<caption id="18D6661669059E17FF6AFA5082F16DEB" ID-DOI="http://doi.org/10.5281/zenodo.4609065" ID-Zenodo-Dep="4609065" httpUri="https://zenodo.org/record/4609065/files/figure.png" pageId="8" pageNumber="328" startId="8.[160,225,1482,1505]" targetBox="[163,1153,142,1451]" targetPageId="8">
<paragraph id="4C16369E69059E17FF6AFA5082F16DEB" blockId="8.[160,1155,1482,1535]" pageId="8" pageNumber="328">
<emphasis id="7EDDEA8C69059E17FF6AFA5083AF6DF5" bold="true" box="[160,253,1482,1505]" pageId="8" pageNumber="328">Figure 3.</emphasis>
<taxonomicName id="8BA94D1D69059E17FEC2FA5081FA6DF5" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[264,680,1482,1505]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="328" phylum="Arthropoda" rank="species" species="cuneatus" subGenus="Pertinax">
<emphasis id="7EDDEA8C69059E17FEC2FA50820A6DF5" box="[264,344,1482,1505]" italics="true" pageId="8" pageNumber="328">Passalus</emphasis>
(
<emphasis id="7EDDEA8C69059E17FEA0FA5082EA6DF5" box="[362,440,1482,1505]" italics="true" pageId="8" pageNumber="328">Pertinax</emphasis>
)
<emphasis id="7EDDEA8C69059E17FE03FA50814D6DF5" box="[457,543,1482,1505]" italics="true" pageId="8" pageNumber="328">cuneatus</emphasis>
<bibRefCitation id="28384B6F69059E17FDE3FA5081FA6DF5" author="Hincks WD" box="[553,680,1482,1505]" pageId="8" pageNumber="328" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
</taxonomicName>
male holotype: (a) habitus dorsal; (b) habitus ventral; (c) habitus lateral.
</paragraph>
</caption>
<caption id="18D6661669049E16FF6AFAAB80ED6D72" ID-DOI="http://doi.org/10.5281/zenodo.4609067" ID-Zenodo-Dep="4609067" httpUri="https://zenodo.org/record/4609067/files/figure.png" pageId="9" pageNumber="329" startId="9.[160,225,1329,1352]" targetBox="[163,1153,142,1297]" targetPageId="9">
<paragraph id="4C16369E69049E16FF6AFAAB80ED6D72" blockId="9.[160,1155,1328,1382]" pageId="9" pageNumber="329">
<emphasis id="7EDDEA8C69049E16FF6AFAAB83AB6D5C" bold="true" box="[160,249,1329,1352]" pageId="9" pageNumber="329">Figure 4.</emphasis>
<taxonomicName id="8BA94D1D69049E16FECAFAAA81DC6D5C" authority="Hincks, 1950" authorityName="Hincks" authorityYear="1950" box="[256,654,1328,1352]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="329" phylum="Arthropoda" rank="species" species="cuneatus" subGenus="Pertinax">
<emphasis id="7EDDEA8C69049E16FECAFAAA82026D53" box="[256,336,1328,1351]" italics="true" pageId="9" pageNumber="329">Passalus</emphasis>
(
<emphasis id="7EDDEA8C69049E16FE97FAAA82F96D53" box="[349,427,1328,1351]" italics="true" pageId="9" pageNumber="329">Pertinax</emphasis>
)
<emphasis id="7EDDEA8C69049E16FE72FAAA815C6D53" box="[440,526,1328,1351]" italics="true" pageId="9" pageNumber="329">cuneatus</emphasis>
<bibRefCitation id="28384B6F69049E16FDD9FAAB81DC6D5C" author="Hincks WD" box="[531,654,1329,1352]" pageId="9" pageNumber="329" pagination="1033 - 1047" refId="ref13231" refString="Hincks WD. 1950. Systematic and Faunistic notes on American Passalidae (Col.). Ann Mag Nat Hist. 3 (36): 1033 - 1047. doi: 10.1080 / 00222935008654117" type="journal article" year="1950">Hincks, 1950</bibRefCitation>
</taxonomicName>
male holotype: (a) head; (b) mentum; (c) head and pronotum in dorsolateral view; (d) mesosternum; (e) metasternum; (f) aedeagus.
</paragraph>
</caption>
<paragraph id="4C16369E69079E15FF6AFB7982996EAB" blockId="10.[160,1156,1251,1727]" pageId="10" pageNumber="330">
<emphasis id="7EDDEA8C69079E15FF6AFB79827D6CE9" bold="true" box="[160,303,1251,1277]" italics="true" pageId="10" pageNumber="330">Dimensions.</emphasis>
Total length:
<quantity id="8B519B7B69079E15FE17FB7F81606CE9" box="[477,562,1253,1277]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="330" unit="mm" value="34.0">34 mm</quantity>
; cephalic length:
<quantity id="8B519B7B69079E15FCC8FB7F800C6CE9" box="[770,862,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="330" unit="mm" value="2.8">2.8 mm</quantity>
; cephalic width:
<quantity id="8B519B7B69079E15FBEFFB7F87D26CE9" box="[1061,1152,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="330" unit="mm" value="6.8">6.8 mm</quantity>
; mediofrontal area length:
<quantity id="8B519B7B69079E15FE1FFA9D81636D0B" box="[469,561,1287,1311]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="330" unit="mm" value="1.2">1.2 mm</quantity>
; mediofrontal area width:
<quantity id="8B519B7B69079E15FCA7FA9D80986D0B" box="[877,970,1287,1311]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="330" unit="mm" value="2.7">2.7 mm</quantity>
; canthus ocular length:
<quantity id="8B519B7B69079E15FF3DFAB082026D56" box="[247,336,1322,1346]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="330" unit="mm" value="0.9">0.9 mm</quantity>
; canthus ocular width:
<quantity id="8B519B7B69079E15FD90FAB081E16D56" box="[602,691,1322,1346]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="330" unit="mm" value="0.3">0.3 mm</quantity>
; area of the mediofrontal area:
<quantity id="8B519B7B69079E15FBD7FAB087216D56" box="[1053,1139,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="330" unit="mm" value="2.5">2.5 mm</quantity>
<superScript id="BBDC9BD669079E15FBB9FABE872F6D20" attach="left" box="[1139,1149,1316,1332]" fontSize="7" pageId="10" pageNumber="330">2</superScript>
; mandibles external angle: 158º; antennal club length:
<quantity id="8B519B7B69079E15FCEFFAD780D36D71" box="[805,897,1357,1381]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="330" unit="mm" value="1.5">1.5 mm</quantity>
; antennal club width:
<quantity id="8B519B7B69079E15FF6AFAF583AE6D93" box="[160,252,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="330" unit="mm" value="1.6">1.6 mm</quantity>
; distal lamella length:
<quantity id="8B519B7B69079E15FDC0FAF581346D93" box="[522,614,1391,1415]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="330" unit="mm" value="0.6">0.6 mm</quantity>
; medial lamella length:
<quantity id="8B519B7B69079E15FC4EFAF580B26D93" box="[900,992,1391,1415]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="330" unit="mm" value="0.3">0.3 mm</quantity>
; width of the mentum at the lateral scars:
<quantity id="8B519B7B69079E15FE29FA0881696DBE" box="[483,571,1426,1450]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="330" unit="mm" value="3.5">3.5 mm</quantity>
; mentum mediobasal area width:
<quantity id="8B519B7B69079E15FC76FA0887466DBE" box="[956,1044,1426,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="330" unit="mm" value="1.6">1.6 mm</quantity>
; diameter of the mentum lateral scars:
<quantity id="8B519B7B69079E15FE20FA2F81116DD9" box="[490,579,1461,1485]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="330" unit="mm" value="0.7">0.7 mm</quantity>
; pronotal length:
<quantity id="8B519B7B69079E15FCDAFA2F803B6DD9" box="[784,873,1461,1485]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="330" unit="mm" value="7.5">7.5 mm</quantity>
; pronotal width:
<quantity id="8B519B7B69079E15FBE7FA2F87D26DD9" box="[1069,1152,1461,1485]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="330" unit="mm" value="10.0">10 mm</quantity>
; length of the pronotal anterior groove:
<quantity id="8B519B7B69079E15FDA6FA4D81946DFB" box="[620,710,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="330" unit="mm" value="1.3">1.3 mm</quantity>
; elytral length:
<quantity id="8B519B7B69079E15FCB0FA4D80B36DFB" box="[890,993,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="10" pageNumber="330" unit="mm" value="19.8">19.8 mm</quantity>
; elytral width:
<quantity id="8B519B7B69079E15FF6AFA6082566E06" box="[160,260,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="330" unit="mm" value="12.5">12.5 mm</quantity>
; humeral width:
<quantity id="8B519B7B69079E15FE77FA6081416E06" box="[445,531,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="330" unit="mm" value="9.7">9.7 mm</quantity>
; profemur length:
<quantity id="8B519B7B69079E15FD2AFA6080656E06" box="[736,823,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="330" unit="mm" value="4.1">4.1 mm</quantity>
; length of the anterior ventral groove of the profemur:
<quantity id="8B519B7B69079E15FE7EF98781596E21" box="[436,523,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="330" unit="mm" value="1.2">1.2 mm</quantity>
; protibiae width:
<quantity id="8B519B7B69079E15FD04F98780436E21" box="[718,785,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="330" unit="mm" value="1.0">1 mm</quantity>
; protarsal length:
<quantity id="8B519B7B69079E15FC10F98787626E21" box="[986,1072,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="330" unit="mm" value="3.4">3.4 mm</quantity>
; length of the last protarsomer:
<quantity id="8B519B7B69079E15FE72F9A581436E43" box="[440,529,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="330" unit="mm" value="1.1">1.1 mm</quantity>
; mesotibiae width:
<quantity id="8B519B7B69079E15FD38F9A580196E43" box="[754,843,1599,1623]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="330" unit="mm" value="0.8">0.8 mm</quantity>
; metasternal lateral groove width:
<quantity id="8B519B7B69079E15FF3BF9F882196E6E" box="[241,331,1634,1658]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="330" unit="mm" value="0.2">0.2 mm</quantity>
; metasternal disc length:
<quantity id="8B519B7B69079E15FDBDF9F881806E6E" box="[631,722,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="330" unit="mm" value="7.7">7.7 mm</quantity>
; abdomen length:
<quantity id="8B519B7B69079E15FC65F9F887586E6E" box="[943,1034,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="330" unit="mm" value="7.7">7.7 mm</quantity>
; aedeagus length:
<quantity id="8B519B7B69079E15FF36F91F820B6E89" box="[252,345,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="330" unit="mm" value="4.4">4.4 mm</quantity>
; aedeagus width:
<quantity id="8B519B7B69079E15FDFFF91F81C16E89" box="[565,659,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="330" unit="mm" value="2.6">2.6 mm</quantity>
; median lobe length:
<quantity id="8B519B7B69079E15FC55F91F80AE6E89" box="[927,1020,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="330" unit="mm" value="2.3">2.3 mm</quantity>
; paremeres projection lenght:
<quantity id="8B519B7B69079E15FEBAF93D829A6EAB" box="[368,456,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="330" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="04B3651569069E14FF6AFF1783AD691B" pageId="11" pageNumber="331" type="materials_examined">
<paragraph id="4C16369E69069E14FF6AFF1783AD691B" blockId="11.[160,1156,141,271]" pageId="11" pageNumber="331">
<emphasis id="7EDDEA8C69069E14FF6AFF17817D68B3" bold="true" box="[160,559,141,167]" italics="true" pageId="11" pageNumber="331">Material examined: type material.</emphasis>
<typeStatus id="9312883C69069E14FDF4FF1581F768B3" box="[574,677,143,167]" pageId="11" pageNumber="331" type="holotype">Holotype</typeStatus>
(
<specimenCount id="5AAFFD1769069E14FD79FF15819568B3" box="[691,711,143,167]" pageId="11" pageNumber="331" type="male"></specimenCount>
) F2439.50.
<collectingCountry id="34BE760E69069E14FC8FFF1580FC68B3" box="[837,942,143,167]" name="Panama" pageId="11" pageNumber="331">PANAMA</collectingCountry>
:
<collectingRegion id="8E6DF87C69069E14FC7CFF15875C68B3" box="[950,1038,143,167]" country="Panama" name="Chiriqui" pageId="11" pageNumber="331">Chiriquí</collectingRegion>
/
<collectingRegion id="8E6DF87C69069E14FBDCFF15872668B3" box="[1046,1140,143,167]" country="Panama" name="Panama" pageId="11" pageNumber="331">Panama</collectingRegion>
// Ex Staudinger &amp;/Bang Haas, Dresden//
<taxonomicName id="8BA94D1D69069E14FD8EFF28804468DE" authority="" box="[580,790,178,202]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="subGenus" subGenus="Pertinax">Passalus (Pertinax)</taxonomicName>
/
<taxonomicName id="8BA94D1D69069E14FCD4FF28808868DE" authority="Hincks" authorityName="Hincks" authorityYear="1950" box="[798,986,178,202]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="species" species="cuneatus">cuneatus Hincks</taxonomicName>
/Type/det. W.D. Hincks//Manchester Museum/
<typeStatus id="9312883C69069E14FE39FF4E812B68F8" box="[499,633,212,236]" pageId="11" pageNumber="331" type="holotype">HOLOTYPE</typeStatus>
//
<specimenCount id="5AAFFD1769069E14FD4DFF4E81C968F8" box="[647,667,212,236]" pageId="11" pageNumber="331" type="male"></specimenCount>
dissect/S. Boucher det. 2012//F2439.50 (
<collectionCode id="2AB8AE5B69069E14FF6CFF6D83A7691B" box="[166,245,247,271]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="11" pageNumber="331">MMUE</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="04B3651569069E14FF6AFEA182536A16" pageId="11" pageNumber="331" type="discussion">
<paragraph id="4C16369E69069E14FF6AFEA182536A16" blockId="11.[160,1156,315,514]" pageId="11" pageNumber="331">
<emphasis id="7EDDEA8C69069E14FF6AFEA1825E6941" bold="true" box="[160,268,315,341]" italics="true" pageId="11" pageNumber="331">Remarks.</emphasis>
Hincks comments that this species is similar to
<taxonomicName id="8BA94D1D69069E14FCA2FEA182476963" authority="Percheron" authorityName="Percheron" authorityYear="1835" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="7EDDEA8C69069E14FCA2FEA187D16940" box="[872,1155,315,340]" italics="true" pageId="11" pageNumber="331">Passalus punctatostriatus</emphasis>
Percheron
</taxonomicName>
; however, we believe it
<emphasis id="7EDDEA8C69069E14FDFAFEC581676963" box="[560,565,351,375]" italics="true" pageId="11" pageNumber="331"></emphasis>
s more similar to
<taxonomicName id="8BA94D1D69069E14FCC6FEC4874A6963" authority="(Kirsch)" baseAuthorityName="Kirsch" box="[780,1048,350,375]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="7EDDEA8C69069E14FCC6FEC480EA6963" box="[780,952,350,375]" italics="true" pageId="11" pageNumber="331">P. aequatorialis</emphasis>
(Kirsch)
</taxonomicName>
, because both species have large body size (&lt;
<quantity id="8B519B7B69069E14FD80FE1881F0698E" box="[586,674,386,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="331" unit="mm" value="30.0">30 mm</quantity>
), macropterous, smooth frontal area with few punctures and pronotal and metasternal punctuations practically absent. However, the set of diagnostic characters of
<taxonomicName id="8BA94D1D69069E14FDE1FE5C81FB69CB" authorityName="Hincks" authorityYear="1950" box="[555,681,454,479]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="7EDDEA8C69069E14FDE1FE5C81FB69CB" box="[555,681,454,479]" italics="true" pageId="11" pageNumber="331">P. cuneatus</emphasis>
</taxonomicName>
distinguishes it from the other species of
<taxonomicName id="8BA94D1D69069E14FF6AFE7383A96A16" authorityName="Fabricius" authorityYear="1792" box="[160,251,489,514]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="7EDDEA8C69069E14FF6AFE7383A96A16" box="[160,251,489,514]" italics="true" pageId="11" pageNumber="331">Passalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="04B3651569069E14FF6AFDB781436A53" box="[160,529,557,583]" pageId="11" pageNumber="331" type="distribution">
<paragraph id="4C16369E69069E14FF6AFDB781436A53" blockId="11.[160,529,557,583]" box="[160,529,557,583]" pageId="11" pageNumber="331">
<emphasis id="7EDDEA8C69069E14FF6AFDB782616A53" bold="true" box="[160,307,557,583]" italics="true" pageId="11" pageNumber="331">Distribution.</emphasis>
<collectingCountry id="34BE760E69069E14FE89FDB582CF6A53" box="[323,413,559,583]" name="Panama" pageId="11" pageNumber="331">Panama</collectingCountry>
(
<collectingRegion id="8E6DF87C69069E14FE60FDB581556A53" box="[426,519,559,583]" country="Panama" name="Chiriqui" pageId="11" pageNumber="331">Chiriquí</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>