295 lines
24 KiB
XML
295 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.857.29011" ID-GBIF-Dataset="2b50a048-8eb3-4ef4-b2ad-fb21cc83c29c" ID-PMC="PMC6606672" ID-Pensoft-Pub="1313-2970-857-85" ID-PubMed="31303806" ID-ZBK="8B50D262C5074697A93466F977031F1D" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-857-85" ModsDocOrigin="ZooKeys 857" ModsDocTitle="The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species" checkinTime="1561532585439" checkinUser="pensoft" docAuthor="Liu, Zhen & Jiang, Shi-hong" docDate="2019" docId="BD3514B353C2BB699AA19ED3F9A03C2D" docLanguage="en" docName="ZooKeys 857: 85-104" docOrigin="ZooKeys 857" docSource="http://dx.doi.org/10.3897/zookeys.857.29011" docTitle="Scutellathous habenularis Liu & Jiang, sp. nov." docType="treatment" docUuid="FBAA77C9-3CBF-4DED-812E-98F9E3D13910" docUuidSource="ZooBank" docVersion="6" lastPageNumber="85" masterDocId="43751E4C815BFFE5FFE8FF9AFFE2FFF2" masterDocTitle="The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species" masterLastPageNumber="104" masterPageNumber="85" pageNumber="85" updateTime="1668167437205" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Scutellathous Kishii, 1955 (Coleoptera, Elateridae, Dendrometrinae) in China, with description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Zhen</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jiang, Shi-hong</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>857</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>85</mods:start>
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<mods:end>104</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.857.29011</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.857.29011</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-857-85</mods:identifier>
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<mods:identifier type="ZBK">8B50D262C5074697A93466F977031F1D</mods:identifier>
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<mods:identifier type="ZooBank">8B50D262C5074697A93466F977031F1D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="158387733" LSID="urn:lsid:zoobank.org:act:FBAA77C9-3CBF-4DED-812E-98F9E3D13910" httpUri="http://treatment.plazi.org/id/BD3514B353C2BB699AA19ED3F9A03C2D" lastPageNumber="85" pageId="0" pageNumber="85">
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<subSubSection pageId="0" pageNumber="85" type="nomenclature">
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<paragraph pageId="0" pageNumber="85">
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<taxonomicName LSID="http://zoobank.org/FBAA77C9-3CBF-4DED-812E-98F9E3D13910" authority="Liu & Jiang" class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis Liu & Jiang</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
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Figs 1, 2, 3, 4, 5, 12
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="type locality">
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<paragraph pageId="0" pageNumber="85">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
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,
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<collectingCountry name="China">China</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="materials_examined">
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<paragraph pageId="0" pageNumber="85">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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<materialsCitation ID-GBIF-Occurrence="2274024754" collectingDate="2012-07-23" collectionCode="MHBU" collectorName="Ji-shan Xu & Ling-xiao Chang & Shenzhen Polytechnic" elevation="2000" latitude="25.98578" location="Gaoligong Mts" longLatPrecision="1" longitude="98.817055" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus>Holotype</typeStatus>
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:
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<specimenCount>♂</specimenCount>
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(
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<collectionCode>MHBU</collectionCode>
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), labels: 1)
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<collectingRegion country="China" name="Yunnan">Yunnan Prov.</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:BD3514B353C2BB699AA19ED3F9A03C2D:6E6C65E8DC589D847B9DCCBE5FFE0C13" latitude="25.98578" longLatPrecision="1" longitude="98.817055" name="Gaoligong Mts" stateProvince="Yunnan">Gaoligong Mts</location>
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(
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
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</quantity>
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,
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<geoCoordinate degrees="25" direction="north" minutes="59" orientation="latitude" precision="1" seconds="8.81" value="25.98578">25°59'8.81"N</geoCoordinate>
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<geoCoordinate degrees="98" direction="east" minutes="49" orientation="longitude" precision="1" seconds="1.40" value="98.817055">98°49'1.40"E</geoCoordinate>
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) (高黎贡山),
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<date value="2012-07-23">
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<collectingDate value="2012-07-23">2012.VII.23</collectingDate>
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</date>
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, leg.
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<collectorName>Ji-shan Xu</collectorName>
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et
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<collectorName>Ling-xiao Chang</collectorName>
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,
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<collectorName>Shenzhen Polytechnic</collectorName>
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; 2
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</materialsCitation>
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)
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<typeStatus>Holotype</typeStatus>
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,
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<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
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sp. nov., Liu et al. 2019; 3) No. 20180353.
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<materialsCitation ID-GBIF-Occurrence="2274024751" collectingDate="2012-07-23" collectionCode="MHBU" collectorName="Ji-shan Xu & Ling-xiao Chang & Shenzhen Polytechnic" elevation="2000" latitude="25.98578" location="Gaoligong Mts" longLatPrecision="1" longitude="98.817055" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus>Paratype</typeStatus>
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:
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<specimenCount type="female">1♀</specimenCount>
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(
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<collectionCode>MHBU</collectionCode>
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), labels: 1)
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<collectingRegion country="China" name="Yunnan">Yunnan Prov.</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:BD3514B353C2BB699AA19ED3F9A03C2D:00F9D92431A2446FAF60C2F87151634C" latitude="25.98578" longLatPrecision="1" longitude="98.817055" name="Gaoligong Mts" stateProvince="Yunnan">Gaoligong Mts</location>
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(
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
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</quantity>
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,
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<geoCoordinate degrees="25" direction="north" minutes="59" orientation="latitude" precision="1" seconds="8.81" value="25.98578">25°59'8.81"N</geoCoordinate>
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<geoCoordinate degrees="98" direction="east" minutes="49" orientation="longitude" precision="1" seconds="1.40" value="98.817055">98°49'1.40"E</geoCoordinate>
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) (高黎贡山),
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<date value="2012-07-23">
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<collectingDate value="2012-07-23">2012.VII.23</collectingDate>
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</date>
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, leg.
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<collectorName>Ji-shan Xu</collectorName>
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et
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<collectorName>Ling-xiao Chang</collectorName>
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,
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<collectorName>Shenzhen Polytechnic</collectorName>
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</materialsCitation>
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; 2)
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<typeStatus>Paratype</typeStatus>
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,
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<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
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sp.nov., Liu et al. 2019; 3) No. 20180354.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="diagnosis">
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<paragraph pageId="0" pageNumber="85">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="85">Body bright red-brown, shiny; anterior edge of head truncate in dorsal view; antennae reaching apices of pronotal hind angles, nearly cylindrical from antennomere 6 onward, attached apico-dorsally, antennomere 3 1.8 times as long as 2 and nearly as long as 4; pronotum with hind angles divergent posterad, disc flat medially with only trace of a glabrous non-furrowed longitudinal line, shiny with umbillicate punctures; scutellar shield 1.1 times longer than wide; aedeagus with penis reaching to apex of parameres, penis gradually narrowed to pointed apex.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
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is similar to
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<taxonomicName lsidName="S. yamashitai" pageId="0" pageNumber="85" rank="species" species="yamashitai">S. yamashitai</taxonomicName>
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<bibRefCitation author="Arimoto, H" journalOrPublisher="Elytra, Tokyo" pageId="0" pageNumber="85" pagination="73 - 78" title="Two new Elaterid beetles (Coleoptera, Elateridae) from Taiwan." volume="20" year="1992">Arimoto 1992</bibRefCitation>
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, but is distinguished by the glabrous non-furrowed longitudinal line throughout the length of the pronotum (with shallow median impression only on basal half of the pronotum in
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<taxonomicName lsidName="S. yamashitai" pageId="0" pageNumber="85" rank="species" species="yamashitai">S. yamashitai</taxonomicName>
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), the obtuse anterior angles of the pronotum (acute in
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<taxonomicName lsidName="S. yamashitai" pageId="0" pageNumber="85" rank="species" species="yamashitai">S. yamashitai</taxonomicName>
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), and the penis reaching beyond parameres (penis shorter than parameres in
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<taxonomicName lsidName="S. yamashitai" pageId="0" pageNumber="85" rank="species" species="yamashitai">S. yamashitai</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="description">
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<paragraph pageId="0" pageNumber="85">Description.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Male (
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<typeStatus>holotype</typeStatus>
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). Body (Fig. 1
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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) length
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.48" unit="mm" value="14.8">14.8 mm</quantity>
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, width
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
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; bright red-brown on head, pronotum (except hind angles), and elytra near scutellar shield; ventral surfaces and legs brown-black, and antennae brown (except antennomere 1-2 brown-black); dorsal pubescence yellow, suberect, shorter, denser, and pointed anterad on pronotum and head, longer, sparser, semi-recumbant, pointed posterad on elytra, ventral pubescence more recumbent and thinner.
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</paragraph>
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<paragraph pageId="0" pageNumber="85">Head. Anterior edge truncate in dorsal view (Fig. 2e), spaces between punctures shiny and 1-2 puncture diameters wide, punctures larger, denser, coarser in deep triangular depression (Fig. 2c); eye semi-spherical, ocular index 71; last segment of maxillary palpus 1.6 times longer than wide; antennae (Fig. 3e), reaching apices of pronotal hind angles, antennomeres 3 to 10 weakly serrate, nearly cylindrical from antennomere 6 onward, attached apico-dorsally, antennomere 1 robust, longest and subclavate, antennomere 2 shortest, obconic, 1.4 times longer than wide, antennomere 3 elongated triangular, 1.8 times as long as 2 and nearly as long as 4, antennomere 11 oblong, 5.0 times longer than wide, 1.1 times longer than antennomere 3, proportions of antennomeres as follows: 100; 52; 93; 95; 95; 94; 82; 96; 87; 82; 98.</paragraph>
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<paragraph pageId="0" pageNumber="85">Thorax. Pronotum (Fig. 2e) nearly 1.2 times longer than wide, parallel-sided in dorsal view, except abruptly concave at anterior end and strongly concave before hind angles, widest at hind angles; disc flat medially with only trace of a glabrous non-furrowed longitudinal line, shiny with umbillicate punctures, spaces between punctures 1 to 2 puncture diameters wide medially (Fig. 3a), strongly umbillicate with interspaces 0.3 puncture diameter wide laterally and posteriorly; hypomera (Fig. 2d) with spaces between punctures narrower than half puncture diameter wide; hind angles (Fig. 2e) divergent, apex upheaved and obtuse, with carina extending only to basal 1/5 of pronotum along sides; sublateral incisions small. Prosternum shiny and sparsely, irregularly punctate, interspaces 1 to 4 times puncture diameters wide; anterior lobe (Fig. 2a) 2.3 times wider than long, with sparse, fine punctures. Prosternal process gradually narrowed to pointed apex. Meso- and meta-ventrites with stronger and denser punctures than those on prosternum, smooth. Metaventrite furrowed medially on anterior 3/4.</paragraph>
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<paragraph pageId="0" pageNumber="85">Scutellar shield. 1.1 times longer than wide (Fig. 2f), widest posterad, narrowing anterad gradually at sides, hind margin arched, weakly emarginate anteriorly; disc convex with umbillicate punctures medially, spaces between punctures 2-3 puncture diameters wide, strongly rugose-punctate near edges, with short yellow, outwardly-oriented pubescence laterally.</paragraph>
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<paragraph pageId="0" pageNumber="85">Elytra. Slender (Fig. 1a), 2.7 times longer than wide, 2.8 times longer and 1.1 times wider than prothorax respectively, widest at anterior one-fifth, parallel-sided, gradually narrowing to apex from midlength (Fig. 3b) with punctate striae, interpuncture spaces about 2-3 puncture diameters wide, interstriae slightly elevated, mostly smooth with smaller, shallower and sparser punctures than punctures within striae, interpuncture spaces about 3 puncture diameters wide, weakly and transversely rugulose anteriorly.</paragraph>
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<paragraph pageId="0" pageNumber="85">Legs. Slender (Fig. 3f); tarsomere 3 with a larger lobe beneath than tarsomeres 2, tarsomere 4 shortest; metacoxal plate (Fig. 3c) with mesal parts subparallel-sided, then abruptly strongly narrowed into at lateral one-fourth.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Abdomen. Surfaces of sternites
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<normalizedToken originalValue="III–VII">III-VII</normalizedToken>
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like metaventrite, with punctures and pubescence more regular and evenly distributed laterally; sternite VII (Fig. 3d) broadly rounded, elongate, 1.7 times wider than long, punctures becoming bigger posteriorly.
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</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Genitalia. Aedeagus (Fig. 4
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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) slender, weakly sclerotized; penis reaching to apex of parameres, gradually narrowed to pointed apex; parameres thin, strongly sinuate laterally at midlength, tapered to rounded apex.
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</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Female. Like male (Fig. 5a), except larger (length:
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.59" unit="mm" value="15.9">15.9 mm</quantity>
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, width:
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.9" unit="mm" value="3.9">3.9 mm</quantity>
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), antennae shorter (short of pronotal hind angle apices by half length of last antennomere) and pronotum strongly arched laterally and narrowed anteriorly in dorsal view. Bursa copulatrix (Fig. 5b) weakly sclerotized (it dissolved after being macerated in 10% NaOH for 10 hours) with a circular thorny line and without thorny plates. Ovipositor (Fig. 5c) 2.3 times longer than wide, with short styli.
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</paragraph>
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<paragraph pageId="0" pageNumber="85">Larva. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="etymology">
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<paragraph pageId="0" pageNumber="85">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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The name of the new species is derived from the Latin "
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<taxonomicName lsidName="habenularis" pageId="0" pageNumber="85" rank="species" species="habenularis">habenularis</taxonomicName>
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" (Latin for
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<normalizedToken originalValue="“strip”">"strip"</normalizedToken>
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) referring to its hind coxae abruptly and strongly narrowed into a strip at lateral one-fourth part.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="distribution">
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<paragraph pageId="0" pageNumber="85">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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<collectingCountry name="China">China</collectingCountry>
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:
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<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
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(Fig. 12).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="biology_ecology">
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<paragraph pageId="0" pageNumber="85">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Collected around the elevation of
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</quantity>
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in subtropical evergreen forest.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="85" type="remarks">
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<paragraph pageId="0" pageNumber="85">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="85">
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Its bursa copulatrix resembles
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<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
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in sharing a circular thorny line, but differs in lacking thorny plates. Other characters (carina on hind angles extending only basal 1/5 of pronotal length; antennomere 3 2.1 times as long as 2; shape of male genitalia; shape of female pronotum; only tarsomeres 2 and 3 lobed beneath) are consistent with the diagnosis of genus
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<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
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. Structures of the bursa copulatrix are diverse within
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<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
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species - from five thorny plates (
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<taxonomicName lsidName="S. comes" pageId="0" pageNumber="85" rank="species" species="comes">S. comes</taxonomicName>
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,
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<taxonomicName lsidName="S. porrecticollis" pageId="0" pageNumber="85" rank="species" species="porrecticollis">S. porrecticollis</taxonomicName>
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,
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<taxonomicName lsidName="S. sasajii" pageId="0" pageNumber="85" rank="species" species="sasajii">S. sasajii</taxonomicName>
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,
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<taxonomicName lsidName="S. shikokuanus" pageId="0" pageNumber="85" rank="species" species="shikokuanus">S. shikokuanus</taxonomicName>
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) or four (
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<taxonomicName lsidName="S. seinoi" pageId="0" pageNumber="85" rank="species" species="seinoi">S. seinoi</taxonomicName>
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) thorny plates and without thorny line to no thorny plates but with thorny line in
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<taxonomicName lsidName="S. habenularis" pageId="0" pageNumber="85" rank="species" species="habenularis">S. habenularis</taxonomicName>
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. The bursa copulatrix structure of
|
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<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
|
||
(a circular thorny line and two thorny plates) falls within the range of variation observed in
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
|
||
and does not support distinction of
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
|
||
from
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Scutellathous</taxonomicName>
|
||
. Unfortunately, male genitalia of
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
|
||
is unknown. Further phylogenetic study is needed to understand the monophyly of
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Ohirathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ohirathous" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Ohirathous</taxonomicName>
|
||
and the relationship between these two genera.
|
||
</paragraph>
|
||
<caption pageId="0" pageNumber="85">
|
||
<paragraph pageId="0" pageNumber="85">
|
||
Figure 1. Habitus of
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
|
||
sp. nov.,
|
||
<typeStatus>holotype</typeStatus>
|
||
, male a dorsal view b ventral view c lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="85">
|
||
<paragraph pageId="0" pageNumber="85">
|
||
Figure 2.
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
|
||
sp. nov.,
|
||
<typeStatus>holotype</typeStatus>
|
||
, male a head, ventral view b head, antero-dorsal view c head, dorsal view d prothorax, ventral view e pronotum, dorsal view f scutellar shield, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="85">
|
||
<paragraph pageId="0" pageNumber="85">
|
||
Figure 3.
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
|
||
sp. nov.,
|
||
<typeStatus>holotype</typeStatus>
|
||
, male surface of pronotum, dorsal view b surface of elytra, dorsal view c hind coxae, ventral view d sternite VII, ventral view e antenna, lateral view f hind leg, anterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="85">
|
||
<paragraph pageId="0" pageNumber="85">
|
||
Figure 4. Aedeagus of
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
|
||
sp. nov.,
|
||
<typeStatus>holotype</typeStatus>
|
||
, male a dorsal view b ventral view c lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="85">
|
||
<paragraph pageId="0" pageNumber="85">
|
||
Figure 5.
|
||
<taxonomicName class="Insecta" family="Elateridae" genus="Scutellathous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scutellathous habenularis" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="habenularis">Scutellathous habenularis</taxonomicName>
|
||
sp. nov.,
|
||
<typeStatus>paratype</typeStatus>
|
||
, female a habitus, dorsal view b bursa copulatrix (before dissolved macerating in 10% NaOH for 10 hours) c ovipositor, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |