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<document id="B3645B8FAB880310D0D15F9AF316996E" ID-CLB-Dataset="33690" ID-DOI="http://dx.doi.org/10.3897/zookeys.545.6470" ID-GBIF-Dataset="52de69a6-afe1-447f-9ace-cbc6ab610abf" ID-PMC="PMC4714369" ID-Pensoft-Pub="1313-2970-545-89" ID-PubMed="26798296" ID-ZooBank="61C2313F341741698E3559D134F884E0" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-545-89" ModsDocOrigin="ZooKeys 545" ModsDocTitle="Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene" checkinTime="1451243628961" checkinUser="pensoft" docAuthor="Blaeser, Marcel, Krogmann, Lars &amp; Peters, Ralph S." docDate="2015" docId="BF06359B59BF50D47F5769B41694165C" docLanguage="en" docName="ZooKeys 545: 89-100" docOrigin="ZooKeys 545" docSource="http://dx.doi.org/10.3897/zookeys.545.6470" docTitle="Tenuicornus dominicus Blaeser, Krogmann &amp; Peters, 2015, sp. n." docType="treatment" docUuid="8968FC81-D138-43FB-AA79-CD15E283D82C" docUuidSource="ZooBank" docVersion="9" lastPageNumber="96" masterDocId="B67CFFF5FFF0B032DD50FFE4FFF76B6C" masterDocTitle="Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene" masterLastPageNumber="100" masterPageNumber="89" pageNumber="95" updateTime="1732709099810" updateUser="ExternalLinkService">
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<mods:title id="182FD83D2EC2DCE995A93106BDC5424F">Two new fossil genera and species of Cerocephalinae (Hymenoptera, Chalcidoidea, Pteromalidae), including the first record from the Eocene</mods:title>
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<mods:namePart id="69FEFED42D8245000628E5BB94BDF9EB">Blaeser, Marcel</mods:namePart>
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<mods:namePart id="CA59CC7387EFA12317DD00AA1765C4E9">Krogmann, Lars</mods:namePart>
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<mods:namePart id="923DA8AF6688D886060394A04881DCBF">Peters, Ralph S.</mods:namePart>
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<mods:date id="847557CC24E24270C835BC8494DA989F">2015</mods:date>
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<mods:classification id="E18F1046219DEA81C28C26E5341AFD71">journal article</mods:classification>
<mods:identifier id="40106FC6B5293A34EDAFE20607991B6F" type="DOI">http://dx.doi.org/10.3897/zookeys.545.6470</mods:identifier>
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<treatment id="BF06359B59BF50D47F5769B41694165C" ID-GBIF-Taxon="127897705" LSID="urn:lsid:zoobank.org:act:8968FC81-D138-43FB-AA79-CD15E283D82C" httpUri="http://treatment.plazi.org/id/BF06359B59BF50D47F5769B41694165C" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95" scope_family="Pteromalidae" scope_order="Hymenoptera" scope_subFamily="Cerocephalinae" scope_superFamily="Chalcidoidea">
<subSubSection id="C01069FF25152C270CF2037FF3D002D8" pageId="6" pageNumber="95" type="multiple">
<paragraph id="BD0E5F941713983EE54AA98F50B192EC" pageId="6" pageNumber="95">Taxon classification Animalia Hymenoptera Pteromalidae</paragraph>
</subSubSection>
<subSubSection id="26C4D298C367690C9B1E9AC772001465" pageId="6" pageNumber="95" type="nomenclature">
<paragraph id="9EED4E884C9961CA40DDB78C38A6BE55" pageId="6" pageNumber="95">
<taxonomicName id="AA3998E2B2A48135EBE4AE0FE28E14EC" LSID="http://zoobank.org/8968FC81-D138-43FB-AA79-CD15E283D82C" genus="Tenuicornus" lsidName="Tenuicornus dominicus" pageId="6" pageNumber="95" rank="species" species="dominicus">Tenuicornus dominicus</taxonomicName>
<taxonomicNameLabel id="975C988BEF362A73620793AA84F994B0" pageId="6" pageNumber="95">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="810076A042B1BA9D8F65DEC4161B8206" pageId="6" pageNumber="95" type="diagnosis">
<paragraph id="A2296D774A523594F5CC720C2559E9D9" pageId="6" pageNumber="95">Diagnosis.</paragraph>
<paragraph id="461C7D37416A4AAEB5F448C3A621D6F3" pageId="6" pageNumber="95">See genus.</paragraph>
</subSubSection>
<subSubSection id="2B4D26BA47EC9CC38A15BB323E56DCAE" pageId="6" pageNumber="95" type="description">
<paragraph id="468F1C6FB426968A5E4895114E58E6EE" pageId="6" pageNumber="95">Description.</paragraph>
<paragraph id="E4D92C7C89664871F800C9AC567B4024" pageId="6" pageNumber="95">
Female: total body length 2.07 mm; length of mesosoma 0.92 mm, of metasoma 1.02 mm. Body without metallic luster. Head: height 0.39 mm, width 0.38 mm. Face deeply impressed. Shape of head cuboid; mandibles not visible, hidden in facial depression. Margin of facial depression with rough surface and with a thickened seta on each side (length of setae 85
<normalizedToken id="FCA56B627F4155E84B0F3DD768FFC9C7" originalValue="µm">µm</normalizedToken>
). Inner antennal process (iap) positioned distinctly above level of toruli; nail-like, 0.11 mm long, with basis wider than apex, orientated anteroventrally. Eyes large and egg-shaped, 0.25 mm high and 0.2 mm wide. Antennae: inserted at ventral margin of eyes; scape long and slightly curved, length 0.25 mm; pedicel length 75
<normalizedToken id="C603CF9CC20DDBDA1B13A0F53550651B" originalValue="µm">µm</normalizedToken>
, rather short and stout; funicle 6-segmented with first two funicular segments longer than wide (F1: length: 71
<normalizedToken id="0666F0F68ABEBE2662667117D6880B59" originalValue="µm">µm</normalizedToken>
<normalizedToken id="58E4C2496DB04D9DA41AA2EAB492023C" originalValue="×">x</normalizedToken>
width: 52
<normalizedToken id="5C688D130E633886E5AD252A6EAC6830" originalValue="µm">µm</normalizedToken>
; F2: 68
<normalizedToken id="2DA440BA5036FF9992A57DE71EFC9576" originalValue="µm">µm</normalizedToken>
<normalizedToken id="78A25C16750EE583C8578691E4824445" originalValue="×">x</normalizedToken>
60
<normalizedToken id="52F88A3E6AD801ED5ACF7FC4147148F3" originalValue="µm">µm</normalizedToken>
) and F3-F6 wider than long, distinctly broadening distally, club-like (F3: 59
<normalizedToken id="4FA54B97C8F960FF23DBF5DEB3EFE320" originalValue="µm">µm</normalizedToken>
<normalizedToken id="7D8E44293E9B58786D392F561B31F0E4" originalValue="×">x</normalizedToken>
72
<normalizedToken id="419F083972DF7B3586F21DD2E3068D05" originalValue="µm">µm</normalizedToken>
; F4: 60
<normalizedToken id="3D92802597F4D44DDACA0B097DBCE12E" originalValue="µm">µm</normalizedToken>
<normalizedToken id="B45B3C81321752A90DDF7A661C6E352A" originalValue="×">x</normalizedToken>
74
<normalizedToken id="1D315383D36A8E22016EF86CC1B58422" originalValue="µm">µm</normalizedToken>
; F5: 58
<normalizedToken id="D646A0A4CDBE4F21C3ADD0E2A2BD20E5" originalValue="µm">µm</normalizedToken>
<normalizedToken id="28187F901B3781C5C1DEE4C337C1322A" originalValue="×">x</normalizedToken>
80
<normalizedToken id="0A6F5B58FED67D8ECC7118EE9CE20FE0" originalValue="µm">µm</normalizedToken>
; F6: 52
<normalizedToken id="5BF7AAE31DEBCA89BE5D169D246ECA7B" originalValue="µm">µm</normalizedToken>
<normalizedToken id="7D8833AB1365CD03D88E56A13B81654B" originalValue="×">x</normalizedToken>
86
<normalizedToken id="8205ABB3BE0D00BE227EB06102118CCD" originalValue="µm">µm</normalizedToken>
). Clava egg-shaped: 97
<normalizedToken id="63EB13D0CA9CACA960F4E5993758D20D" originalValue="µm">µm</normalizedToken>
long and 77
<normalizedToken id="F17E63253ABF382073E2E3E45BA8B404" originalValue="µm">µm</normalizedToken>
wide. Wings: fore wing long and slender (length 1.53 mm and width 0.55 mm); submarginal vein 0.57 mm long, marginal vein 0.38 mm long, postmarginal vein 0.11 mm long, stigmal vein 99
<normalizedToken id="609A5CF4E9F68AAC2C67B507289EFFF9" originalValue="µm">µm</normalizedToken>
long; stigma slightly thickened and uncus visible. Wing disc bare, setal bases absent. Elongate admarginal setae present. Hind wing length 1.27 mm and width 0.27 mm, three hamuli present. Mesosoma: pronotum and anterior part of mesoscutum with slight traces of strigulate surface sculpture; mesonotum otherwise polished. Notauli foveolate. Length of pronotum 0.37 mm, length of mesoscutum 0.22 mm, length of mesoscutellum 0.22 mm; no frenal line. Axillae medially connected. Mesopleuron (in lateral view) height 0.29 mm, width 0.36 mm. Prepectus enlarged and triangular in shape (0.14 mm
<normalizedToken id="BB2AA9A2515D1F49EBFC56FBC0BE51EB" originalValue="×">x</normalizedToken>
0.17 mm). Length of metanotum 39
<normalizedToken id="95418C0F0B59B42F9DBBF4C4376BBDFB" originalValue="µm">µm</normalizedToken>
; length of propodeum 72
<normalizedToken id="BDDBCC25D823C021DAD43907524DEBDE" originalValue="µm">µm</normalizedToken>
. Propodeum without a median carina or plicae. Legs: coxae strong and stout. Femora and tibiae long and slender (Fe1: 0.45 mm; Fe2: 0.32 mm; Fe3: 0.41 mm; Ti1: 0.39 mm; Ti2: 0.49 mm; Ti3: 0.42 mm). Metasoma: petiole (Mt1) short and transverse, hardly visible. Metasomal tergites Mt2 to Mt9 smooth, Mt2: 0.28 mm, Mt3: 0.17 mm, Mt4: 0.15 mm, Mt5: 0.16 mm, Mt6: 0.11 mm, Mt7 60
<normalizedToken id="DB39AFB5243D163E4B1BB363A5132D4F" originalValue="µm">µm</normalizedToken>
, Mt8/9: 71
<normalizedToken id="23EF1A9051C9615692C68991689E5884" originalValue="µm">µm</normalizedToken>
. Ovipositor 0.23 mm (top view) exerted beyond end of gaster.
</paragraph>
</subSubSection>
<subSubSection id="1CC33D281475B905BDB5BECDDAD7564D" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95" type="taxonomic remarks">
<paragraph id="C090748BEF0650D4EFA7676E268507FD" pageId="6" pageNumber="95">Taxonomic remarks.</paragraph>
<paragraph id="41F4E478EAFB4AD6E2CBA7278F3A8268" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95">
The two genera that are most similar to
<taxonomicName id="386853B6A698525C0EDFB7681A49F975" genus="Tenuicornus" lsidName="Tenuicornus" pageId="6" pageNumber="95" rank="genus">Tenuicornus</taxonomicName>
are
<taxonomicName id="000C6015330EE32380FA5011EA929CF8" class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
and
<taxonomicName id="9F5FE464ED25A95EA78D06FFD007F0D3" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
.
<taxonomicName id="75FD644E7D0592D14C2BED75CF0A5CE5" genus="Tenuicornus" lsidName="Tenuicornus" pageId="6" pageNumber="95" rank="genus">Tenuicornus</taxonomicName>
runs to
<taxonomicName id="A7BDB1C4CCE4AD41DB2EFDF89C9D6753" class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
in the key of
<bibRefCitation id="7236134CF4A71FE52774FADF9C3FAA38" author="Krogmann, L" journalOrPublisher="Historical Biology" pageId="10" pageNumber="99" pagination="43 - 49" title="First fossil record of cerocephaline wasps with a description of new genus and species from Dominican amber (Hymenoptera: Chalcidoidea: Pteromalidae: Cerocephalinae)." url="10.1080/08912963.2012.685729" volume="25" year="2013">Krogmann (2013)</bibRefCitation>
but differs from this genus in various features:
<taxonomicName id="BC8AF4D7EC2D6365096235F91B9480E6" genus="Tenuicornus" lsidName="Tenuicornus" pageId="6" pageNumber="95" rank="genus">Tenuicornus</taxonomicName>
possesses a head that is deeply impressed (convex in
<taxonomicName id="31E6637200D42C1A0D2F41B4FA6E3814" class="Insecta" family="Pteromalidae" genus="Neocalosoter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocalosoter" order="Hymenoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">Neocalosoter</taxonomicName>
), the dark transverse bands on
<pageBreakToken id="D0A584DF69AA40A100559A28B4BCFCBB" pageId="7" pageNumber="96" start="start">its</pageBreakToken>
fore wing are absent, and the propodeum has no traces of a median carina or plicae (Fig. 2B). The tentative similarity between
<taxonomicName id="CF804718A7D9A133C54B6699739739CD" genus="Tenuicornus" lsidName="Tenuicornus" pageId="7" pageNumber="96" rank="genus">Tenuicornus</taxonomicName>
and
<taxonomicName id="103ACB185F054C4A014B6ED8EE68C238" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
is based on the deeply impressed face. However,
<taxonomicName id="06727BCB6001B1EF5445B87852350E6D" genus="Tenuicornus" lsidName="Tenuicornus" pageId="7" pageNumber="96" rank="genus">Tenuicornus</taxonomicName>
differs from
<taxonomicName id="85DA37F4A11B405EFF26AEB865088FE9" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
by the absence of a tuft of setae on the parastigma on the fore wings (Fig. 2E), one of the most distinct characters for the generic classification of
<taxonomicName id="4E6A3B2DF3BEED66FFACF6A9489D9FFE" lsidName="" pageId="7" pageNumber="96" rank="subFamily" subFamily="Cerocephalinae">Cerocephalinae</taxonomicName>
.
<taxonomicName id="A9ECCC34B81537D91C2FD5BD403C7D86" genus="Tenuicornus" lsidName="Tenuicornus" pageId="7" pageNumber="96" rank="genus">Tenuicornus</taxonomicName>
further differs from
<taxonomicName id="F2767C3F09ACC9FBF85B495C0FA10B92" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
by the shape and position of the inner antennal process. In
<taxonomicName id="81F2774FE5123CF7B2F84F7472DDD11F" genus="Tenuicornus" lsidName="Tenuicornus" pageId="7" pageNumber="96" rank="genus">Tenuicornus</taxonomicName>
, the process is shaped like a nail and positioned above the level of the toruli at the upper margin of the facial depression (Fig. 2C). In
<taxonomicName id="8A9B6581C67E7D528CA9E00A2F36B9C6" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
, the process is broader and stouter and positioned at or slightly below the level of the toruli. Furthermore,
<taxonomicName id="58845D5F416D505817B017D13D0B1771" class="Insecta" family="Pteromalidae" genus="Cerocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocephala" order="Hymenoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">Cerocephala</taxonomicName>
is described to have all funicular segments longer than wide (
<bibRefCitation id="7C16673311457A9570AF13B58AB46507" author="Westwood, JO" journalOrPublisher="Philosophical Magazine" pageId="10" pageNumber="99" title="Descriptions of several new British forms amongst the parasitic hymenopterous insects." volume="1" year="1832">Westwood 1832</bibRefCitation>
), while in
<taxonomicName id="E324A9239EB614F5D042C893DCFF28AD" genus="Tenuicornus" lsidName="Tenuicornus" pageId="7" pageNumber="96" rank="genus">Tenuicornus</taxonomicName>
only the first two funicular segments are longer than wide. The funicular segments three to six are wider than long (Fig. 2D).
</paragraph>
</subSubSection>
<subSubSection id="BA2FD15ACE7860AB7EFD5143E4842A42" pageId="7" pageNumber="96" type="etymology">
<paragraph id="B997959717A930B24680C9252D125FCA" pageId="7" pageNumber="96">Etymology.</paragraph>
<paragraph id="2DD2D39AB8BF942AD9D5F2FFDCE21F23" pageId="7" pageNumber="96">The name dominicus is derived from the amber deposit in which the fossil was found.</paragraph>
</subSubSection>
</treatment>
</document>