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<document id="6090C5F6A48ECE0542B308A47C311CCC" ID-DOI="10.3897/zookeys.1223.122412" ID-ZooBank="5FCDC07F-5C80-4B2D-92E1-0ECAB3B36108" ID-publisher-id="122412" URI-arpha="E6E8C3B0-208D-536E-BB51-AA758643C2D9" URI-zoobank="http://zoobank.org/5FCDC07F-5C80-4B2D-92E1-0ECAB3B36108" article-type="research-article" checkinTime="1736187725735" checkinUser="pensoft" docAuthor="Jiang, Ri-Xin &amp; Chen, Xiang-Sheng" docDate="2025" docId="C0ED14C230FD5FCA94F97B2C9B797812" docLanguage="en" docName="ZooKeys 1223: 57-67" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/122412/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sinonychus luodianensis Jiang &amp; Chen 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="67" masterDocId="E6E8C3B0208D536EBB51AA758643C2D9" masterDocTitle="Two new species of the genus Sinonychus (Coleoptera, Elmidae) from Guizhou, China" masterLastPageNumber="67" masterPageNumber="57" pageNumber="57" updateTime="1736188092793" updateUser="ExternalLinkService">
<mods:mods id="56DBD4F0ED6AC0304E17A7F917F73E65" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EEB38633E021BBCD3678550C90180CDB">
<mods:title id="C7157ADB87D8A44B9D8D98D09BCEDCC8">Two new species of the genus Sinonychus (Coleoptera, Elmidae) from Guizhou, China</mods:title>
</mods:titleInfo>
<mods:name id="3DC54120AA224ED5AFC786C62015FEB5" type="personal">
<mods:role id="B1923A64A2F8F3AF1ECF5D0A750CCF66">
<mods:roleTerm id="D33A7C970403EA84C5C439A4FFC450B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D5083A5720C423FE4D5384F45680B28">Jiang, Ri-Xin</mods:namePart>
<mods:affiliation id="67DBCE9111877E404201C71E96DC060E">Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, China &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, China</mods:affiliation>
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</mods:role>
<mods:namePart id="299F06EC6FB696F5A6B5943DD41420FE">Chen, Xiang-Sheng</mods:namePart>
<mods:affiliation id="C40AFE8D568905BC7FBF585288EB9B08">Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, China &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, China</mods:affiliation>
</mods:name>
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<mods:title id="132F8E1749C6FCAECB247AD41448C041">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="C3E1A5F6873F80D684312F5F631FB48C">
<mods:date id="F8D221E256F3185692FFF67ABA079313">2025</mods:date>
<mods:detail id="CC8A5A01BEB3E896C953333FAF3EFD46" type="pubDate">
<mods:number id="2FDB403230749732956B60DB62B8DD14">2025-01-06</mods:number>
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<mods:number id="4C55E5E936E0508689762599A0F997CA">1223</mods:number>
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<mods:identifier id="B9BDE55DE0F541F98414FF58F1194E15" type="DOI">10.3897/zookeys.1223.122412</mods:identifier>
<mods:identifier id="6794ED44A53B93D0C94858E32BBE37B0" type="ZooBank">5FCDC07F-5C80-4B2D-92E1-0ECAB3B36108</mods:identifier>
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<treatment id="C0ED14C230FD5FCA94F97B2C9B797812" ID-DOI="http://doi.org/10.5281/zenodo.14606277" ID-Zenodo-Dep="14606277" ID-arpha="C0ED14C2-30FD-5FCA-94F9-7B2C9B797812" ID-zoobank="https://zoobank.org/EC1B7D2E-24F8-4320-82A4-3F50B1E6D07C" LSID="urn:lsid:plazi:treatment:C0ED14C230FD5FCA94F97B2C9B797812" LSID-ZBK="urn:lsid:zoobank.org:act:EC1B7D2E-24F8-4320-82A4-3F50B1E6D07C" httpUri="http://treatment.plazi.org/id/C0ED14C230FD5FCA94F97B2C9B797812">
<subSubSection id="E02AF3935271A9C4057E596E5D38699A" type="nomenclature">
<paragraph id="F36524541D78963DDA99F17E326C6590">
<taxonomicName id="98494897EE730088C38A063E02F5641A" authority="Jiang &amp; Chen" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="luodianensis" status="sp. nov.">
<emphasis id="2C4DFA002A456572C6BF2B1A503D63EE" italics="true">Sinonychus luodianensis</emphasis>
Jiang &amp; Chen
</taxonomicName>
<taxonomicNameLabel id="225B3133BF2548BF3B09CCD34961692D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5578FA100A632622467C414C468DEAB1" type="description">
<paragraph id="8C323062A41BD1137563E6B44EFC29F4">
<figureCitation id="3EED8DBD2F3819A8FC13A788746AF95D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A dorsal habitus of Sinonychus lipinae sp. nov. B dorsal habitus of S. luodianensis sp. nov. C antenna of S. lipinae sp. nov. D antenna of S. luodianensis sp. nov. Scale bars: 0.5 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1223.122412.figure1" httpUri="https://binary.pensoft.net/fig/1213469">Figs 1 B, D</figureCitation>
,
<figureCitation id="6357410EC3B9FCF10BD934D10EADA1DE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 A F</figureCitation>
,
<figureCitation id="E52220F75EA49BCBE5EC2C47BF8A6CA6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genital features of Sinonychus species: A G Sinonychus lipinae sp. nov. H N, Sinonychus luodianensis sp. nov. A, H aedeagus, dorsal view B, J ditto, ventral C, I ditto, median lobe D, K ditto lateral view E, L sternite IX F, M ovipositor G, N ditto, apical part. Scale bars: 0.05 mm (G, N); 0.1 mm (A F, H M)." figureDoi="10.3897/zookeys.1223.122412.figure4" httpUri="https://binary.pensoft.net/fig/1213472">4 H N (罗甸华溪泥甲</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDGAE" type="type material">
<paragraph id="E3B0ED517E41AD1F983FECC00E42DE2A">
<heading id="61C699A10779EBF8DCE752CC188667B8" reason="title">
<typeStatus id="672D2B2E87C89860C422C8294ED3746B">Type</typeStatus>
material.
</heading>
</paragraph>
<paragraph id="A98898CD6CC92910F39AA04420BB7E5E">
41 exs:
<specimenCount id="42F29B0987D850EFDB0FFA7B5E790582" count="11" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="9715DEFC038961349331D6E33F1A8001" count="10" type="female">10 ♀♀</specimenCount>
,
<specimenCount id="FA3274966EFB117F1D87CFDD086B8885" count="20" type="generic">20 exs.</specimenCount>
, sex undetermined.
<emphasis id="CD7BA33F893399748C4CC1DDD8FAD24F" bold="true" italics="true">
<typeStatus id="038AD1A250E598A92FFE9DA78C649D00" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<materialsCitation id="C29B0DE0AD94ED498BA5D0FA52ABD273" collectingDate="2022-11-09" collectionCode="GUGC" collectorName="Jiang Ri-Xin &amp; Paratypes" country="China" county="Luodian County" elevation="6" latitude="25.328611" location="Qiannan Buyi" longLatPrecision="21" longitude="106.641106" municipality="China" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<collectingCountry id="A5BBABC07133871D5A8382435E88994F" name="China">CHINA</collectingCountry>
:
<specimenCount id="D6550A30BD4B1EC1AC306F8F83CC65AC" count="1" type="male"></specimenCount>
, labeled “
<collectingMunicipality id="BAC44608269FE52AF55B1D5755D4A7E6">China</collectingMunicipality>
:
<collectingRegion id="F545A9CAF462E6560EE3A1435CB85BEC" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<location id="CA0EA7089EFF7E986E38A8CC7F6C3264" LSID="urn:lsid:plazi:treatment:C0ED14C230FD5FCA94F97B2C9B797812:CA0EA7089EFF7E986E38A8CC7F6C3264" country="China" county="Luodian County" latitude="25.328611" longLatPrecision="21" longitude="106.641106" municipality="China" name="Qiannan Buyi" stateProvince="Guizhou">Qiannan Buyi</location>
and
<location id="98B01AB27BBABB1A1FD6AC1F64BCF151" LSID="urn:lsid:plazi:treatment:C0ED14C230FD5FCA94F97B2C9B797812:98B01AB27BBABB1A1FD6AC1F64BCF151" country="China" county="Luodian County" latitude="25.328611" longLatPrecision="21" longitude="106.641106" municipality="China" name="Miao Autonomous Prefecture" stateProvince="Guizhou">Miao Autonomous Prefecture</location>
(黔南布依族苗族自治州),
<collectingCounty id="63F436FDBC7312596A0821A5F97E5953">Luodian County</collectingCounty>
(罗甸县),
<location id="5CBADFAB5832736549339EACFD67CFC7" LSID="urn:lsid:plazi:treatment:C0ED14C230FD5FCA94F97B2C9B797812:5CBADFAB5832736549339EACFD67CFC7" country="China" county="Luodian County" latitude="25.328611" longLatPrecision="21" longitude="106.641106" municipality="China" name="Luokun Town" stateProvince="Guizhou">Luokun Town</location>
(罗悃镇),
<location id="57C0C93CC95BFE27851150BE597548A5" LSID="urn:lsid:plazi:treatment:C0ED14C230FD5FCA94F97B2C9B797812:57C0C93CC95BFE27851150BE597548A5" country="China" county="Luodian County" latitude="25.328611" longLatPrecision="21" longitude="106.641106" municipality="China" name="Xiangshui Village" stateProvince="Guizhou">Xiangshui Village</location>
(响水村),
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPGAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.641111,25.328611]}">
<geoCoordinate id="FE02A3B8B7D6E4F25CD9CAD434FB68B5" degrees="25" direction="north" minutes="19" orientation="latitude" precision="15" seconds="43" value="25.328611">25 ° 19 ' 43 &quot; N</geoCoordinate>
<geoCoordinate id="B072811AE70B52E9981AE059EACD3859" degrees="106" direction="east" minutes="38" orientation="longitude" precision="15" seconds="28" value="106.641106">106 ° 38 ' 28 &quot; E</geoCoordinate>
</named-content>
, H: 666.10 ±
<quantity id="E3706CCA2A802FFA8F7B96AA45D1FA1F" metricMagnitude="0" metricUnit="m" metricValue="6.4" unit="m" value="6.4">
<elevation id="69A47BA42D1A7D69ED0644259BED34FF" metricMagnitude="0" metricUnit="m" metricValue="6.4" unit="m" value="6.4">6.40 m</elevation>
</quantity>
,
<date id="3E119426577D3B23680EEBD0D0336159" value="2022-11-09">
<collectingDate id="B8F20F7C390F95304E31A7ADA012AEB4" value="2022-11-09">09. XI. 2022</collectingDate>
</date>
,
<collectorName id="F92EBB36F2ADC8DF2FBF555A45B3C227">Jiang Ri-Xin</collectorName>
leg. ” (
<abbrev content-type="institution" id="ABBRID0EUGAE" xlink_title="Institute of Entomology, Guizhou University, Guiyang, China">
<collectionCode id="077C54C7DC0D30A7E6CB1173BCEF3E88">GUGC</collectionCode>
</abbrev>
).
<emphasis id="BB8FC0EC7213C92A15AB7DCE7EB1E7D3" bold="true" italics="true">
<typeStatus id="5ED2827167143115E7EA381A5376B402" type="paratype">
<collectorName id="62757E985A6F2EEFE1B5483B88D9BA16">Paratypes</collectorName>
</typeStatus>
</emphasis>
</materialsCitation>
: •
<materialsCitation id="59ADD968544D510F93612A4193620935" collectingDate="2022-11-09" collectionCode="GUGC" collectorName="Jiang Ri-Xin &amp; Paratypes" country="China" county="Luodian County" elevation="6" latitude="25.328611" location="Qiannan Buyi" longLatPrecision="21" longitude="106.641106" municipality="China" specimenCount="40" specimenCount-female="10" specimenCount-male="10" stateProvince="Guizhou" typeStatus="holotype">
<specimenCount id="DFAEEA9F9146B8AD1B8BC2DE74E4A764" count="10" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="7A69343E8773E40106DE208DEC333D2E" count="10" type="female">10 ♀♀</specimenCount>
,
<specimenCount id="E8373FB42011E6FF063AE1D7670EFE7E" count="20" type="generic">20 exs.</specimenCount>
, sex undetermined, with same label data as the holotype (
<abbrev content-type="institution" id="ABBRID0E3GAE" xlink_title="Institute of Entomology, Guizhou University, Guiyang, China">
<collectionCode id="9027538BFABB44E4FA5C91EEFF58E482">GUGC</collectionCode>
</abbrev>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBHAE" type="diagnosis">
<paragraph id="7290A4E4BB51041F72897768D9229F21">
<heading id="9F24B422D3D926EB64008D82823E7C5F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="673BFDD0724D6D7F470E60455F92CB37">Body long-oval, mostly black; pronotum (except basal part), antennae, base of tibia and tarsi (including claws) light brown. Plastron setae scaly. Elytral intervals 5, 6 and 7 with granulate carinae. Aedeagus with apex of median lobe rounded; median lobe with two pairs of long, elongate sclerotizations located at basal 1 / 2; a very thin and long sclerotization extends from middle of median lobe to well beyond the apex, where it is curved.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGHAE" type="description">
<paragraph id="A49FB5A6FBEE861FEB08AB07DB7C7563">
<heading id="290A528082205C411901F9BDD89758D8" reason="title">Description.</heading>
</paragraph>
<paragraph id="A89D2AE2059725352B9AF5CA41298A1C">
Body long-oval (Fig.
<figureCitation id="54F58031E0465A38F7B187344CB65B0C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A dorsal habitus of Sinonychus lipinae sp. nov. B dorsal habitus of S. luodianensis sp. nov. C antenna of S. lipinae sp. nov. D antenna of S. luodianensis sp. nov. Scale bars: 0.5 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1223.122412.figure1" httpUri="https://binary.pensoft.net/fig/1213469">1 B</figureCitation>
); black with pronotum (except basal part), antennae, base of tibia and tarsi (including claws) light brown. Plastron setae confined to following areas: head (both dorsal and ventral surfaces, including disc of clypeus); pronotum (lateral areas); elytra (lateral areas, including epipleura); prosternum, mesoventrite, metaventrite and abdomen (lateral areas); and femora.
</paragraph>
<paragraph id="450C90BF94B3CA6E8CD6EBC2C8BA332B">
Head (Fig.
<figureCitation id="B9D714147D436A3EDCFCE6A7F09B0448" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 A</figureCitation>
), wider than long, surface covered with plastron setae and sparse, long setae and granules. Clypeus with disc covered with plastron setae; anterior and lateral areas without plastron setae, covered with sparse, long setae. Labrum transverse, narrower than clypeus, lateral margins with long bristles. Antenna (Fig.
<figureCitation id="048EC9A20CEAB4DE105E9076D1D1A1A0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A dorsal habitus of Sinonychus lipinae sp. nov. B dorsal habitus of S. luodianensis sp. nov. C antenna of S. lipinae sp. nov. D antenna of S. luodianensis sp. nov. Scale bars: 0.5 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1223.122412.figure1" httpUri="https://binary.pensoft.net/fig/1213469">1 D</figureCitation>
) 7 - segmented with apical antennomere clubbed.
</paragraph>
<caption id="86999383AF54672EC02504A0D61F8F53" ID-DOI="10.3897/zookeys.1223.122412.figure3" ID-arpha="3F01F47A-787A-5F56-99EC-3D3B3089BA1F" httpUri="https://binary.pensoft.net/fig/1213471" startId="F3">
<paragraph id="F6AAF0A344F8378E2DC590A8E7C53B46">
<label id="8C623FC184CAE2482A7FCB4731EAD411">Figure 3.</label>
</paragraph>
<paragraph id="22C2C135BFF3E880D0C7A90149BD1677">
Diagnostic features of
<taxonomicName id="359A1235A48706D62C8F6F3B526CE8B3" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="luodianensis" status="sp. nov.">
<emphasis id="8C516047257322C6996EDF3E61E44210" italics="true">Sinonychus luodianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="691D3C948671C795E4075BEB59B93D19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9426BE291E54817A37ABD19217F1825F" bold="true">A</emphasis>
head
<emphasis id="16C773094F773815A46E5F19A06213D7" bold="true">B</emphasis>
pronotum
<emphasis id="789ABB6C3CA931C4B6E8916D9C76D7E7" bold="true">C</emphasis>
prosternal process
<emphasis id="B4BB6A1CB96CF4155DD76E07A4319AA9" bold="true">D</emphasis>
metaventrite
<emphasis id="AD0EAB174F4752E12C6D3D4E05AA787F" bold="true">E</emphasis>
abdomen
<emphasis id="8CEB19328A671529A116DADB5A297E80" bold="true">F</emphasis>
elytron. Scale bars: 0.1 mm (
<emphasis id="3A43AC9DA4A736C6EC0640312B82DB82" bold="true">A</emphasis>
); 0.2 mm (
<emphasis id="0B64F04545AF87502BD96CABA51DDB3B" bold="true">B F</emphasis>
).
</paragraph>
</caption>
<paragraph id="94C44B156C8992C4556E07B5C1666EC5">
Pronotum (Fig.
<figureCitation id="2DE6040F6A08698624A3F4B571C8F001" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 B</figureCitation>
) slightly wider than long, widest at basal 2 / 5, gradually narrowed from basal 2 / 5 to apex. Surface finely punctate; covered with sparse, long setae and with plastron setae laterally. Median longitudinal sulcus distinct and long, extending from base nearly to anterior 1 / 4, with a pair of small round fovea near base. Sublateral grooves straight, short, less than 1 / 3 length of pronotum. Anterior margin distinctly curved, anterior angles not produced. Lateral margins gradually curved. Basal margin trisinuate, emarginate anterior to scutellum, posterior angles nearly orthogonal. Prosternal process (Fig.
<figureCitation id="63B5AC6676D4264B1A508661C25115BC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 C</figureCitation>
) very wide, with apex broadly rounded; surface distinctly microreticulate, glabrous.
</paragraph>
<paragraph id="9DDF2E073566314171F36609C4045F58">
Scutellum (Fig.
<figureCitation id="6C6962FFB6DA1A894F5E4F68420D3010" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 B</figureCitation>
) cordate, longer than wide, widest at basal 1 / 4; surface microreticulate and glabrous. Anterior margin strongly curved, lateral margins weakly curved, apex acutangular.
</paragraph>
<paragraph id="AA579C334DF7948B4917EA5050FCA3B5">
Elytra (Fig.
<figureCitation id="E34C37AD5481F7D5F6AE1D0D45F801C3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 F</figureCitation>
) about 1.5 times as long as wide, widest near in median 1 / 3. Strial punctures large in basal 2 / 3 of elytra, separated by about twice a diameter; much smaller and more widely separated in other parts of elytra. Elytral intervals 5, 6, 7 with granulate carinae extending from base of elytra nearly to apex. Interval 5 to lateral margins covered with plastron setae. Hing wings reduced.
</paragraph>
<paragraph id="C93903BC82B9D708212DA4672532B134">
Metaventrite (Fig.
<figureCitation id="6F9D023AE6A8CB7E63B72B3996D87DFE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 D</figureCitation>
) with disc distinctly microreticulate and covered with sparse, long setae; lateral areas with plastron setae. Median sulcus shallow and indistinct, less than half the length of metaventrite.
</paragraph>
<paragraph id="38DAAB69590E577EAA71A221E27123A8">
Admedian carinae of abdominal ventrite 1 indistinct, straight, extending from base to near apex. Median areas of ventrites 14 (Fig.
<figureCitation id="737EC870C47EDAC33E57F2A6D4172563" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Diagnostic features of Sinonychus luodianensis sp. nov. A head B pronotum C prosternal process D metaventrite E abdomen F elytron. Scale bars: 0.1 mm (A); 0.2 mm (B F)." figureDoi="10.3897/zookeys.1223.122412.figure3" httpUri="https://binary.pensoft.net/fig/1213471">3 E</figureCitation>
) and anteromedial area of ventrite 5 microreticulate, covered with sparse, long setae; remaining surface of ventrites 15 covered with plastron setae and mixed with sparse, long setae.
</paragraph>
<paragraph id="D0A0A4A21CA1B7F453ECC6F549C02DE3">Legs simple, surface granulate (except tarsi). Surface of femora surface covered with plastron setae; inner side of tibia with cleaning fringes; tarsi slightly shorter than tibiae; tarsal claws simple.</paragraph>
<paragraph id="BF887C60BE40ED292E68084151C8F067">
Aedeagus (Fig.
<figureCitation id="BA8D7C35F05908801A9F1BAA1FD3A73D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genital features of Sinonychus species: A G Sinonychus lipinae sp. nov. H N, Sinonychus luodianensis sp. nov. A, H aedeagus, dorsal view B, J ditto, ventral C, I ditto, median lobe D, K ditto lateral view E, L sternite IX F, M ovipositor G, N ditto, apical part. Scale bars: 0.05 mm (G, N); 0.1 mm (A F, H M)." figureDoi="10.3897/zookeys.1223.122412.figure4" httpUri="https://binary.pensoft.net/fig/1213472">4 H K</figureCitation>
) slender and elongate. Median lobe symmetrical, widest at base, weakly narrowed from base to apex; apex rounded; with two pairs of long, elongate sclerotizations located at basal 1 / 2; a very thin, very long sclerotization extends from middle of median lobe to well beyond the apex, where it is curved. Sternite IX (Fig.
<figureCitation id="7E015D74C8939FB97C7BADD14A56E814" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genital features of Sinonychus species: A G Sinonychus lipinae sp. nov. H N, Sinonychus luodianensis sp. nov. A, H aedeagus, dorsal view B, J ditto, ventral C, I ditto, median lobe D, K ditto lateral view E, L sternite IX F, M ovipositor G, N ditto, apical part. Scale bars: 0.05 mm (G, N); 0.1 mm (A F, H M)." figureDoi="10.3897/zookeys.1223.122412.figure4" httpUri="https://binary.pensoft.net/fig/1213472">4 L</figureCitation>
) with apical margin weakly emarginate, without setae, median strut curved at middle with base curved.
</paragraph>
<caption id="0751E1A6F4F0B1AD87F25A049FB9EC04" ID-DOI="10.3897/zookeys.1223.122412.figure4" ID-arpha="2C26C76D-1FF7-5E14-B8A9-255294EB090C" httpUri="https://binary.pensoft.net/fig/1213472" startId="F4">
<paragraph id="68ACEC34BB85C4F6212A0053A80908D7">
<label id="EF5486252E6FA42F31FD3578A4C110A0">Figure 4.</label>
</paragraph>
<paragraph id="74644A2059CA5CB0ED889A07AFF6D81C">
Genital features of
<taxonomicName id="0D1BEAFEF923BBC25E49F96C96A45BFE">
<emphasis id="9AEAB29CD7DFC38C04EC666BDDE5E4C5" italics="true">Sinonychus</emphasis>
species
</taxonomicName>
:
<emphasis id="B5D3FB700F67274D84DD68A10E5EEA00" bold="true">A G</emphasis>
<taxonomicName id="72E2C48CEA5C9690153538DD5BF56A04" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="lipinae" status="sp. nov.">
<emphasis id="28D6AF0160EECD0F762441819D14EB53" italics="true">Sinonychus lipinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFAEDF6D46CAE82F8940086AD77DFF01" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5A9D35ACFCC524B994DB1ED0C74D152F" bold="true">H N</emphasis>
,
<taxonomicName id="475ECBB50B5364C53FE9ABE0FA15183B" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="luodianensis" status="sp. nov.">
<emphasis id="727CD64E5B1735DB7059A7A5201E61D5" italics="true">Sinonychus luodianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41BBFB8BE608E02C414D6FF3FFEDEEEC" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C86098F3CFF8B85E8FAD3FA4B030A978" bold="true">A, H</emphasis>
aedeagus, dorsal view
<emphasis id="24CFFE6C17B709D71F62CBE31D5B1288" bold="true">B, J</emphasis>
ditto, ventral
<emphasis id="C18EF9F65F62BDCF532B2D275DA9F5CF" bold="true">C, I</emphasis>
ditto, median lobe
<emphasis id="F87A36B093B67F69C992DD226D0DCBBD" bold="true">D, K</emphasis>
ditto lateral view
<emphasis id="4C574F1DD0B847312F8AE3B3290F61F1" bold="true">E, L</emphasis>
sternite IX
<emphasis id="8865795CAB7818EAFD16C24F73C0B99E" bold="true">F, M</emphasis>
ovipositor
<emphasis id="DD5D541048C7A311FA3D18B40B8CB6AF" bold="true">G, N</emphasis>
ditto, apical part. Scale bars: 0.05 mm (
<emphasis id="9D69CCCC07722D7D112A36DACC3EDE2E" bold="true">G, N</emphasis>
); 0.1 mm (
<emphasis id="BD0515FDF71A20704440EBD629754F96" bold="true">A F, H M</emphasis>
).
</paragraph>
</caption>
<paragraph id="0A6176904D91E89B8B6B3386CDBB18CA">
Measurements:
<abbrev id="ABBRID0ERMAE" xlink_title="sum of PL + EL">CL</abbrev>
:
<quantity id="AB6D1427BFE68F35D35B0F8CD2119255" metricMagnitude="-3" metricUnit="m" metricValue="1.07" metricValueMax="1.11" metricValueMin="1.03" unit="mm" value="1.07" valueMax="1.11" valueMin="1.03">1.031.11 mm</quantity>
;
<abbrev id="ABBRID0EVMAE" xlink_title="length of pronotum along midline">
<collectionCode id="9A423BE3D106A10C3016396F71D40316" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
</abbrev>
:
<quantity id="4F13F20673B60891AAA23061B0812068" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.330.35 mm</quantity>
,
<abbrev id="ABBRID0EZMAE" xlink_title="maximum width of pronotum">
<collectionCode id="3597C391169574DF7118983CAB20ED9C" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
:
<quantity id="BC13355328720EF5877A8DBB01292E64" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" unit="mm" value="0.43" valueMax="0.44" valueMin="0.42">0.420.44 mm</quantity>
;
<abbrev id="ABBRID0E4MAE" xlink_title="length of elytra along suture">EL</abbrev>
:
<quantity id="4517C779E2F71BD4DFF2102FDBF7BF68" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="7.8" metricValueMin="7.0" unit="mm" value="0.74" valueMax="0.78" valueMin="0.7">0.700.78 mm</quantity>
,
<abbrev id="ABBRID0EBNAE" xlink_title="maximum width of elytra">EW</abbrev>
:
<quantity id="2071D5FE40786CB3F788CA0FB59DD4E8" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" unit="mm" value="0.52" valueMax="0.54" valueMin="0.5">0.500.54 mm</quantity>
.
</paragraph>
<paragraph id="2793198D9B6278929D95AFBA8A77681E">
Female externally similar to the male, averaging larger. Ovipositor as in Fig.
<figureCitation id="A4F7519E8FF138851393BBE5D9533F3A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genital features of Sinonychus species: A G Sinonychus lipinae sp. nov. H N, Sinonychus luodianensis sp. nov. A, H aedeagus, dorsal view B, J ditto, ventral C, I ditto, median lobe D, K ditto lateral view E, L sternite IX F, M ovipositor G, N ditto, apical part. Scale bars: 0.05 mm (G, N); 0.1 mm (A F, H M)." figureDoi="10.3897/zookeys.1223.122412.figure4" httpUri="https://binary.pensoft.net/fig/1213472">4 M, N</figureCitation>
: valvifer about twice as long as coxite, distinctly expanded at base; coxite apex strongly expanded, roundly broadened at outer margin; stylus short, weakly curved.
</paragraph>
<paragraph id="447C58ECBB5992879FF047D62F7C9E2E">
Measurements:
<abbrev id="ABBRID0ENNAE" xlink_title="sum of PL + EL">CL</abbrev>
:
<quantity id="0110F415BA2E1E6A15FC50F0E809970C" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.17" metricValueMin="1.04" unit="mm" value="1.105" valueMax="1.17" valueMin="1.04">1.041.17 mm</quantity>
;
<abbrev id="ABBRID0ERNAE" xlink_title="length of pronotum along midline">
<collectionCode id="64822B4AB901A6DF9F9EB5E7061FBED3" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
</abbrev>
:
<quantity id="EF47FB601277EA6EE74E06D599BE8F84" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.330.38 mm</quantity>
,
<abbrev id="ABBRID0EVNAE" xlink_title="maximum width of pronotum">
<collectionCode id="9F16018A1031E7F8B770A26400522215" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
:
<quantity id="912D1B521CC6FC5ECC6262D29044B251" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" unit="mm" value="0.415" valueMax="0.44" valueMin="0.39">0.390.44 mm</quantity>
;
<abbrev id="ABBRID0EZNAE" xlink_title="length of elytra along suture">EL</abbrev>
:
<quantity id="3710B937A498CA885352FBECF37F2F7A" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="7.8" metricValueMin="7.199999999999999" unit="mm" value="0.75" valueMax="0.78" valueMin="0.72">0.720.78 mm</quantity>
,
<abbrev id="ABBRID0E4NAE" xlink_title="maximum width of elytra">EW</abbrev>
:
<quantity id="EBD5780EDF7B096329A9CB6D82F40757" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.699999999999999" metricValueMin="4.9" unit="mm" value="0.53" valueMax="0.57" valueMin="0.49">0.490.57 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBOAE" type="distribution">
<paragraph id="7F1D5D7B0EB739062D9D26DF1D15465E">
<heading id="F99667D0D4C059367D0AD7F34F77194D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="5C65347683DE34C142331E91D4B585A8">
<collectingCountry id="748F876AE667ABE4ABDE41CB86493707" name="China">China</collectingCountry>
. Only known from the
<typeStatus id="376EAD7E24A0C657A90E24965FB7A33F">type</typeStatus>
locality in Luodian County, Qiannan Buyi and Miao Autonomous Prefecture,
<collectingRegion id="E6717CFE884AB9C599E748DC2262E1C3" country="China" name="Guizhou">Guizhou Province</collectingRegion>
.
</paragraph>
<caption id="8E94DE01AC4E8E9ECEB54BC392BCA998" ID-DOI="10.3897/zookeys.1223.122412.figure5" ID-arpha="6DCBF731-87D6-586C-81BC-68B1B338D2FA" httpUri="https://binary.pensoft.net/fig/1213473" startId="F5">
<paragraph id="E32F897B09C4E22D182BABDC5D1089A8">
<label id="BE9FE9DB5E006821100D1B0C8769DDE9">Figure 5.</label>
</paragraph>
<paragraph id="3AFFAEE9AA1FCF208ED7CB5DE828A287">
Habitat of
<taxonomicName id="5DD185D63B7BF21D615D9D374A52BF2D" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="lipinae" status="sp. nov.">
<emphasis id="9C01BEF1FDF9591791C5D27568145D34" italics="true">Sinonychus lipinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="935E0661CDF4FCB9BB1468D10F16159F" rank="species">sp. nov.</taxonomicNameLabel>
at the type locality
<emphasis id="CA1C5A435CF02128A4CE689D23F44374" bold="true">A</emphasis>
general environment
<emphasis id="9CF2776CDF11224050078CD357D78FB0" bold="true">B</emphasis>
microenvironment
<emphasis id="0C60EB353FEA6179993FEC2F723FF693" bold="true">C</emphasis>
living adult
<emphasis id="80827662540BD05698547B50D523D64D" bold="true">D</emphasis>
collectors, left: Miss Yu-Hao Zhang, right: Dr Yin-Lin Mu
<emphasis id="D0D5084C49D263D4C42DE7ABC839AC3F" bold="true">E</emphasis>
collectors, left: Dr Hai-Tao Li, right: Dr Pin Li.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIPAE" type="biology">
<paragraph id="354464A96FD3C69D4D1C4D4B2EABC273">
<heading id="ECED802C892A3B6F4E3A82F22EA67D3B" reason="title">Biology.</heading>
</paragraph>
<paragraph id="461A4C9C0D72476AA03E8805E88114C8">
All adults were collected from gravel on the bottom of a small stream in a ravine (Fig.
<figureCitation id="04C33391E4ADC2C1C8C1BF5CE4F655B1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitat of Sinonychus luodianensis sp. nov. at the type locality A general environment B microenvironment C living adult." figureDoi="10.3897/zookeys.1223.122412.figure6" httpUri="https://binary.pensoft.net/fig/1213474">6 A C</figureCitation>
).
</paragraph>
<caption id="EB8A8E664071088506DA7EE63E28A395" ID-DOI="10.3897/zookeys.1223.122412.figure6" ID-arpha="ED385404-8950-5A4D-833A-97AFC338A635" httpUri="https://binary.pensoft.net/fig/1213474" startId="F6">
<paragraph id="1C018BE4FC0D0961CD362D1BEAE47D2D">
<label id="D095B4AAEEEE8A1C48488166F7A7DFB5">Figure 6.</label>
</paragraph>
<paragraph id="3C89C0441BBE2D241F865A7FB9781214">
Habitat of
<taxonomicName id="999B8001CA5926C88C1A0FEEE6E6D05E" authorityName="Jiang &amp; Chen" authorityYear="2025" class="Insecta" family="Elmidae" genus="Sinonychus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="luodianensis" status="sp. nov.">
<emphasis id="08854509335C8A02A26BB0F2423499E8" italics="true">Sinonychus luodianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A99E6DDC65DA21C2F2C2CB416A63D5" rank="species">sp. nov.</taxonomicNameLabel>
at the type locality
<emphasis id="91A9B2EDDC5334C83FC0288A40D5630A" bold="true">A</emphasis>
general environment
<emphasis id="6934DFD900609D575CF813F2A3BAEAD5" bold="true">B</emphasis>
microenvironment
<emphasis id="02DF1560A9BF77A85570A2E9C4AE6825" bold="true">C</emphasis>
living adult.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQQAE" type="etymology">
<paragraph id="2F2BC93F50DD77FE51E891E402631936">
<heading id="FE80B5CFC069FF68C5EDCF6CECE9A5E4" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="744F56045DE2D7FBAB5C62E706BFE2E7">
The specific epithet “
<emphasis id="855617A83936D1302457A4A615F62B09" italics="true">luodianensis</emphasis>
” refers to the
<typeStatus id="5B57E923E548DCAF548B24540644CB65">type</typeStatus>
locality, Luodian County, Qiannan Buyi and Miao Autonomous Prefecture,
<collectingRegion id="46C05575622C60E1A9C9EE1AB35978DC" country="China" name="Guizhou">Guizhou Province</collectingRegion>
; the name is treated as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EARAE" type="Comparative diagnosis">
<paragraph id="D780C2564973E708E725477A5D3E2F0B">
<heading id="53A4261688C83490523509EB97BFF5A2" reason="title">Comparative diagnosis.</heading>
</paragraph>
<paragraph id="DB5887AC9EB8BACA5EE746ACBBA92597">
The new species can be easily distinguished from other
<taxonomicName id="70BE3E11552306EFCFDA07A02AB9251A">
<emphasis id="EE2C1B4D759F2361B0541ABA85BA9902" italics="true">Sinonychus</emphasis>
species
</taxonomicName>
by its scaly plastron setae, and males by the aedeagus with very long, thread-like sclerotization extending from the middle of the median lobe and exceeding its apex.
</paragraph>
</subSubSection>
</treatment>
</document>