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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.70.762" ID-GBIF-Dataset="b8fceb5a-9f78-4dc4-8fa2-92fe20bb2c42" ID-PMC="PMC3088446" ID-Pensoft-Pub="1313-2970-70-1" ID-PubMed="21594041" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-70-1" ModsDocOrigin="ZooKeys 70" ModsDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an “oceanic” island" checkinTime="1451250880545" checkinUser="pensoft" docAuthor="Rowson, B, Warren, B. H. &amp; Ngereza, C. F." docDate="2010" docId="BCBE16CFA84DEC82BF22C1738F425B7D" docLanguage="en" docName="ZooKeys 70: 1-39" docOrigin="ZooKeys 70" docSource="http://dx.doi.org/10.3897/zookeys.70.762" docTitle="&quot;Gulella&quot; radius Preston 1910" docType="treatment" docVersion="5" lastPageNumber="20" masterDocId="FFB2FFB6591BFFBAFFFF0472FFE6A56D" masterDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island" masterLastPageNumber="39" masterPageNumber="1" pageNumber="20" updateTime="1668164835731" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rowson, B</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Warren, B. H.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ngereza, C. F.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>70</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>39</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.70.762</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.70.762</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-70-1</mods:identifier>
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<treatment ID-GBIF-Taxon="159360819" LSID="urn:lsid:plazi:treatment:BCBE16CFA84DEC82BF22C1738F425B7D" httpUri="http://treatment.plazi.org/id/BCBE16CFA84DEC82BF22C1738F425B7D" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection pageId="19" pageNumber="20" type="nomenclature">
<paragraph pageId="19" pageNumber="20">
28.
<taxonomicName authority="Preston, 1910" authorityName="Preston" authorityYear="1910" lsidName="&quot;Gulella&quot; radius" pageId="19" pageNumber="20" rank="species" species="radius">
<normalizedToken originalValue="“Gulella”">&quot;Gulella&quot;</normalizedToken>
radius (Preston, 1910)
</taxonomicName>
Figs 60-61
</paragraph>
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<subSubSection pageId="19" pageNumber="20" type="reference_group">
<paragraph pageId="19" pageNumber="20">
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ennea radius" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="radius">Ennea radius</taxonomicName>
<bibRefCitation author="Preston, HB" journalOrPublisher="Annals and Magazine of Natural History" pageId="23" pageNumber="24" pagination="526 - 536" title="Additions to the non-marine molluscan fauna of British and German East Africa and Lake Albert Edward" volume="8" year="1910">Preston 1910</bibRefCitation>
: 529; pl. VII, fig. 8
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="notes">
<paragraph pageId="19" pageNumber="20">Notes.</paragraph>
<paragraph pageId="19" pageNumber="20">
As noted by
<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of Conchology" pageId="24" pageNumber="25" pagination="109 - 121" title="New taxa of Gulella L. Pfr. and Ptychotrema Moerch (Mollusca, Streptaxidae) from Eastern Africa." volume="32" year="1985">Verdcourt (1985)</bibRefCitation>
there is either substantial shell variation in the species
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ennea radius" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="radius">Ennea radius</taxonomicName>
Preston, 1910, or it comprises a complex of related taxa. This cannot be resolved without a thorough revision.
<normalizedToken originalValue="Prestons">Preston's</normalizedToken>
(1910) type from the Shimba Hills is said to be 3.25mm high and is strongly acuminate.
<bibRefCitation author="Verdcourt, B" journalOrPublisher="Journal of Conchology" pageId="24" pageNumber="25" pagination="109 - 121" title="New taxa of Gulella L. Pfr. and Ptychotrema Moerch (Mollusca, Streptaxidae) from Eastern Africa." volume="32" year="1985">Verdcourt (1985)</bibRefCitation>
figured a specimen from Diani Beach, Kenya which is much less acuminate but measures 4.23mm (calculated from his drawing). At up to 5.45mm, Pemba specimens are larger still, but resemble the type in being strongly acuminate and with less tumid whorls than the Diani Beach material. Specimens referred to
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gulella radius" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="radius">Gulella radius</taxonomicName>
from Unguja (
<bibRefCitation author="Rowson, B" journalOrPublisher="Journal of Conchology" pageId="24" pageNumber="25" pagination="425 - 466" title="Land molluscs of Zanzibar Island (Unguja), Tanzania with the description of a new species of Gulella (Pulmonata: Streptaxidae)." volume="39" year="2007">Rowson 2007</bibRefCitation>
) are small (to 3.0mm) and not strongly acuminate. The peristomal teeth are in the same basic pattern in each of these populations, but vary in their size and complexity, none being quite as different as some of the other nominal species (discussed in
<bibRefCitation author="Rowson, B" journalOrPublisher="African Invertebrates" pageId="24" pageNumber="25" pagination="21 - 32" title="Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya." volume="48" year="2007">Rowson and Lange 2007</bibRefCitation>
). What does appear relatively constant is the size, shape, and sculpture of Pemba specimens, which occur throughout the island (Table 2). They may form an island taxon worthy of subspecies or species status, which is given consideration in the discussions on endemism in the present paper. Recent systematic work (
<bibRefCitation author="Rowson, B" journalOrPublisher="University of Cardiff, Cardiff, UK" pageId="24" pageNumber="25" title="Systematics and diversity of the Streptaxidae, with particular reference to the East African region. Unpublished PhD thesis" year="2010">Rowson 2010</bibRefCitation>
) indicates
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gulella radius" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="radius">
<normalizedToken originalValue="“Gulella”">&quot;Gulella&quot;</normalizedToken>
radius
</taxonomicName>
does not belong in the genus
<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gulella" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">Gulella</taxonomicName>
L. Pfeiffer and a genus-level revision is in progress.
</paragraph>
</subSubSection>
</treatment>
</document>