366 lines
52 KiB
XML
366 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-PMC="PMC3689139" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-PubMed="23794925" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="BDD7E7F5F66E8D0DF032734B2018D9BD" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Lonchodraco giganteus Rodrigues & Kellner, 2013, comb. n." docType="treatment" docVersion="6" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="18" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="15" updateTime="1668155938761" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044964" LSID="urn:lsid:plazi:treatment:BDD7E7F5F66E8D0DF032734B2018D9BD" httpUri="http://treatment.plazi.org/id/BDD7E7F5F66E8D0DF032734B2018D9BD" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
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<subSubSection pageId="14" pageNumber="15" type="nomenclature">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName LSID="BDD7E7F5-F66E-8D0D-F032-734B2018D9BD" authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Lonchodraco giganteus" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Lonchodraco giganteus</taxonomicName>
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<taxonomicNameLabel pageId="14" pageNumber="15">comb. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchodraco giganteus comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation). A-F articulated anterior parts of the rostrum and mandible A right lateral view B respective line drawing C left lateral view D respective line drawing E anterior view F respective line drawing G associated scapulocoracoid in posterior view. Abbreviations: ch - choanae, cor - coracoid, d - dentary, dcr - dentary crest, m - maxillae, pl - palatine, pm - premaxillae, pmcr - premaxillaery crest, prid - palatal ridge, sca - scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22072" pageId="14" pageNumber="15">Fig. 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="reference_group">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Pterodactylus giganteus</taxonomicName>
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Bowerbank:
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<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="7 - 8" publicationUrl="10.1144/GSL.JGS.1846.002.01-02.05" refId="B11" refString="Bowerbank, JS, 1846. On a new species of pterodactyl found in the Upper Chalk of Kent. Quarterly Journal of the Geological Society 2: 7 - 8, 10.1144/GSL.JGS.1846.002.01-02.05" title="On a new species of pterodactyl found in the Upper Chalk of Kent." url="10.1144/GSL.JGS.1846.002.01-02.05" volume="2" year="1846">Bowerbank 1846</bibRefCitation>
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: p. 8, fig. 1, 2, 5.
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Pterodactylus giganteus</taxonomicName>
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Bowerbank:
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<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="2 - 10" publicationUrl="10.1144/GSL.JGS.1848.004.01-02.07" refId="B12" refString="Bowerbank, JS, 1848. Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals. Quarterly Journal of the Geological Society 4: 2 - 10, 10.1144/GSL.JGS.1848.004.01-02.07" title="Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals." url="10.1144/GSL.JGS.1848.004.01-02.07" volume="4" year="1848">Bowerbank 1848</bibRefCitation>
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: pl. I, fig. 1.
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="conirostris" status="comb. n.">Pterodactylus conirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Dixon, F" journalOrPublisher="Longman, Brown, Green, and Longmans, London" pageId="65" pageNumber="66" refId="B18" refString="Dixon, F, 1850. The geology and fossils of the Tertiary and Cretaceous Formations of Sussex. Longman, Brown, Green, and Longmans, London, 423 pp." title="The geology and fossils of the Tertiary and Cretaceous Formations of Sussex." year="1850">Owen in Dixon 1850</bibRefCitation>
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: p. 401, pl. XXXVIII, fig. 4-7
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Pterodactylus giganteus</taxonomicName>
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Bowerbank:
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<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="65" pageNumber="66" pagination="14 - 20" publicationUrl="10.1111/j.1096-3642.1851.tb01125.x" refId="B13" refString="Bowerbank, JS, 1851. On the pterodactyles of the Chalk Formation. Proceedings of the Zoological Society of London 19: 14 - 20, 10.1111/j.1096-3642.1851.tb01125.x" title="On the pterodactyles of the Chalk Formation." url="10.1111/j.1096-3642.1851.tb01125.x" volume="19" year="1851">Bowerbank 1851</bibRefCitation>
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: p. 19
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authority="" authorityName="Rodrigues & Kellner" authorityYear="2013" class="Aves" genus="Cimoliornis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="diomedaeus" status="comb. n." subSpecies="sic">Cimoliornis diomedaeus [sic]</taxonomicName>
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(Gervais):
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<bibRefCitation author="Owen, R" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="68" pageNumber="69" pagination="21 - 34" publicationUrl="10.1111/j.1096-3642.1851.tb01126.x" refId="B64" refString="Owen, R, 1851b. On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species. Proceedings of the Zoological Society of London 19: 21 - 34, 10.1111/j.1096-3642.1851.tb01126.x" title="On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species." url="10.1111/j.1096-3642.1851.tb01126.x" volume="19" year="1851 b">Owen 1851b</bibRefCitation>
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: p. 21
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Pterodactylus giganteus</taxonomicName>
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Bowerbank: Owen 1851a</bibRefCitation>
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: p. 91, pl. XXXI, fig. 1-9, 12-13
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ornithochirus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="sic" status="comb. n." subSpecies="giganteus">Ornithochirus [sic](?) giganteus</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Lydekker, R" journalOrPublisher="W. Part I. Trustees of the British Museum, London" pageId="67" pageNumber="68" refId="B48" refString="Lydekker, R, 1888. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S. W. Part I. Trustees of the British Museum, London, 309 pp." title="Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S." year="1888">Lydekker 1888</bibRefCitation>
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: p. 12
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Lonchodectes giganteus</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 538
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Ornithodesmidae" genus="Ornithodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Ornithodesmus(?) giganteus</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Arthaber, G" journalOrPublisher="Palaeontologische Zeitschrift" pageId="64" pageNumber="65" pagination="1 - 47" refId="B2" refString="Arthaber, G, 1922. Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1 - 47" title="Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier." volume="4" year="1922">Arthaber 1922</bibRefCitation>
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: p. 20, fig. 10
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Ornithocheirus giganteus</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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: p. 57, fig. 28
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="giganteus" status="comb. n.">Lonchodectes giganteus</taxonomicName>
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(Bowerbank):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: p. 210
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="lectotype">
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<paragraph pageId="14" pageNumber="15">Lectotype.</paragraph>
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<paragraph pageId="14" pageNumber="15">
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NHMUK PV 39412, anterior portions of the rostrum and mandible, incomplete scapulocoracoid, proximal ends of the humerus and ulna, and a partial wing phalanx (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchodraco giganteus comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation). A-F articulated anterior parts of the rostrum and mandible A right lateral view B respective line drawing C left lateral view D respective line drawing E anterior view F respective line drawing G associated scapulocoracoid in posterior view. Abbreviations: ch - choanae, cor - coracoid, d - dentary, dcr - dentary crest, m - maxillae, pl - palatine, pm - premaxillae, pmcr - premaxillaery crest, prid - palatal ridge, sca - scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22072" pageId="14" pageNumber="15">Fig. 4A-G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="type locality">
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<paragraph pageId="14" pageNumber="15">Type locality.</paragraph>
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<paragraph pageId="14" pageNumber="15">Near Maidstone, Burham, Kent, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="type horizon">
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<paragraph pageId="14" pageNumber="15">Type horizon.</paragraph>
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<paragraph pageId="14" pageNumber="15">Chalk Formation (Cenomanian / Turonian).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph pageId="14" pageNumber="15">Diagnosis.</paragraph>
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<paragraph lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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Lonchodraconid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): anterior portion of the premaxillae rounded; anterior portion of the
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<pageBreakToken pageId="15" pageNumber="16" start="start">dentaries</pageBreakToken>
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rounded; divergent alveolar margins of the anterior end of the upper and lower jaws; presence of a premaxillary crest; short, low, blade-like dentary crest*; approximately6 alveoli per 3 cm of jaw margin*.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="description">
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<paragraph pageId="16" pageNumber="17">
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<pageBreakToken pageId="16" pageNumber="17" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="16" pageNumber="17">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="16" pageNumber="17">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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was briefly described by
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<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="7 - 8" publicationUrl="10.1144/GSL.JGS.1846.002.01-02.05" refId="B11" refString="Bowerbank, JS, 1846. On a new species of pterodactyl found in the Upper Chalk of Kent. Quarterly Journal of the Geological Society 2: 7 - 8, 10.1144/GSL.JGS.1846.002.01-02.05" title="On a new species of pterodactyl found in the Upper Chalk of Kent." url="10.1144/GSL.JGS.1846.002.01-02.05" volume="2" year="1846">Bowerbank (1846)</bibRefCitation>
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, and then in more detail by
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Owen (1851a)</bibRefCitation>
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. The lectotype, NHMUK PV 39412, includes the anterior parts of the rostrum and mandible preserved, and, contra
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="7 - 8" publicationUrl="10.1144/GSL.JGS.1846.002.01-02.05" refId="B11" refString="Bowerbank, JS, 1846. On a new species of pterodactyl found in the Upper Chalk of Kent. Quarterly Journal of the Geological Society 2: 7 - 8, 10.1144/GSL.JGS.1846.002.01-02.05" title="On a new species of pterodactyl found in the Upper Chalk of Kent." url="10.1144/GSL.JGS.1846.002.01-02.05" volume="2" year="1846">Bowerbank (1846)</bibRefCitation>
|
||
and subsequent authors (
|
||
<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Martill, DM" journalOrPublisher="Cretaceous Research" pageId="67" pageNumber="68" pagination="236 - 243" publicationUrl="10.1016/j.cretres.2010.12.008" refId="B53" refString="Martill, DM, 2011. A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil. Cretaceous Research 32: 236 - 243, 10.1016/j.cretres.2010.12.008" title="A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil." url="10.1016/j.cretres.2010.12.008" volume="32" year="2011">Martill 2011</bibRefCitation>
|
||
), does not include the anterior portion of the nasoantorbital fenestra because what appears to be the anterior margin of the fenestra is not present on both sides of the specimen and most likely represents breakage. The lectotype of
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Lonchodraco giganteus</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguishable from pterosaurs from other Cretaceous deposits in Britain. Owen (in
|
||
<bibRefCitation author="Dixon, F" journalOrPublisher="Longman, Brown, Green, and Longmans, London" pageId="65" pageNumber="66" refId="B18" refString="Dixon, F, 1850. The geology and fossils of the Tertiary and Cretaceous Formations of Sussex. Longman, Brown, Green, and Longmans, London, 423 pp." title="The geology and fossils of the Tertiary and Cretaceous Formations of Sussex." year="1850">Dixon 1850</bibRefCitation>
|
||
) described it as deep-jawed and cone-beaked. The tips of the jaws are dorsoventrally flattened, and there is no upward curvature of the palate. The alveolar margins of the upper and lower jaw are divergent even in their anterior portions. The premaxilla is tall and triangular in lateral view, indicating the presence of a crest. The crest is not thin as seen in
|
||
<taxonomicName class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Anhanguera</emphasis>
|
||
</taxonomicName>
|
||
or thick as in
|
||
<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Coloborhynchus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Fastnacht, M" journalOrPublisher="Longman, Brown, Green, and Longmans, London" pageId="65" pageNumber="66" refId="B19" refString="Fastnacht, M, 2001. First record of Coloborhynchus (Pterosauria) from the Santana Formation (Lower Cretaceous) of the Chapada do Araripe, Brazil. Palaeontologische Zeitschrift 75: 23-36." title="First record of Coloborhynchus (Pterosauria) from the Santana Formation (Lower Cretaceous) of the Chapada do Araripe, Brazil. Palaeontologische Zeitschrift 75: 23 - 36." year="2001">Fastnacht 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner 2008</bibRefCitation>
|
||
). The mandibular symphysis also has a distinctive crest from that in anhanguerids because it does not start at the tip of the mandible. The crest is blade-like, short, and located medially in the relatively wide symphysis. Unfortunately, incomplete preparation of the specimen precludes more detailed observations of its oral region, including the palatal ridge. The mandibular groove appears to be deep but cannot be accurately measured. The teeth are conical and elongated, smaller than the ones in anhanguerids; similarly, the alveoli are small and oval to round.
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Lonchodraco giganteus</emphasis>
|
||
</taxonomicName>
|
||
has a shorter mandibular crest and a larger tooth density than
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="machaerorhynchus">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Lonchodraco machaerorhynchus</emphasis>
|
||
</taxonomicName>
|
||
and a tall rostrum as opposed to the elongated premaxillae and maxillae in
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="" pageId="16" pageNumber="17" rank="species" species="polyodon">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">
|
||
<normalizedToken originalValue="‘Ornithocheirus’">'Ornithocheirus'</normalizedToken>
|
||
polyodon
|
||
</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="remarks">
|
||
<paragraph pageId="16" pageNumber="17">Remarks.</paragraph>
|
||
<paragraph pageId="17" pageNumber="18">
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">
|
||
<pageBreakToken pageId="17" pageNumber="18" start="start">Lonchodraco</pageBreakToken>
|
||
giganteus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
has a complex taxonomic history. The species was named
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="7 - 8" publicationUrl="10.1144/GSL.JGS.1846.002.01-02.05" refId="B11" refString="Bowerbank, JS, 1846. On a new species of pterodactyl found in the Upper Chalk of Kent. Quarterly Journal of the Geological Society 2: 7 - 8, 10.1144/GSL.JGS.1846.002.01-02.05" title="On a new species of pterodactyl found in the Upper Chalk of Kent." url="10.1144/GSL.JGS.1846.002.01-02.05" volume="2" year="1846">Bowerbank (1846)</bibRefCitation>
|
||
. He referred several specimens to the species, including both cranial and postcranial material. Some of these specimens were found associated (NHMUK PV 39412), whereas others were not found associated but came from the same locality as the associated material; additional material was collected at different localities. It is unclear which specimen was considered the holotype.
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="2 - 10" publicationUrl="10.1144/GSL.JGS.1848.004.01-02.07" refId="B12" refString="Bowerbank, JS, 1848. Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals. Quarterly Journal of the Geological Society 4: 2 - 10, 10.1144/GSL.JGS.1848.004.01-02.07" title="Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals." url="10.1144/GSL.JGS.1848.004.01-02.07" volume="4" year="1848">Bowerbank (1848)</bibRefCitation>
|
||
described the paleohistology of some bones that he referred to
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
, including the cranial material (NHMUK PV 39412;
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="2 - 10" publicationUrl="10.1144/GSL.JGS.1848.004.01-02.07" refId="B12" refString="Bowerbank, JS, 1848. Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals. Quarterly Journal of the Geological Society 4: 2 - 10, 10.1144/GSL.JGS.1848.004.01-02.07" title="Microscopical observations on the structure of the bones of Pterodactylus giganteus and other fossil animals." url="10.1144/GSL.JGS.1848.004.01-02.07" volume="4" year="1848">Bowerbank 1848</bibRefCitation>
|
||
: fig. (1). Owen (in
|
||
<bibRefCitation author="Dixon, F" journalOrPublisher="Longman, Brown, Green, and Longmans, London" pageId="65" pageNumber="66" refId="B18" refString="Dixon, F, 1850. The geology and fossils of the Tertiary and Cretaceous Formations of Sussex. Longman, Brown, Green, and Longmans, London, 423 pp." title="The geology and fossils of the Tertiary and Cretaceous Formations of Sussex." year="1850">Dixon 1850</bibRefCitation>
|
||
) proposed a new name,
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="conirostris">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus conirostris</emphasis>
|
||
</taxonomicName>
|
||
, for NHMUK PV 39412, because he argued that the specimen was not gigantic in size and thus deemed the specific epithet
|
||
<emphasis italics="true" pageId="17" pageNumber="18">giganteus</emphasis>
|
||
inappropriate.
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="65" pageNumber="66" pagination="14 - 20" publicationUrl="10.1111/j.1096-3642.1851.tb01125.x" refId="B13" refString="Bowerbank, JS, 1851. On the pterodactyles of the Chalk Formation. Proceedings of the Zoological Society of London 19: 14 - 20, 10.1111/j.1096-3642.1851.tb01125.x" title="On the pterodactyles of the Chalk Formation." url="10.1111/j.1096-3642.1851.tb01125.x" volume="19" year="1851">Bowerbank (1851)</bibRefCitation>
|
||
responded that at the time of the description larger pterosaurs were unknown, that modifications of the names of species based on them being inappropriate would cause much instability, and refused to adopt
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="conirostris">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus conirostris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="65" pageNumber="66" pagination="14 - 20" publicationUrl="10.1111/j.1096-3642.1851.tb01125.x" refId="B13" refString="Bowerbank, JS, 1851. On the pterodactyles of the Chalk Formation. Proceedings of the Zoological Society of London 19: 14 - 20, 10.1111/j.1096-3642.1851.tb01125.x" title="On the pterodactyles of the Chalk Formation." url="10.1111/j.1096-3642.1851.tb01125.x" volume="19" year="1851">Bowerbank (1851)</bibRefCitation>
|
||
cited the Law of Priority of the British Association Code (also known as the Strickland Code, published in 1843), which was approved by a committee that included Owen (
|
||
<bibRefCitation author="International, Commission on Zoological Nomenclature" journalOrPublisher="Bulletin of the Natural History Museum: Geology Series" pageId="66" pageNumber="67" refId="B28" refString="International, Commission on Zoological Nomenclature, 1999. International Code of Zoological Nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature." title="International Code of Zoological Nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature." year="1999">International Commission on Zoological Nomenclature 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Dayrat, B" editor="Polaszek, A" journalOrPublisher="CRC, London" pageId="65" pageNumber="66" pagination="186 - 239" publicationUrl="10.1201/EBK1420095012-c17" refId="B17" refString="Dayrat, B, 2010. Celebrating 250 dynamic years of nomenclatural debates. In: Polaszek, A, Ed., System Naturae 250: The Linnean Ark. CRC, London: 186 - 239, 10.1201/EBK1420095012-c17" title="Celebrating 250 dynamic years of nomenclatural debates." url="10.1201/EBK1420095012-c17" volumeTitle="System Naturae 250: The Linnean Ark." year="2010">Dayrat 2010</bibRefCitation>
|
||
; see also
|
||
<bibRefCitation author="Martill, DM" editor="Moody, RTJ" journalOrPublisher="Geological Society Special Publications" pageId="67" pageNumber="68" pagination="287 - 311" refId="B52" refString="Martill, DM, 2010. The early history of pterosaur discovery in Great Britain. In: Moody, RTJ, Buffetaut, E, Naish, D, Martill, DM, Eds., Dinosaurs and other extinct saurians: a historical perspective. Geological Society Special Publications 343: 287 - 311" title="The early history of pterosaur discovery in Great Britain." volume="343" volumeTitle="Dinosaurs and other extinct saurians: a historical perspective." year="2010">Martill 2010</bibRefCitation>
|
||
). The Law of Priority stated that the first name of a species should be the one considered valid.
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Owen (1851a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="68" pageNumber="69" pagination="21 - 34" publicationUrl="10.1111/j.1096-3642.1851.tb01126.x" refId="B64" refString="Owen, R, 1851b. On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species. Proceedings of the Zoological Society of London 19: 21 - 34, 10.1111/j.1096-3642.1851.tb01126.x" title="On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species." url="10.1111/j.1096-3642.1851.tb01126.x" volume="19" year="1851 b">b</bibRefCitation>
|
||
) answered Bowerbank that he had understood that the name
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
was proposed for a bone from the Chalk Formation that he (
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Proceedings of the Geologists Association of London" pageId="68" pageNumber="69" publicationUrl="10.1144/transgslb.6.2.411" refId="B60" refString="Owen, R, 1842. Description of the remains of a bird, tortoise, and lizard from the Chalk of Kent. Transactions of the Geological Society of London, series 2, 6: 411-413., 10.1144/transgslb.6.2.411" title="Description of the remains of a bird, tortoise, and lizard from the Chalk of Kent. Transactions of the Geological Society of London, series 2, 6: 411 - 413." url="10.1144/transgslb.6.2.411" year="1842">Owen 1842</bibRefCitation>
|
||
) had previously described as avian. He also pointed out that, among the material described and referred as
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Bowerbank, JS" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="65" pageNumber="66" pagination="7 - 8" publicationUrl="10.1144/GSL.JGS.1846.002.01-02.05" refId="B11" refString="Bowerbank, JS, 1846. On a new species of pterodactyl found in the Upper Chalk of Kent. Quarterly Journal of the Geological Society 2: 7 - 8, 10.1144/GSL.JGS.1846.002.01-02.05" title="On a new species of pterodactyl found in the Upper Chalk of Kent." url="10.1144/GSL.JGS.1846.002.01-02.05" volume="2" year="1846">Bowerbank (1846)</bibRefCitation>
|
||
, there were at least two individuals, of very different size, the smaller one (NHMUK PV 39412) being osteologically mature (based on the fusion between scapula and coracoid), and the other one much larger.
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Owen (1851a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="68" pageNumber="69" pagination="21 - 34" publicationUrl="10.1111/j.1096-3642.1851.tb01126.x" refId="B64" refString="Owen, R, 1851b. On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species. Proceedings of the Zoological Society of London 19: 21 - 34, 10.1111/j.1096-3642.1851.tb01126.x" title="On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species." url="10.1111/j.1096-3642.1851.tb01126.x" volume="19" year="1851 b">b</bibRefCitation>
|
||
) assumed the larger individual to be the one referred as
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
and thus designated
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="conirostris">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus conirostris</emphasis>
|
||
</taxonomicName>
|
||
for the cranial material and the bones associated with it. He also brought up several rules of the British Association Code on which he based his designations, including exceptions to the Law of Priority in relation to inappropriate names (
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Owen 1851a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="68" pageNumber="69" pagination="21 - 34" publicationUrl="10.1111/j.1096-3642.1851.tb01126.x" refId="B64" refString="Owen, R, 1851b. On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species. Proceedings of the Zoological Society of London 19: 21 - 34, 10.1111/j.1096-3642.1851.tb01126.x" title="On a new species of pterodactyle (Pterodactylus compressirostris, Owen) from the Chalk; with some remarks on the nomenclature of the previously described species." url="10.1111/j.1096-3642.1851.tb01126.x" volume="19" year="1851 b">b</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Dayrat, B" editor="Polaszek, A" journalOrPublisher="CRC, London" pageId="65" pageNumber="66" pagination="186 - 239" publicationUrl="10.1201/EBK1420095012-c17" refId="B17" refString="Dayrat, B, 2010. Celebrating 250 dynamic years of nomenclatural debates. In: Polaszek, A, Ed., System Naturae 250: The Linnean Ark. CRC, London: 186 - 239, 10.1201/EBK1420095012-c17" title="Celebrating 250 dynamic years of nomenclatural debates." url="10.1201/EBK1420095012-c17" volumeTitle="System Naturae 250: The Linnean Ark." year="2010">Dayrat 2010</bibRefCitation>
|
||
), but finally accepted the name
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
for the material (
|
||
<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B63" refString="Owen, R, 1851a. A monograph on the fossil Reptilia of the Cretaceous formations. Palaeontographical Society, London, 118 pp." title="A monograph on the fossil Reptilia of the Cretaceous formations." year="1851 a">Owen 1851a</bibRefCitation>
|
||
). The name
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="conirostris">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus conirostris</emphasis>
|
||
</taxonomicName>
|
||
has never been used since, but the question as to which material was the holotype of
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
remained overlooked for several years.
|
||
<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley (1914: 538)</bibRefCitation>
|
||
reviewed the species based only on the cranial material (NHMUK PV 39412). Finally,
|
||
<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer (1978: 57)</bibRefCitation>
|
||
, in his review, designated NHMUK PV 39412 as the lectotype of
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
, citing only the skull material and not the associated bones.
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus giganteus</emphasis>
|
||
</taxonomicName>
|
||
Bowerbank, 1846 and
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="conirostris">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Pterodactylus conirostris</emphasis>
|
||
</taxonomicName>
|
||
Owen, 1850 clearly are objective synonyms because they are founded on the same type specimen, and the former binomen has priority over the latter.
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/22072" pageId="17" pageNumber="18" start="Figure 4" startId="F4">
|
||
<paragraph pageId="17" pageNumber="18">
|
||
<emphasis bold="true" pageId="17" pageNumber="18">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="17" pageNumber="18">Lonchodraco giganteus</emphasis>
|
||
</taxonomicName>
|
||
comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation).
|
||
<emphasis bold="true" pageId="17" pageNumber="18">A-F</emphasis>
|
||
articulated anterior parts of the rostrum and mandible
|
||
<emphasis bold="true" pageId="17" pageNumber="18">A</emphasis>
|
||
right lateral view
|
||
<emphasis bold="true" pageId="17" pageNumber="18">B</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="17" pageNumber="18">C</emphasis>
|
||
left lateral view
|
||
<emphasis bold="true" pageId="17" pageNumber="18">D</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="17" pageNumber="18">E</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" pageId="17" pageNumber="18">F</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="17" pageNumber="18">G</emphasis>
|
||
associated scapulocoracoid in posterior view. Abbreviations:
|
||
<emphasis bold="true" pageId="17" pageNumber="18">ch</emphasis>
|
||
- choanae,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">cor</emphasis>
|
||
- coracoid,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">d</emphasis>
|
||
- dentary,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">dcr</emphasis>
|
||
- dentary crest,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">m</emphasis>
|
||
- maxillae,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">pl</emphasis>
|
||
- palatine,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">pm</emphasis>
|
||
- premaxillae,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">pmcr</emphasis>
|
||
- premaxillaery crest,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">prid</emphasis>
|
||
- palatal ridge,
|
||
<emphasis bold="true" pageId="17" pageNumber="18">sca</emphasis>
|
||
- scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |