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<document id="54EBC6B60EC96D4964C0525F908C14EA" ID-CLB-Dataset="301504" ID-DOI="10.1093/zooldnnean/zlad146" ID-GBIF-Dataset="5e66aeaf-44c5-4be9-924d-cc13680ace6a" ID-ISSN="0024-4082" ID-Zenodo-Dep="13315269" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722845279467" checkinUser="plazi" docAuthor="Lumen, Ryan &amp; Kamdńskd, Marcdn J." docDate="2024" docId="C27387A7FFE7FFC89A8807DFFE5C7174" docLanguage="en" docName="zlad146.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Biolus sauroides, comb. nov." docType="treatment" docVersion="6" lastPageNumber="379" masterDocId="3E4AFFDFFFF2FFDD9A72053DFF83770F" masterDocTitle="evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae" masterLastPageNumber="386" masterPageNumber="358" pageNumber="379" updateTime="1732206558056" updateUser="felipe">
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<mods:title id="04364CF9CED69E887FC3C6D7433D5BE9">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="F0D601D36F65F3578FFE1321E3E903D6">Lumen, Ryan</mods:namePart>
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<mods:date id="39EF10D471116BEC52B56FEA060BCF5C">2024</mods:date>
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<treatment id="C27387A7FFE7FFC89A8807DFFE5C7174" ID-DOI="http://doi.org/10.5281/zenodo.13323872" ID-GBIF-Taxon="238560663" ID-Zenodo-Dep="13323872" LSID="urn:lsid:plazi:treatment:C27387A7FFE7FFC89A8807DFFE5C7174" httpUri="http://treatment.plazi.org/id/C27387A7FFE7FFC89A8807DFFE5C7174" lastPageNumber="379" pageId="21" pageNumber="379">
<subSubSection id="02C0653AFFE7FFC89A8807DFFD1375F3" box="[250,656,738,765]" pageId="21" pageNumber="379" type="nomenclature">
<paragraph id="4A6536B1FFE7FFC89A8807DFFD1375F3" blockId="21.[250,656,738,765]" box="[250,656,738,765]" pageId="21" pageNumber="379">
<heading id="112D81DDFFE7FFC89A8807DFFD1375F3" box="[250,656,738,765]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="379" reason="4">
<emphasis id="78AEEAA3FFE7FFC89A8807DFFD1375F3" bold="true" box="[250,656,738,765]" pageId="21" pageNumber="379">
8.
<taxonomicName id="8DDA4D32FFE7FFC89B6307DFFD9975F3" authority="(Koch)" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[273,538,738,765]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="sauroides" status="comb. nov.">
<emphasis id="78AEEAA3FFE7FFC89B6307DFFE4B75F2" bold="true" box="[273,456,738,765]" italics="true" pageId="21" pageNumber="379">Biolus sauroides</emphasis>
(Koch)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE7FFC8986D07DFFD1375F3" box="[543,656,738,764]" pageId="21" pageNumber="379" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE7FFC89B2F061DFC897450" pageId="21" pageNumber="379" type="description">
<paragraph id="4A6536B1FFE7FFC89B2F061DFDAD7435" blockId="21.[129,778,800,863]" box="[349,558,800,827]" pageId="21" pageNumber="379">
<emphasis id="78AEEAA3FFE7FFC89B2F061DFDAD7435" bold="true" box="[349,558,800,827]" pageId="21" pageNumber="379">
(
<figureCitation id="D2E12A34FFE7FFC89B15061CFE3A7434" box="[359,441,801,827]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.species.A, Biolus analisetosus (Koch) comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey) stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey) comb. nov. Lectotype; E, B. graafi (Koch) comb. nov. Holotype; F,B. granulatus (Fabricius) comb. nov. Lectotype; G, B. omeri (Koch) comb.nov.Holotype; H, B. sauroides (Koch) comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." figureDoi="http://doi.org/10.5281/zenodo.13315277" httpUri="https://zenodo.org/record/13315277/files/figure.png" pageId="21" pageNumber="379">Figs 2H</figureCitation>
,
<figureCitation id="D2E12A34FFE7FFC89BB1061CFE757434" box="[451,502,801,827]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used in the cladistic analysis.Arrows indicate particular states of given characters from the morphological matrix [e.g." figureDoi="http://doi.org/10.5281/zenodo.13315289" httpUri="https://zenodo.org/record/13315289/files/figure.png" pageId="21" pageNumber="379">6A, I</figureCitation>
,
<figureCitation id="D2E12A34FFE7FFC89872061CFDA07434" box="[512,547,801,827]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotina species.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres highlighted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="21" pageNumber="379">8H</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="4A6536B1FFE7FFC89AF30675FC897450" blockId="21.[129,778,800,863]" box="[129,778,839,863]" pageId="21" pageNumber="379">
<taxonomicName id="8DDA4D32FFE7FFC89AF30675FF627450" ID-CoL="BF4MM" box="[129,225,840,863]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="21" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE7FFC89AF30675FF627450" box="[129,225,840,863]" italics="true" pageId="21" pageNumber="379">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE7FFC89A9C067AFDAF7450" authority="Koch 1955: 420" authorityName="Koch" authorityPageNumber="420" authorityYear="1955" box="[238,556,839,863]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE7FFC89A9C067AFEA47450" box="[238,295,839,863]" italics="true" pageId="21" pageNumber="379">Biolus</emphasis>
)
<emphasis id="78AEEAA3FFE7FFC89B47067AFE097450" box="[309,394,839,863]" italics="true" pageId="21" pageNumber="379">sauroides</emphasis>
Koch 1955: 420
</taxonomicName>
. Kamiński 2016: 235.
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE7FFC89AF306BCFE5C7174" pageId="21" pageNumber="379" type="diagnosis">
<paragraph id="4A6536B1FFE7FFC89AF306BCFF78737B" blockId="21.[129,778,897,1140]" pageId="21" pageNumber="379">
<emphasis id="78AEEAA3FFE7FFC89AF306BCFE067496" box="[129,389,897,921]" italics="true" pageId="21" pageNumber="379">
Diagnosis:
<taxonomicName id="8DDA4D32FFE7FFC89A9906BCFE067496" authorityName="Koch" authorityYear="1955" box="[235,389,897,921]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="sauroides">Biolus sauroides</taxonomicName>
</emphasis>
is sharply separable from all other spe - cies by the extremely coarse sculpturing dorsally and ventrally (large, knobby tubercles extending to the disc and base of the elytra and large closely spaced punctures on abdominal under - side). Kochs (1955) description outlines most of the species morphology sufficiently. Here we provide only additional char - acters we found in a description below to supplement Kochs description.
</paragraph>
<paragraph id="4A6536B1FFE7FFC89AF3019CFF4B727B" blockId="21.[129,778,1185,1396]" pageId="21" pageNumber="379">
<emphasis id="78AEEAA3FFE7FFC89AF3019CFD0073B6" box="[129,643,1185,1209]" italics="true" pageId="21" pageNumber="379">Supplementary description (see: Koch 1955):</emphasis>
Abdomen: Abdominal ventrite V with setigerous punctures. Females without median apical notch. Legs: Male midtibia with small spurical projection at apex. Male terminalia: Parameres weakly tapering with tips sharply tapering and directed medially. Female Terminalia: Bursa copulatrix without sclerites or accessory pouch.
</paragraph>
<paragraph id="4A6536B1FFE7FFC89AF3009CFEFB72F7" blockId="21.[129,778,1441,1590]" pageId="21" pageNumber="379">
<emphasis id="78AEEAA3FFE7FFC89AF3009CFEC272B6" box="[129,321,1441,1465]" italics="true" pageId="21" pageNumber="379">Material examined:</emphasis>
<typeStatus id="95618813FFE7FFC89B35009CFE2972B6" box="[327,426,1441,1465]" pageId="21" pageNumber="379" type="holotype">Holotype</typeStatus>
(TMNH): Garies Namaqualand; Museum Staff
<date id="3E641071FFE7FFC89B6600FCFEFC72D6" box="[276,383,1473,1497]" pageId="21" pageNumber="379" value="1930-06">June 1930</date>
;
<taxonomicName id="8DDA4D32FFE7FFC89BFA00FDFDAA72D7" authority="Koch" authorityName="Koch" box="[392,553,1472,1496]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="sauroddes">sauroides Koch</taxonomicName>
;
<typeStatus id="95618813FFE7FFC8984100FCFD1B72D6" box="[563,664,1473,1497]" pageId="21" pageNumber="379" type="paratype">Paratypus</typeStatus>
<taxonomicName id="8DDA4D32FFE7FFC898EF00FCFEB872F7" authority="C. Koch" authorityName="C. Koch" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="21" pageNumber="379" phylum="Arthropoda" rank="species" species="sauroddes" status="sp. nov.">Eurynotus sauroides C. Koch</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE7FFC89B3200DDFEF272F7" box="[320,369,1504,1528]" pageId="21" pageNumber="379" rank="species">sp. n.</taxonomicNameLabel>
.
</paragraph>
<paragraph id="4A6536B1FFE7FFC89AEE00C2FEAD7139" blockId="21.[129,778,1441,1590]" pageId="21" pageNumber="379">(See Supporting Information, Appendix S1 for additional ma - terial examined.)</paragraph>
<paragraph id="4A6536B1FFE7FFC89AF3035EFE5C7174" blockId="21.[129,479,1635,1659]" box="[129,479,1635,1659]" pageId="21" pageNumber="379">
<emphasis id="78AEEAA3FFE7FFC89AF3035EFF7D7174" box="[129,254,1635,1659]" italics="true" pageId="21" pageNumber="379">Distribution:</emphasis>
South Africa (Fig. 9).
</paragraph>
</subSubSection>
</treatment>
</document>