109 lines
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109 lines
13 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner & Keupp, Helmut" docDate="2022" docId="BF22FCEFE1625BFEB90A1735C48C8812" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Leptolepis steberae Schwarzhans & Keupp 2022, sp. nov." docType="treatment" docUuid="35891115-EFFB-42C3-A196-CA22CEF8799D" docUuidSource="ZooBank" docVersion="4" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageNumber="51" updateTime="1732488386774" updateUser="ExternalLinkService">
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<mods:titleInfo id="76CB4073161C9D0EDECAABBD57D0BDDC">
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<mods:title id="B60227AD4907044CB883918EABA6A32B">Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
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</mods:titleInfo>
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<mods:name id="1C64B3B47A8BAD81FB235ED57C1A9D49" type="personal">
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<mods:roleTerm id="2D6762B00A6EB19FE80BDE380AC55177">Author</mods:roleTerm>
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<mods:namePart id="2C3105D9E21004EEFD13FE62B8291A63">Schwarzhans, Werner</mods:namePart>
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<mods:nameIdentifier id="60ECC03BCA35524A7D1A64A88821CE85" type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
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<mods:affiliation id="AB6E5138C0E83DDC2578F96C8E024042">Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="3EE3899B01255C0B09B13F3C3AF580A3" type="email">wwschwarz@aol.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="351019E41E4D564170090DE42F11C2B9">Keupp, Helmut</mods:namePart>
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<mods:affiliation id="B70EDF058C1E9EC948417462C2A1AA54">Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
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<mods:typeOfResource id="DB313E851F41ED6DA2DD88AB68310AFC">text</mods:typeOfResource>
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<mods:title id="CE4E5CEE72676B5A6D647D05E45F6B2C">Zitteliana</mods:title>
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<mods:part id="0B8148428AB417D597B4DA621A4186F7">
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<mods:date id="7B270D810A8AB920C661A48DCB2F7403">2022</mods:date>
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<mods:number id="0334CEB4161E545FF67BAD70F7A224FE">2022-04-12</mods:number>
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<mods:detail id="B375FA46FDC63C5E3EAD4C9B2A1FAA2B" type="volume">
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<mods:number id="F169827AADE132DD95734956AEB18283">96</mods:number>
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<mods:start id="A179C2170CD6F190C7B5739F4FCDB518">51</mods:start>
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<mods:end id="D8C8A609BCA535936BEBDD2F5851BFF4">67</mods:end>
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<mods:url id="65B856465955BF343D3A4F4F93B1D6DE">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:url>
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<mods:classification id="C6D655A79381E21298420EB07CE07939">journal article</mods:classification>
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<mods:identifier id="F93F1C244AFADE4570EBCC0ECBA4B3DC" type="DOI">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:identifier>
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<treatment id="BF22FCEFE1625BFEB90A1735C48C8812" ID-GBIF-Taxon="194588019" LSID="urn:lsid:zoobank.org:act:35891115-EFFB-42C3-A196-CA22CEF8799D" httpUri="http://treatment.plazi.org/id/BF22FCEFE1625BFEB90A1735C48C8812" lastPageNumber="51" pageId="0" pageNumber="51" scope_phylum="Teleostei">
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<subSubSection id="6798A7EFE1C847AB1E8989EB02E0A976" pageId="0" pageNumber="51" type="nomenclature">
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<paragraph id="8700ECF16E0BAAD5EB8E687F13E4D502" pageId="0" pageNumber="51">
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<taxonomicName id="ACE78D13824A0C105665E525EEF073AD" LSID="http://zoobank.org/35891115-EFFB-42C3-A196-CA22CEF8799D" authority="Schwarzhans & Keupp, 2022" authorityName="Schwarzhans & Keupp" authorityYear="2022" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis steberae" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="steberae" status="sp. nov.">Leptolepis steberae</taxonomicName>
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<taxonomicNameLabel id="2F5A60BDB8473DBFF051A266FAB01347" pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="BF917F66EAD7C2A05D905DE8E4CFBBE5" pageId="0" pageNumber="51" type="description">
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<paragraph id="9FB081B4BB6240A663C3D359BDE537C7" pageId="0" pageNumber="51">
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<figureCitation id="E41CCA76D415CA815FB472C109E861F7" captionStart="Plate 2" captionStartId="F4" captionText="Plate 2. Otoliths of Leptolepis buttenheimensis and L. steberae. figs 1 - 7 Leptolepis buttenheimensis sp. nov. from the late Pliensbachian of Buttenheim; 1: holotype SNSB-BSPG 2022 IV 4, EPH horizon; 2 - 7: paratypes SNSB-BSPG 2022 IV 5 - 6; 2 - 5, 7: EPH horizon; 6: Apyrenum subzone. figs 8 - 12: Leptolepis steberae sp. nov. from the Pliensbachian of Buttenheim, EPH horizon; 8: holotype SNSB-BSPG 2022 IV 13; 9 - 12: paratypes SNSB-BSPG 2022 IV 14." figureDoi="10.3897/zitteliana.96.81737.plate2" httpUri="https://binary.pensoft.net/fig/671632" pageId="0" pageNumber="51">Plate 2, figs 8-12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0EE5E8D3C24A6D160939629C6AB2CA54" pageId="0" pageNumber="51" type="etymology">
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<paragraph id="BFE071B1036674232F9DB867B9BA7067" pageId="0" pageNumber="51">Etymology.</paragraph>
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<paragraph id="DBBAB6314E2A11570F0501F1F1CC541C" pageId="0" pageNumber="51">Named after Birgit Steber (Leibersdorf), who has intensely collected from the Buttenheim clay pit and provided specimens of this species which were instrumental for its recognition.</paragraph>
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</subSubSection>
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<subSubSection id="18C95FA1FB1C70DA99D893D35EFB2452" pageId="0" pageNumber="51" type="holotype">
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<paragraph id="B040451EFFFE3809407D8E36283C21DF" pageId="0" pageNumber="51">Holotype.</paragraph>
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<paragraph id="7FEE9CBE68F7C2F59FA02E1873BFCE6D" pageId="0" pageNumber="51">
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SNSB-BSPG 2022 IV 13 (Plate
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<figureCitation id="4F0955989AA2DEA02D8F7420948AD9C1" captionStart="Plate 2" captionStartId="F4" captionText="Plate 2. Otoliths of Leptolepis buttenheimensis and L. steberae. figs 1 - 7 Leptolepis buttenheimensis sp. nov. from the late Pliensbachian of Buttenheim; 1: holotype SNSB-BSPG 2022 IV 4, EPH horizon; 2 - 7: paratypes SNSB-BSPG 2022 IV 5 - 6; 2 - 5, 7: EPH horizon; 6: Apyrenum subzone. figs 8 - 12: Leptolepis steberae sp. nov. from the Pliensbachian of Buttenheim, EPH horizon; 8: holotype SNSB-BSPG 2022 IV 13; 9 - 12: paratypes SNSB-BSPG 2022 IV 14." figureDoi="10.3897/zitteliana.96.81737.plate2" httpUri="https://binary.pensoft.net/fig/671632" pageId="0" pageNumber="51">2</figureCitation>
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, fig. 8), Clay pit near Buttenheim, upper Pliensbachian,
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<taxonomicName id="C5A8ECC9ECE9BBF50AD745FD0DE3742B" genus="Hawkerense" lsidName="Hawkerense" pageId="0" pageNumber="51" rank="genus">
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<emphasis id="F6C2F94900B3D11DB1A38F264BEEC72A" italics="true" pageId="0" pageNumber="51">Hawkerense</emphasis>
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</taxonomicName>
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subzone, EPH horizon.
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</paragraph>
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</subSubSection>
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<subSubSection id="9E6DD235BA02194C05563276FA559836" pageId="0" pageNumber="51" type="paratypes">
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<paragraph id="8305B3B143552E184D4B1DE3AE9142FB" pageId="0" pageNumber="51">Paratypes.</paragraph>
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<paragraph id="424BE430C8A744DB2486F3E9DC1934D1" pageId="0" pageNumber="51">
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9 specimens same data as holotype (SNSB-BSPG 2022 IV 14, Plate
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<figureCitation id="A0D7E9784182437CF7BA235396A10750" captionStart="Plate 2" captionStartId="F4" captionText="Plate 2. Otoliths of Leptolepis buttenheimensis and L. steberae. figs 1 - 7 Leptolepis buttenheimensis sp. nov. from the late Pliensbachian of Buttenheim; 1: holotype SNSB-BSPG 2022 IV 4, EPH horizon; 2 - 7: paratypes SNSB-BSPG 2022 IV 5 - 6; 2 - 5, 7: EPH horizon; 6: Apyrenum subzone. figs 8 - 12: Leptolepis steberae sp. nov. from the Pliensbachian of Buttenheim, EPH horizon; 8: holotype SNSB-BSPG 2022 IV 13; 9 - 12: paratypes SNSB-BSPG 2022 IV 14." figureDoi="10.3897/zitteliana.96.81737.plate2" httpUri="https://binary.pensoft.net/fig/671632" pageId="0" pageNumber="51">2</figureCitation>
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, figs 9-12).
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</paragraph>
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</subSubSection>
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<subSubSection id="DBDEC0B05204E2B81BCC8552F6D4EB9B" pageId="0" pageNumber="51" type="diagnosis">
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<paragraph id="09706F5C8C3D2538FB29EC268A2227B9" pageId="0" pageNumber="51">Diagnosis.</paragraph>
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<paragraph id="4C9C7F6E4973A141AC6029CD15E7226A" pageId="0" pageNumber="51">OL:OH = 1.1-1.25. Rostrum short, its length 12-25% of OL. Ventral and dorsal rims regularly curving. Ventral rim of ostium shallow, often fading, no or very weak distinction of ostium and cauda.</paragraph>
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</subSubSection>
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<subSubSection id="64B1499E2969C6A86B4C49744B37D7D7" pageId="0" pageNumber="51" type="description">
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<paragraph id="05207A5B35EB15FA788493A7BCBAD094" pageId="0" pageNumber="51">Description.</paragraph>
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<paragraph id="D1C8AEE3613D30F6F4DD3A7764439BDA" pageId="0" pageNumber="51">Nearly round, thin otoliths up to 2.5 mm in length (holotype 2.2 mm); OH:OT = 3.5-4.5. Ventral rim deeply and regularly curved, and smooth or intensely and finely crenulated. Rostrum short, blunt. Dorsal rim behind ostial relatively long, elevated, rounded, broad and irregularly undulating. Posterior rim broadly rounded.</paragraph>
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<paragraph id="D1A06170963CC7D86932380A4A175CDC" pageId="0" pageNumber="51">Inner face distinctly bent with slightly supramedian sulcus. Ostium short, indistinctly separated from cauda, slightly expanding backward from rostrum and antirostral notch, about as long as cauda. Ventral margin of sulcus often indistinct, gradual, relatively straight with no or very feeble indication of differentiation into ostium and cauda. Cauda straight, terminating close to posterior tip of otolith. Dorsal depression very small, restricted to position above anterior part of cauda. No ventral furrow. Outer face flat to slightly concave, with many short radial furrows particularly along ventral margin of otolith. No or very feeble postcentral umbo.</paragraph>
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</subSubSection>
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<subSubSection id="A8159C87D940AA02CCCFFF616C2332AE" pageId="0" pageNumber="51" type="discussion">
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<paragraph id="EB8616C70912AC0A071D1C36D77A211C" pageId="0" pageNumber="51">Discussion.</paragraph>
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<paragraph id="7DB669CDA78CA8401B6FC44102E6A860" pageId="0" pageNumber="51">
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<taxonomicName id="24B2B6D4F751B889A45D10FAB00C05B1" authorityName="Schwarzhans & Keupp" authorityYear="2022" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis steberae" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="steberae">
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<emphasis id="E1291012B1B19A2A6823A951DD6F68A4" italics="true" pageId="0" pageNumber="51">Leptolepis steberae</emphasis>
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</taxonomicName>
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is a relatively rare and small species in the late Pliensbachian of Buttenheim. It is recognized by its compressed, rounded outline with only a moderately projecting and rather blunt rostrum. Its OL:OH ratio is less than in contemporaneous congeners (1.1-1.25 vs. 1.25-175). The rostrum is shorter than in coeval
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<taxonomicName id="CFFD01C10D681B305E8BD818D8B8DBB7" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="genus">
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<emphasis id="53189D3DD65A443F74132357D5DC9F1E" italics="true" pageId="0" pageNumber="51">Leptolepis</emphasis>
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</taxonomicName>
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species (12-25% of OL vs. 30-45%). Another typical characteristic is the nearly straight ventral margin of the sulcus, which is also often rather gradational.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |