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<mods:title id="1D515B4A73178E73329A2A5EAE2314BA">Notes on the genus Astenus Dejean, 1833 in the Mediterranean and adjacent regions. A new species from Turkey, seven new synonyms, and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<taxonomicName id="F0FE4D27FFCDFFDCFF4DFB309070551E" ID-CoL="HP2H" authority="(Gravenhorst, 1806)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[151,620,1276,1302]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="058AEAB6FFCDFFDCFF4DFB309375551E" bold="true" box="[151,361,1276,1302]" italics="true" pageId="1" pageNumber="252">Astenus procerus</emphasis>
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<bibRefCitation id="536F4B55FFCDFFDCFEA2FB309078551E" author="Gravenhorst, J. L. C." box="[376,612,1276,1302]" pageId="1" pageNumber="252" pagination="12" refId="ref10308" refString="Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 236 + [12] pp. https: // doi. org / 10.5962 / bhl. title. 67769" type="journal article" year="1806">Gravenhorst, 1806</bibRefCitation>
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<figureCitation id="AFC52A21FFCDFFDCFDA6FB3090E0551E" box="[636,764,1276,1302]" captionStart-0="FIGURES 112" captionStart-1="FIGURES 1324" captionStartId-0="2.[151,264,1029,1053]" captionStartId-1="2.[151,264,1916,1940]" captionTargetBox-0="[271,1310,237,997]" captionTargetBox-1="[265,1317,1158,1889]" captionTargetId-0="figure-47@2.[267,1320,234,1003]" captionTargetId-1="figure-92@2.[263,1323,1150,1893]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURES 112. Astenus procerus (Gravenhorst, 1806), intraspecific variation of aedeagus in ventral and lateral view: 12) Turkey, northeastern Anatolia; 34) Turkey, central Anatolia; 56) Turkey, southern Anatolia; 78) Turkey, western Anatolia; 910) Azerbaijan, Ordubad; 1112) Iran, Mazandran. Scale bars: 112) 0.2 mm." captionText-1="FIGURES 1324. Astenus procerus (Gravenhorst, 1806), intraspecific variation of aedeagus in ventral and lateral view: 1314) Syria; 1516) Iraq, Baghdad; 1718) Cyprus; 1920) Bulgaria, Strandza Mts.; 2122) Italy, Sicilia; 2324) Russia, Krasnodar. Scale bars: 1324) 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4418581" figureDoi-1="http://doi.org/10.5281/zenodo.4418583" httpUri-0="https://zenodo.org/record/4418581/files/figure.png" httpUri-1="https://zenodo.org/record/4418583/files/figure.png" pageId="1" pageNumber="252">Figs. 124</figureCitation>
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<treatmentCitation id="B65F10B5FFCDFFDCFF4DFAF79051555B" author="Gravenhorst, J. L. C." box="[151,589,1339,1363]" page="141" pageId="1" pageNumber="252" year="1806">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFAF79051555B" authority="Gravenhorst, 1806: 141" authorityName="Gravenhorst" authorityPageNumber="141" authorityYear="1806" box="[151,589,1339,1363]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="058AEAB6FFCDFFDCFF4DFAF7934A555B" box="[151,342,1339,1363]" italics="true" pageId="1" pageNumber="252">Paederus procerus</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFE87FAF79051555B" author="Gravenhorst, J. L. C." box="[349,589,1339,1363]" pageId="1" pageNumber="252" pagination="12" refId="ref10308" refString="Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 236 + [12] pp. https: // doi. org / 10.5962 / bhl. title. 67769" type="journal article" year="1806">Gravenhorst, 1806: 141</bibRefCitation>
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<treatmentCitationGroup id="17EE118AFFCDFFDCFF4DFA969047557A" box="[151,603,1370,1394]" pageId="1" pageNumber="252">
<treatmentCitation id="B65F10B5FFCDFFDCFF4DFA969013557A" author="Reitter, E." box="[151,527,1370,1394]" page="150" pageId="1" pageNumber="252" year="1909">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFA969013557A" authority="Reitter, 1909: 150" authorityName="Reitter" authorityPageNumber="150" authorityYear="1909" box="[151,527,1370,1394]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="058AEAB6FFCDFFDCFF4DFA96934D557A" box="[151,337,1370,1394]" italics="true" pageId="1" pageNumber="252">Astenus rufopacus</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFE82FA969013557A" author="Reitter, E." box="[344,527,1370,1394]" pageId="1" pageNumber="252" pagination="41 - 80" refId="ref10504" refString="Reitter, E. (1909) Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. II Band. Fauna Germanica. Schriften des Deutschen Lehrervereins fur Naturkunde 24. K. G. Lutz, Stuttgart, 392 pp., pls. 41 - 80." type="book chapter" year="1909">Reitter, 1909: 150</bibRefCitation>
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<emphasis id="058AEAB6FFCDFFDCFDCFFA969047557A" bold="true" box="[533,603,1370,1394]" pageId="1" pageNumber="252">
<taxonomicNameLabel id="1EB957CDFFCDFFDCFDCFFA969047557A" box="[533,603,1370,1394]" pageId="1" pageNumber="252">syn. n.</taxonomicNameLabel>
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<paragraph id="374136A4FFCDFFDCFF4DFAB591905599" blockId="1.[151,908,1339,1517]" box="[151,908,1400,1425]" pageId="1" pageNumber="252">
<treatmentCitationGroup id="17EE118AFFCDFFDCFF4DFAB591905599" box="[151,908,1400,1425]" pageId="1" pageNumber="252">
<treatmentCitation id="B65F10B5FFCDFFDCFF4DFAB590AE5598" author="Jarrige, J." box="[151,690,1400,1424]" page="125" pageId="1" pageNumber="252" year="1952">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFAB590AE5598" ID-CoL="HP7D" authority="Jarrige, 1952: 125" authorityName="circumflexus Jarrige" authorityPageNumber="125" authorityYear="1952" box="[151,690,1400,1424]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="subSpecies" species="circumflexus" subSpecies="circumflexus">
<emphasis id="058AEAB6FFCDFFDCFF4DFAB593EF5598" box="[151,499,1400,1424]" italics="true" pageId="1" pageNumber="252">Astenus circumflexus circumflexus</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFE20FAB490AE5598" author="Jarrige, J." box="[506,690,1400,1424]" pageId="1" pageNumber="252" pagination="117 - 139" refId="ref10349" refString="Jarrige, J. (1952) Brachelytres nouveaux ou mal connus de la faune circamediterraneenne. Annales de la Societe Entomologique de France, 119, 117 - 139." type="journal article" year="1952">Jarrige, 1952: 125</bibRefCitation>
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<taxonomicNameLabel id="1EB957CDFFCDFFDCFD62FAB590E25599" box="[696,766,1401,1425]" pageId="1" pageNumber="252">syn. n.</taxonomicNameLabel>
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(
<figureCitation id="AFC52A21FFCDFFDCFCD6FAB4919A5599" box="[780,902,1400,1425]" captionStart="FIGURES 2530" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[473,1120,747,1668]" captionTargetId="figure-280@4.[468,1120,741,1673]" captionTargetPageId="4" captionText="FIGURES 2530. Details of Astenus circumflexus scutellaris Coiffait 1971 (2527) and Astenus circumflexus circumflexus Jarrige, 1952 (2830): 25) habitus, holotype; 26) forebody, holotype; 27) holotype labels; 28) aedeagus, lateral view; 29) aedeagus, ventral view; 30) labels. Scale bars: 2526) 0.5 mm, 2829) 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4418585" httpUri="https://zenodo.org/record/4418585/files/figure.png" pageId="1" pageNumber="252">Figs. 2830</figureCitation>
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<paragraph id="374136A4FFCDFFDCFF4DFA5B916355A7" blockId="1.[151,908,1339,1517]" box="[151,895,1431,1455]" pageId="1" pageNumber="252">
<treatmentCitationGroup id="17EE118AFFCDFFDCFF4DFA5B916355A7" box="[151,895,1431,1455]" pageId="1" pageNumber="252">
<treatmentCitation id="B65F10B5FFCDFFDCFF4DFA5B90B855A7" author="Coiffait, H." box="[151,676,1431,1455]" page="184" pageId="1" pageNumber="252" year="1971">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFA5B90B855A7" authority="Coiffait, 1971: 184" authorityName="Coiffait" authorityPageNumber="184" authorityYear="1971" box="[151,676,1431,1455]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="subSpecies" species="circumflexus" subSpecies="scutellaris">
<emphasis id="058AEAB6FFCDFFDCFF4DFA5B93C155A7" box="[151,477,1431,1455]" italics="true" pageId="1" pageNumber="252">Astenus circumflexus scutellaris</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFE39FA5B90B855A7" author="Coiffait, H." box="[483,676,1431,1455]" pageId="1" pageNumber="252" pagination="177 - 199" refId="ref10141" refString="Coiffait, H. (1971) Nouveaux Astenus de la region palearctique occidentale. Nouvelle Revue d'Entomologie, 1, 177 - 199." type="journal article" year="1971">Coiffait, 1971: 184</bibRefCitation>
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<emphasis id="058AEAB6FFCDFFDCFD70FA5B90EC55A7" bold="true" box="[682,752,1431,1455]" pageId="1" pageNumber="252">
<taxonomicNameLabel id="1EB957CDFFCDFFDCFD70FA5B90EC55A7" box="[682,752,1431,1455]" pageId="1" pageNumber="252">syn. n.</taxonomicNameLabel>
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(
<figureCitation id="AFC52A21FFCDFFDCFD24FA5B916455A7" box="[766,888,1431,1455]" captionStart="FIGURES 2530" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[473,1120,747,1668]" captionTargetId="figure-280@4.[468,1120,741,1673]" captionTargetPageId="4" captionText="FIGURES 2530. Details of Astenus circumflexus scutellaris Coiffait 1971 (2527) and Astenus circumflexus circumflexus Jarrige, 1952 (2830): 25) habitus, holotype; 26) forebody, holotype; 27) holotype labels; 28) aedeagus, lateral view; 29) aedeagus, ventral view; 30) labels. Scale bars: 2526) 0.5 mm, 2829) 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4418585" httpUri="https://zenodo.org/record/4418585/files/figure.png" pageId="1" pageNumber="252">Figs. 2527</figureCitation>
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<treatmentCitation id="B65F10B5FFCDFFDCFF4DFA7A901E55C6" author="Jarrige, J." box="[151,514,1462,1486]" page="126" pageId="1" pageNumber="252" year="1952">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFA7A901E55C6" ID-CoL="HPBY" authority="Jarrige, 1952: 126" authorityName="Jarrige" authorityPageNumber="126" authorityYear="1952" box="[151,514,1462,1486]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="058AEAB6FFCDFFDCFF4DFA7A935F55C6" box="[151,323,1462,1486]" italics="true" pageId="1" pageNumber="252">Astenus obliquus</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFE90FA7A901E55C6" author="Jarrige, J." box="[330,514,1462,1486]" pageId="1" pageNumber="252" pagination="117 - 139" refId="ref10349" refString="Jarrige, J. (1952) Brachelytres nouveaux ou mal connus de la faune circamediterraneenne. Annales de la Societe Entomologique de France, 119, 117 - 139." type="journal article" year="1952">Jarrige, 1952: 126</bibRefCitation>
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<emphasis id="058AEAB6FFCDFFDCFDD2FA7A905255C6" bold="true" box="[520,590,1462,1486]" pageId="1" pageNumber="252">
<taxonomicNameLabel id="1EB957CDFFCDFFDCFDD2FA7A905255C6" box="[520,590,1462,1486]" pageId="1" pageNumber="252">syn. n.</taxonomicNameLabel>
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<paragraph id="374136A4FFCDFFDCFF4DFA1990AA55E5" blockId="1.[151,908,1339,1517]" box="[151,694,1492,1517]" pageId="1" pageNumber="252">
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<treatmentCitation id="B65F10B5FFCDFFDCFF4DFA1993C755E5" author="Coiffait, H." box="[151,475,1492,1517]" page="94" pageId="1" pageNumber="252" year="1960">
<taxonomicName id="F0FE4D27FFCDFFDCFF4DFA1993C755E5" authority="Coiffait, 1960: 94" authorityName="Coiffait" authorityPageNumber="94" authorityYear="1960" box="[151,475,1492,1517]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="058AEAB6FFCDFFDCFF4DFA19933C55E4" box="[151,288,1492,1516]" italics="true" pageId="1" pageNumber="252">Astenus baali</emphasis>
<bibRefCitation id="536F4B55FFCDFFDCFEFDFA1893C755E5" author="Coiffait, H." box="[295,475,1492,1517]" pageId="1" pageNumber="252" pagination="49 - 99" refId="ref10105" refString="Coiffait, H. (1960) Les Astenus d'Europe et de la region mediterraneenne (Coleopteres, Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 95, 49 - 99." type="journal article" year="1960">Coiffait, 1960: 94</bibRefCitation>
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<emphasis id="058AEAB6FFCDFFDCFE3BFA19903B55E5" bold="true" box="[481,551,1493,1517]" pageId="1" pageNumber="252">
<taxonomicNameLabel id="1EB957CDFFCDFFDCFE3BFA19903B55E5" box="[481,551,1493,1517]" pageId="1" pageNumber="252">syn. n.</taxonomicNameLabel>
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(
<figureCitation id="AFC52A21FFCDFFDCFDEFFA1890B255E5" box="[565,686,1492,1517]" captionStart="FIGURES 3141" captionStartId="5.[151,264,1374,1398]" captionTargetBox="[497,1091,599,1346]" captionTargetId="figure-227@5.[497,1091,597,1351]" captionTargetPageId="5" captionText="FIGURES 3141. Details of Astenus baali Coiffait, 1960: 31) habitus, holotype; 32) forebody, holotype; 33) habitus, paratype; 34) forebody, paratype; 35) holotype aedeagus, lateral view; 36) holotype aedeagus, ventral view; 37) paratype aedeagus, lateral view; 38) holotype labels; 3941) paratype labels. Scale bars: 3134) 0.5 mm, 3537) 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4418587" httpUri="https://zenodo.org/record/4418587/files/figure.png" pageId="1" pageNumber="252">Figs. 3141</figureCitation>
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