190 lines
21 KiB
XML
190 lines
21 KiB
XML
<document id="C25D5305F7CB35AA8B1C34364DE9125D" ID-CLB-Dataset="22233" ID-DOI="10.11646/zootaxa.4777.1.1" ID-GBIF-Dataset="75de85af-39c5-41ca-9ccc-4ff7b743eefa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821174" ID-ZooBank="BF4149EF-478F-4FE4-BE7B-B497272F99C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267332095" checkinUser="plazi" docAuthor="Shen, Jia-Wei & Leschen, Richard A. B." docDate="2020" docId="C0438794D865FF87FF01735AFD9CFFFD" docLanguage="en" docName="zootaxa.4777.1.1.pdf.imf" docOrigin="Zootaxa 4777 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eupines (Byraxis) illustris" docType="treatment" docVersion="6" lastPageNumber="41" masterDocId="3C7AFFECD842FFAFFF967608FFA4FE0A" masterDocTitle="Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)" masterLastPageNumber="84" masterPageNumber="1" pageNumber="40" updateTime="1698839235073" updateUser="ExternalLinkService">
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<mods:titleInfo id="A72ABBF9D6138D84D96002B182D001E6">
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<mods:title id="BA4A2A8084AA7777FB1366CAEA251F83">Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)</mods:title>
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<mods:namePart id="30C88B24F12887F493EBAC954BD4F84E">Shen, Jia-Wei</mods:namePart>
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<mods:namePart id="E05F86F915C8EEF3435306CC6A14202B">Leschen, Richard A. B.</mods:namePart>
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<mods:title id="2919E270D0A78011CAE3B6611DF72C0E">Zootaxa</mods:title>
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<mods:part id="9976C18B4E8273BC39CAA28BC52880A8">
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<mods:date id="49D508841C3BDAEA32CC878D9B182B29">2020</mods:date>
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<mods:number id="D108AB7E0EC2547C0D07967ED5791078">4777</mods:number>
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<mods:identifier id="41F4FDC13DD54630E187D40914868FAC" type="DOI">10.11646/zootaxa.4777.1.1</mods:identifier>
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<treatment id="C0438794D865FF87FF01735AFD9CFFFD" ID-DOI="http://doi.org/10.5281/zenodo.3844528" ID-GBIF-Taxon="164123587" ID-Zenodo-Dep="3844528" LSID="urn:lsid:plazi:treatment:C0438794D865FF87FF01735AFD9CFFFD" httpUri="http://treatment.plazi.org/id/C0438794D865FF87FF01735AFD9CFFFD" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection id="00F06509D865FF88FF01735AFD2CFB61" box="[151,648,1361,1388]" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="48553682D865FF88FF01735AFD2CFB61" blockId="39.[151,648,1361,1424]" box="[151,648,1361,1388]" pageId="39" pageNumber="40">
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<heading id="131D81EED865FF88FF01735AFD2CFB61" bold="true" box="[151,648,1361,1388]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
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<taxonomicName id="8FEA4D01D865FF88FF01735AFD2CFB61" ID-CoL="6HC6Z" authority="(Broun, 1914)" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[151,648,1361,1388]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="illustris" subGenus="Byraxis">
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<emphasis id="7A9EEA90D865FF88FF01735AFD2CFB61" bold="true" box="[151,648,1361,1388]" pageId="39" pageNumber="40">
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<emphasis id="7A9EEA90D865FF88FF01735AFF5EFB66" bold="true" box="[151,250,1362,1388]" italics="true" pageId="39" pageNumber="40">Eupines</emphasis>
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||
(
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<emphasis id="7A9EEA90D865FF88FE9A735AFECDFB66" bold="true" box="[268,361,1362,1388]" italics="true" pageId="39" pageNumber="40">Byraxis</emphasis>
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)
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<emphasis id="7A9EEA90D865FF88FEEC7359FE71FB61" bold="true" box="[378,469,1361,1387]" italics="true" pageId="39" pageNumber="40">illustris</emphasis>
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||
(Broun, 1914)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="00F06509D865FF88FF01737DFED8FB9A" box="[151,380,1397,1424]" pageId="39" pageNumber="40" type="description">
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<paragraph id="48553682D865FF88FF01737DFED8FB9A" blockId="39.[151,648,1361,1424]" box="[151,380,1397,1424]" pageId="39" pageNumber="40">
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<figureCitation id="D0D12A07D865FF88FF01737DFF4EFB9A" box="[151,234,1397,1424]" captionStart="FIGURE 25" captionStartId="40.[151,250,1652,1677]" captionTargetBox="[167,1417,533,1621]" captionTargetId="figure@40.[163,1425,525,1627]" captionTargetPageId="40" captionText="FIGURE 25. Diagnostic characters of E. (B.) illustris (Broun, 1914). A) Habitus. B) Antenna, in dorsal view. C) Abdomen, in ventral view. Scale bars: A = 1 mm, B–C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821226" httpUri="https://zenodo.org/record/3821226/files/figure.png" pageId="39" pageNumber="40">Fig. 25</figureCitation>
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; map
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<figureCitation id="D0D12A07D865FF88FEBB737DFED8FB9A" box="[301,380,1397,1424]" captionStart="FIGURE 51" captionStartId="77.[151,250,1945,1970]" captionTargetBox="[205,1383,201,1918]" captionTargetId="figure@77.[191,1397,181,1920]" captionTargetPageId="77" captionText="FIGURE 51. Geographic distributions of E. (B.) glabrata (Broun, 1886), E. (B.) graceae sp. n., E. (B.) halli (Broun, 1921), E. (B.) hectori (Broun, 1895), E. (B.) hoarei sp. n., E. (B.) huizhenae sp. n., and E. (B.) illustris (Broun, 1914) in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.3821280" httpUri="https://zenodo.org/record/3821280/files/figure.png" pageId="39" pageNumber="40">Fig. 51</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="00F06509D865FF88FF0173B3FE57F805" pageId="39" pageNumber="40" type="reference_group">
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<paragraph id="48553682D865FF88FF0173B3FC9BFBD8" blockId="39.[151,1436,1465,1551]" box="[151,831,1465,1490]" pageId="39" pageNumber="40">
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<treatmentCitationGroup id="68FA11ACD865FF88FF0173B3FC9BFBD8" box="[151,831,1465,1490]" pageId="39" pageNumber="40">
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<treatmentCitation id="C94B1093D865FF88FF0173B3FDA4FBD8" author="Broun, T." box="[151,512,1465,1490]" page="168" pageId="39" pageNumber="40" year="1914">
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<taxonomicName id="8FEA4D01D865FF88FF0173B3FDA4FBD8" ID-CoL="9NSXC" authority="Broun, 1914 b: 168" authorityName="Broun" authorityPageNumber="168" authorityYear="1914" box="[151,512,1465,1490]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="illustris">
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<emphasis id="7A9EEA90D865FF88FF0173B3FE9CFBD8" box="[151,312,1466,1490]" italics="true" pageId="39" pageNumber="40">Byraxis illustris</emphasis>
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<bibRefCitation id="2C7B4B73D865FF88FEA973B2FDA4FBD8" author="Broun, T." box="[319,512,1465,1490]" pageId="39" pageNumber="40" pagination="143 - 266" refId="ref58289" refString="Broun, T. (1914 b) Descriptions of new genera and species of Coleoptera. Part III. Bulletin of the New Zealand Institute Wellington, 1 (3), 143 - 266." type="journal article" year="1914">Broun, 1914b: 168</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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<treatmentCitation id="C94B1093D865FF88FD9C73B1FD63FBD8" author="Hudson, G. V." box="[522,711,1465,1490]" page="364" pageId="39" pageNumber="40" year="1923">
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<bibRefCitation id="2C7B4B73D865FF88FD9C73B1FD63FBD8" author="Hudson, G. V." box="[522,711,1465,1490]" pageId="39" pageNumber="40" pagination="353 - 399" refId="ref58506" refString="Hudson, G. V. (1923) An index to New Zealand beetles. Transactions of the New Zealand Institute, 54, 353 - 399." type="journal article" year="1923">Hudson 1923: 364</bibRefCitation>
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</treatmentCitation>
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,
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||
<treatmentCitation id="C94B1093D865FF88FD4473B2FC9FFBD8" author="Hudson, G. V." box="[722,827,1465,1490]" page="185" pageId="39" pageNumber="40" year="1934">
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<bibRefCitation id="2C7B4B73D865FF88FD4473B2FC9FFBD8" author="Hudson, G. V." box="[722,827,1465,1490]" pageId="39" pageNumber="40" refId="ref58535" refString="Hudson, G. V. (1934) New Zealand beetles and their larvae. An elementary introduction to the study of our native Coleoptera. Wellington Ferguson & Osborn Ltd., Wellington, 236 pp., 17 pls." type="book" year="1934">1934: 185</bibRefCitation>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="48553682D865FF88FF0173D1FE57F805" blockId="39.[151,1436,1465,1551]" pageId="39" pageNumber="40">
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<treatmentCitationGroup id="68FA11ACD865FF88FF0173D1FE57F805" pageId="39" pageNumber="40">
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<taxonomicName id="8FEA4D01D865FF88FF0173D1FE06FBFA" ID-CoL="6HC6Z" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[151,418,1496,1521]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="illustris" subGenus="Byraxis">
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<emphasis id="7A9EEA90D865FF88FF0173D1FF4DFBFA" box="[151,233,1497,1520]" italics="true" pageId="39" pageNumber="40">Eupines</emphasis>
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||
(
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<emphasis id="7A9EEA90D865FF88FF6E73D1FEE1FBFA" box="[248,325,1497,1520]" italics="true" pageId="39" pageNumber="40">Byraxis</emphasis>
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||
)
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||
<emphasis id="7A9EEA90D865FF88FEC373D0FE06FBFA" box="[341,418,1496,1520]" italics="true" pageId="39" pageNumber="40">illustris</emphasis>
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||
</taxonomicName>
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||
: combination and subgeneric assignment by
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<treatmentCitation id="C94B1093D865FF88FCF873D1FB1CFBFB" author="Nomura, S. & Leschen, R. A. B." box="[878,1208,1496,1521]" page="255" pageId="39" pageNumber="40" year="2006">
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<bibRefCitation id="2C7B4B73D865FF88FCF873D1FB1CFBFB" author="Nomura, S. & Leschen, R. A. B." box="[878,1208,1496,1521]" pageId="39" pageNumber="40" pagination="239 - 272" refId="ref58967" refString="Nomura, S. & Leschen, R. A. B. (2006) Faunistic review on the pselaphine species known from New Zealand (Insecta, Coleoptera, Staphylinidae). National Science Museum Monographs, 34, 239 - 272." type="journal article" year="2006">Nomura and Leschen 2006: 255</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="C94B1093D865FF88FB5473D0FA3CFBFB" author="Maddison, P." box="[1218,1432,1496,1521]" page="417" pageId="39" pageNumber="40" year="2010">
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<bibRefCitation id="2C7B4B73D865FF88FB5473D0FA3CFBFB" author="Maddison, P." box="[1218,1432,1496,1521]" pageId="39" pageNumber="40" pagination="233 - 467" refId="ref58776" refString="Maddison, P. (2010) Checklist of New Zealand Hexapoda. Order Coleoptera. Chapter Nine: Phylum Arthropoda: Subphylum Hexapoda: Protura, springtails, Diplura, and insects. In: Gordon, D. P. (Ed.), New Zealand inventory of biodiversity. Vol. 2. Kingdom Animalia: Chaetognatha, Ecdysozoa, Ichnofossils. Canterbury University Press, Christchurch, pp. 233 - 467." type="book chapter" year="2010">Maddison 2010: 417</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C94B1093D865FF88FF5173FEFE4BF805" author="Shen, J. - W. & Leschen, R. A. B." box="[199,495,1526,1551]" httpUri="http://treatment.plazi.org/id/2D4AB362C80D8D42FF61FD3F4096E192" page="267" pageId="39" pageNumber="40" year="2019">
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<bibRefCitation id="2C7B4B73D865FF88FF5173FEFE4BF805" author="Shen, J. - W. & Leschen, R. A. B." box="[199,495,1526,1551]" pageId="39" pageNumber="40" pagination="255 - 302" refId="ref59638" refString="Shen, J. - W. & Leschen, R. A. B. (2019) Catalogue and type designations for New Zealand Goniaceritae (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4614 (2), 255 - 302. https: // doi. org / 10.11646 / zootaxa. 4614.2.2" type="journal article" year="2019">Shen and Leschen 2019: 267</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="00F06509D865FF88FF017036FD21F876" pageId="39" pageNumber="40" type="materials_examined">
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<paragraph id="48553682D865FF88FF017036FD0AF85D" blockId="39.[151,1436,1597,2020]" box="[151,686,1597,1624]" pageId="39" pageNumber="40">
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||
<materialsCitation id="F8823CDFD865FF88FF017036FD0EF85D" ID-GBIF-Occurrence="2612164346" box="[151,682,1597,1624]" collectionCode="MC" location="Broken River" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Canterbury" typeStatus="holotype">
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||
<emphasis id="7A9EEA90D865FF88FF017036FE9DF852" bold="true" box="[151,313,1597,1624]" pageId="39" pageNumber="40">
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||
<typeStatus id="97518820D865FF88FF017036FF76F852" box="[151,210,1598,1624]" pageId="39" pageNumber="40">Type</typeStatus>
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||
locality:
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||
</emphasis>
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||
[
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||
<collectionCode id="2EFBAE47D865FF88FED17035FEDCF852" box="[327,376,1597,1624]" country="0" name="Museo de Cipolleti" pageId="39" pageNumber="40">MC</collectionCode>
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||
]
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||
<location id="4D356059D865FF88FE137035FDBBF85D" LSID="urn:lsid:plazi:treatment:C0438794D865FF87FF01735AFD9CFFFD:4D356059D865FF88FE137035FDBBF85D" box="[389,543,1597,1623]" name="Broken River" pageId="39" pageNumber="40" stateProvince="Canterbury">Broken River</location>
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||
,
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||
<collectingRegion id="8A2EF860D865FF88FDBC7035FD0EF85D" box="[554,682,1597,1623]" country="New Zealand" name="Canterbury" pageId="39" pageNumber="40">Canterbury</collectingRegion>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="48553682D865FF88FF51706AFD21F876" blockId="39.[151,1436,1597,2020]" box="[199,645,1633,1660]" pageId="39" pageNumber="40">
|
||
<materialsCitation id="F8823CDFD865FF88FF51706AFD25F876" ID-GBIF-Occurrence="2612164386" box="[199,641,1633,1660]" collectionCode="BMNH" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="7A9EEA90D865FF88FF51706AFE34F876" bold="true" box="[199,400,1633,1660]" pageId="39" pageNumber="40">
|
||
<typeStatus id="97518820D865FF88FF51706AFEA6F876" box="[199,258,1634,1660]" pageId="39" pageNumber="40">Type</typeStatus>
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||
depository:
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||
</emphasis>
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||
<collectionCode id="2EFBAE47D865FF88FE01706AFE4FF876" box="[407,491,1634,1660]" pageId="39" pageNumber="40">BMNH</collectionCode>
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LT
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<specimenCount id="5EECFD0BD865FF88FD8E706AFD8BF876" box="[536,559,1634,1660]" pageId="39" pageNumber="40" type="male">♂</specimenCount>
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||
, 1 PLT
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00F06509D865FF88FF51708DFAA6F925" pageId="39" pageNumber="40" type="diagnosis">
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||
<paragraph id="48553682D865FF88FF51708DFAA6F925" blockId="39.[151,1436,1597,2020]" pageId="39" pageNumber="40">
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||
<emphasis id="7A9EEA90D865FF88FF51708DFEE0F8AA" bold="true" box="[199,324,1669,1696]" pageId="39" pageNumber="40">Diagnosis:</emphasis>
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Head with vertexal foveae punctiform; antennae 10-segmented with 3-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A9 large and asymmetrical, strongly transverse and obliquely truncate at apex, with lateral acute spine, A10 largest and conical, with basal excavation. Abdominal ventrite 2 with pair of large discal protuberances, inserted with strongly curved large apical trichomes; ventrite 6 unknown. Protrochanters with minute ventral spine, about one-tenth of protrochanteral width; protibiae and mesotibiae unmodified.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00F06509D865FF87FF517131FB5BFF6D" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
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||
<paragraph id="48553682D865FF87FF517131FB5BFF6D" blockId="39.[151,1436,1597,2020]" lastBlockId="40.[151,1437,153,503]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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||
<emphasis id="7A9EEA90D865FF88FF517131FED3F95E" bold="true" box="[199,375,1849,1876]" pageId="39" pageNumber="40">Redescription:</emphasis>
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Body length
|
||
<quantity id="8F129B67D865FF88FD837131FDDAF95E" box="[533,638,1849,1876]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="39" pageNumber="40" unit="mm" value="1.64">1.64 mm</quantity>
|
||
. body colour reddish brown with lighter coloured tarsi.
|
||
<emphasis id="7A9EEA90D865FF88FB697132FA9BF95E" bold="true" box="[1279,1343,1850,1876]" pageId="39" pageNumber="40">Head</emphasis>
|
||
slightly wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, with about 31 facets; vertexal foveae punctiform; antennal tubercles weak; postantennal notches weak; postantennal cavities large, about as long as antennal tubercles in dorsal view; antennae 10-segmented with 3-segmented antennal club; A4, A5, A6, A7 and A8 unmodified, A9 large and asymmetrical, strongly transverse and obliquely truncate at apex, with lateral acute spine, A10 largest and conical, with basal excavation.
|
||
<emphasis id="7A9EEA90D86AFF87FB137691FB5AFEB9" bold="true" box="[1157,1278,153,179]" pageId="40" pageNumber="41">Pronotum</emphasis>
|
||
about as long as wide, wider than head at eye level; lacking lateral antebasal foveae.
|
||
<emphasis id="7A9EEA90D86AFF87FC3276B5FC54FEDD" bold="true" box="[932,1008,189,215]" pageId="40" pageNumber="41">Elytra</emphasis>
|
||
as long as wide, widest at about apical three-fourths; humeral calli slightly prominent; sutural stria distinctively extending from base to apex.
|
||
<emphasis id="7A9EEA90D86AFF87FABE76E8FA39FEF1" bold="true" box="[1320,1437,224,251]" pageId="40" pageNumber="41">Abdomen</emphasis>
|
||
with tergite 1 about twice length of tergite 2; ventrite 2 with pair of large discal protuberances, inserted with strongly curved large apical trichomes; ventrite 6 unknown.
|
||
<emphasis id="7A9EEA90D86AFF87FD777721FCBDFF49" bold="true" box="[737,793,297,323]" pageId="40" pageNumber="41">Legs</emphasis>
|
||
with protrochanters modified with minute ventral spine, about one-tenth of protrochanteral width; protibiae and mesotibiae unmodified.
|
||
<emphasis id="7A9EEA90D86AFF87FB837744FB2CFF6D" bold="true" box="[1045,1160,332,359]" pageId="40" pageNumber="41">Aedeagus</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00F06509D86AFF87FF517778FE71FFD9" pageId="40" pageNumber="41" type="discussion">
|
||
<paragraph id="48553682D86AFF87FF517778FE71FFD9" blockId="40.[151,1437,153,503]" pageId="40" pageNumber="41">
|
||
<emphasis id="7A9EEA90D86AFF87FF517778FEF6FF81" bold="true" box="[199,338,368,395]" pageId="40" pageNumber="41">Comments:</emphasis>
|
||
Only two type specimens are known. We did not dissect
|
||
<typeStatus id="97518820D86AFF87FC477779FB97FF81" box="[977,1075,369,395]" pageId="40" pageNumber="41" type="syntype">syntypes</typeStatus>
|
||
since males of this species have distinctive A9 modifications and two tufts of large trichomes on ventrite 2 (
|
||
<figureCitation id="D0D12A07D86AFF87FC75779DFB9AFFA5" box="[995,1086,404,431]" captionStart="FIGURE 25" captionStartId="40.[151,250,1652,1677]" captionTargetBox="[167,1417,533,1621]" captionTargetId="figure@40.[163,1425,525,1627]" captionTargetPageId="40" captionText="FIGURE 25. Diagnostic characters of E. (B.) illustris (Broun, 1914). A) Habitus. B) Antenna, in dorsal view. C) Abdomen, in ventral view. Scale bars: A = 1 mm, B–C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821226" httpUri="https://zenodo.org/record/3821226/files/figure.png" pageId="40" pageNumber="41">Figs. 25</figureCitation>
|
||
B–C) that can immediately differentiate it from congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00F06509D86AFF87FF5177D4FD9CFFFD" box="[199,568,476,503]" pageId="40" pageNumber="41" type="distribution">
|
||
<paragraph id="48553682D86AFF87FF5177D4FD9CFFFD" blockId="40.[151,1437,153,503]" box="[199,568,476,503]" pageId="40" pageNumber="41">
|
||
<emphasis id="7A9EEA90D86AFF87FF5177D4FEC7FFFD" bold="true" box="[199,355,476,503]" pageId="40" pageNumber="41">Distribution:</emphasis>
|
||
South Island. MC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |