103 lines
12 KiB
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103 lines
12 KiB
XML
<document id="7C8EB4876EB948286DD11A7926973256" ID-CLB-Dataset="35377" ID-DOI="http://dx.doi.org/10.3897/zookeys.416.7706" ID-GBIF-Dataset="47852319-33aa-4966-89af-68e6352e05ae" ID-PMC="PMC4109512" ID-Pensoft-Pub="1313-2970-416-113" ID-PubMed="25061349" ID-ZooBank="CE0561FB5EE4498BA2C2EDF9B14F241D" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-416-113" ModsDocOrigin="ZooKeys 416" ModsDocTitle="An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion)" checkinTime="1451245749490" checkinUser="pensoft" docAuthor="Maddison, David R." docDate="2014" docId="C2262CE55E9982E0182EBDF4A39CDF3C" docLanguage="en" docName="ZooKeys 416: 113-155" docOrigin="ZooKeys 416" docSource="http://dx.doi.org/10.3897/zookeys.416.7706" docTitle="Bembidium (Notholopha) Jeannel 1962" docType="treatment" docVersion="6" lastPageNumber="131" masterDocId="7643F6149609155E6212FF95A85FFFBF" masterDocTitle="An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion)" masterLastPageNumber="155" masterPageNumber="113" pageNumber="131" updateTime="1732677897023" updateUser="ExternalLinkService">
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<mods:mods id="30C9A50904B289CB666973119C336BA7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F8DC071DE00BEF6C8107AD7E16BD4F8F">
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<mods:title id="2BB97A72FFA383631D045841CEAA75F3">An unexpected clade of South American ground beetles (Coleoptera, Carabidae, Bembidion)</mods:title>
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<mods:name id="F54BCB74DA305CCBFC219760458578B8" type="personal">
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<mods:roleTerm id="11AA46E1B7362E0E0D9F1FCC6964FB3D">Author</mods:roleTerm>
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<mods:namePart id="F47E8B4CB64087BF5F261756D85392ED">Maddison, David R.</mods:namePart>
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<mods:typeOfResource id="175F31ACA10B340C8D9E69EE47F14B31">text</mods:typeOfResource>
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<mods:title id="5521AFD7A9A669575CCF356B8B836F36">ZooKeys</mods:title>
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<mods:part id="B8A2088DDB7D0E47E53B8D181B7EE4A6">
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<mods:date id="D1746117D241464F1848499BB30DDDF7">2014</mods:date>
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<mods:detail id="69F749FF0C366A5625842F42162C0799" type="volume">
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<mods:number id="1432F957982207EAA4670C2A759A464F">416</mods:number>
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<mods:start id="0F7DD4B1D2214C1CEC0B2BFE2BACDED3">113</mods:start>
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<mods:location id="A1D33A35F4B050EF35A12FBF0022EC63">
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<mods:url id="9BD55F2D2142FA609C193FDD698604DB">http://dx.doi.org/10.3897/zookeys.416.7706</mods:url>
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</mods:location>
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<mods:classification id="EC19F543945B59DDC1C1A134B68C6731">journal article</mods:classification>
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<mods:identifier id="D4713F6578E37D1DB412D94FB6BB9FFD" type="DOI">http://dx.doi.org/10.3897/zookeys.416.7706</mods:identifier>
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<mods:identifier id="74B1386ACD8464537411166EF5F93D35" type="Pensoft-Pub">1313-2970-416-113</mods:identifier>
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<mods:identifier id="D1ED5D3561B0193E0440899F038B3796" type="ZooBank">CE0561FB5EE4498BA2C2EDF9B14F241D</mods:identifier>
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</mods:mods>
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<treatment id="C2262CE55E9982E0182EBDF4A39CDF3C" ID-GBIF-Taxon="152053574" LSID="urn:lsid:plazi:treatment:C2262CE55E9982E0182EBDF4A39CDF3C" httpUri="http://treatment.plazi.org/id/C2262CE55E9982E0182EBDF4A39CDF3C" lastPageNumber="131" pageId="18" pageNumber="131" scope_family="Carabidae" scope_genus="Antiperyphanes" scope_order="Coleoptera">
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<subSubSection id="B576CBDD3563CF65EBDEB1FF4CF78F78" pageId="18" pageNumber="131" type="multiple">
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<paragraph id="F6A25AF083A11217AC873AC3C23ADBF2" pageId="18" pageNumber="131">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection id="3765C93FC43CB27838CCF1CE0919D738" pageId="18" pageNumber="131" type="nomenclature">
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<paragraph id="32D1105531797E9B23511B3A11408E06" pageId="18" pageNumber="131">
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Subgenus
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<taxonomicName id="ADCFF881002C099D994A0977806B7288" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidium (Notholopha)" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="subGenus" subGenus="Notholopha">Notholopha Jeannel, 1962</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="547611A77B92DED3D8BE3C97966FDB5D" pageId="18" pageNumber="131" type="reference_group">
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<paragraph id="49290B1DDC3E3AC412395A488C2CF113" pageId="18" pageNumber="131">
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<taxonomicName id="A18EE02967562CD1D00F7EDABADC32A0" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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Jeannel, 1962; type species
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<taxonomicName id="74128B1CAA6A19A6373B504D8994363C" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidium punctigerum" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="punctigerum">Bembidium punctigerum</taxonomicName>
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Solier, by original designation.
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</paragraph>
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<paragraph id="BB2CBCE3B5C475A2AF3EABB72D8CCAB5" pageId="18" pageNumber="131">
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<taxonomicName id="B8E0AFC9159B10D0F758A14A76B17762" genus="Pacmophena" lsidName="Pacmophena" pageId="18" pageNumber="131" rank="genus">Pacmophena</taxonomicName>
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Jeannel, 1962; type species
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<taxonomicName id="7AF36C0DC3409DF83EF7C8B61EA90597" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidium scitulum" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="scitulum">Bembidium scitulum</taxonomicName>
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Erichson, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="14D29B47F33761C9765BAADA1F48A0B6" pageId="18" pageNumber="131" type="remarks">
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<paragraph id="642E47ABE54B37124C683DCBD57C59D8" pageId="18" pageNumber="131">Remarks.</paragraph>
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<paragraph id="755024131B91550C8EB6B030F5AE1BFD" pageId="18" pageNumber="131">
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<taxonomicName id="DC1739EA9D6484C0E0C900F196925DDF" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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consists of 11 described species (
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<bibRefCitation id="96B719DF2B53B27BFF892036D7731E4D" author="Toledano, L" journalOrPublisher="Koleopterologische Rundschau" pageId="27" pageNumber="140" pagination="1 - 14" title="Nomenclatorial revision of the supraspecific taxa of Bembidiini s. str. of South America described by Jeannel (1962) and related taxa with some descriptions of the fauna of South America (Coleoptera: Carabidae)." volume="72" year="2002">Toledano 2002</bibRefCitation>
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;
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<bibRefCitation id="4F22EC7888FE085705E596C2E6E9F821" author="Toledano, L" journalOrPublisher="Memoirs on Biodiversity" pageId="27" pageNumber="140" pagination="81 - 130" title="Systematic notes on the Bembidiina of the northern Andes with particular reference to the fauna of Ecuador (Coleoptera, Carabidae)." volume="1" year="2008">Toledano 2008</bibRefCitation>
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), and several undescribed (two of which are sequenced here). These are small beetles with large, protruding eyes (Fig. 2B, C), with small flagella in the internal sac of the male genitalia, and with brush sclerites. They have the general appearance when running of a member of the Holarctic subgenus
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<taxonomicName id="1027FF6D41EDFEDC67AFF09E6F0D5AC8" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Bembidion</taxonomicName>
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. Some frequent habitats similar to those of subgenus
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<taxonomicName id="C349ACE08E5E10AE97AA6814AC19345A" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Bembidion</taxonomicName>
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, including dry habitats far from water (
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<taxonomicName id="0C45B9653CFBFE548DC98E05B2819A51" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion stricticolle" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="stricticolle">Bembidion stricticolle</taxonomicName>
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), or upper banks of creeks (e.g.,
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<taxonomicName id="575285E0C68FCF15A2ABEC8FA97E3DC6" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Bembidion</taxonomicName>
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sp.
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<normalizedToken id="8B398804D5612EF8D58B753372F4A9A7" originalValue="“Nahuelbuta”">"Nahuelbuta"</normalizedToken>
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). Others occur at high elevation near small rivulets in open, alpine areas (e.g.,
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<taxonomicName id="2F0DB233F0ADFBE07541E107FF0DB84D" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion rugosellum" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="rugosellum">Bembidion rugosellum</taxonomicName>
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and
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<taxonomicName id="2F1F0DC7A4467153CFB02A4ADA63FE5E" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion melanopodum" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="melanopodum">Bembidion melanopodum</taxonomicName>
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), or at the edges of snowfields (
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<taxonomicName id="2896F2E0DD3755CEC05FE85C51D48457" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion sexfoveolatum" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="sexfoveolatum">Bembidion sexfoveolatum</taxonomicName>
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).
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</paragraph>
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<paragraph id="4CF169CED7CDC09D5C3F0A64292CE30F" pageId="18" pageNumber="131">
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Jeannel considered
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<taxonomicName id="74E0B0ECE136CF37DF4867F11D8208FD" genus="Pacmophena" lsidName="Pacmophena" pageId="18" pageNumber="131" rank="genus">Pacmophena</taxonomicName>
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and
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<taxonomicName id="D7B706726FEFC58F2022029D037B44F5" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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to be two subgenera within the genus
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<taxonomicName id="C89BD124691C267B8C0F2D013CFFB779" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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. As the characters that Jeannel used to distinguish the two are minor characters such as surface texture and antennal length, and as it appears that
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<taxonomicName id="4E5884CBB7E0F12B1FF793F44B64F798" genus="Pacmophena" lsidName="Pacmophena" pageId="18" pageNumber="131" rank="genus">Pacmophena</taxonomicName>
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is paraphyletic with respect to the
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<taxonomicName id="AAF5DF849B9A70BA13BD67318311F86C" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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(s. str.) (Fig. 7), I consider them synonymous, with
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<taxonomicName id="27F85CEB52284ACE2CB51E3B258D7C21" class="Insecta" family="Carabidae" genus="Notholopha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notholopha" order="Coleoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="genus">Notholopha</taxonomicName>
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as the valid name.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |