treatments-xml/data/C0/7A/73/C07A73CCDA665F3BB3E9E8FBC045A737.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="C07A73CCDA665F3BB3E9E8FBC045A737" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila szekeresi Pall-Gergely &amp; Hunyadi 2015" docType="treatment" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737" httpUri="http://treatment.plazi.org/id/C07A73CCDA665F3BB3E9E8FBC045A737" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="66QMX" LSID="C07A73CC-DA66-5F3B-B3E9-E8FBC045A737" authority="Pall-Gergely &amp; Hunyadi, 2015" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">
Angustopila szekeresi 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, 2015
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Synoptic view of shells from different Angustopila szekeresi Pall-Gergely &amp; Hunyadi, 2015 populations." figureDoi="10.3897/zookeys.1147.93824.figure56" httpUri="https://binary.pensoft.net/fig/808865" pageId="0" pageNumber="1">Fig. 56</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">Angustopila szekeresi</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.523.6114" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="31 - 64" refId="B39" refString="Pall-Gergely, B, Hunyadi, A, Jochum, A, Asami, T, 2015a. Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys 523: 31 - 64, DOI: https://doi.org/10.3897/zookeys.523.6114" title="Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)." url="https://doi.org/10.3897/zookeys.523.6114" volume="523" year="2015 a">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al., 2015a
</bibRefCitation>
: 42-45, fig. 5.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;China, Guangxi (广西), Hechi Shi (河池市), Bama Xian (巴马县), cliffs at the southern edge of Jiaole Cun (交乐村), 590 m, 
<geoCoordinate degrees="24" direction="north" minutes="7.045" orientation="latitude" precision="1" value="24.117416">24°7.045'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="7.847" orientation="longitude" precision="1" value="107.13078">107°7.847'E</geoCoordinate>
.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2003-09-21" collectingDateMax="2020-02-19" collectingDateMin="2003-09-21" collectorName="W. J. M. Maassen, Thanh District, H, A. Hunyadi, Xa Chi, On, B, Tr, Hang Nh, Nhung, Province, H, District, H, Lien, Ba Nang, Lam" country="Vietnam" county="Pu Puong National Park" elevation="20" latitude="21.6985" location="Thanh District" longLatPrecision="12" longitude="106.36283" municipality="Am" specimenCount="383" specimenCount-adult="382" specimenCount-juvenile="1" stateProvince="Thanh Hoa Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 1 figured + 
<specimenCount type="juvenile">1 juvenile</specimenCount>
shell; 
<collectingRegion country="Vietnam" name="Thanh Hoa">Thanh Hoa Province</collectingRegion>
, 
<collectingCounty>Pu Puong National Park</collectingCounty>
, limestone hill near small native village 
<collectingMunicipality>Am</collectingMunicipality>
(locality code: WMVT.0344); 
<geoCoordinate degrees="20" direction="north" minutes="27.39" orientation="latitude" precision="9" value="20.4565">20°27.39'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="13.65" orientation="longitude" precision="9" value="105.2275">105°13.65'E</geoCoordinate>
; 
<collectingDate value="2003-09-21">21 Sep. 2003</collectingDate>
; 
<collectorName>W.J.M. Maassen</collectorName>
leg.; RMNH.5006720 • 
<specimenCount type="adult">59 adult</specimenCount>
+ 36 j/b shells; 
<collectingRegion country="Vietnam" name="Thanh Hoa">
Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province
</collectingRegion>
, 
<normalizedToken originalValue="Như">Như</normalizedToken>
<collectorName>
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:FD254F521F4DAA6D83F5E019C8AEEE19" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Thanh District" stateProvince="Thanh Hoa Province">Thanh District</location>
, H
</collectorName>
ải 
<normalizedToken originalValue="Vân">
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:CC27E8CB47F660E4FDC53BE05F7B2478" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Van" stateProvince="Thanh Hoa Province">Van</location>
</normalizedToken>
, 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:B597E8CBC9DBD4A8F41EEE6E5670901D" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Hang Lo Cao Khang Chi" stateProvince="Thanh Hoa Province">
Hang 
<normalizedToken originalValue="Lò">Lo</normalizedToken>
Cao 
<normalizedToken originalValue="Kháng">Khang</normalizedToken>
Chi
</location>
ến, vicinity of the cave (locality code: 2020/41); 
<geoCoordinate degrees="19" direction="north" minutes="37.08" orientation="latitude" precision="9" value="19.618">19°37.08'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="34.63" orientation="longitude" precision="9" value="105.57716">105°34.63'E</geoCoordinate>
; 
<elevation metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-14">14 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 
<specimenCount type="adult">26 adult</specimenCount>
shells; 
<collectingRegion country="Vietnam" name="Thanh Hoa">
Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province
</collectingRegion>
, 
<normalizedToken originalValue="Như">Như</normalizedToken>
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:8BA91692A272FF85C9D084CEAE68399E" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Thanh District" stateProvince="Thanh Hoa Province">Thanh District</location>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
south from 
<normalizedToken originalValue="Xuân">Xuan</normalizedToken>
Khang along road no. 45, around the cave temple (locality code: 2020/40); 
<geoCoordinate degrees="19" direction="north" minutes="40.35" orientation="latitude" precision="9" value="19.6725">19°40.35'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="31.07" orientation="longitude" precision="9" value="105.51783">105°31.07'E</geoCoordinate>
; 
<elevation metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">80 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-14">14 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 
<specimenCount type="adult">288 adult</specimenCount>
+ 122 j/b shells; 
<collectingRegion country="Vietnam" name="Thanh Hoa">
Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province
</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:801AC47615B69FE81995C4F4AAC26D9C" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Lang Chanh District" stateProvince="Thanh Hoa Province">
Lang 
<normalizedToken originalValue="Chánh">Chanh</normalizedToken>
District
</location>
, 
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<normalizedToken originalValue="Lương">Lương</normalizedToken>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.4" unit="km" value="9.4">9.4 km</quantity>
northeast from centre of Lang 
<normalizedToken originalValue="Chánh">Chanh</normalizedToken>
towards 
<normalizedToken originalValue="Làng">Lang</normalizedToken>
Thung (locality code: 2020/32); 
<geoCoordinate degrees="20" direction="north" minutes="11.52" orientation="latitude" precision="9" value="20.192">20°11.52'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="15.59" orientation="longitude" precision="9" value="105.259834">105°15.59'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-12">12 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 3 shells; 
<collectingRegion country="Vietnam" name="Son La">
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province
</collectingRegion>
, Mộc 
<normalizedToken originalValue="Châu">Chau</normalizedToken>
, Hang 
<normalizedToken originalValue="Dơi">Dơi</normalizedToken>
, around the entrance of the cave (locality code: 2020/26); 
<geoCoordinate degrees="20" direction="north" minutes="50.96" orientation="latitude" precision="9" value="20.849333">20°50.96'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="38.34" orientation="longitude" precision="9" value="104.639">104°38.34'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">865 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-11">11 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 12 shells; 
<collectingRegion country="Vietnam" name="Hoa Binh">
<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
<normalizedToken originalValue="Bình">Binh</normalizedToken>
Province
</collectingRegion>
, Kim 
<normalizedToken originalValue="Bȏi">Bȏi</normalizedToken>
District, Cao 
<normalizedToken originalValue="Dương">Dương</normalizedToken>
, north of 
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ồng 
<normalizedToken originalValue="Phú">Phu</normalizedToken>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.8" unit="km" value="58.0">58 km</quantity>
from Nho Quan towards 
<collectingRegion country="Vietnam" name="Ha Noi">Hanoi</collectingRegion>
on the Hồ 
<normalizedToken originalValue="Chí">Chi</normalizedToken>
Minh road (locality code: 2020/47); 
<geoCoordinate degrees="20" direction="north" minutes="42.59" orientation="latitude" precision="9" value="20.709833">20°42.59'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="39.30" orientation="longitude" precision="9" value="105.655">105°39.30'E</geoCoordinate>
; 
<elevation metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-16">16 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 
<specimenCount type="adult">9 adult</specimenCount>
+ 10 j/b shells; 
<collectingRegion country="Vietnam" name="Son La">
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province
</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:779BE8EC7FAB01AC93E98E0ECBC9292A" country="Vietnam" county="Pu Puong National Park" latitude="21.6985" longLatPrecision="12" longitude="106.36283" municipality="Am" name="Yen Chau District" stateProvince="Thanh Hoa Province">
<normalizedToken originalValue="Yên">Yen</normalizedToken>
<normalizedToken originalValue="Châu">Chau</normalizedToken>
District
</location>
, 
<collectorName>
<normalizedToken originalValue="Xã">Xa</normalizedToken>
Chi
</collectorName>
ềng 
<collectorName>On, B</collectorName>
ản 
<collectorName>Tr</collectorName>
ạm Hốc, 
<collectorName>Hang Nh</collectorName>
ả 
<collectorName>Nhung</collectorName>
, around the cave (locality code: 2020/20); 
<geoCoordinate degrees="20" direction="north" minutes="59.48" orientation="latitude" precision="9" value="20.991333">20°59.48'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="11.27" orientation="longitude" precision="9" value="104.187836">104°11.27'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">970 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-09">9 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 shell; Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
<collectorName>Province, H</collectorName>
ữu 
<normalizedToken originalValue="Lũng">Lũng</normalizedToken>
<collectorName>District, H</collectorName>
ữu 
<normalizedToken originalValue="Liên">
<collectorName>Lien</collectorName>
</normalizedToken>
, cross of roads no. 1B-241, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.35" unit="km" value="33.5">33.5 km</quantity>
towards 
<collectorName>
Ba 
<normalizedToken originalValue="Nàng">Nang</normalizedToken>
</collectorName>
, left side of road (locality code: 2020/52); 
<geoCoordinate degrees="21" direction="north" minutes="41.06" orientation="latitude" precision="9" value="21.684334">21°41.06'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="22.87" orientation="longitude" precision="9" value="106.381165">106°22.87'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-19">19 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA (with unusually high parietal tooth) • 1 shell; Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
<collectorName>Province, H</collectorName>
ữu 
<normalizedToken originalValue="Lũng">Lũng</normalizedToken>
<collectorName>District, H</collectorName>
ữu 
<normalizedToken originalValue="Liên">
<collectorName>Lien</collectorName>
</normalizedToken>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
west from 
<normalizedToken originalValue="Đȏng">Đȏng</normalizedToken>
<normalizedToken originalValue="Lâm">
<collectorName>Lam</collectorName>
</normalizedToken>
along road no. 241 (locality code: 2020/53); 
<geoCoordinate degrees="21" direction="north" minutes="41.91" orientation="latitude" precision="9" value="21.6985">21°41.91'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="21.77" orientation="longitude" precision="9" value="106.36283">106°21.77'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">210 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-19">19 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA (with unusually high parietal tooth)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2019-10-07" collectorName="A. Hunyadi" country="Laos" county="Na Mor" elevation="635" latitude="20.872833" location="Tham Nathong" longLatPrecision="12" longitude="101.783" municipality="Ban Nathong" specimenCount="141" specimenCount-adult="141" stateProvince="Udomxai Province">
<collectingCountry name="Laos">Laos</collectingCountry>
• 
<specimenCount type="adult">141 adult</specimenCount>
+ 42 j/b shells; 
<collectingRegion country="Laos" name="Oudomxai">Udomxai Province</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.5" unit="km" value="6.5">6.5 km</quantity>
southeast from centre of 
<collectingCounty>Na Mor</collectingCounty>
towards 
<collectingRegion country="Laos" name="Oudomxai">Udomxai</collectingRegion>
, 
<collectingMunicipality>Ban Nathong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:336DF831A999F1C0524C152293943850" country="Laos" county="Na Mor" latitude="20.872833" longLatPrecision="12" longitude="101.783" municipality="Ban Nathong" name="Tham Nathong" stateProvince="Udomxai Province">Tham Nathong</location>
, below cave spring (locality code: 2019/118); 
<geoCoordinate degrees="20" direction="north" minutes="52.37" orientation="latitude" precision="9" value="20.872833">20°52.37'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="46.98" orientation="longitude" precision="9" value="101.783">101°46.98'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.35" unit="m" value="635.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.35" unit="m" value="635.0">635 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-07">7 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2019-10-07" collectorName="A. Hunyadi" country="Laos" elevation="660" latitude="20.8895" location="Na Xay" longLatPrecision="12" longitude="101.816" municipality="Na Mor" specimenCount="75" specimenCount-adult="75" stateProvince="Udomxai Province">
<specimenCount type="adult">75 adult</specimenCount>
shells; 
<collectingRegion country="Laos" name="Oudomxai">Udomxai Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
south of centre of 
<collectingMunicipality>Na Mor</collectingMunicipality>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.8" unit="km" value="3.8">3.8 km</quantity>
east-southeast of 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:F58400C8E99CF13F6E77CB8EB8E864E3" country="Laos" latitude="20.8895" longLatPrecision="12" longitude="101.816" municipality="Na Mor" name="Na Xay" stateProvince="Udomxai Province">Na Xay</location>
, rock wall facing north (locality code: 2019/119); 
<geoCoordinate degrees="20" direction="north" minutes="53.37" orientation="latitude" precision="9" value="20.8895">20°53.37'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="48.96" orientation="longitude" precision="9" value="101.816">101°48.96'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.6" unit="m" value="660.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.6" unit="m" value="660.0">660 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-07">7 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
• 
<materialsCitation collectingDate="2019-10-05" collectorName="A. Hunyadi" country="Laos" county="Nong Khiaw" latitude="20.543167" location="Ban Huai Lek" longLatPrecision="12" longitude="102.686" municipality="Pak Xeng" specimenCount="12" specimenCount-adult="12" stateProvince="Luang Prabang Province">
<specimenCount type="adult">12 adult</specimenCount>
+ 6 j/b shells; 
<collectingRegion country="Laos" name="Louangphabang">Luang Prabang Province</collectingRegion>
, 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
from centre of 
<collectingCounty>Nong Khiaw</collectingCounty>
towards 
<collectingMunicipality>Pak Xeng</collectingMunicipality>
, northeast from 
<location LSID="urn:lsid:plazi:treatment:C07A73CCDA665F3BB3E9E8FBC045A737:BB492F32D0134E2DC2FCA0A764825206" country="Laos" county="Nong Khiaw" latitude="20.543167" longLatPrecision="12" longitude="102.686" municipality="Pak Xeng" name="Ban Huai Lek" stateProvince="Luang Prabang Province">Ban Huai Lek</location>
, left side of the road (locality code: 2019/115); 
<geoCoordinate degrees="20" direction="north" minutes="32.59" orientation="latitude" precision="9" value="20.543167">20°32.59'N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="41.16" orientation="longitude" precision="9" value="102.686">102°41.16'E</geoCoordinate>
; 
<collectingDate value="2019-10-05">5 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg., coll. HA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A small to medium-sized 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with conical to slightly conical-globular shell (shell height variable), an adnate peristome, and a weak parietal tooth (sometimes absent).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.82-0.96, D = 0.77-0.9, H/D*100 = 96.6-126.5 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 50, newly collected populations), RUD = 18.1-25.3 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 11); H = 0.88-1.03, D = 0.77-0.89, H/D*100 = 82.5-92.1 (shells from the type locality, 
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Distinguishable from all congeners (including the widespread 
<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
</taxonomicName>
, which also lives in sympatry) based on the conical-globular shell, the relatively large aperture, and the deeply situated parietal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila szekeresi</emphasis>
</taxonomicName>
was described from Guangxi Province, China. Here we report this species from several new localities in northern Laos and northern Vietnam (Fig. 
<figureCitation captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. Distribution of Angustopila szekeresi Pall-Gergely &amp; Hunyadi, 2015." figureDoi="10.3897/zookeys.1147.93824.figure57" httpUri="https://binary.pensoft.net/fig/808866" pageId="0" pageNumber="1">57</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure56" httpUri="https://binary.pensoft.net/fig/808865" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
Synoptic view of shells from different 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila szekeresi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, 2015 populations.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure57" httpUri="https://binary.pensoft.net/fig/808866" pageId="0" pageNumber="1" start="Figure 57" startId="F57">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 57.</emphasis>
Distribution of 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila szekeresi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, 2015.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Most of the newly discovered populations show little variability in terms of shell characters. The only notable samples are 2020/52 and 2020/53 (distance between them is only 2.5 km) in Vietnam, which have an unusually high parietal tooth and a conical (not conical-globular) shell. More material is necessary to infer the taxonomic position of the latter populations because, for both samples, single shells are all we have. For the time being, they are referred to as 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila szekeresi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="szekeresi">Angustopila cf. szekeresi</taxonomicName>
. We do not assign these two populations to 
<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
</taxonomicName>
because they are both sympatric with 
<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
</taxonomicName>
(distinguishable based on the larger, proportionally wider shell).
</paragraph>
<paragraph pageId="0" pageNumber="1">The shells from the type locality are taller than the newly collected ones. However, no other important shell characters are found to be different.</paragraph>
</subSubSection>
</treatment>
</document>