725 lines
81 KiB
XML
725 lines
81 KiB
XML
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<mods:title id="6ACE3AE44A28AC28BC13CF4B85E499E0">Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China</mods:title>
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<mods:affiliation id="C52B6A7F29ACF7BD8155F06EE071902C">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
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<mods:affiliation id="8452A699EDE21F716E5D4BA1A0742FBF">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
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<mods:title id="E92829CB0A86B715F0EE22181D8296EB">Vertebrate Zoology</mods:title>
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<mods:date id="4AE9587E0E2BDFFB3B4791F19C3D5A4B">2022</mods:date>
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<taxonomicName id="BB77A324BF1D5A3750B23BE74FC7EEE3" ID-CoL="3MQZP" LSID="BFD1E4E6-F782-59D2-9A9F-27662F224A79" authority="(Gray, 1831)" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura" status="comb. nov.">Occidozyga obscura (Gray, 1831)</taxonomicName>
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<taxonomicNameLabel id="581372968F1574289408470671FE0C75" pageId="0" pageNumber="495">comb. nov.</taxonomicNameLabel>
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<taxonomicName id="905648BB1D97B66E3CB02C5691BE2B79" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Houlema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Houlema obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura" status="comb. nov.">Houlema obscura</taxonomicName>
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<bibRefCitation id="207B66FFD1DAE91214069BBFDB27A22D" author="Gray, JE" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="495" refId="B20" refString="Gray, JE, 1831. Description of two new genera of frogs discovered by John Reeves, Esq. in China. Zoological Miscellany, Part 1: 38." title="Description of two new genera of frogs discovered by John Reeves, Esq. in China. Zoological Miscellany, Part 1: 38." year="1831">Gray 1831</bibRefCitation>
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<paragraph id="CE3279DC7DB39FE9EEA78267DC14EC94" pageId="0" pageNumber="495">
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<taxonomicName id="DBD5F4E06559D82B67D764B8293B63FA" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Oxyglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyglossa lima subsp. var. var. chinens" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="variety" species="lima" status="comb. nov." subSpecies="var." variety="chinens">Oxyglossa lima var. chinens</taxonomicName>
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<bibRefCitation id="EFC44B5F7FDA3498D45702738E220B85" author="Mueller, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="495" pagination="560 - 709" refId="B35" refString="Mueller, F, 1878. Katalog der im Museum und Universitaetskabinat zu Basel aufgestellten Amphibien und Reptilien nebst Anmerkungen. Verhandlungen der Naturforschenden Gesellschaft in Basel 6: 560 - 709" title="Katalog der im Museum und Universitaetskabinat zu Basel aufgestellten Amphibien und Reptilien nebst Anmerkungen." volume="6" year="1878">
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<normalizedToken id="B574835C3C77B672F2B09A0F95928488" originalValue="Müller">Mueller</normalizedToken>
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1878
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<paragraph id="EDDA7AC3CF08AC477C05A9694C16C984" pageId="0" pageNumber="495">
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<taxonomicName id="B662AD46BF151ECED22E6681C814E1F4" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Oxydozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxydozyga lima" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lima" status="comb. nov.">Oxydozyga lima</taxonomicName>
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-
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<bibRefCitation id="B34BB723C32C1CF0F02EF3AFC57CE1AA" author="Stejneger, L" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="495" pagination="1 - 115" refId="B46" refString="Stejneger, L, 1925. Chinese amphibians and reptiles in the United States National Museum. Proceedings of the United States National Museum 66: 1 - 115" title="Chinese amphibians and reptiles in the United States National Museum." volume="66" year="1925">Stejneger 1925</bibRefCitation>
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</paragraph>
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<paragraph id="DBEFA31FC749B4E010BCF3E67305C0E0" pageId="0" pageNumber="495">
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<taxonomicName id="B5158B7DD08209BB3DADDEEE718A2FCA" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Osteosternum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osteosternum amoyense" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="amoyense" status="comb. nov.">Osteosternum amoyense</taxonomicName>
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<bibRefCitation id="BA324C3AD44AEE89E0C196A5A4531A1B" author="Wu, HW" journalOrPublisher="Contributions from the Biological Laboratory of the Science Society of China" pageId="0" pageNumber="495" pagination="1 - 9" refId="B52" refString="Wu, HW, 1929. Osteosternum amoyense, a new frog from Amoy. Contributions from the Biological Laboratory of the Science Society of China 5: 1 - 9" title="Osteosternum amoyense, a new frog from Amoy." volume="5" year="1929">Wu 1929</bibRefCitation>
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</paragraph>
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<paragraph id="09214818B912430E6ABBD65316243EE7" pageId="0" pageNumber="495">
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<taxonomicName id="2C7B6FAADA3E3FAAA91E1A361C9DDC64" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Ooeidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ooeidozyga lima" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lima" status="comb. nov.">Ooeidozyga lima</taxonomicName>
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-
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<bibRefCitation id="35CFC49BA6806F8AC0E0AA6A5A5DE7BB" author="Pope, CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="495" pagination="397 - 611" refId="B38" refString="Pope, CH, 1931. Notes on amphibians from Fukien, Hainan and other parts of China. Bulletin of the American Museum of Natural History 61: 397 - 611" title="Notes on amphibians from Fukien, Hainan and other parts of China." volume="61" year="1931">Pope 1931</bibRefCitation>
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;
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<bibRefCitation id="DEE8D004BAA52EF61058BEA8FF1BC2F6" author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="495" pagination="3 - 32" refId="B45" refString="Smith, MA, 1931. The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft. Bulletin of the Raffles Museum 5: 3 - 32" title="The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft." volume="5" year="1931">Smith 1931</bibRefCitation>
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; Liu & Hu 1961
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</paragraph>
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<paragraph id="E96B2420981299933DE5A1DB880B6EE6" pageId="0" pageNumber="495">
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<taxonomicName id="79AE637B7BD8979FE1E8B159A9C0FC3C" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lima" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lima" status="comb. nov.">Occidozyga lima</taxonomicName>
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-
|
||
<bibRefCitation id="6993057F4E7D133FF1F2B7304EBDA72D" author="Dubois, A" journalOrPublisher="Monitore Zoologico Italiano, Nuova Serie, Supplemento" pageId="0" pageNumber="495" pagination="225 - 284" refId="B7" refString="Dubois, A, 1981. Liste des genres et sous-genres nominaux de Ranoidea (Amphibiens Anoures) du monde, avec identification de leurs especes types; consequences nomenclaturales. Monitore Zoologico Italiano, Nuova Serie, Supplemento 15: 225 - 284" title="Liste des genres et sous-genres nominaux de Ranoidea (Amphibiens Anoures) du monde, avec identification de leurs especes types; consequences nomenclaturales." volume="15" year="1981">Dubois 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="02439EE7B6F9610B973E25087F2E64DB" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B12" refString="Fei, L, Ye, CY, Huang, YZ, 1990. Key to Chinese Amphibians. Publishing House for Scientific and Technological Literature, Chongqing, China." title="Key to Chinese Amphibians. Publishing House for Scientific and Technological Literature, Chongqing, China." year="1990">Fei et al. 1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="48AB5DABCA65232CE7F30576342A3B9B" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B11" refString="Fei, L, Hu, SQ, Ye, CY, Huang, YZ, 2009. Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." title="Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." year="2009">2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E08522C3CBC29F8724FEFC6FB249CE1D" author="Fei, L" journalOrPublisher="Herpetologica Sinica" pageId="0" pageNumber="495" pagination="1 - 43" refId="B13" refString="Fei, L, Ye, CY, Jiang, JP, 2010. Phylogenetic systematics of Ranidae. Herpetologica Sinica 12: 1 - 43" title="Phylogenetic systematics of Ranidae." volume="12" year="2010">2010</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="DAE3F9D277996EBE112D7F8E5A3032CC" author="Fei, L" journalOrPublisher="Herpetologica Sinica" pageId="0" pageNumber="495" refId="B14" refString="Fei, L, Ye, CY, Jiang, JP, 2012. Colored Atlas of Chinese Amphibians and Their Distributions. Sichuan Publishing House of Science & Technology, Chengdu, China." title="Colored Atlas of Chinese Amphibians and Their Distributions. Sichuan Publishing House of Science & Technology, Chengdu, China." year="2012">2012</bibRefCitation>
|
||
; Zhao & Adler 1993;
|
||
<bibRefCitation id="294A0CD450B657DF69B5A17D5C7EE64B" author="Li, ZC" journalOrPublisher="Algemeene Konst-en Letter-Bode" pageId="0" pageNumber="495" refId="B29" refString="Li, ZC, Xiao, Z, Liu, SR, 2011. Amphibians and reptiles of Guangdong. Guangdong Science and Technology Press, Guangzhou, China." title="Amphibians and reptiles of Guangdong. Guangdong Science and Technology Press, Guangzhou, China." year="2011">Li et al. 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8A41C801B36B98C62787CC8A39CA6899" author="Mo, YM" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="495" refId="B34" refString="Mo, YM, Wei, ZY, Chen, WC, 2014. Colored Atlas of Guangxi Amphibians. Guangxi Science and Technology Publishing House, Nanning, China." title="Colored Atlas of Guangxi Amphibians. Guangxi Science and Technology Publishing House, Nanning, China." year="2014">Mo et al. 2014</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8AAE9518F170E8F456E8EB121FDAC320" pageId="0" pageNumber="495" type="type material">
|
||
<paragraph id="9D6D9E5A28AECFA7DCDDD9D66988F550" pageId="0" pageNumber="495">Type materials.</paragraph>
|
||
<paragraph id="38EBFADEBF8E59741458C5630309AC6E" pageId="0" pageNumber="495">
|
||
<emphasis id="60CC7F047192D4C7C74540FF3882BAC7" bold="true" pageId="0" pageNumber="495">Holotype.</emphasis>
|
||
BMNH 1932.5.1.2, adult female, collected from China (discussion for the exact locality see Remarks below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FFD92D3D9F56FFE6A9D77AF28129796C" pageId="0" pageNumber="495" type="materials_examined">
|
||
<paragraph id="FCB4EC632543FE7728B8EF082D509F23" pageId="0" pageNumber="495">Specimens examined.</paragraph>
|
||
<paragraph id="021EBA59548E7FA30D035D917A251E39" pageId="0" pageNumber="495">
|
||
<materialsCitation id="F0A86B45F980D2E640742527C6A7B170" accessionNumber="CIB 44475" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="China" location="Guangzhou" specimenCount="three" specimenCount-adult="1" stateProvince="Guangdong Province">
|
||
Seven adult males and
|
||
<specimenCount id="F9C84BC2B00B6A8070F75D84F1E539BD" type="adult">three adult</specimenCount>
|
||
females. Males
|
||
<accessionNumber id="0FDEED0B41ADC0E48E52BC5D963A808B" httpUri="http://www.ncbi.nlm.nih.gov/protein/CIB44475">CIB 44475</accessionNumber>
|
||
-44476, from
|
||
<location id="8F11EFBBA2759F17A70B268991FB87DF" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:8F11EFBBA2759F17A70B268991FB87DF" country="China" name="Guangzhou" stateProvince="Guangdong Province">Guangzhou</location>
|
||
,
|
||
<collectingRegion id="5B3AA37F38D20B6EB98D24456EB31C94" country="China" name="Guangdong">Guangdong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="FB8AA008B42716D39B2CB454F76217AE" name="China">China</collectingCountry>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="7DD8C0925121C5A82A46CA04D7443F1B" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="China" county="Renhua County" latitude="24.3818" location="Mt Danxia" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" specimenCount="1" stateProvince="Guangdong">
|
||
male SYS a000534, from
|
||
<location id="C710CAD937AA1447BCD82365A90503F4" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:C710CAD937AA1447BCD82365A90503F4" country="China" county="Renhua County" latitude="24.3818" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" name="Mt Danxia" stateProvince="Guangdong">Mt Danxia</location>
|
||
(
|
||
<geoCoordinate id="DBC98870B48B3F2CEF8803BD6216139F" degrees="25.0347" direction="north" orientation="latitude" precision="5" value="25.0347">25.0347 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AC19689C5924E73F9EBF39AAAA3BBB8A" degrees="113.7407" direction="east" orientation="longitude" precision="5" value="113.7407">113.7407 E</geoCoordinate>
|
||
),
|
||
<collectingCounty id="BAEEF23A921EB78F2EE12498DF961D15">Renhua County</collectingCounty>
|
||
,
|
||
<collectingRegion id="A1E9882F4E6BD1E15E7864A5615ED5A0" country="China" name="Guangdong">Guangdong</collectingRegion>
|
||
; males SYS a008120-8121, 8123, 8125, and females SYS a008122, 8124, from
|
||
<location id="765FE727FBB9D8F6A3F00D89A1EE94DB" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:765FE727FBB9D8F6A3F00D89A1EE94DB" country="China" county="Renhua County" latitude="24.3818" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" name="Shimentai Nature Reserve" stateProvince="Guangdong">Shimentai Nature Reserve</location>
|
||
(
|
||
<geoCoordinate id="6FF5BD243C4FA19538A0C1D82DE0BF1A" degrees="24.3818" direction="north" orientation="latitude" precision="5" value="24.3818">24.3818 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ECE65AE221BB6FB83682C032D52ADA6B" degrees="113.3927" direction="east" orientation="longitude" precision="5" value="113.3927">113.3927 E</geoCoordinate>
|
||
),
|
||
<location id="C2428DFE6A306EE3F541F84AAEB83210" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:C2428DFE6A306EE3F541F84AAEB83210" country="China" county="Renhua County" latitude="24.3818" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" name="Yingde City" stateProvince="Guangdong">Yingde City</location>
|
||
,
|
||
<collectingRegion id="6A99891163C24DD7265A046B383D464C" country="China" name="Guangdong">Guangdong</collectingRegion>
|
||
; female SYS a000488, from
|
||
<location id="264DB1EB3B830D52AF92021BC52BFBF7" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:264DB1EB3B830D52AF92021BC52BFBF7" country="China" county="Renhua County" latitude="24.3818" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" name="Mt Luoyang" stateProvince="Guangdong">Mt Luoyang</location>
|
||
,
|
||
<location id="27C935F424EE131E688EA6BD641EF273" LSID="urn:lsid:plazi:treatment:BFD1E4E6F78259D29A9F27662F224A79:27C935F424EE131E688EA6BD641EF273" country="China" county="Renhua County" latitude="24.3818" longLatPrecision="7" longitude="113.3927" municipality="Autonomous Region" name="Lingshan County" stateProvince="Guangdong">Lingshan County</location>
|
||
,
|
||
<collectingRegion id="0D84DE7A4955CC437D21B62FA7F9C109" country="China" name="Guangxi">Guangxi Zhuang</collectingRegion>
|
||
<collectingMunicipality id="DBBF7E566B7205604C2CBE9BA4C3D5D9">Autonomous Region</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="56786B1D3618DE4FA7BB9623998ABE02" name="China">China</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="34778BDC66B54DC4A9C5D46F762C7B51" pageId="0" pageNumber="495" type="etymology">
|
||
<paragraph id="F7ABEC2B6326C1280D83548BA218DABE" pageId="0" pageNumber="495">Etymology.</paragraph>
|
||
<paragraph id="80A5A2CFA702EFD9012AC3924FD249E2" pageId="0" pageNumber="495">
|
||
The specific name
|
||
<emphasis id="ADA5159F8D46780A0122A7673743EBA8" italics="true" pageId="0" pageNumber="495">obscura</emphasis>
|
||
means
|
||
<normalizedToken id="3F54BAB9DDC97E65B337DF533C829C77" originalValue="‘obscure’">'obscure'</normalizedToken>
|
||
in Latin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE6C7AF2D4C95C605E40D2E4A489EA54" pageId="0" pageNumber="495" type="common name">
|
||
<paragraph id="F22BC1A8B9F70F12D5B9E6F128030337" pageId="0" pageNumber="495">Common name.</paragraph>
|
||
<paragraph id="0110F11716368A591602E4702373C7B5" pageId="0" pageNumber="495">
|
||
"Chinese floating frog" in English / "中国浮蛙 (zhōng
|
||
<normalizedToken id="69053886B704DFB5E198D5F654C85948" originalValue="guó">guo</normalizedToken>
|
||
<normalizedToken id="4364B69C2850C0A349F41D36823F48CD" originalValue="fú">fu</normalizedToken>
|
||
<normalizedToken id="AE144E06120FF3195791A9EC10A72443" originalValue="wā)”">wa)"</normalizedToken>
|
||
in Chinese.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F0BABBF98B0937F3620BE1A0499085D0" pageId="0" pageNumber="495" type="diagnosis">
|
||
<paragraph id="29088AABAF3A34858E7BD306D95409DC" pageId="0" pageNumber="495">Diagnosis.</paragraph>
|
||
<paragraph id="1D0DB269ECA96F29A6BF9D045ADB5212" pageId="0" pageNumber="495">(1) Body stocky, size small, SVL 24.2-27.5 mm in adult males (n=7) and 31.5-32.2 mm in adult females (n=3); (2) snout short triangular shaped; (3) nostrils located dorsally; (4) eye orientation vertically; (5) loreal region oblique, not concave or convex; (6) interorbital space narrower than internarial distance; (7) tongue narrow and slender, unnotched, pointed distally, lingual papilla absent; (8) vomerine ridge and vomerine teeth absent; (9) supratympanic fold distinct, raised, and granulated, curved on the temporal region; (10) tympanum hidden, edge invisible; (11) fingers with rudimentary webs, toes with full webs; (12) heels not meeting, tibio-tarsal articulation reaching between the posterior and anterior of the eye.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D234E8453D874071A22750A4E7A4DC18" pageId="0" pageNumber="495" type="comparisons">
|
||
<paragraph id="0DBF4010401A57A2B3568B5394846BC4" pageId="0" pageNumber="495">Comparisons.</paragraph>
|
||
<paragraph id="B73A3080284F47FFE6C06E7267A7099D" pageId="0" pageNumber="495">
|
||
<taxonomicName id="5AF93032BCC7CA5778665B528288C49B" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="8C962729EEBA935CF707891661577EF6" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. was previously synonymized with
|
||
<taxonomicName id="7F49A6F2020DD8723D338FD108DD6C5C" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="FCE4999C7F506E26DF7C4B0D50E46FD3" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
. These two species are most similar to each other. However,
|
||
<taxonomicName id="465076EC0ECB2802EF13CF6C822DE155" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="39ECA94BE60E43FDC1A858D6A596B6E6" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. can be distinguished by the combination of the following characteristics: loreal region oblique, not concave or convex (vs. slightly concave in
|
||
<taxonomicName id="C9C041F32DD834D047F8D417ADFF36A6" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="B085A56F80E3B9B9A6D20BE49B1ADDC7" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
), interorbital space narrower than internarial distance (vs. subequal), supratympanic fold distinct, raised, and granulated (vs. indistinct), inner metatarsal tubercle large and raised (vs. weakly projecting), and tibio-tarsal articulation reaching between the posterior and anterior of the eye (vs. reaching tip of nostril).
|
||
</paragraph>
|
||
<paragraph id="A85F0BC9A2DE5FFED821F3A6E6A9FD74" pageId="0" pageNumber="495">
|
||
<taxonomicName id="0F3CCB9B722F74D30247F44D4FC780F4" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="EE11E1631A8907D277A598294A0A53DA" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. furthers differs from
|
||
<taxonomicName id="25CDF31EFDCEEA4176C6ABCB239F21CE" lsidName="O. berbeza" pageId="0" pageNumber="495" rank="species" species="berbeza">
|
||
<emphasis id="63C43B3618C1A2D6B711231A59683138" italics="true" pageId="0" pageNumber="495">O. berbeza</emphasis>
|
||
</taxonomicName>
|
||
by the supratympanic fold curved on the temporal region (vs. straight), fingers with rudimentary webs (vs. unwebbed), and outer metatarsal tubercle present (vs. absent).
|
||
<taxonomicName id="898E4781714495C99BFB5A2A155BAD50" lsidName="O. obscura" pageId="0" pageNumber="495" rank="species" species="obscura">
|
||
<emphasis id="71A6AC257E02893034C83B38ECC50106" italics="true" pageId="0" pageNumber="495">O. obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. distinctly differs from
|
||
<taxonomicName id="DBFBC08918B46EEFDA9DB881975C2E51" lsidName="O. rhacoda" pageId="0" pageNumber="495" rank="species" species="rhacoda">
|
||
<emphasis id="080F86BA2B7FE55FDA4D7C65EB93C5DA" italics="true" pageId="0" pageNumber="495">O. rhacoda</emphasis>
|
||
</taxonomicName>
|
||
by the dorsolateral fold absent (vs. present), and fingers with rudimentary webs (vs. unwebbed).
|
||
</paragraph>
|
||
<paragraph id="9D262771792ECBBB6A7C53CE5E3DD5DD" pageId="0" pageNumber="495">
|
||
<taxonomicName id="D8936A970D36A80845C4131D4E5622AF" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="71D9FF36126CBF3F313A06EE92B0AC1B" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. can be easily distinguished from the remaining 13 congeners
|
||
<taxonomicName id="9D781F545CAE94C067D3D2B1B4DAD76B" lsidName="O. baluensis" pageId="0" pageNumber="495" rank="species" species="baluensis">
|
||
<emphasis id="1EB4C020DB408D3BF5DE0A88268419F3" italics="true" pageId="0" pageNumber="495">O. baluensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1A80E2C5C3A5B0350A9254F4A4EECC18" lsidName="O. celebensis" pageId="0" pageNumber="495" rank="species" species="celebensis">
|
||
<emphasis id="E96682D26BD7C44110B390600894D3A4" italics="true" pageId="0" pageNumber="495">O. celebensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C1A018A8175CD7DB78C9DB265C195A06" lsidName="O. diminutiva" pageId="0" pageNumber="495" rank="species" species="diminutiva">
|
||
<emphasis id="2995E62166FC99792FA0F574069E2705" italics="true" pageId="0" pageNumber="495">O. diminutiva</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="092124FCE4082DD814B05E7BAB1A39C7" lsidName="O. floresiana" pageId="0" pageNumber="495" rank="species" species="floresiana">
|
||
<emphasis id="8E363989D8F4ADBA72F033B128D66A88" italics="true" pageId="0" pageNumber="495">O. floresiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="896A63A03565AB314FD72751759CCDFC" lsidName="O. laevis" pageId="0" pageNumber="495" rank="species" species="laevis">
|
||
<emphasis id="8D0FC38407CE6C7812F1F3E5F55520C3" italics="true" pageId="0" pageNumber="495">O. laevis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="23BDF4EC10470EF63C01896BF2E69B1E" lsidName="O. magnapustulosa" pageId="0" pageNumber="495" rank="species" species="magnapustulosa">
|
||
<emphasis id="BB3C3CE6D4B427AD7E9724C0548DA8F5" italics="true" pageId="0" pageNumber="495">O. magnapustulosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B1F8DF481D2DAA64350B452D2F906C47" lsidName="O. martensii" pageId="0" pageNumber="495" rank="species" species="martensii">
|
||
<emphasis id="8EAB01392E4070C2731C96C996B9FC99" italics="true" pageId="0" pageNumber="495">O. martensii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF5B05C903D38A1923CC0DD1502FA4FC" lsidName="O. myanhessei" pageId="0" pageNumber="495" rank="species" species="myanhessei">
|
||
<emphasis id="08A4741013C82E7B8CA61D5B1A52F9E7" italics="true" pageId="0" pageNumber="495">O. myanhessei</emphasis>
|
||
</taxonomicName>
|
||
comb. nov.,
|
||
<taxonomicName id="CA4773CAF7C37C36D9F7B4A8150D3FF7" lsidName="O. semipalmata" pageId="0" pageNumber="495" rank="species" species="semipalmata">
|
||
<emphasis id="C9481B6AA9B9EFDDCFE5B49C137DBB63" italics="true" pageId="0" pageNumber="495">O. semipalmata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8F017D86F5E6738F06FB794E1ED9FAF3" lsidName="O. sumatrana" pageId="0" pageNumber="495" rank="species" species="sumatrana">
|
||
<emphasis id="0C11EF06DEE0FC1EACB72C7AF4827C68" italics="true" pageId="0" pageNumber="495">O. sumatrana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DB7335B4331AC7CF45EBFEE874592F74" lsidName="O. shiwandashanensis" pageId="0" pageNumber="495" rank="species" species="shiwandashanensis">
|
||
<emphasis id="0D595CBC22C29F8AF9C9D8A5F73A8AAD" italics="true" pageId="0" pageNumber="495">O. shiwandashanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6BF1B81B3BC0D2A7C458665AC91AD890" lsidName="O. swanbornorum" pageId="0" pageNumber="495" rank="species" species="swanbornorum">
|
||
<emphasis id="58D981DDACED356E693CC5864781C4D4" italics="true" pageId="0" pageNumber="495">O. swanbornorum</emphasis>
|
||
</taxonomicName>
|
||
comb. nov., and
|
||
<taxonomicName id="50F6C933445169F7023C0D85052545BC" lsidName="O. tompotika" pageId="0" pageNumber="495" rank="species" species="tompotika">
|
||
<emphasis id="812B198AD5456F04F643F23C4EAB25F1" italics="true" pageId="0" pageNumber="495">O. tompotika</emphasis>
|
||
</taxonomicName>
|
||
, by the tongue narrow and slender (vs. wide and swollen in all of these 13 species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C04CA009FD5F575BCBC50F939FF4DE14" pageId="0" pageNumber="495" type="description">
|
||
<paragraph id="5DED4ABC2AECB6F62AD273818DB0FA93" pageId="0" pageNumber="495">Re-description.</paragraph>
|
||
<paragraph id="7EEF9339EF69797E8847E4E1122E4034" pageId="0" pageNumber="495">
|
||
Based on the examined specimens (n=10). All specimens were similar in morphology. The measurements are given in Table
|
||
<tableCitation id="30596BC89C2D0141A66C10EB4FFDB54D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements (in mm) of the examined specimens of Occidozyga obscura comb. nov." httpUri="http://table.plazi.org/id/C547C9E22A55445AFAD5613EC2764DE1" pageId="0" pageNumber="495" tableUuid="C547C9E22A55445AFAD5613EC2764DE1">3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="011F2B298ACC94432404440205F277D7" pageId="0" pageNumber="495">Body stocky, small-sized, SVL 24.2-27.5 mm (n=7) in males and 31.5-32.2 mm (n=3) in females. Head flat above, almost as wide as long (HDW/HDL 0.97-1.09, n=10); pineal ocellus absent; snout short triangular shaped, distinctly protruding beyond lower jaw, tip of snout rounded in dorsal view and profile; canthus rostralis absent, loreal region oblique, not concave or convex; nostril rounded, located dorsally, distinctly raised, closer to the tip of snout than to the eye; eye orientation vertically, pupil diamond shaped; interorbital space narrower than internarial distance; tympanum hidden, edge invisible; vomerine ridge and teeth absent; tongue narrow and slender, unnotched, pointed distally, lingual papilla absent.</paragraph>
|
||
<caption id="C547C9E22A55445AFAD5613EC2764DE1" ID-Table-UUID="C547C9E22A55445AFAD5613EC2764DE1" httpUri="http://table.plazi.org/id/C547C9E22A55445AFAD5613EC2764DE1" pageId="0" pageNumber="495" start="Table 3" startId="T3">
|
||
<paragraph id="20EA0558A9EDEA5D8680C1D1CECE3BA5" pageId="0" pageNumber="495">
|
||
<emphasis id="E04E63BBE3D85C8046148BBB7B99CAEE" bold="true" pageId="0" pageNumber="495">Table 3.</emphasis>
|
||
Measurements (in mm) of the examined specimens of
|
||
<taxonomicName id="E37447243B7C4D9285B77AA48626C838" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="D1EE7794726A6433A13974A131068AE3" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BFECAB2AD4BED9399F3A932786D27B7A" pageId="0" pageNumber="495">
|
||
<table id="3445D7532E1E30D0E4D2CC0E4E9F411C" pageId="0" pageNumber="495">
|
||
<tr id="236AF91CE19FC02CA67CD1E421CB427A" pageId="0" pageNumber="495">
|
||
<td id="9BB94C602B3ED0DDFE1A70E192A6C707" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="7A06BEC96A49365568FA083CED15BC6F" bold="true" pageId="0" pageNumber="495">Specimen</emphasis>
|
||
</td>
|
||
<td id="24EAD3E7E8840381D1C69DF8DFF8682F" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="E8DAB5B8344C8D1AE5092EEBB78736D4" bold="true" pageId="0" pageNumber="495">CIB 44475</emphasis>
|
||
</td>
|
||
<td id="26E0EE9486A72913E5CC96AB96F55DA1" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="0F1B65B568F87A85B60BC2B6DE0C0667" bold="true" pageId="0" pageNumber="495">CIB 44476</emphasis>
|
||
</td>
|
||
<td id="FFC674F356075335B5F797245D6F373E" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="37D4679A321E6F32D719B842E0715745" bold="true" pageId="0" pageNumber="495">SYS a000534</emphasis>
|
||
</td>
|
||
<td id="AA5E136C5E4EB21E02C4A1EDBFC68899" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="665E8629DB37191CBD701294A4610F1F" bold="true" pageId="0" pageNumber="495">SYS a008120</emphasis>
|
||
</td>
|
||
<td id="347FF33867DB4786C2E2D51D5E4DA2E2" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="C3092650C9516FD93147622FB766DC17" bold="true" pageId="0" pageNumber="495">SYS a008121</emphasis>
|
||
</td>
|
||
<td id="F620C12A70715D7CBB9BA6967628DE03" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="B8E273B382BB8F51460A79B958F90560" bold="true" pageId="0" pageNumber="495">SYS a008123</emphasis>
|
||
</td>
|
||
<td id="FA1B2F3DE422A637411EF460AEBDA5F4" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="55A10E923F76E5E512F2E5415585D283" bold="true" pageId="0" pageNumber="495">SYS a008125</emphasis>
|
||
</td>
|
||
<td id="49864E0C338B29D3DFAF1610549FC140" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="BCB637FDBB707B070D0EF10EFB7A9E68" bold="true" pageId="0" pageNumber="495">SYS a008122</emphasis>
|
||
</td>
|
||
<td id="0AB65B0CA0446709EEB85C476AFD85C0" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="BE82B95F437B534C23A14D09B361EB02" bold="true" pageId="0" pageNumber="495">SYS a008124</emphasis>
|
||
</td>
|
||
<td id="A4E5E0A458A54BD74EC5AB413C9CF088" colspan="1" pageId="0" pageNumber="495" rowspan="1">
|
||
<emphasis id="E4A90460360D36F0FFED99BA03F40ABA" bold="true" pageId="0" pageNumber="495">SYS a000488</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="598045143BC0EB1FD3571D80554DD60B" pageId="0" pageNumber="495">
|
||
<td id="8493481470C45FCA09626BD50B38348F" colspan="1" pageId="0" pageNumber="495" rowspan="1">Sex</td>
|
||
<td id="040C2531DEA8143619ABDB800CEC9F72" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="A908788363A7D3BB0F2E9360B59C17E1" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="F1C317FAABFC316CCD5A958A0CE14A24" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="E3F98586C338708CB831B3693C2044FA" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="01C1B7F936AF12839BDE4C9279289E84" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="59623DF3123FEE3E1A577754B278FF40" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="7EE40F7C191B881D57FF572EA5D6D2E6" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
|
||
<td id="945D9C1F7C595CD91A55F2B8AE3B4E2A" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
|
||
<td id="6446C4A3F958F51D0E24CFA1822C0FB5" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
|
||
<td id="F74A78BABA57E29F74CBF8D7501161D8" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
|
||
</tr>
|
||
<tr id="FD62A2438BCE3AFE0A82D7F8222389C9" pageId="0" pageNumber="495">
|
||
<td id="D82DE0F5F950BFC091C678A137E9A06D" colspan="1" pageId="0" pageNumber="495" rowspan="1">SVL</td>
|
||
<td id="12DA5A94AD19A61B1237613B0F8527FD" colspan="1" pageId="0" pageNumber="495" rowspan="1">24.2</td>
|
||
<td id="9A1A8D01FD86DC7D83AC2A532D2BF114" colspan="1" pageId="0" pageNumber="495" rowspan="1">24.7</td>
|
||
<td id="14DDA0C2EC23BEEC504062B24B87E8E8" colspan="1" pageId="0" pageNumber="495" rowspan="1">27.5</td>
|
||
<td id="54DD33E5FA0366BDD28A1BA683FA19BE" colspan="1" pageId="0" pageNumber="495" rowspan="1">24.3</td>
|
||
<td id="F43CF6451D8E2538B75FAE4429E47DDC" colspan="1" pageId="0" pageNumber="495" rowspan="1">26.2</td>
|
||
<td id="DBB2B9A1D604441CF4C1E793886F4491" colspan="1" pageId="0" pageNumber="495" rowspan="1">27.0</td>
|
||
<td id="9B15C910D159DA38D9D8CEDF6E81D00B" colspan="1" pageId="0" pageNumber="495" rowspan="1">24.8</td>
|
||
<td id="342D74ADD9235D67961EA6AA78DF0210" colspan="1" pageId="0" pageNumber="495" rowspan="1">32.2</td>
|
||
<td id="362DBF9DC3CE9EB6781B9DFB569164E7" colspan="1" pageId="0" pageNumber="495" rowspan="1">32.0</td>
|
||
<td id="CE3E07C4F74308900E53E4FDD8215CED" colspan="1" pageId="0" pageNumber="495" rowspan="1">31.5</td>
|
||
</tr>
|
||
<tr id="BFEA4FBFA2B34A14474D7D31763FD6B3" pageId="0" pageNumber="495">
|
||
<td id="0E638D0835441616A7D22AA420DDD49A" colspan="1" pageId="0" pageNumber="495" rowspan="1">HDL</td>
|
||
<td id="7DEFFD50BFBC3F52B0BC60E893DCD64C" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.1</td>
|
||
<td id="FAF30F79DF7A46E19811E3CC22D5538A" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.4</td>
|
||
<td id="6262DDB6894864FE8B2B190AE9906B22" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.9</td>
|
||
<td id="1D968FC28AE82F90CA2DB561933F761F" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.8</td>
|
||
<td id="81CD3D34E25A45240F0E84DF76906665" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.4</td>
|
||
<td id="E9D811541A3DCE3DDCF38B4B991AB75C" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.4</td>
|
||
<td id="0EAB8311B2D83A2786A49EEE48D63E68" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.8</td>
|
||
<td id="39EA6CDB3F9857B4F54FED395E885838" colspan="1" pageId="0" pageNumber="495" rowspan="1">10.4</td>
|
||
<td id="8B97B19B4917CD927AF58DF7E861C739" colspan="1" pageId="0" pageNumber="495" rowspan="1">10.4</td>
|
||
<td id="B27FE504340C011E461C670DF03216B5" colspan="1" pageId="0" pageNumber="495" rowspan="1">10.1</td>
|
||
</tr>
|
||
<tr id="25191EF2659BADE05EB628034CF77D69" pageId="0" pageNumber="495">
|
||
<td id="1ED4BA989DEFCB206E5611CBF3A5099F" colspan="1" pageId="0" pageNumber="495" rowspan="1">HDW</td>
|
||
<td id="642543B00A28E033F81094820767580B" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.8</td>
|
||
<td id="15D13CEDDEE593710EB88A2AFDC13608" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.6</td>
|
||
<td id="AA2CBC62651213020A4CFA87875356E9" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.7</td>
|
||
<td id="795C13A442B3F1DE37B9E1B4921EB376" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.6</td>
|
||
<td id="27DEDAD171D236BB005E680BA68441E4" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.4</td>
|
||
<td id="A0C6BEC5C7D7A5C690E65777148073BD" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.3</td>
|
||
<td id="3F8E642827DEC158995E5F4FE2B24E96" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.6</td>
|
||
<td id="3F608A257AE1990D5AB0E54E35480398" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.0</td>
|
||
<td id="843458E550C441E8D7DBB74CEEE1A11B" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.4</td>
|
||
<td id="5830712A9D93BD78AB90EEBFA1EED8F0" colspan="1" pageId="0" pageNumber="495" rowspan="1">10.0</td>
|
||
</tr>
|
||
<tr id="C61AE3A45E8D290DB516BC3A8537DD14" pageId="0" pageNumber="495">
|
||
<td id="CFD3AFDA12C2498B89959E67027A03DA" colspan="1" pageId="0" pageNumber="495" rowspan="1">SNT</td>
|
||
<td id="DE29CBBBDB61EB6A5926BD2AA2593BE0" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.7</td>
|
||
<td id="6D77FE41C7FF19D2C155BF008D045745" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.1</td>
|
||
<td id="28DA08A94B70E30ED8F273B29351DD02" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.3</td>
|
||
<td id="5C02B33015FE3BA09B441CF9B83B5E5D" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.8</td>
|
||
<td id="E1D0535F9A1A94C9BB21F2B07EEE45C5" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.3</td>
|
||
<td id="7C5B4B4950C8E642D069BDA2D5E6B3A9" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.1</td>
|
||
<td id="D10CD2556B683E7CC4E1092153B24509" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.2</td>
|
||
<td id="19D5116200AA67A85F7F78EA9E768ED2" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.2</td>
|
||
<td id="FAD03D78EAB9D357CF954608433BB04C" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.0</td>
|
||
<td id="D1BDE53D7CF1C97C2373C2B4F6929E25" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.7</td>
|
||
</tr>
|
||
<tr id="432016EDEFBECB11A86B0A6D3439AEE4" pageId="0" pageNumber="495">
|
||
<td id="BF105D90E15F33B94C4DB0BF77AC80C8" colspan="1" pageId="0" pageNumber="495" rowspan="1">IND</td>
|
||
<td id="35E6D5A192B769123AD15672D3E2886B" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="3B56C268976C51A61B758A22849876A1" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.3</td>
|
||
<td id="64BE1EB788F1608E86D03E732FEB2B67" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
|
||
<td id="C5989CD7EC483E5284626D5BF3271AF3" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
<td id="0B646AC2E28DFA96244E35440206E11A" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
<td id="E083D9016CA3BD8192AAD9F393C83EAF" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="EBC5B3D502882320542C987E15E2E390" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.2</td>
|
||
<td id="F8AFB770FF35C606B6792AB720293778" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
|
||
<td id="92B8096AEA0701FEE0522BDF72B4CADC" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="A4020AC6F710A29858684D8259E3385A" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
|
||
</tr>
|
||
<tr id="7651271B2FE704CD4B9670C4313E068B" pageId="0" pageNumber="495">
|
||
<td id="CB3E7EC396B02E1E55BC459D62208938" colspan="1" pageId="0" pageNumber="495" rowspan="1">IOD</td>
|
||
<td id="9EE96B2B3D937ED6128F06AD275CD849" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="8EB3E595BCD0F063460645B59350B1EA" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.2</td>
|
||
<td id="CBB8C9EDAA1C43E1FE4581DFD21984BD" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="F711C889B40A204CAB266E6065D9EE16" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.3</td>
|
||
<td id="441E9AF2CC97E301C9C865162740AF97" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
<td id="C35FB9055E2FD0756CB87E3A28CE7EB2" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
<td id="C68F5D17DEB866509BAE2B47377917D1" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.1</td>
|
||
<td id="E51A1EE03470CB4F3FE0009797AEAE45" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
|
||
<td id="461CCE6048BFBCA2146F8E7EC9E42927" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
<td id="7B748102D53FE3A6DC77C7C396229CD6" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
|
||
</tr>
|
||
<tr id="67ABF4A8EF9A3228F2AC78CA5A3F17BF" pageId="0" pageNumber="495">
|
||
<td id="5EB6856718FEBEA120A7BFDA9ABF2890" colspan="1" pageId="0" pageNumber="495" rowspan="1">ED</td>
|
||
<td id="EFC9F4289E539D0A8E4E4C3E22867B0B" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.2</td>
|
||
<td id="8CCC2567E6695C3A83DE673991CF881A" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.8</td>
|
||
<td id="9226D0CCE3C422F6AB9C6F0A191F9CD7" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.7</td>
|
||
<td id="07ECD35B3FCEE31F7A2F7977411C6BA4" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.7</td>
|
||
<td id="D3AF7765FD87BC9F6721489F0C88F193" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.8</td>
|
||
<td id="F86B8E9441FE7511A91099FE75C2B9A8" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.9</td>
|
||
<td id="F059ABA5A930DC343B831CF3458A6ECA" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.6</td>
|
||
<td id="22AF22A90E0C10B10E800B2F84FD3C5C" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.1</td>
|
||
<td id="F57EB7F6F4CADBC6623C72D058EBB4D5" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.0</td>
|
||
<td id="02450F0FD6C9C5C97B95BF22E908D4DC" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.2</td>
|
||
</tr>
|
||
<tr id="81EAFC25679B18AD1030F23911A840B7" pageId="0" pageNumber="495">
|
||
<td id="1B22D4118B08C0F9035F4EF002F10908" colspan="1" pageId="0" pageNumber="495" rowspan="1">HND</td>
|
||
<td id="DC8BB7D91C20E0CF4F2FE7E69CB71B39" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.7</td>
|
||
<td id="51FCA217D5A2F3EE3F42A59D4A7E010E" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.5</td>
|
||
<td id="52C2ADE9A45D96ED7559DACF5F44BA31" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.8</td>
|
||
<td id="5CD8A83B6C465EF989A86B8C751049B8" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.1</td>
|
||
<td id="0F577D80D169B2DB7B63E0D020770901" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.3</td>
|
||
<td id="9CA67F42E601412590187F8C5C86F05B" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.4</td>
|
||
<td id="DB17CE14B73B82AAB2C39F5F6FC0CD4C" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.6</td>
|
||
<td id="F1176DD39293B3BEDDB70A78AF2ECD88" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.2</td>
|
||
<td id="780EAB3385BFD1CC9CC0255670A3DCD6" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.8</td>
|
||
<td id="578A83160B8A764D3F25FA69B1B3136E" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.5</td>
|
||
</tr>
|
||
<tr id="886CAD261033DB2AE3620ACC750AD9A5" pageId="0" pageNumber="495">
|
||
<td id="661CC49F7816F00920C6F2829C0F74E8" colspan="1" pageId="0" pageNumber="495" rowspan="1">RAD</td>
|
||
<td id="3208CB621BBE2B2AB54E840F0F18694D" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.0</td>
|
||
<td id="F3E7ECA5BCAAABB64868FC465A837A4D" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.5</td>
|
||
<td id="EF8FF5D7CE5556B1057A54A283871C6F" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.6</td>
|
||
<td id="7F58759F0F5FD2FE8B51878DBB94D8E6" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.1</td>
|
||
<td id="404DD205D6DCAC0F6FD5CF5EC5847B76" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.4</td>
|
||
<td id="23B3F023AA1D6E3F61B186055409E222" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.0</td>
|
||
<td id="27A338F3279142EAC3C15099650EED79" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.9</td>
|
||
<td id="2EB6E024937F6DC67876AF4AA1F15F79" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.9</td>
|
||
<td id="AD1852F25BB710117582DF6B18DE6E63" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.1</td>
|
||
<td id="A7609E81B788C187A2853A9337A67004" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.9</td>
|
||
</tr>
|
||
<tr id="CE196E332189E2507783E306CCD1493B" pageId="0" pageNumber="495">
|
||
<td id="56DDFBE8C9120440CA599AF1F7DD3CC9" colspan="1" pageId="0" pageNumber="495" rowspan="1">FTL</td>
|
||
<td id="9254ED39416B6D2B3C88191C5DD44BB3" colspan="1" pageId="0" pageNumber="495" rowspan="1">19.3</td>
|
||
<td id="2498A20BA5087F17C5000B24ABB598BF" colspan="1" pageId="0" pageNumber="495" rowspan="1">18.4</td>
|
||
<td id="33BAEA151B66C698E3C13A5F6C5727DE" colspan="1" pageId="0" pageNumber="495" rowspan="1">18.4</td>
|
||
<td id="D66E702C305B8AA72785F5FF7BC6E17D" colspan="1" pageId="0" pageNumber="495" rowspan="1">18.0</td>
|
||
<td id="8B256B7DB3E2FCA7CFC1FA11EE40F91D" colspan="1" pageId="0" pageNumber="495" rowspan="1">18.8</td>
|
||
<td id="86064FFDE9C592877FB4886A1541B5A5" colspan="1" pageId="0" pageNumber="495" rowspan="1">18.8</td>
|
||
<td id="32A348B7A8A7A81E29564DAD69672147" colspan="1" pageId="0" pageNumber="495" rowspan="1">17.5</td>
|
||
<td id="5C4D3FD0A84B403D914DF9EC84025600" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.3</td>
|
||
<td id="0E641951A724A10C0F4F299CFF94F2F8" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.5</td>
|
||
<td id="64148F203F8392AD788F58D20AB6C650" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.6</td>
|
||
</tr>
|
||
<tr id="2D4ACDB53CF43A4C74172008B4FFDE22" pageId="0" pageNumber="495">
|
||
<td id="84DA8A6C029DE6F6D331E6E581FE02CF" colspan="1" pageId="0" pageNumber="495" rowspan="1">TIB</td>
|
||
<td id="3224CD8D1BADA901EA438EBB9A5613C7" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.3</td>
|
||
<td id="F6AE13635F3537C650BA53B90793DF6F" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.4</td>
|
||
<td id="B681B8FC28F07D02E0F64914C345D5EB" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.2</td>
|
||
<td id="C03DE26B08E118214B3E618A067565C1" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.1</td>
|
||
<td id="583429ABF3BD4F3C1138EB143288E8BF" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.7</td>
|
||
<td id="10036E756E8D2E9E497DC1AEE11AC694" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.5</td>
|
||
<td id="627410AA680C0B061D2342782477D9F6" colspan="1" pageId="0" pageNumber="495" rowspan="1">10.9</td>
|
||
<td id="819FB827729136128B6E35A2C6A03451" colspan="1" pageId="0" pageNumber="495" rowspan="1">13.4</td>
|
||
<td id="43A08E22D9A9776CA1FD7AED611F1357" colspan="1" pageId="0" pageNumber="495" rowspan="1">13.6</td>
|
||
<td id="A049755E9E92C8BA022041A37C063AF8" colspan="1" pageId="0" pageNumber="495" rowspan="1">12.8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="40AE5FE3BF55E56A3E30FA587EDDED7D" pageId="0" pageNumber="495">Forelimbs short, lower arm 13-18% (n=10) of SVL and hand 26-32% (n=10) of SVL; fingers distinctly thin and long, relative finger lengths I<II<IV<III; tips of fingers pointed, not dilated, and without disks; distinct lateral fringes on inner and outer sides of each finger, fingers with rudimentary webs, more distinct between fingers I and II; subarticular tubercles present at the bases of each finger, prominent and rounded; supernumerary tubercles absent; inner and outer palmar tubercles prominent and rounded.</paragraph>
|
||
<paragraph id="377259FF4C247ACC540EFDDD22676AE8" pageId="0" pageNumber="495">Hind limbs robust, tibia 41-47% (n=8) of SVL and foot 66-80% (n=8) of SVL; heels not meeting when hind limbs flexed at right angles to the axis of the body; tibio-tarsal articulation reaching between the posterior and anterior of the eye when hind limb is stretched along the side of the body; toes distinctly long and thin, relative lengths I<II<III<V<IV; tips of toes pointed, dilated into pear-shaped disks; toes with full webs, metatarsal web present, distinct lateral fringes on lateral edges of toes I and V; subarticular tubercles rounded, prominent; inner metatarsal tubercle large and long-elliptic, slightly raised, length twice the width; outer metatarsal tubercle relatively smaller than the inner metatarsal tubercle, distinctly raised and pointed, length slightly larger than width; inner tarsal fold relatively flat, in contact with the inner metatarsal tubercle; tarsal tubercle large and distinctly raised, close to the tibio-tarsal articulation.</paragraph>
|
||
<paragraph id="173343BDB72D28204C2E1A293E7637CF" pageId="0" pageNumber="495">Dorsal surface relatively rough, transverse wrinkles and dense tubercles on the dorsum, head, flanks, and limbs; small granules on the dorsal rears of hands and tarsi; not bearing spinules on the dorsal skin; supratympanic fold distinct, raised and granulated, extending from the posterior corner of the eye, curved on the temporal region, to the previous shoulder; dorsolateral fold absent. Ventral surface with large flattened tubercles, denser on the throat and thighs; dense granules on the ventral feet and tarsi.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="427E7B1D7B2E0986CECCD32E433A6B47" pageId="0" pageNumber="495" type="coloration">
|
||
<paragraph id="7F1BEAB1AD4E3BF553302AD169BFEAA8" pageId="0" pageNumber="495">Coloration.</paragraph>
|
||
<paragraph id="8CAF09744F1BA3AB671E3084AB94C92A" pageId="0" pageNumber="495">
|
||
In life (Fig.
|
||
<figureCitation id="BCFA59F73DC241B36EDE6E3DDDBEF8B8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological features of Occidozyga obscura comb. nov. in life. A - D dorsolateral view, ventral view, left hand, and right foot of SYS a 008122 E dorsolateral view of SYS a 008123 F dorsolateral view of an uncaptured individual in situ from Shimentai Nature Reserve." figureDoi="10.3897/vz.72.e80019.figure3" httpUri="https://binary.pensoft.net/fig/709316" pageId="0" pageNumber="495">3</figureCitation>
|
||
), dorsal surfaces vary from olive brown to dark brown; dorsum and flanks interspersed with irregular black speckles; mid-dorsal stripe grass-green with distinct border; lateral line system grayish brown to yellowish brown. Pupil bordered with yellow, iris beige to brown. Supratympanic fold olive-brown to dark brown. Skin of throat, chest, and belly uniform creamy white; gular with a pair of distinct or indistinct longitudinal dark stripes; skin of limbs dark gray, tubercles on ventral thighs and tibias creamy white.
|
||
</paragraph>
|
||
<paragraph id="5229E03B6BC2907A287085C0DF73186C" pageId="0" pageNumber="495">
|
||
In preservative (Fig.
|
||
<figureCitation id="B60AD6D7AAB7B2912EDBF672A9537495" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of Occidozyga obscura comb. nov. and Occidozyga lingnanica sp. nov. in preservative. Occidozyga obscura comb. nov.: A adult male SYS a 008121 B adult female SYS a 008122; Occidozyga lingnanica sp. nov.: C adult male paratype SYS a 007373 D adult female paratype SYS a 007375. 1 dorsal view 2 ventral view." figureDoi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495">4A, B</figureCitation>
|
||
), dorsum light gray; black speckles on dorsal surface light brown; mid-dorsal stripe fainted and the border indistinct; ventral skin grayish white; tubercles on ventral surface more distinct.
|
||
</paragraph>
|
||
<caption id="36C5BA55FFECF237AF33952F696D7F81" doi="10.3897/vz.72.e80019.figure3" httpUri="https://binary.pensoft.net/fig/709316" pageId="0" pageNumber="495" start="Figure 3" startId="F3">
|
||
<paragraph id="731DA542AB1ABA10B4B2113C2E053B3B" pageId="0" pageNumber="495">
|
||
<emphasis id="0A1C776CE4B6E047B59F4CDDAFFBE559" bold="true" pageId="0" pageNumber="495">Figure 3.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName id="768C687ABFF67906605807BBB8DCB00D" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="5D2CC9B23C6BA3B1EAB049B44BB1B666" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. in life.
|
||
<emphasis id="E99C7B0D5628DFBDA318817726C26F47" bold="true" pageId="0" pageNumber="495">A</emphasis>
|
||
-
|
||
<emphasis id="D91E3E9C48C5CFF21F72DB8428B6DCA6" bold="true" pageId="0" pageNumber="495">D</emphasis>
|
||
dorsolateral view, ventral view, left hand, and right foot of SYS a008122
|
||
<emphasis id="668428DD7530375B52A7EB6475FC34FC" bold="true" pageId="0" pageNumber="495">E</emphasis>
|
||
dorsolateral view of SYS a008123
|
||
<emphasis id="B234F03A82FC8B474D7B579B170FEE30" bold="true" pageId="0" pageNumber="495">F</emphasis>
|
||
dorsolateral view of an uncaptured individual in situ from Shimentai Nature Reserve.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F8852CB177F6023A70A354A0B2D76C6F" doi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495" start="Figure 4" startId="F4">
|
||
<paragraph id="14DABCFE8E66697DEC84D67C3737CB7C" pageId="0" pageNumber="495">
|
||
<emphasis id="297CE069847015E548321E468E38DBB7" bold="true" pageId="0" pageNumber="495">Figure 4.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName id="CCD178C0A328AF8081940DDD60778A67" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="074CE16F3E6DF26E378172BBC9321E3D" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. and
|
||
<taxonomicName id="4CDB208ECFD0C288AA34EE850D92F197" authorityName="Lyu & Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
|
||
<emphasis id="4B19C01CE8A053AC5489FCDB1F3430EC" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in preservative.
|
||
<taxonomicName id="DFBE167973EB08F5DBC3FBB233340CCB" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="1D489DC551B984429FA3DC537E900728" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov.:
|
||
<emphasis id="725DD2AD054C88A303D3DDE9AC74F3A3" bold="true" pageId="0" pageNumber="495">A</emphasis>
|
||
adult male SYS a008121
|
||
<emphasis id="CAB088468B0AEF2D1232F4663296BB02" bold="true" pageId="0" pageNumber="495">B</emphasis>
|
||
adult female SYS a008122;
|
||
<taxonomicName id="18F02CB190348107F2C047FC912B6236" authorityName="Lyu & Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
|
||
<emphasis id="078E98A92DAF8E2D234DAE06F1AEF2F5" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.:
|
||
<emphasis id="B761DAE12CCA66D23032D57DCB200734" bold="true" pageId="0" pageNumber="495">C</emphasis>
|
||
adult male paratype SYS a007373
|
||
<emphasis id="3749D35D82CEF442B8CAC7C525E15D10" bold="true" pageId="0" pageNumber="495">D</emphasis>
|
||
adult female paratype SYS a007375.
|
||
<emphasis id="5B1127C507F0549A5B852B24A66600A1" bold="true" pageId="0" pageNumber="495">1</emphasis>
|
||
dorsal view
|
||
<emphasis id="42DCEB3392887830441D2F33DAEB55E9" bold="true" pageId="0" pageNumber="495">2</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="AD85BF328CCAC67B34383C313554EF2C" pageId="0" pageNumber="495" type="description">
|
||
<paragraph id="521BC1889792EEC94A15CBD661696B79" pageId="0" pageNumber="495">Male secondary sexual characteristics.</paragraph>
|
||
<paragraph id="78C3F7BE5995C59748BB5A99DDB24172" pageId="0" pageNumber="495">
|
||
Male with a single subgular vocal sac; in breeding season, a single, light grey nuptial pad on the dorsal surface of finger I, nuptial spinules invisible. Males (SVL 24.2-27.5 mm) distinctly smaller than females (SVL 31.5-32.2 mm) (Fig.
|
||
<figureCitation id="BFC853022A6F1BA318895B84AD95CC26" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of Occidozyga obscura comb. nov. and Occidozyga lingnanica sp. nov. in preservative. Occidozyga obscura comb. nov.: A adult male SYS a 008121 B adult female SYS a 008122; Occidozyga lingnanica sp. nov.: C adult male paratype SYS a 007373 D adult female paratype SYS a 007375. 1 dorsal view 2 ventral view." figureDoi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495">4A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FDB603CE96A372D7155662E5FAB1F02" pageId="0" pageNumber="495" type="ecology">
|
||
<paragraph id="60C4996DB8197B0BE16E8B7E4F8CFD28" pageId="0" pageNumber="495">Ecology.</paragraph>
|
||
<paragraph id="160416BF8104D282AF2FDB41FE6BCFAD" pageId="0" pageNumber="495">
|
||
This frog inhabits natural or artificial ponds and paddy fields in plain areas. They quickly dive underwater after being disturbed during the daytime, while become relatively insensitive at night. Males call in the water surface or waterside grass from dusk to dawn, more active during the rain. The breeding season is from April to August (this study;
|
||
<bibRefCitation id="692A0EC92B0CAC0364FBA9BF9A4E9D95" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B11" refString="Fei, L, Hu, SQ, Ye, CY, Huang, YZ, 2009. Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." title="Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." year="2009">Fei et al. 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8DE32EF91A4CA84031CC11D94FC2D789" pageId="0" pageNumber="495" type="distribution">
|
||
<paragraph id="F9FF5630278FA63EAAB0855ACE42409E" pageId="0" pageNumber="495">Distribution.</paragraph>
|
||
<paragraph id="81F0A4E887A8804BE3DEFA817BCC5CAA" pageId="0" pageNumber="495">
|
||
<taxonomicName id="D70298D7480A9306208AE5A5F59EAB8F" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="C88B278E3445A10D4A3F6AF430419401" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. can be recognized from several localities of Guangdong and Guangxi, southeastern China (Figs
|
||
<figureCitation id="40ED322975E95C02B37F51D831D06BAF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic trees of the genus Occidozyga sensu lato based on mitochondrial genes 16 S and CO 1. Bayesian posterior probabilities (BPP) and bootstrap supports (BS) are labeled above and below the nodes respectively. Numbers at the terminal of the lineages correspond to the IDs in Table 1." figureDoi="10.3897/vz.72.e80019.figure1" httpUri="https://binary.pensoft.net/fig/709314" pageId="0" pageNumber="495">1</figureCitation>
|
||
,
|
||
<figureCitation id="59C0C2CFF244BC767A8D82D040E47228" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the floating frogs Occidozyga obscura comb. nov., Occidozyga lingnanica sp. nov., and their closest congeners. Numbers correspond to the IDs in Table 1." figureDoi="10.3897/vz.72.e80019.figure2" httpUri="https://binary.pensoft.net/fig/709315" pageId="0" pageNumber="495">2</figureCitation>
|
||
), based on the analyzed vouchers in this work. The previous records of
|
||
<taxonomicName id="5DE291AB8C2C1E4E68F30F95FF98FC92" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="17D7387430BB3890C0560D7823D83917" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
from Guangdong, Guangxi, Hong Kong, Macao, and Fujian, southeastern China are reassigned to
|
||
<taxonomicName id="0C1928CE8D8D41FF43102A1DA4BAADD1" lsidName="O. obscura" pageId="0" pageNumber="495" rank="species" species="obscura">
|
||
<emphasis id="943923EBF5854824CE5BD93DE28F0EEF" italics="true" pageId="0" pageNumber="495">O. obscura</emphasis>
|
||
</taxonomicName>
|
||
currently (see Remarks below). The records of
|
||
<taxonomicName id="C7DD0A5E654804D1E59686183251D30F" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="10B727DA7F081863F3CA492EF76D6249" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
from Yunnan require further clarifications with molecular and morphological vouchers (which might be close to the lineage of
|
||
<taxonomicName id="806906301418B70AED35E90E4DFEC3B3" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">O. cf. lima</taxonomicName>
|
||
based on the biogeographical similarity).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07C9D0DFB406A7158F0558C885AA20C3" pageId="0" pageNumber="495" type="conservation status">
|
||
<paragraph id="0DC0F6396289EBE9DDA62A01F4DB67E1" pageId="0" pageNumber="495">Conservation status.</paragraph>
|
||
<paragraph id="7459D9B7900D432BEEFE46FED5F9D893" pageId="0" pageNumber="495">
|
||
This species was previously reported as common and widespread species in southeastern China under the nomen
|
||
<taxonomicName id="2F3B3735E1F51EAA5E1ED569E024B69E" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="AD0865B97F6DF0421CBB119E53BE465E" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
, but its population quantity is found rapidly declining due to the influence of human activities such as pesticide abuse and urban construction. The populations in Hong Kong, Macao, Shenzhen, Guangzhou, and Xiamen might disappear, as no more reports and vouchers in nearly 20 years (this study;
|
||
<bibRefCitation id="69430276CFDC23C6D8B920ED7D21DC4E" author="Chan, HK" journalOrPublisher="Annals and Magazine of Natural History, Series six" pageId="0" pageNumber="495" refId="B5" refString="Chan, HK, 2013. Phylogeography and cryptic diversity of Occidozyga lima (Gravenhorst 1829). Master dissertation at the University of Hong Kong, Hong Kong, China." title="Phylogeography and cryptic diversity of Occidozyga lima (Gravenhorst 1829). Master dissertation at the University of Hong Kong, Hong Kong, China." year="2013">Chan 2013</bibRefCitation>
|
||
). We recommend
|
||
<taxonomicName id="784BBA93672693CFF7847E3C6609E1DB" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="7E8B883A81288202217DC71C50DD1186" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. to be listed as Endangered (EN) [IUCN Red List criteria A2cd].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0172EA0542569B54ECE0D4242D779961" pageId="0" pageNumber="495" type="remarks">
|
||
<paragraph id="0C3C3E3596551DA5D5E2BE927B87D5A1" pageId="0" pageNumber="495">Remarks.</paragraph>
|
||
<paragraph id="3E35C40F99009D058C322C9266E172A5" pageId="0" pageNumber="495">
|
||
The type specimen of
|
||
<taxonomicName id="3514461B21FF0FBF2461F8D2FF352B44" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="C7D8A3AE07D338AA9A018EF7FB40CD59" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. was collected by John Reeves (1774-1856) but the exact type locality was not given in the original description (
|
||
<bibRefCitation id="419836B46EF339BC814BB5787A90E18E" author="Gray, JE" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="495" refId="B20" refString="Gray, JE, 1831. Description of two new genera of frogs discovered by John Reeves, Esq. in China. Zoological Miscellany, Part 1: 38." title="Description of two new genera of frogs discovered by John Reeves, Esq. in China. Zoological Miscellany, Part 1: 38." year="1831">Gray 1831</bibRefCitation>
|
||
). Zhao & Adler (1993) speculated that the type specimens may be from somewhere in southeastern China, especially Macao or Canton (= Guangzhou) in Pearl River Delta, where John Reeves lived and worked in.
|
||
</paragraph>
|
||
<paragraph id="DE876AC477B69133F91F2483F5723FFC" pageId="0" pageNumber="495">
|
||
Moreover, there were two historic species currently regarded as synonymies of
|
||
<taxonomicName id="0FDA8ED008AA23E9F724BBB29F002058" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
|
||
<emphasis id="26BC30BEE14A0F42310C90F40D4A43B0" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
|
||
</taxonomicName>
|
||
, namely
|
||
<taxonomicName id="F62BEC4F43134F16A037A1A5AAB4B394" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Oxyglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyglossa lima subsp. var. var. chinens" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="variety" species="lima" subSpecies="var." variety="chinens">Oxyglossa lima var. chinens</taxonomicName>
|
||
and
|
||
<taxonomicName id="D13C2C9FD70A89951557F6BADDF9FFDF" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Osteosternum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osteosternum amoyense" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="amoyense">
|
||
<emphasis id="3ADEC4F5C872B26C10DE00008B8B9240" italics="true" pageId="0" pageNumber="495">Osteosternum amoyense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="6CC46187A1D4C8EF560A4C50EC49347E" author="Stejneger, L" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="495" pagination="1 - 115" refId="B46" refString="Stejneger, L, 1925. Chinese amphibians and reptiles in the United States National Museum. Proceedings of the United States National Museum 66: 1 - 115" title="Chinese amphibians and reptiles in the United States National Museum." volume="66" year="1925">Stejneger 1925</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8B6A415A784128C589AA5AC9EC092342" author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="495" pagination="3 - 32" refId="B45" refString="Smith, MA, 1931. The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft. Bulletin of the Raffles Museum 5: 3 - 32" title="The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft." volume="5" year="1931">Smith 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BA3EE7A5A519B007456D01E27FE21BC3" author="Pope, CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="495" pagination="397 - 611" refId="B38" refString="Pope, CH, 1931. Notes on amphibians from Fukien, Hainan and other parts of China. Bulletin of the American Museum of Natural History 61: 397 - 611" title="Notes on amphibians from Fukien, Hainan and other parts of China." volume="61" year="1931">Pope 1931</bibRefCitation>
|
||
). The type locality for
|
||
<taxonomicName id="BC9CFFBD59E3304248DB808816ECEFB2" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Oxyglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyglossa lima subsp. var. var. chinens" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="variety" species="lima" subSpecies="var." variety="chinens">Oxyglossa lima var. chinens</taxonomicName>
|
||
is Lilong, Canton (= Lilang, Shenzhen, Guangdong), and for
|
||
<taxonomicName id="573687C54D464AEF1F51C5154DFF966E" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" baseAuthorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Osteosternum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osteosternum amoyense" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="amoyense">
|
||
<emphasis id="9989694085F1D5415D92353B908687C8" italics="true" pageId="0" pageNumber="495">Osteosternum amoyense</emphasis>
|
||
</taxonomicName>
|
||
is Amoy (= Xiamen, Fujian). During our field surveys, we are unable to observe any frogs that resembled these two historic species from Shenzhen or Xiamen, possibly due to the dramatic urbanization of these two cities. Nevertheless, according to their original descriptions and distributions, we propose to transfer these two taxa to be the synonymy of
|
||
<taxonomicName id="455C0F7E747EA2DAE4755E4F7F481AA5" authorityName="Lyu & Wang & Zeng & Luo & Zhang & Guo & Ren & Qi & Mo & Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
|
||
<emphasis id="4A7D7E188800B665B1B9BC1756BA1022" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
|
||
</taxonomicName>
|
||
comb. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |