171 lines
19 KiB
XML
171 lines
19 KiB
XML
<document id="A06301719ED78EA4239D395DB753BE94" ID-DOI="10.11646/zootaxa.2150.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5311432" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629626226088" checkinUser="felipe" docAuthor="Kruse, James J. & Powell, Jerry A." docDate="2009" docId="BD2F87FBFFA8F419FF70FF74FAC7F9B1" docLanguage="en" docName="zt02150p078.pdf" docOrigin="Zootaxa 2150 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2150.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Sparganothoides polymitariana Kruse and Powell 2009, new species" docType="treatment" docVersion="5" lastPageNumber="53" masterDocId="4116FF83FF9CF42DFFE7FFE3FFE5FFDA" masterDocTitle="Systematics of Sparganothoides Lambert and Powell, 1986 (Lepidoptera: Tortricidae: Sparganothini)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="53" updateTime="1699085985238" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="91F1FB4AF93E50E971550E72A93CA9CF">
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<mods:title id="BD6569F78A8F056585A825EA4145B1F8">Systematics of Sparganothoides Lambert and Powell, 1986 (Lepidoptera: Tortricidae: Sparganothini)</mods:title>
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<mods:namePart id="7EFD9AD93762D0A5FF4BEBFCF3FB02EC">Kruse, James J.</mods:namePart>
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<mods:roleTerm id="F58A7B6848761E810CD2FE36B22DFA42">Author</mods:roleTerm>
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<mods:namePart id="C06F8D6CC08E993145CE9C0C69E573ED">Powell, Jerry A.</mods:namePart>
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<mods:typeOfResource id="FA1D21E4281C95F70F1D755AE0B1A2C3">text</mods:typeOfResource>
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<mods:title id="CC127D4F0DA09B0E67484918380D7914">Zootaxa</mods:title>
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<mods:part id="F31E622E08AE86D7775D0CB27E3CDFB9">
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<mods:date id="0057CFF2284C48DF28BC9DF9E59AC01D">2009</mods:date>
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<mods:number id="43DBE72C970CB70971C7F184225290C8">2009-07-06</mods:number>
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<mods:number id="08F8B4A148E612F137F9883EBAC74C55">2150</mods:number>
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<mods:url id="E0A7A0FEFA3FB5BE41B35534156D58CB">https://biotaxa.org/Zootaxa/article/view/zootaxa.2150.1.1</mods:url>
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<mods:classification id="1F7694876A3921B0B45717F52EFDA956">journal article</mods:classification>
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<mods:identifier id="F06B7DF5797F96A5D86F5C03FC6E3F38" type="DOI">10.11646/zootaxa.2150.1.1</mods:identifier>
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<mods:identifier id="292890EE3C39E46575F0607043CA3441" type="ISSN">11755334</mods:identifier>
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<treatment id="BD2F87FBFFA8F419FF70FF74FAC7F9B1" ID-DOI="http://doi.org/10.5281/zenodo.5318895" ID-Zenodo-Dep="5318895" LSID="urn:lsid:plazi:treatment:BD2F87FBFFA8F419FF70FF74FAC7F9B1" httpUri="http://treatment.plazi.org/id/BD2F87FBFFA8F419FF70FF74FAC7F9B1" lastPageNumber="53" pageId="52" pageNumber="53">
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<subSubSection id="7D9C6566FFA8F419FF70FF74FC27FF6B" box="[151,962,151,177]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="353936EDFFA8F419FF70FF74FC27FF6B" blockId="52.[151,962,151,212]" box="[151,962,151,177]" pageId="52" pageNumber="53">
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<heading id="6E718181FFA8F419FF70FF74FC27FF6B" bold="true" box="[151,962,151,177]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<emphasis id="07F2EAFFFFA8F419FF70FF74FC27FF6B" bold="true" box="[151,962,151,177]" pageId="52" pageNumber="53">
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28.
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<taxonomicName id="F2864D6EFFA8F419FF24FF74FCC0FF6B" ID-CoL="B42YW" authority="Kruse and Powell" authorityName="Kruse and Powell" authorityYear="2009" box="[195,805,151,177]" class="Insecta" family="Tortricidae" genus="Sparganothoides" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="polymitariana" status="sp. nov.">
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<emphasis id="07F2EAFFFFA8F419FF24FF74FDDBFF6B" bold="true" box="[195,574,151,177]" italics="true" pageId="52" pageNumber="53">Sparganothoides polymitariana</emphasis>
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Kruse and Powell
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</taxonomicName>
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,
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<taxonomicNameLabel id="1CC15784FFA8F419FCD5FF74FC27FF6B" box="[818,962,151,177]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7D9C6566FFA8F419FF70FF59FEC4FF0E" box="[151,289,186,212]" pageId="52" pageNumber="53" type="description">
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<paragraph id="353936EDFFA8F419FF70FF59FEC4FF0E" blockId="52.[151,962,151,212]" box="[151,289,186,212]" pageId="52" pageNumber="53">
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<figureCitation id="ADBD2A68FFA8F419FF70FF59FF1DFF0E" box="[151,248,186,212]" captionStart="FIGURES 31–36" captionStartId="48.[151,269,1958,1982]" captionTargetBox="[173,1424,303,1930]" captionTargetId="figure-67@48.[151,1436,271,1934]" captionTargetPageId="48" captionText="FIGURES 31–36. Male genitalia of Sparganothoides; valvae spread, aedeagus removed. 31. S. albescens. 32. S. torusana. 33. S. prolesana. 34. S. polymitariana. 35. S. silaceana. 36. S. aciculana." figureDoi="http://doi.org/10.5281/zenodo.5311452" httpUri="https://zenodo.org/record/5311452/files/figure.png" pageId="52" pageNumber="53">Figs. 34</figureCitation>
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,
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<figureCitation id="ADBD2A68FFA8F419FEE3FF59FEC4FF0E" box="[260,289,186,212]" captionStart="FIGURES 79–94" captionStartId="61.[151,269,1692,1716]" captionTargetBox="[175,1419,186,1665]" captionTargetId="figure-172@61.[175,1419,180,1674]" captionTargetPageId="61" captionText="FIGURES 79–94. Adults of Sparganothoides. 79. S. probolosana (paratype: Mexico: Durango: 24 mi W La Ciudad). 80. S. licrosana (holotype). 81. S. canorisana (paratype: Mexico: Tamaulipas: Gómez Farias, 1000m, 29–31.vii.1988, V. Becker and M. Solis). 82. S. capitiornata (paratype). 83. S. arcuatana (paratype, copper form: Mexico: Veracruz: 22 km W Cd. Mendoza, 2150m, 13.viii.1987, J. Brown and J. Powell, JAP lot 87H7.6, emerged 12.i.1988). 84. S. arcuatana (paratype, gold form: Mexico: Jalisco: Nevado de Colima, 10.7 mi. N Hwy 54, 8200', 17.ix.1986, N. Bloomfield). 85. S. lentiginosana (USA: FL: Escambia County, Pensacola, 3.v.1962, S. Hills). 86. S. morata (Venezuela: Aragua, Rancho Grande, 1100', 21–25.i.1966, S. and W. Duckworth). 87. S. albescens (holotype). 88. S. torusana (holotype). 89. S. prolesana (holotype). 90. S. polymitariana (holotype). 91. S. silaceana (paratype: Costa Rica: Alajuela: Volcán Poás, 2350m, 25.viii.1987, J. Brown and J. Powell). 92. S. aciculana (holotype). 93. S. broccusana (holotype). 94. S. cornutana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5345532" httpUri="https://zenodo.org/record/5345532/files/figure.png" pageId="52" pageNumber="53">90</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7D9C6566FFA8F419FF70FEE4FB23FE92" pageId="52" pageNumber="53" type="diagnosis">
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<paragraph id="353936EDFFA8F419FF70FEE4FB23FE92" blockId="52.[151,1437,263,1643]" pageId="52" pageNumber="53">
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<emphasis id="07F2EAFFFFA8F419FF70FEE4FEEAFEFB" bold="true" box="[151,271,263,289]" pageId="52" pageNumber="53">Diagnosis</emphasis>
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. This is the only known species of
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<taxonomicName id="F2864D6EFFA8F419FD21FEEBFC6EFEFB" authorityName="Lambert and Powell" authorityYear="1986" box="[710,907,264,289]" class="Insecta" family="Tortricidae" genus="Sparganothoides" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="07F2EAFFFFA8F419FD21FEEBFC6EFEFB" box="[710,907,264,289]" italics="true" pageId="52" pageNumber="53">Sparganothoides</emphasis>
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</taxonomicName>
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in which the enlarged apices of the socius/ gnathos arms are strongly asymmetrically bilobed with the longer lobe bilobed secondarily.
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</paragraph>
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</subSubSection>
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<subSubSection id="7D9C6566FFA8F419FF21FEB6FE4EFA87" pageId="52" pageNumber="53" type="description">
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||
<paragraph id="353936EDFFA8F419FF21FEB6FA6FFAEC" blockId="52.[151,1437,263,1643]" pageId="52" pageNumber="53">
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||
<emphasis id="07F2EAFFFFA8F419FF21FEB6FEB2FEB5" bold="true" box="[198,343,341,367]" pageId="52" pageNumber="53">Description</emphasis>
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.
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<emphasis id="07F2EAFFFFA8F419FE8CFEB6FE4FFEB5" bold="true" box="[363,426,341,367]" pageId="52" pageNumber="53">Male</emphasis>
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||
.
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<emphasis id="07F2EAFFFFA8F419FE5AFEB6FE1AFEB5" bold="true" box="[445,511,341,367]" pageId="52" pageNumber="53">Head</emphasis>
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||
: Frons brownish yellow, smooth scaled, deeply concave; vertex roughened laterally, yellowish brown; two large exoskeletal protuberances between mesal-posterior margins of compound eyes, angled toward middle; a laterally broad, dorsoventrally flattened, bilobed protuberance between antennae, projected anteriorly; broad area of densely packed, short, yellowish brown scales between anterior and posterior protuberances; short, erect, yellowish brown scales between posterior protuberances. Labial palpus yellowish brown mesally, yellowish brown laterally. Antennal scaling yellowish brown.
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<emphasis id="07F2EAFFFFA8F419FF70FDDEFF11FD8D" bold="true" box="[151,244,573,599]" pageId="52" pageNumber="53">Thorax</emphasis>
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: Dorsum smooth scaled, yellowish brown with grayish tinge on tegulae, scales long and dense, parted at center, obscuring wide median channel in notum filled with dense row of short white scales mesally and long thin white scales laterally; tegula with clump of small orange or brown pointed scales at apex giving tegula truncate appearance. Forewing length
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||
<quantity id="F27E9B08FFA8F419FD57FD52FCE9FD11" box="[688,780,689,715]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="52" pageNumber="53" unit="mm" value="7.4">7.4 mm</quantity>
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(n = 1). Forewing costal fold extending less than onehalf wing length (dark grayish brown scales within fold); forewing ground color yellowish brown with grayish tinge basally; ill-defined orange and dark brown transverse striae throughout; five dark brown spots along costa beyond costal fold; distinct, broad, grayish brown pretornal patch from discal cell dorsad, lined with orange and brown scales on basal margin creating sharply contrasting border; indistinct dark brown transverse strigulae in subterminal and terminal areas. Fringe yellowish brown. Hindwing yellowish gray basally and near anal angle; gray beyond, with very faint gray transverse striae visible only in yellowish gray portion near anal angle; patch of gray scales on anal angle.
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<emphasis id="07F2EAFFFFA8F419FCBFFC5CFC37FC03" bold="true" box="[856,978,959,985]" pageId="52" pageNumber="53">Abdomen</emphasis>
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: Genitalia (
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<figureCitation id="ADBD2A68FFA8F419FBB8FC23FB52FC00" box="[1119,1207,960,986]" captionStart="FIGURES 31–36" captionStartId="48.[151,269,1958,1982]" captionTargetBox="[173,1424,303,1930]" captionTargetId="figure-67@48.[151,1436,271,1934]" captionTargetPageId="48" captionText="FIGURES 31–36. Male genitalia of Sparganothoides; valvae spread, aedeagus removed. 31. S. albescens. 32. S. torusana. 33. S. prolesana. 34. S. polymitariana. 35. S. silaceana. 36. S. aciculana." figureDoi="http://doi.org/10.5281/zenodo.5311452" httpUri="https://zenodo.org/record/5311452/files/figure.png" pageId="52" pageNumber="53">Fig. 34</figureCitation>
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; EME slide #5613;
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<collectingCountry id="4D91767DFFA8F419FF70FC05FEF9FBDA" box="[151,284,998,1024]" name="Costa Rica" pageId="52" pageNumber="53">Costa Rica</collectingCountry>
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,
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<collectingRegion id="F742F80FFFA8F419FECBFC05FE55FBDA" box="[300,432,998,1024]" country="Costa Rica" name="Puntarenas" pageId="52" pageNumber="53">Puntarenas</collectingRegion>
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||
, Estación Biol. Las Cruces,
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||
<quantity id="F27E9B08FFA8F419FCEBFC05FCAFFBDA" box="[780,842,998,1024]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="km" value="6.0">6 km</quantity>
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SE San Vito, Río Jaba; n = 1) with uncus long, curved, widened and triangular subapically with a small subrectangular process apically, with long setae dorsally and short setae in two patches ventrolaterally on widened area; tegumen raised, broadly rounded at base of uncus; socii subtriangular posteriorly, secondary arms long, slender, curved before enlarged, strongly asymmetrically bilobed apices with longer lobe bilobed secondarily; transtilla strongly sclerotized, straight, spines short, numerous over most of posterior margin, enlarged anterior process unreinforced at middle; valva elongate, ovate, without a sclerotized crease, costa convex, sacculus convex, extending to apex of valva, pulvinus absent; phallus pistol-shaped, aedeagus shorter than phallobase, gently curved, convexly curved dorsally, tip attenuate apically, attached to juxta by a short process, vesica with dense bundle of>30 cornuti.
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</paragraph>
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<paragraph id="353936EDFFA8F419FF21FAA1FE4EFA87" blockId="52.[151,1437,263,1643]" box="[198,427,1346,1373]" pageId="52" pageNumber="53">
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<emphasis id="07F2EAFFFFA8F419FF21FAA1FEC4FA86" bold="true" box="[198,289,1346,1372]" pageId="52" pageNumber="53">Female</emphasis>
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. Unknown.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="7D9C6566FFA8F419FF21FA8BFDC4FA0A" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="353936EDFFA8F419FF21FA8BFCBEFA70" blockId="52.[151,1437,263,1643]" pageId="52" pageNumber="53">
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<emphasis id="07F2EAFFFFA8F419FF21FA8BFE92FA58" bold="true" box="[198,375,1384,1410]" pageId="52" pageNumber="53">Type material</emphasis>
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||
.
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<materialsCitation id="85EE3CB0FFA8F419FE6EFA8AFCBEFA70" collectingDate="1993-01-20" collectingDateMax="1993-01-21" collectingDateMin="1993-01-20" collectionCode="EME" collectorName="J. Powell" country="Costa Rica" elevation="1150" location="Rio Jaba" municipality="Male" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
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||
<typeStatus id="EA3D884FFFA8F419FE6EFA8AFE1FFA59" box="[393,506,1385,1411]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
|
||
:
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||
<collectingMunicipality id="D55DAC97FFA8F419FDEFFA8AFDADFA59" box="[520,584,1385,1411]" pageId="52" pageNumber="53">Male</collectingMunicipality>
|
||
:
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||
<collectingCountry id="4D91767DFFA8F419FDB1FA8AFCECFA59" box="[598,777,1385,1411]" name="Costa Rica" pageId="52" pageNumber="53">COSTA RICA</collectingCountry>
|
||
:
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||
<collectingRegion id="F742F80FFFA8F419FCF1FA8AFC3FFA59" box="[790,986,1385,1411]" country="Costa Rica" name="Puntarenas" pageId="52" pageNumber="53">PUNTARENAS</collectingRegion>
|
||
: Estación Biol. Las Cruces,
|
||
<locationDeviation id="248B2FCFFFA8F419FAD5FA8AFF1AFA70" location="San Vito" pageId="52" pageNumber="53">
|
||
<quantity id="F27E9B08FFA8F419FAD5FA8AFA8AFA59" box="[1330,1391,1385,1411]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="km" value="6.0">6 km</quantity>
|
||
SE San Vito
|
||
</locationDeviation>
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||
,
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||
<location id="30596036FFA8F419FEECFA73FE91FA70" LSID="urn:lsid:plazi:treatment:BD2F87FBFFA8F419FF70FF74FAC7F9B1:30596036FFA8F419FEECFA73FE91FA70" box="[267,372,1424,1450]" country="Costa Rica" municipality="Male" name="Rio Jaba" pageId="52" pageNumber="53" stateProvince="Puntarenas">Río Jaba</location>
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||
,
|
||
<quantity id="F27E9B08FFA8F419FE98FA73FE3CFA70" box="[383,473,1424,1450]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="52" pageNumber="53" unit="m" value="1150.0">
|
||
<elevation id="BEABD1DEFFA8F419FE98FA73FE3CFA70" box="[383,473,1424,1450]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="52" pageNumber="53" unit="m" value="1150.0">1150 m</elevation>
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||
</quantity>
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||
,
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||
<date id="4138102DFFA8F419FE01FA73FD67FA70" box="[486,642,1424,1450]" pageId="52" pageNumber="53" value="1993-01-20" valueMax="1993-01-21" valueMin="1993-01-20">
|
||
<collectingDate id="517CE9C5FFA8F419FE01FA73FD67FA70" box="[486,642,1424,1450]" pageId="52" pageNumber="53" value="1993-01-20" valueMax="1993-01-21" valueMin="1993-01-20">20–21.i.1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="9873533BFFA8F419FD68FA73FD1FFA70" box="[655,762,1424,1450]" pageId="52" pageNumber="53">J. Powell</collectorName>
|
||
(
|
||
<collectionCode id="5397AE28FFA8F419FCEDFA73FCABFA70" box="[778,846,1424,1450]" pageId="52" pageNumber="53">EME</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="353936EDFFA8F419FF21FA55FDC4FA0A" blockId="52.[151,1437,263,1643]" box="[198,545,1462,1488]" pageId="52" pageNumber="53">
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||
<emphasis id="07F2EAFFFFA8F419FF21FA55FE72FA0A" bold="true" box="[198,407,1462,1488]" pageId="52" pageNumber="53">Immature stages</emphasis>
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||
. Unknown.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="7D9C6566FFA8F419FF21FA3EFC36FA2D" box="[198,979,1501,1527]" pageId="52" pageNumber="53" type="biology_ecology">
|
||
<paragraph id="353936EDFFA8F419FF21FA3EFC36FA2D" blockId="52.[151,1437,263,1643]" box="[198,979,1501,1527]" pageId="52" pageNumber="53">
|
||
<emphasis id="07F2EAFFFFA8F419FF21FA3EFEC7FA2D" bold="true" box="[198,290,1501,1527]" pageId="52" pageNumber="53">Biology</emphasis>
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||
. This species was collected in secondary forest in January.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="7D9C6566FFA8F419FF21F9E0FB80F99F" pageId="52" pageNumber="53" type="discussion">
|
||
<paragraph id="353936EDFFA8F419FF21F9E0FB80F99F" blockId="52.[151,1437,263,1643]" pageId="52" pageNumber="53">
|
||
<emphasis id="07F2EAFFFFA8F419FF21F9E0FEDDF9C7" bold="true" box="[198,312,1539,1565]" pageId="52" pageNumber="53">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="F2864D6EFFA8F419FEADF9E7FD24F9C7" authorityName="Kruse and Powell" authorityYear="2009" box="[330,705,1540,1565]" class="Insecta" family="Tortricidae" genus="Sparganothoides" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="polymitariana">
|
||
<emphasis id="07F2EAFFFFA8F419FEADF9E7FD24F9C7" box="[330,705,1540,1565]" italics="true" pageId="52" pageNumber="53">Sparganothoides polymitariana</emphasis>
|
||
</taxonomicName>
|
||
is one of the most derived species in the genus, possessing several autapomorphies; however, it is convincingly assigned to this species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7D9C6566FFA8F419FF21F9B2FAC7F9B1" box="[198,1314,1617,1643]" pageId="52" pageNumber="53" type="etymology">
|
||
<paragraph id="353936EDFFA8F419FF21F9B2FAC7F9B1" blockId="52.[151,1437,263,1643]" box="[198,1314,1617,1643]" pageId="52" pageNumber="53">
|
||
<emphasis id="07F2EAFFFFA8F419FF21F9B2FEAEF9B1" bold="true" box="[198,331,1617,1643]" pageId="52" pageNumber="53">Etymology</emphasis>
|
||
. The name is derived from the Latin “polymitarius” (= highly wrought or finished).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |