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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1166.96193" ID-Pensoft-Pub="1313-2970-1166-91" ID-Pensoft-UUID="4632CBCBEB0659009F53366E2553A21C" ID-ZooBank="4B15C8CDDFF94ACEAD2A749D3AB5C5AD" ModsDocID="1313-2970-1166-91" checkinTime="1686086550896" checkinUser="pensoft" docAuthor="Dvorak, Libor, Fogasova, Katarina, Obona, Jozef, Toeroek, Edina &amp; Manko, Peter" docDate="2023" docId="BFF28901CF5653939BCF41D24D443143" docLanguage="en" docName="ZooKeys 1166: 91-102" docOrigin="ZooKeys 1166" docPubDate="2023-06-06" docSource="http://dx.doi.org/10.3897/zookeys.1166.96193" docTitle="Ptychoptera staryi Dvorak, Obona &amp; Manko 2023, sp. nov." docType="treatment" docUuid="21E4C563-97EA-497A-92C6-02A6A8621746" docUuidSource="ZooBank" docVersion="2" id="4632CBCBEB0659009F53366E2553A21C" lastPageNumber="91" masterDocId="4632CBCBEB0659009F53366E2553A21C" masterDocTitle="Two new Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from the Western Palaearctic" masterLastPageNumber="102" masterPageNumber="91" pageNumber="91" updateTime="1686086854992" updateUser="ExternalLinkService">
<mods:mods id="D8FDAA7771A644CBA5E0839C570F41F0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A37FD25ADFBDE2E08255D9F7FCD821C8">
<mods:title id="C8840CF13A2ABE3D32014E565822FB71">Two new Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from the Western Palaearctic</mods:title>
</mods:titleInfo>
<mods:name id="BABDDBF6209E58BEF86DC78F17376711" type="personal">
<mods:role id="776612A025CBFC901E663386EFEC6AD9">
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<mods:namePart id="9C30DB73A3F8296340AD9FCC6C2D60FE">Dvorak, Libor</mods:namePart>
<mods:nameIdentifier id="5DC21F7EEDBAC3F643F02F9B2DD75088" type="ORCID">https://orcid.org/0000-0002-4712-3679</mods:nameIdentifier>
<mods:affiliation id="4298FA49DB0D2F6F8B306B86EDA3F303">Tri Sekery 21, CZ- 35301 Marianske Lazne, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="DEA24695E67B1AB398536BADC37E8A24" type="personal">
<mods:role id="E9BF4AF4819AA30E06A20BB5B2CF9CC4">
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</mods:role>
<mods:namePart id="EF318FB1DC130140A0A30D9BE9660F11">Fogasova, Katarina</mods:namePart>
<mods:nameIdentifier id="7255AF290E12CD46F279067896DE574F" type="ORCID">https://orcid.org/0009-0003-9604-1885</mods:nameIdentifier>
<mods:affiliation id="FFAE9B14BFE012581868F20CF76546C0">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
<mods:nameIdentifier id="CDA44F92609011590467D885F737C74E" type="email">katarina.fogasova@smail.unipo.sk</mods:nameIdentifier>
</mods:name>
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<mods:role id="E2FE23B37F852BB8B0A02A19C64CDF99">
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</mods:role>
<mods:namePart id="E072ED74E889298E85ECFDBE9F007859">Obona, Jozef</mods:namePart>
<mods:nameIdentifier id="FC9B54FC32E7C37BA760E5B1AE1D1250" type="ORCID">https://orcid.org/0000-0002-1185-658X</mods:nameIdentifier>
<mods:affiliation id="37D13F8E894B0FF6A0A221815B57C22C">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
</mods:name>
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<mods:role id="AEC6295D4F3DAB8D99BE80CFD2475FD3">
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</mods:role>
<mods:namePart id="E634C2AD4A2FC32E9D6CE17FF2B5802D">Toeroek, Edina</mods:namePart>
<mods:nameIdentifier id="92B53AF3F86F4F02C13343DD27D01A52" type="ORCID">https://orcid.org/0000-0001-5982-7078</mods:nameIdentifier>
<mods:affiliation id="B1D37C74E2C99AB2922D411ACC8DB589">&quot; Lenduelet &quot; Landscape and Conservation Ecology Research Group, Institute of Ecology and Botany, Centre for Ecological Research, Alkotmany str. 2 - 4, Vacratot, H- 2163, Hungary</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="1A5A823665ECF2EF72A7828F12FCFB07">Manko, Peter</mods:namePart>
<mods:nameIdentifier id="6379FF02A2B09F2CE8085F3D19AF1D49" type="ORCID">https://orcid.org/0000-0003-1862-9117</mods:nameIdentifier>
<mods:affiliation id="8945ADF7B2CBAA650A0C6410B08C9185">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
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<mods:title id="A6B21739E055D2CDD4175577A000D971">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="555E886B5572592C54165106C1EAC631">
<mods:date id="2AA43CE062F5F95817AEE8BFEFDABDE9">2023</mods:date>
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<mods:number id="D2D2AEC116E4B5F6D9372B2F0A42A429">2023-06-06</mods:number>
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<mods:number id="B622862D71B9A3AD69295F79DA3CE317">1166</mods:number>
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<mods:start id="EBE9042E904C07FA06625977CF239FB8">91</mods:start>
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<mods:classification id="695D1A33297EF50BC34B38AB152696C2">journal article</mods:classification>
<mods:identifier id="CC0D8D7D77B1F0CCAD1FB3D5867FA7B6" type="DOI">http://dx.doi.org/10.3897/zookeys.1166.96193</mods:identifier>
<mods:identifier id="4EC74B1294FB488388AEE8FC231040E8" type="Pensoft-Pub">1313-2970-1166-91</mods:identifier>
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<subSection id="392EA0827BC94B12EA29ED49C3A355E7" pageId="0" pageNumber="91" type="taxonomic account">
<treatment id="BFF28901CF5653939BCF41D24D443143" LSID="urn:lsid:zoobank.org:act:21E4C563-97EA-497A-92C6-02A6A8621746" httpUri="http://treatment.plazi.org/id/BFF28901CF5653939BCF41D24D443143" lastPageNumber="91" pageId="0" pageNumber="91">
<subSubSection id="2B3B6D6F786D66B42F50BF8E57FADB23" pageId="0" pageNumber="91" type="nomenclature">
<paragraph id="390760CB10870811969C11DC8EC7B31D" pageId="0" pageNumber="91">
<taxonomicName id="B34E4E5020DC97B46DD33691E9FC80FA" LSID="https://zoobank.org/21E4C563-97EA-497A-92C6-02A6A8621746" authority="Dvořák, Oboňa &amp; Manko" authorityName="Dvorak, Obona &amp; Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi" status="sp. nov.">
Ptychoptera staryi
<normalizedToken id="3C24E8661F53CAD2AAAF2A6CCB857965" originalValue="Dvořák">Dvorak</normalizedToken>
,
<normalizedToken id="DC083D7EB7FB8F1DDD95101981D8A38E" originalValue="Oboňa">Obona</normalizedToken>
&amp; Manko
</taxonomicName>
<taxonomicNameLabel id="C71911AC798C4F5266E98E48C217D5BE" pageId="0" pageNumber="91">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0A26B680709914C1065D583FFDEA1129" pageId="0" pageNumber="91" type="description">
<paragraph id="2C317AE4D4F2FD4812F8B67181BB37C2" pageId="0" pageNumber="91">
<figureCitation id="4FB66BAE3CA7020AE8D327CC9410665F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ptychoptera staryi sp. nov. a habitus of male, lateral view b wing." figureDoi="10.3897/zookeys.1166.96193.figure4" httpUri="https://binary.pensoft.net/fig/860368" pageId="0" pageNumber="91">Figs 4</figureCitation>
<figureCitation id="C00B0E2B6649F134196AA3B7FF16C281" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epandrium, gonocoxite and gonostylus, and hypandrium of P. staryi sp. nov., P. incognita, and P. albimana. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GCT = gonocoxite, GAS = apical stylus of gonostylus, GAT = tertiary lobe of apical stylus of gonostylus, PPA = apical process of paramere, HBD = basal division of hypandrium, HMW = membranous window of terminal division of hypandrium, HTD = terminal division of hypandrium." figureDoi="10.3897/zookeys.1166.96193.figure5" httpUri="https://binary.pensoft.net/fig/860369" pageId="0" pageNumber="91">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="26B8B8F545CF54AFCA2B8570AA7E52A9" pageId="0" pageNumber="91" type="type material">
<paragraph id="498A1015B02C022B86AEBBDD00B1B641" pageId="0" pageNumber="91">Type material.</paragraph>
<paragraph id="03C19CB1F60D90A9ED80EF51F56AC272" pageId="0" pageNumber="91">
<emphasis id="C0859F3436621ED3084FD259A01AA35C" bold="true" italics="true" pageId="0" pageNumber="91">Holotype</emphasis>
: 1 ♂: Bulgaria, Rhodopes, Yundola, 1 300 m a. s. l.,
<geoCoordinate id="18AAC0D5E6FE4849D37EB3D68BB41F4E" degrees="42" direction="north" minutes="3" orientation="latitude" precision="15" seconds="47" value="42.063053">42°3'47&quot;N</geoCoordinate>
,
<geoCoordinate id="C0D1FD71B30F22985E150D363DAFED5B" degrees="23" direction="east" minutes="51" orientation="longitude" precision="15" seconds="17" value="23.854723">23°51'17&quot;E</geoCoordinate>
, 30.VI.2016, leg. M.
<normalizedToken id="BC04BD378CD95F62064B84253A0D68F4" originalValue="Barták">Bartak</normalizedToken>
et
<normalizedToken id="0E0A3EF488E67790D9C1FE87F6B08862" originalValue="Š">S</normalizedToken>
.
<normalizedToken id="7CC9D03428335978547AC357B6A807AD" originalValue="Kubík">Kubik</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="FAEA6F523D6A02AF7FF8E26B63FE1018" pageId="0" pageNumber="91" type="description">
<paragraph id="FECD459D830B5530A074C6B2C2D6057E" pageId="0" pageNumber="91">Description.</paragraph>
<paragraph id="A254A4FB4C1290629333779E0B9E7A1E" pageId="0" pageNumber="91">
<emphasis id="72169A4A684465014497357A81EBC2AF" bold="true" pageId="0" pageNumber="91">
Male.
<emphasis id="51E9AD4F81D243692DFCD619A7C1421F" italics="true" pageId="0" pageNumber="91">Head</emphasis>
</emphasis>
: Frons, vertex, and occiput black, mouthparts including palpi pale yellow, scape and pedicel yellowish orange, antennal flagellomeres greyish.
</paragraph>
<paragraph id="892A4F16B6B601FFB4C0B63E6F084990" pageId="0" pageNumber="91">
<emphasis id="A802E1F60A058A173C01FB0976976CEE" bold="true" italics="true" pageId="0" pageNumber="91">Thorax</emphasis>
: Predominantly black with silvery pubescent pleurae. Pronotum, epimeron 3 and metanotum 3 yellow. Fore and mid coxae and trochanters yellow, hind coxae brownish black basally, yellow apically, coxae also yellow. Almost all legs are missing. Halteres whitish yellow with a darker knob.
</paragraph>
<paragraph id="FF7492A1D639DB6F2F19A6BE8B74C7DB" pageId="0" pageNumber="91">
Wing length 12 mm (Fig.
<figureCitation id="2C43055779F39D4E381429D1E47BEC3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ptychoptera staryi sp. nov. a habitus of male, lateral view b wing." figureDoi="10.3897/zookeys.1166.96193.figure4" httpUri="https://binary.pensoft.net/fig/860368" pageId="0" pageNumber="91">4b</figureCitation>
). Wing with yellowish veins and infuscated spots on fork vein Rs+R, all cross-veins, end of R1 vein up to its fork with R2 vein, on fork vein R4+R5 and on fork vein M1+M2. Legs: femur pale, darker in apical ¹⁄5, almost black on extreme apex; tibia pale brown in basal
<normalizedToken id="BDCF34C892648CAFF30A8DAB2D22D00E" originalValue="½">1/2</normalizedToken>
, darker apically and almost black on extreme apex; tarsomere 1 almost black, tarsomeres 2 and 3 dark brown, tarsomeres 4 and 5 pale brown.
</paragraph>
<caption id="8AF9409CAEED1995D428C1CA43F9CD4D" doi="10.3897/zookeys.1166.96193.figure4" httpUri="https://binary.pensoft.net/fig/860368" pageId="0" pageNumber="91" start="Figure 4" startId="F4">
<paragraph id="3525451980248C1A5EA5AD559E355CCA" pageId="0" pageNumber="91">
<emphasis id="3E2A5583D9BE20C23BB5F6CBC70D64E6" bold="true" pageId="0" pageNumber="91">Figure 4.</emphasis>
<taxonomicName id="B75D6BD4FC871AB612EBF38B07BA86E4" authorityName="Dvorak, Obona &amp; Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="694F6D64A1F202E124972782AFADDB2F" italics="true" pageId="0" pageNumber="91">Ptychoptera staryi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="6D4373DA8803808C74099994B770CEFD" bold="true" pageId="0" pageNumber="91">a</emphasis>
habitus of male, lateral view
<emphasis id="6CFE424D06B17F299FF5C523840B05A9" bold="true" pageId="0" pageNumber="91">b</emphasis>
wing.
</paragraph>
</caption>
<paragraph id="5B79A80FCC25E11A478E3E40EBB2893C" pageId="0" pageNumber="91">
<emphasis id="CFD67B7717E041F2FEC2C02EB504EA04" bold="true" italics="true" pageId="0" pageNumber="91">Abdomen</emphasis>
: Tergum 1, tergum 2, apical ¹⁄5 of tergum 3, almost whole tergum 4 except base, and whole terga 5-7 black; sternum 1 black, sternum 2 brown, sterna 5-7 black; the remainder orange yellow.
</paragraph>
<paragraph id="F1873BE66E535C361DCEA32C2895D4FC" pageId="0" pageNumber="91">
<emphasis id="923E525593C8DE842C52B38271A8A146" bold="true" italics="true" pageId="0" pageNumber="91">Male genitalia</emphasis>
: similar to
<taxonomicName id="EB1198C6E30897C347FC6C79612C7B70" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="AABD992C74507EE569AF0D36FC5CDD41" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
. Epandrial clasper slightly curved outwards with simple obtuse apex; anterior projection of ventromesal lobe sharp, posterior projection bow-shaped backwards; space between both projections is rounded, almost semi-circular. Gonocoxite and gonostylus: apical process of paramere with a U-shaped dark structure with thick edges; paramere base rounded, convex; width to height ratio of dorsal gonocoxal lobe 0.5; dorsal gonocoxal lobe with dense tiny dark hairs. Aedeagus: sides of lateral ejaculatory process distinctly convex, basal projections markedly convergent; transition to lateral ejaculatory process smooth, undulate. Hypandrium: width to length ratio 1.2; apex of hypogynial valves start under basal division of hypandrium. See also differential diagnosis.
</paragraph>
<paragraph id="57BD9BD693653CB3AA4A4C60E37DCF7E" pageId="0" pageNumber="91">
<emphasis id="58C13868951E197301108EC89AEDD87E" bold="true" pageId="0" pageNumber="91">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="E289082C1CDBEB68486947BA159C3C42" pageId="0" pageNumber="91" type="etymology">
<paragraph id="466B1716741D75F8264AFD947792A40E" pageId="0" pageNumber="91">Etymology.</paragraph>
<paragraph id="A7FE581930449C8DEA45F3EE45614D1D" pageId="0" pageNumber="91">
The name is dedicated to our colleague Jaroslav
<normalizedToken id="6A06D262C17E8AA8C9F61804F7C5B3BE" originalValue="Starý">Stary</normalizedToken>
and his life jubilee. (Jaroslav discovered the holotype of new species in his own material and provided it for the description).
</paragraph>
</subSubSection>
<subSubSection id="023879205F54DE3D073B14EC082397C6" pageId="0" pageNumber="91" type="differential diagnosis">
<paragraph id="78FFB0D5074099E53FCB2AD5652CBA57" pageId="0" pageNumber="91">Differential diagnosis.</paragraph>
<paragraph id="37C4FE1074AF207FDF44546CB494E116" pageId="0" pageNumber="91">
The new species is very similar to
<taxonomicName id="880CA4A7F99BE10A59DF11FBFD27C943" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="C22FBFE4B482BCDA0132C1B5A3FBFA27" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
<normalizedToken id="2EA9B2B71F83FC76404A5829A3BCBDDC" originalValue="Török">Toeroek</normalizedToken>
,
<normalizedToken id="BD3EA6C9138F00C5887057B2849212D2" originalValue="Kolcsár">Kolcsar</normalizedToken>
&amp; Keresztes, 2015 (see also Table
<tableCitation id="58EDA777B7E25D2FFD27C22E9A5ACA74" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Diagnostics of P. staryi sp. nov., P. incognita, and P. albimana based on male genitalia (see also Fig. 5)." httpUri="http://table.plazi.org/id/802D144D7AD7887680F19499911CA410" pageId="0" pageNumber="91" tableUuid="802D144D7AD7887680F19499911CA410">2</tableCitation>
). After comparing the holotype of
<taxonomicName id="3FEE9A63E2DE5B7B24F29B0E83092C53" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="953069A6B4DC584CD01EE97466401E89" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
sp. nov. with individuals of
<taxonomicName id="97AB521A12DDA65CE8A94BFF66662804" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="B170E23F3FB8C89A6071F468D68B4CFC" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
(material used in
<bibRefCitation id="34DAA795F4496C47993603AAAFD6150C" author="Toeroek, E" journalOrPublisher="North-Western Journal of Zoology" pageId="0" pageNumber="91" pagination="304 - 315" refId="B17" refString="Toeroek, E, Kolcsar, L-P, Denes, A-L, Keresztes, L, 2015. Morphologies tells more than molecules in the case of the European widespread Ptychoptera albimana (Fabricius, 1787) (Diptera, Ptychopteridae). North-Western Journal of Zoology 11 (2): 304 - 315" title="Morphologies tells more than molecules in the case of the European widespread Ptychoptera albimana (Fabricius, 1787) (Diptera, Ptychopteridae)." volume="11" year="2015">
<normalizedToken id="7D289D1B8D90A9E5132ACD9219ED74AF" originalValue="Török">Toeroek</normalizedToken>
et al. 2015
</bibRefCitation>
: 2 individuals from Bulgaria, 17 individuals from Romania), we found that a diagnosis was possible on the basis of differences in male genitalia (Fig.
<figureCitation id="BEC928043105A720030DF61FC74D07C1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epandrium, gonocoxite and gonostylus, and hypandrium of P. staryi sp. nov., P. incognita, and P. albimana. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GCT = gonocoxite, GAS = apical stylus of gonostylus, GAT = tertiary lobe of apical stylus of gonostylus, PPA = apical process of paramere, HBD = basal division of hypandrium, HMW = membranous window of terminal division of hypandrium, HTD = terminal division of hypandrium." figureDoi="10.3897/zookeys.1166.96193.figure5" httpUri="https://binary.pensoft.net/fig/860369" pageId="0" pageNumber="91">5</figureCitation>
, marked with arrows), namely: (i) shape of the plate on ventral parts of epandrium is of a different shape and orientated at a different angle in
<taxonomicName id="96979E7C36BCBB0926BCAF92FBF3263A" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="E7E36AEBD2DBE54BB814439B82F52A27" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
; (ii) chitinisation (sclerotisation) of the proximo-lateral processes of the gonocoxite is not developed and these processes are light coloured in
<taxonomicName id="5E0C4E7B5091F6F1A36D10B36DB9459B" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="7F957480F2A02EE5437BE93E6B8026EF" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
in contrast to
<taxonomicName id="B5E0AC9CE3C6509D88ABC8BBAC9D3DED" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="F2B02661E590A13314B6BEFB60B5D888" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
with strong chitinisation and dark colouration; (iii) the hairs of margins of gonocoxite, gonostylus, and epandrium are much less dense and finer in contrast to
<taxonomicName id="155B7B2FF0D6AA08206C0B448EB95170" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="FECBA2CC3737ECAFB9B7C777A5DEFC22" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
(not visible in Fig.
<figureCitation id="39A60BD023E64DA75D4A1DB4BDD05C5E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epandrium, gonocoxite and gonostylus, and hypandrium of P. staryi sp. nov., P. incognita, and P. albimana. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GCT = gonocoxite, GAS = apical stylus of gonostylus, GAT = tertiary lobe of apical stylus of gonostylus, PPA = apical process of paramere, HBD = basal division of hypandrium, HMW = membranous window of terminal division of hypandrium, HTD = terminal division of hypandrium." figureDoi="10.3897/zookeys.1166.96193.figure5" httpUri="https://binary.pensoft.net/fig/860369" pageId="0" pageNumber="91">5</figureCitation>
).
</paragraph>
<caption id="B2F8F415C92004A21D9A35C51D0C3A81" doi="10.3897/zookeys.1166.96193.figure5" httpUri="https://binary.pensoft.net/fig/860369" pageId="0" pageNumber="91" start="Figure 5" startId="F5">
<paragraph id="0812E2DA40FBE8CEDB6958F7F8B2660C" pageId="0" pageNumber="91">
<emphasis id="47816C30128EE08F89F727342363070F" bold="true" pageId="0" pageNumber="91">Figure 5.</emphasis>
Epandrium, gonocoxite and gonostylus, and hypandrium of
<taxonomicName id="B31714EEFBAB6A88B772BC7409602549" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="2A188E5247F077553D2366B88C521406" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="DB2DDAA4251AAE2236BEC43CBE6BA14C" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="C206BA2017F113FC90F06038D078B5CA" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
, and
<taxonomicName id="EBF7917185C0B239683FB99D91EEFECC" class="Insecta" kingdom="Animalia" lsidName="P. albimana." order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="albimana.">
<emphasis id="B94FA6F1A20D59FE62700C41780FD8F8" italics="true" pageId="0" pageNumber="91">P. albimana.</emphasis>
</taxonomicName>
Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GCT = gonocoxite, GAS = apical stylus of gonostylus, GAT = tertiary lobe of apical stylus of gonostylus, PPA = apical process of paramere, HBD = basal division of hypandrium, HMW = membranous window of terminal division of hypandrium, HTD = terminal division of hypandrium.
</paragraph>
</caption>
<caption id="802D144D7AD7887680F19499911CA410" ID-Table-UUID="802D144D7AD7887680F19499911CA410" httpUri="http://table.plazi.org/id/802D144D7AD7887680F19499911CA410" pageId="0" pageNumber="91" start="Table 2" startId="T2">
<paragraph id="49C56677183335F9D680F9E74BF38B35" pageId="0" pageNumber="91">
<emphasis id="ED7B99DEC31CA6821EF6BE55C53FDCCE" bold="true" pageId="0" pageNumber="91">Table 2.</emphasis>
Diagnostics of
<taxonomicName id="CC174365A35F89DF672D2AE8757B2739" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="E5F8FE2EE94CD1630C3423F28457449F" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="B5CC4D7AFFDC01158EC916595CD61089" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="D62475C88581878C647BD24D731F143B" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
, and
<taxonomicName id="782DB28B6286F98363FC4852606FD430" class="Insecta" kingdom="Animalia" lsidName="P. albimana" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="0BCD70F7505AFC02B2F32645BDE8A205" italics="true" pageId="0" pageNumber="91">P. albimana</emphasis>
</taxonomicName>
based on male genitalia (see also Fig.
<figureCitation id="C06853D178ADF7A179979BDA44DCAAD8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epandrium, gonocoxite and gonostylus, and hypandrium of P. staryi sp. nov., P. incognita, and P. albimana. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GCT = gonocoxite, GAS = apical stylus of gonostylus, GAT = tertiary lobe of apical stylus of gonostylus, PPA = apical process of paramere, HBD = basal division of hypandrium, HMW = membranous window of terminal division of hypandrium, HTD = terminal division of hypandrium." figureDoi="10.3897/zookeys.1166.96193.figure5" httpUri="https://binary.pensoft.net/fig/860369" pageId="0" pageNumber="91">5</figureCitation>
).
</paragraph>
</caption>
<paragraph id="C03A648E270B01B9A6D41E96E54D2E7A" pageId="0" pageNumber="91">
<table id="983E2894CB4AB1DD610D9AE20DB6E58E" pageId="0" pageNumber="91">
<tr id="85C0F69C37BB45453CC4F2400271F663" pageId="0" pageNumber="91">
<th id="979F8CA1C72F6C02BED9223C407675AB" colspan="1" isEmpty="true" pageId="0" pageNumber="91" rowspan="1">-</th>
<th id="F9B4153C253FA68296A314E9875C34B4" colspan="1" pageId="0" pageNumber="91" rowspan="1">
<taxonomicName id="83A9D697133698A1AA015F4A2B9091A4" class="Insecta" kingdom="Animalia" lsidName="P. staryi" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="staryi">
<emphasis id="F11B79BE8B04C46AFF679C90BA0BF287" italics="true" pageId="0" pageNumber="91">P. staryi</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="DCDDF09AA82CBC052CB0F9A9E7344299" colspan="1" pageId="0" pageNumber="91" rowspan="1">
<taxonomicName id="D7124481E2568846DD5A13B3E50C691A" class="Insecta" kingdom="Animalia" lsidName="P. incognita" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="76AE73C4483EEA1E8FD9BFCE08B4CE60" italics="true" pageId="0" pageNumber="91">P. incognita</emphasis>
</taxonomicName>
</th>
<th id="EFC25BB4DF67C7AD26A94516080D879E" colspan="1" pageId="0" pageNumber="91" rowspan="1">
<taxonomicName id="E15BABE460B3BA77F57DADF090AD32A3" class="Insecta" kingdom="Animalia" lsidName="P. albimana" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="albimana">
<emphasis id="98467BB8969B037414E8A9579AF30A51" italics="true" pageId="0" pageNumber="91">P. albimana</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="03FFF8E686862C4D16ECBA58C1E01AD4" pageId="0" pageNumber="91">
<td id="FE15CEC8DD443180F1C81B7FC9B591BE" colspan="1" pageId="0" pageNumber="91" rowspan="1">Epandrium</td>
<td id="FA1F48E6D132B371FAA0929FE5A9C751" colspan="1" pageId="0" pageNumber="91" rowspan="1">Shape of the plate on ventral site and the space between projections rounded, almost semi-circular.</td>
<td id="2337819866AFE2AE45B4A4C1DE6C07E1" colspan="1" pageId="0" pageNumber="91" rowspan="1">Projections of the plate on ventral site in about right angle, space between projections quadrangular.</td>
<td id="A465BD933A20A554665B606EB69102B4" colspan="1" pageId="0" pageNumber="91" rowspan="1">Projections of the plate on ventral site in obtuse angle, space between projections rounded, more than semicircle.</td>
</tr>
<tr id="1C9EACFD7B3EEA99D13923E9B23322EC" pageId="0" pageNumber="91">
<td id="622753DFB45781EE5E89EC26AD247D36" colspan="1" pageId="0" pageNumber="91" rowspan="1">Gonocoxite and gonostylus</td>
<td id="F0D9F58A5342656E4D2976A20F8B6188" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width to height ratio of dorsal gonocoxal lobe 0.5; dorsal gonocoxal lobe with dense tiny dark hairs. The hairs of margins of gonocoxite and gonostylus much less dense and finer. Chitinisation of the proximo-lateral processes of the gonocoxite not developed, processes light coloured.</td>
<td id="9AF285AB20160064E712DBA00B148D56" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width to height ratio of dorsal gonocoxal lobe 0.6; dorsal gonocoxal lobe with sparse small hairs. The hairs of margins of gonocoxite and gonostylus dense and stronger. Chitinisation of the proximo-lateral processes of the gonocoxite strongly chitinised, dark.</td>
<td id="986CF1278BC47C047074C9552D0C500A" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width to height ratio of dorsal gonocoxal lobe 0.4; dorsal gonocoxal lobe with tiny dark hairs at the top. The hairs of margins of gonocoxite and gonostylus dense and stronger. Chitinisation of the proximo-lateral processes of the gonocoxite not developed, processes pale coloured.</td>
</tr>
<tr id="DD070801074385B95AAA33FFBC16B0A8" pageId="0" pageNumber="91">
<td id="B177F0CC61E45395CCB32218CA52C8E7" colspan="1" pageId="0" pageNumber="91" rowspan="1">Aedeagus</td>
<td id="E86BD58A83F4E4CC72E436F55F1F172F" colspan="1" pageId="0" pageNumber="91" rowspan="1">Sides of lateral ejaculatory process distinctly convex, basal projections markedly convergent; transition to lateral ejaculatory process smooth, undulate.</td>
<td id="AC61E8E8B8D59D493003C1959DCF7E3D" colspan="1" pageId="0" pageNumber="91" rowspan="1">Sides of lateral ejaculatory process slightly convex, basal projections divergent; transition to lateral ejaculatory process steep, in a right or acute angle to the axis of aedeagus.</td>
<td id="CF71FA33221E64B5BA07607E5903065A" colspan="1" pageId="0" pageNumber="91" rowspan="1">Sides of lateral ejaculatory process slightly convex, basal projections almost parallel; transition to lateral ejaculatory process steep, in an obtuse angle to the axis of aedeagus.</td>
</tr>
<tr id="BA8A8D88DB80629A651ABED514773974" pageId="0" pageNumber="91">
<td id="95CCD508CD6D4A695FAB8A14117B2081" colspan="1" pageId="0" pageNumber="91" rowspan="1">Hypandrium</td>
<td id="07FBF7242ACCEB886AA58314A45E7DEB" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width and length ratio 1.2; apex of hypogynial valves start under basal division of hypandrium.</td>
<td id="F728C53E1C3662BEA918784DBB8EDE08" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width and length ratio 0.9; apex of hypogynial valves start on bases of basal division of hypandrium.</td>
<td id="0921652F2983DB9F7AE90A77332F7BC3" colspan="1" pageId="0" pageNumber="91" rowspan="1">Width and length ratio 0.9; apex of hypogynial valves start over basal division of hypandrium.</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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