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<document id="231A4D7A0D44AA3F911AE759BD324F7A" ID-CLB-Dataset="301504" ID-DOI="10.1093/zooldnnean/zlad146" ID-GBIF-Dataset="5e66aeaf-44c5-4be9-924d-cc13680ace6a" ID-ISSN="0024-4082" ID-Zenodo-Dep="13315269" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722845279467" checkinUser="plazi" docAuthor="Lumen, Ryan &amp; Kamdńskd, Marcdn J." docDate="2024" docId="C27387A7FFE4FFCA99D0006CFC7770B4" docLanguage="en" docName="zlad146.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Eurynotus asperatus Mulsant and Rey" docType="treatment" docVersion="4" lastPageNumber="381" masterDocId="3E4AFFDFFFF2FFDD9A72053DFF83770F" masterDocTitle="evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae" masterLastPageNumber="386" masterPageNumber="358" pageNumber="380" updateTime="1732206558056" updateUser="felipe">
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<mods:title id="384C3ADB465965AF7BCDD33BD5342A34">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="595164939BFEF1340CE21050E5727B24">Lumen, Ryan</mods:namePart>
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<mods:date id="6AC3BEA954BBEA60271EF75CD96B5260">2024</mods:date>
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<treatment id="C27387A7FFE4FFCA99D0006CFC7770B4" ID-DOI="http://doi.org/10.5281/zenodo.13323874" ID-GBIF-Taxon="238560652" ID-Zenodo-Dep="13323874" LSID="urn:lsid:plazi:treatment:C27387A7FFE4FFCA99D0006CFC7770B4" httpUri="http://treatment.plazi.org/id/C27387A7FFE4FFCA99D0006CFC7770B4" lastPageId="23" lastPageNumber="381" pageId="22" pageNumber="380">
<subSubSection id="02C0653AFFE4FFCB99D0006CFAB87264" box="[930,1339,1361,1387]" pageId="22" pageNumber="380" type="nomenclature">
<paragraph id="4A6536B1FFE4FFCB99D0006CFAB87264" blockId="22.[930,1339,1361,1387]" box="[930,1339,1361,1387]" pageId="22" pageNumber="380">
<heading id="112D81DDFFE4FFCB99D0006CFAB87264" box="[930,1339,1361,1387]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="380" reason="4">
<emphasis id="78AEEAA3FFE4FFCB99D0006CFAB87264" bold="true" box="[930,1339,1361,1387]" pageId="22" pageNumber="380">
2.
<taxonomicName id="8DDA4D32FFE4FFCB99C8006CFAB87264" authority="Mulsant and Rey" authorityName="Mulsant and Rey" box="[954,1339,1361,1387]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB99C8006CFB067264" bold="true" box="[954,1157,1361,1387]" italics="true" pageId="22" pageNumber="380">Eurynotus asperatus</emphasis>
Mulsant and Rey
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE4FFCB998700BDFB6A7295" box="[1013,1257,1408,1434]" pageId="22" pageNumber="380" type="description">
<paragraph id="4A6536B1FFE4FFCB998700BDFB6A7295" blockId="22.[1013,1257,1408,1434]" box="[1013,1257,1408,1434]" pageId="22" pageNumber="380">
<heading id="112D81DDFFE4FFCB998700BDFB6A7295" box="[1013,1257,1408,1434]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="380" reason="3">
<emphasis id="78AEEAA3FFE4FFCB998700BDFB6A7295" bold="true" box="[1013,1257,1408,1434]" pageId="22" pageNumber="380">
(
<figureCitation id="D2E12A34FFE4FFCB998D00BDFBE97295" box="[1023,1130,1408,1434]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.species. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabricius) Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." figureDoi="http://doi.org/10.5281/zenodo.13315279" httpUri="https://zenodo.org/record/13315279/files/figure.png" pageId="22" pageNumber="380">Figs 3B, C</figureCitation>
,
<figureCitation id="D2E12A34FFE4FFCB9E0600BDFB127295" box="[1140,1169,1408,1434]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used in the cladistic analysis.Arrows indicate particular states of given characters from the morphological matrix [e.g." figureDoi="http://doi.org/10.5281/zenodo.13315289" httpUri="https://zenodo.org/record/13315289/files/figure.png" pageId="22" pageNumber="380">6B</figureCitation>
,
<figureCitation id="D2E12A34FFE4FFCB9EEE00BDFB3E7295" box="[1180,1213,1408,1434]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovivipary, and female reproductive structures of Eurynotina.A, Eurynotus barbosai, 1st instar larva dissected from bursa copulatrix. B, Biolus omeri, genital tubes. C, B. brunoi, spermatheca and ovipositor.D, E. capensis, genital tubes of two individuals.E, B.analisetosus, sclerite and egg dissected from bursa copulatrix. F, B. granulatus, sclerite from bursa copulatrix.G, E. asperatus, paired sclerites on distal part of bursa copulatrix. H, Oncotus cedrimontis, bursa copulatrix. Abbreviations: c1–c4, lobes of coxities. Numbers in parentheses represent characters and states used in analysis." figureDoi="http://doi.org/10.5281/zenodo.13323862" httpUri="https://zenodo.org/record/13323862/files/figure.png" pageId="22" pageNumber="380">7G</figureCitation>
,
<figureCitation id="D2E12A34FFE4FFCB9EB500BDFB5D7295" box="[1223,1246,1408,1434]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotina species.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres highlighted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="22" pageNumber="380">8I</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE4FFCB99580095FC7C7133" pageId="22" pageNumber="380" type="reference_group">
<paragraph id="4A6536B1FFE4FFCB99580095FA1472F2" blockId="22.[810,1459,1447,1596]" pageId="22" pageNumber="380">
<treatmentCitationGroup id="6ACA119FFFE4FFCB99580095FA1472F2" pageId="22" pageNumber="380">
<taxonomicName id="8DDA4D32FFE4FFCB99580095FC0D72B0" ID-CoL="BF4MM" authorityName="Kdrby" authorityYear="1819" box="[810,910,1448,1471]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE4FFCB99580095FC0D72B0" box="[810,910,1448,1471]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE4FFCB99D10095FA2C72B0" authority="Mulsant and Rey, 1854 a: 23" authorityName="Mulsant and Rey" authorityPageNumber="23" authorityYear="1854" box="[931,1455,1447,1471]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB99D10095FB8472B0" box="[931,1031,1448,1471]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
)
<emphasis id="78AEEAA3FFE4FFCB9E690095FBF572B0" box="[1051,1142,1448,1471]" italics="true" pageId="22" pageNumber="380">asperatus</emphasis>
Mulsant and Rey, 1854a: 23
</taxonomicName>
. Mulsant and Rey 1854b: 167; Lacordaire 1859: 239;
<treatmentCitation id="CB7B10A0FFE4FFCB9F1A00FBFC1672F1" author="Gebden H" page="414" pageId="22" pageNumber="380" year="1938">
<bibRefCitation id="2E4B4B40FFE4FFCB9F1A00FBFC1672F1" author="Gebden H" pageId="22" pageNumber="380" pagination="49 - 80" refId="ref26000" refString="Gebden H. Katalog der Tenebrioniden. Teil II. Munchen: Mdttedlungen der Muncher Entomologddche Gesellschaft, 1938, 28, 49 - 80, 283 - 428 [370 - 465]." type="journal article" year="1938">Gebien 1938: 414</bibRefCitation>
</treatmentCitation>
; Koch 1954a: 283; 1954b: 2; Kamiński 2016: 236.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4A6536B1FFE4FFCB99340338FC7C7133" blockId="22.[810,1459,1447,1596]" pageId="22" pageNumber="380">
=
<taxonomicName id="8DDA4D32FFE4FFCB99260338FC3B7113" ID-CoL="BF4MM" authorityName="Kdrby" authorityYear="1819" box="[852,952,1541,1564]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE4FFCB99260338FC3B7113" box="[852,952,1541,1564]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE4FFCB99BB0338FAC57112" authority="Koch 1955: 423" authorityName="Koch" authorityPageNumber="423" authorityYear="1955" box="[969,1350,1541,1565]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="rudebecki" status="syn. nov.">
<emphasis id="78AEEAA3FFE4FFCB99BB0338FBAE7113" box="[969,1069,1541,1564]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
)
<emphasis id="78AEEAA3FFE4FFCB9E4C0338FB197112" box="[1086,1178,1541,1565]" italics="true" pageId="22" pageNumber="380">rudebecki</emphasis>
Koch 1955: 423
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE4FFCB9F3E0338FA257113" box="[1356,1446,1541,1565]" pageId="22" pageNumber="380">syn. nov.</taxonomicNameLabel>
– Kamiński 2016: 237.
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE4FFCB99580363FB5C71BB" pageId="22" pageNumber="380" type="discussion">
<paragraph id="4A6536B1FFE4FFCB99580363FB5C71BB" blockId="22.[810,1459,1629,1716]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB99580363FCE2717A" box="[810,865,1630,1653]" italics="true" pageId="22" pageNumber="380">Note:</emphasis>
The description provided below supplements Koch’s (1954a, 1955) description of
<taxonomicName id="8DDA4D32FFE4FFCB9E270340FB49719A" box="[1109,1226,1661,1685]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="rudebecki">
<emphasis id="78AEEAA3FFE4FFCB9E270340FB49719A" box="[1109,1226,1661,1685]" italics="true" pageId="22" pageNumber="380">E. rudebecki</emphasis>
</taxonomicName>
to include new charac - ters and variation observed in this revision.
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE4FFCA995803EBFC7770B4" lastPageId="23" lastPageNumber="381" pageId="22" pageNumber="380" type="diagnosis">
<paragraph id="4A6536B1FFE4FFCB995803EBFA3070A6" blockId="22.[810,1460,1749,1961]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB995803EBFC1371E2" box="[810,912,1750,1773]" italics="true" pageId="22" pageNumber="380">Diagnosis:</emphasis>
One of the most commonly collected and identified species in collections observed in this study (alongside
<taxonomicName id="8DDA4D32FFE4FFCB9F3D03C8FCFB7023" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9F3D03C8FCFB7023" italics="true" pageId="22" pageNumber="380">Eurynotus capensis</emphasis>
</taxonomicName>
). Can be separated from
<taxonomicName id="8DDA4D32FFE4FFCB9EF90228FB767023" box="[1163,1269,1813,1836]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9EF90228FB767023" box="[1163,1269,1813,1836]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
based on size (
<taxonomicName id="8DDA4D32FFE4FFCB9FED0228FCFB7044" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9FED0228FCFB7044" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
: length
<quantity id="8D229B54FFE4FFCB99BD020EFBB87044" box="[975,1083,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="22" pageNumber="380" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20mm</quantity>
, width
<quantity id="8D229B54FFE4FFCB9EFB020EFB697044" box="[1161,1258,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="22" pageNumber="380" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10mm</quantity>
;
<taxonomicName id="8DDA4D32FFE4FFCB9E870209FAE87044" box="[1269,1387,1844,1867]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9E870209FAE87044" box="[1269,1387,1844,1867]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
length
<quantity id="8D229B54FFE4FFCB9958026EFC2A7064" box="[810,937,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="1.1125" metricValueMax="1.325" metricValueMin="0.9" pageId="22" pageNumber="380" unit="mm" value="11.125" valueMax="13.25" valueMin="9.0">9–13.25mm</quantity>
, width
<quantity id="8D229B54FFE4FFCB9983026EFBE37064" box="[1009,1120,1875,1899]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="22" pageNumber="380" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5mm</quantity>
), and characteristics of the males (modified hind tibiae in
<taxonomicName id="8DDA4D32FFE4FFCB9E5C024EFB1B7085" box="[1070,1176,1907,1930]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9E5C024EFB1B7085" box="[1070,1176,1907,1930]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
) and females (
<taxonomicName id="8DDA4D32FFE4FFCB9F49024EFA307085" box="[1339,1459,1907,1930]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9F49024EFA307085" box="[1339,1459,1907,1930]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
bursa copulatrix with sclerites;
<emphasis id="78AEEAA3FFE4FFCB9E0902AFFB6470A6" box="[1147,1255,1938,1961]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
with characteristic
</paragraph>
<paragraph id="4A6536B1FFE5FFCA9EEC0577FA417750" blockId="23.[1182,1474,74,96]" box="[1182,1474,74,96]" pageId="23" pageNumber="381">
Revision of
<taxonomicName id="8DDA4D32FFE5FFCA9F7A0576FAE3776F" ID-CoL="BF4MM" authorityName="Kdrby" authorityYear="1819" box="[1288,1376,75,96]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE5FFCA9F7A0576FAE3776F" box="[1288,1376,75,96]" italics="true" pageId="23" pageNumber="381">Eurynotus</emphasis>
</taxonomicName>
<emphasis id="78AEEAA3FFE5FFCA9F0E0576FA417750" bold="true" box="[1404,1474,74,96]" pageId="23" pageNumber="381">• 381</emphasis>
</paragraph>
<caption id="1EA56639FFE5FFCA9AF30126FEBA7341" ID-DOI="http://doi.org/10.5281/zenodo.13315297" ID-Zenodo-Dep="13315297" httpUri="https://zenodo.org/record/13315297/files/figure.png" pageId="23" pageNumber="381" startId="23.[129,194,1051,1075]" targetBox="[132,1473,145,1021]" targetPageId="23" targetType="figure">
<paragraph id="4A6536B1FFE5FFCA9AF30126FEBA7341" blockId="23.[129,1422,1051,1103]" pageId="23" pageNumber="381">
<emphasis id="78AEEAA3FFE5FFCA9AF30126FF66733C" bold="true" box="[129,229,1051,1075]" pageId="23" pageNumber="381">Figure 10.</emphasis>
Distribution and selected habits of representatives of the genus
<taxonomicName id="8DDA4D32FFE5FFCA994F0121FC19733C" authorityName="Kdrby" authorityYear="1819" box="[829,922,1052,1075]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE5FFCA994F0121FC19733C" box="[829,922,1052,1075]" italics="true" pageId="23" pageNumber="381">Eurynotus</emphasis>
</taxonomicName>
. Sample photographs of habitat for
<taxonomicName id="8DDA4D32FFE5FFCA9E980121FAC4733C" authorityName="Kdrby" authorityYear="1819" box="[1258,1351,1052,1075]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE5FFCA9E980121FAC4733C" box="[1258,1351,1052,1075]" italics="true" pageId="23" pageNumber="381">Eurynotus</emphasis>
</taxonomicName>
species marked by A and B.
</paragraph>
</caption>
<paragraph id="4A6536B1FFE5FFCA9AF30142FF657162" blockId="23.[129,778,1151,1645]" pageId="23" pageNumber="381">
spermatheca and constricted, bilobed bursa copulatrix). From other
<taxonomicName id="8DDA4D32FFE5FFCA9AB301A2FEA673B9" authorityName="Kdrby" authorityYear="1819" box="[193,293,1183,1206]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE5FFCA9AB301A2FEA673B9" box="[193,293,1183,1206]" italics="true" pageId="23" pageNumber="381">Eurynotus</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE5FFCA9B4101A2FE2873B9" box="[307,427,1183,1206]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE5FFCA9B4101A2FE2873B9" box="[307,427,1183,1206]" italics="true" pageId="23" pageNumber="381">E. asperatus</emphasis>
</taxonomicName>
can also be separated by size:
<taxonomicName id="8DDA4D32FFE5FFCA988701A2FF7173DA" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFE5FFCA988701A2FF7173DA" italics="true" pageId="23" pageNumber="381">E. graniderma</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE5FFCA9B500183FE0D73DA" box="[290,398,1213,1237]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE5FFCA9B500183FE0D73DA" box="[290,398,1213,1237]" italics="true" pageId="23" pageNumber="381">E. barbosai</emphasis>
</taxonomicName>
are smaller (length
<quantity id="8D229B54FFE5FFCA982A0180FD4773DA" box="[600,708,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="23" pageNumber="381" unit="mm" value="8.75" valueMax="9.5" valueMin="8.0">8–9.5 mm</quantity>
; width
<quantity id="8D229B54FFE5FFCA9AF301E0FE8773FA" box="[129,260,1245,1269]" metricMagnitude="-3" metricUnit="m" metricValue="5.125" metricValueMax="5.5" metricValueMin="4.75" pageId="23" pageNumber="381" unit="mm" value="5.125" valueMax="5.5" valueMin="4.75">4.75–5.5 mm</quantity>
); however,
<taxonomicName id="8DDA4D32FFE5FFCA9B0D01E0FD8473FA" box="[383,519,1245,1269]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFE5FFCA9B0D01E0FD8473FA" box="[383,519,1245,1269]" italics="true" pageId="23" pageNumber="381">E. thermarum</emphasis>
</taxonomicName>
(while generally smaller) specimens may be similar in size in large specimens (which can then be differentiated based on the width of the epipleura and presence of microtuberculation on the pronotum and elytra). The presence of additional, larger setigerous punctures on ab - dominal ventrite V (absent in
<taxonomicName id="8DDA4D32FFE5FFCA9BB50047FDC3729E" box="[455,576,1402,1425]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE5FFCA9BB50047FDC3729E" box="[455,576,1402,1425]" italics="true" pageId="23" pageNumber="381">E. asperatus</emphasis>
</taxonomicName>
) can also be diag - nostic.
<taxonomicName id="8DDA4D32FFE5FFCA9AB900A4FE1872BE" box="[203,411,1433,1457]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="privisolum">
<emphasis id="78AEEAA3FFE5FFCA9AB900A4FE1872BE" box="[203,411,1433,1457]" italics="true" pageId="23" pageNumber="381">Eurynotus privisolum</emphasis>
</taxonomicName>
, while similar at first glance, can be differentiated based on the sculpture of the pronotum and head (close, coarse punctation; versus
<taxonomicName id="8DDA4D32FFE5FFCA9BAC00E5FDD572E0" box="[478,598,1496,1519]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE5FFCA9BAC00E5FDD572E0" box="[478,598,1496,1519]" italics="true" pageId="23" pageNumber="381">E. asperatus</emphasis>
</taxonomicName>
with finer, more widely separated punctation). Additional separation of species is possible using the lustre of the cuticle (dull or opaque in
<taxonomicName id="8DDA4D32FFE5FFCA9887032AFF5F7142" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE5FFCA9887032AFF5F7142" italics="true" pageId="23" pageNumber="381">E. asperatus</emphasis>
</taxonomicName>
versus the shinier
<taxonomicName id="8DDA4D32FFE5FFCA9BD1030BFDA57142" box="[419,550,1589,1613]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="privisolum">
<emphasis id="78AEEAA3FFE5FFCA9BD1030BFDA57142" box="[419,550,1589,1613]" italics="true" pageId="23" pageNumber="381">E. privisolum</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE5FFCA9846030BFD3A7142" box="[564,697,1589,1613]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="sulcimargo">
<emphasis id="78AEEAA3FFE5FFCA9846030BFD3A7142" box="[564,697,1589,1613]" italics="true" pageId="23" pageNumber="381">E. sulcimargo</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE5FFCA9887030BFF577162" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE5FFCA9887030BFF577162" italics="true" pageId="23" pageNumber="381">E. barbosai</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4A6536B1FFE5FFCA9AF303BAFC8A70B7" blockId="23.[129,778,1671,1976]" pageId="23" pageNumber="381">
<emphasis id="78AEEAA3FFE5FFCA9AF303BAFDEA7190" box="[129,617,1671,1695]" italics="true" pageId="23" pageNumber="381">Description (to supplement Koch 1954a, 1955):</emphasis>
Dorsal surface dull with microgranulation.
<emphasis id="78AEEAA3FFE5FFCA9BC0039BFE7671B1" bold="true" box="[434,501,1702,1726]" pageId="23" pageNumber="381">Head:</emphasis>
Punctures spaced by ≥1 puncture diameter. Mentum without apical notch. Eye with im - pressed sulcus around margin. Prothorax: Pronotum finely punc - tate, punctures separated by&gt;1 puncture diameter. Hypomeron nearly smooth, at most with fine and shallow punctation and light wrinkling. Pterothorax: Elytral interval X terminating between hind coxae and apex of elytra. Epipleuron subequal to width of elytral intervals 9 + 10. Elytral intervals obscured by epipleuron in ventral view. Abdomen: Punctate. Abdominal ventrite V finely
</paragraph>
<paragraph id="4A6536B1FFE5FFCA994B0142FADF727D" blockId="23.[825,1475,1150,1394]" pageId="23" pageNumber="381">punctate, punctures separated by ≥1 puncture diameter without larger, setigerous punctures intermixed. Females without me - dian apical notch. Legs: Male and female legs unmodified. Tarsi with black or dark setae. Males: pro/meso tarsi greatly expanded, with large, ventral, tomentose pads. Male terminalia: Parameres rounded and slightly tapering apically, rounded at apex. Female terminalia: Bursa copulatrix sac -like and with paired sclerites, spermatheca thick and coiled, accessory pouch absent.</paragraph>
<paragraph id="4A6536B1FFE5FFCA994B00AEFA8E7167" blockId="23.[825,1475,1427,1827]" pageId="23" pageNumber="381">
<emphasis id="78AEEAA3FFE5FFCA994B00AEFB4872A4" box="[825,1227,1427,1451]" italics="true" pageId="23" pageNumber="381">
Material examined:
<taxonomicName id="8DDA4D32FFE5FFCA9E7600A9FB4872A4" box="[1028,1227,1428,1451]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">Eurynotus asperatus</taxonomicName>
</emphasis>
:
<materialsCitation id="FAB23CECFFE5FFCA9EA900A9FA8E7167" ID-GBIF-Occurrence="4922112302" collectionCode="MNHN" pageId="23" pageNumber="381" specimenCount="3" typeStatus="lectotype">
Lectotype, here desig - nated (
<collectionCode id="2CCBAE74FFE5FFCA99F2008EFC5072C4" box="[896,979,1459,1483]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="381">MNHN</collectionCode>
):
<collectorName id="E72F5367FFE5FFCA9982008FFB9C72C5" box="[1008,1055,1458,1482]" pageId="23" pageNumber="381">Blue</collectorName>
circle label ‘
<taxonomicName id="8DDA4D32FFE5FFCA9EEC008EFA2B72C4" authority="Cap." authorityName="Cap." box="[1182,1448,1459,1483]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="asperatus">Eurynotus asperatus Cap.</taxonomicName>
<collectorName id="E72F5367FFE5FFCA9FDF008EFCD872E5" pageId="23" pageNumber="381">B. Sp.</collectorName>
?; type
<collectorName id="E72F5367FFE5FFCA99D000EFFC7972E5" box="[930,1018,1490,1514]" pageId="23" pageNumber="381">Mulsant</collectorName>
; asperatus’ red label with handwritten ‘Type’.
<emphasis id="78AEEAA3FFE5FFCA994B00CFFCF37106" box="[825,880,1522,1545]" italics="true" pageId="23" pageNumber="381">
<collectorName id="E72F5367FFE5FFCA994B00CFFCE97106" box="[825,874,1522,1545]" pageId="23" pageNumber="381">Note</collectorName>
:
</emphasis>
<collectorName id="E72F5367FFE5FFCA990400CCFC127106" box="[886,913,1521,1545]" pageId="23" pageNumber="381">As</collectorName>
the original descriptions lack information defining the number or specificity of specimens used, a representative from the presumed syntype series is here designated as lectotype to fix the taxonomic status of the species and genus.
</materialsCitation>
</paragraph>
<paragraph id="4A6536B1FFE5FFCA99270352FACB71EB" blockId="23.[825,1475,1427,1827]" pageId="23" pageNumber="381">
<taxonomicName id="8DDA4D32FFE5FFCA99270352FB9C7189" box="[853,1055,1646,1670]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="species" species="rudebecki">
<emphasis id="78AEEAA3FFE5FFCA99270352FB9C7189" box="[853,1055,1646,1670]" italics="true" pageId="23" pageNumber="381">Eurynotus rudebecki</emphasis>
</taxonomicName>
<materialsCitation id="FAB23CECFFE5FFCA9E590352FACB71EB" ID-GBIF-Occurrence="4922112308" collectionCode="KOCH" pageId="23" pageNumber="381" specimenCount="2" typeStatus="lectotype">
<typeStatus id="95618813FFE5FFCA9E590352FB177188" box="[1067,1172,1647,1671]" pageId="23" pageNumber="381" type="lectotype">Lectotype</typeStatus>
, here designated (TMNH): SwedishSouthAfricaExpedition 1950–1951
<collectorName id="E72F5367FFE5FFCA9E8903B3FAC671A9" box="[1275,1349,1678,1702]" pageId="23" pageNumber="381">Brunck</collectorName>
-
<collectorName id="E72F5367FFE5FFCA9F2003B3FA3871A9" box="[1362,1467,1678,1702]" pageId="23" pageNumber="381">Rudebeck</collectorName>
/
<collectorName id="E72F5367FFE5FFCA994B0390FBE671CA" box="[825,1125,1709,1733]" pageId="23" pageNumber="381">S. Afr. Cape Prov Mosselbaai.</collectorName>
8–9.1.51.
<collectorName id="E72F5367FFE5FFCA9EBC0393FB7771CA" box="[1230,1268,1710,1733]" pageId="23" pageNumber="381">No.</collectorName>
127/ Holotype
<collectorName id="E72F5367FFE5FFCA9FE80393FA3D71CA" box="[1434,1470,1710,1733]" pageId="23" pageNumber="381">No</collectorName>
: 123?
<taxonomicName id="8DDA4D32FFE5FFCA991D03F0FC5971EA" authorityName="Kdrby" authorityYear="1819" box="[879,986,1741,1765]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="381" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
rudebecki
<collectionCode id="2CCBAE74FFE5FFCA9E3B03F1FB1A71EB" box="[1097,1177,1740,1764]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13442" lsid="urn:lsid:biocol.org:col:13442" name="Kochi University" pageId="23" pageNumber="381" type="Herbarium">KOCH</collectionCode>
/ rudebecki koch.
</materialsCitation>
</paragraph>
<paragraph id="4A6536B1FFE5FFCA992703D1FC64702C" blockId="23.[825,1475,1427,1827]" pageId="23" pageNumber="381">(See Supporting Information, Appendix S1 for additional ma - terial examined.)</paragraph>
<paragraph id="4A6536B1FFE5FFCA994B0278FC7770B4" blockId="23.[825,1474,1860,1979]" pageId="23" pageNumber="381">
<emphasis id="78AEEAA3FFE5FFCA994B0278FCF37053" box="[825,880,1861,1884]" italics="true" pageId="23" pageNumber="381">Note:</emphasis>
As the original descriptions specify the number of speci - mens used to over 100, a representative from the
<typeStatus id="95618813FFE5FFCA9F400259FA007073" box="[1330,1411,1892,1916]" pageId="23" pageNumber="381" type="syntype">syntype</typeStatus>
series is here designated as lectotype to fix the taxonomic status of the species and genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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