1261 lines
165 KiB
XML
1261 lines
165 KiB
XML
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<mods:title id="8CD11ACCA26BBCEA8CC85AB7753533AC">Taxonomic notes on Alpheus inopinatus Holthuis & Gottlieb, 1958 and Alpheus cf lobidens De Haan, 1849 from Kuwait (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<mods:namePart id="8D4420FE5BED29A0C33D4F07BA24DF64">Anker, Arthur</mods:namePart>
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<mods:affiliation id="7F0E567F86DEF5785206C6DB3CFB8938">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiânia, Goiás, Brazil. arthuranker 7 @ gmail; https: // orcid. org / 0000 - 0002 - 5350 - 4267</mods:affiliation>
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<mods:nameIdentifier id="821EE6A2917B891432312402A9D50EB5" type="email">arthuranker7@gmail</mods:nameIdentifier>
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<mods:namePart id="BBA6B867A358FBDD3A756401E2667A02">Al-Kandari, Manal</mods:namePart>
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<mods:affiliation id="F4E8E176600C68AC8EA0DE2668E387EC">Ecosystem-Based Management of Marine Resources, Environment and Life Sciences Research Center, Kuwait Institute of Scientific Research, Hamad Al-Mubarak Street, Building 900004, Area 1, Raas Salmiya, Kuwait.</mods:affiliation>
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<mods:namePart id="78378A09D9859446A5DACD549020CF6C">Grave, Sammy De</mods:namePart>
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<taxonomicName id="F3D075A6FFF8FFBFFF2FFDF920C3D51B" ID-CoL="85LK3" authority="Holthuis & Gottlieb, 1958" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[151,717,546,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="inopinatus">
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<emphasis id="06A4D237FFF8FFBFFF2FFDF9238DD51A" bold="true" box="[151,387,546,573]" italics="true" pageId="1" pageNumber="190">Alpheus inopinatus</emphasis>
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<bibRefCitation id="504173D4FFF8FFBFFE32FDF920C3D51B" author="Holthuis, L. B. & Gottlieb, E." box="[394,717,546,572]" pageId="1" pageNumber="190" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb, 1958</bibRefCitation>
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<paragraph id="346F0E25FFF8FFBFFF2FFD9C2349D546" blockId="1.[151,717,546,609]" box="[151,327,583,609]" pageId="1" pageNumber="190">
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(
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<figureCitation id="ACEB12A0FFF8FFBFFF27FD9C22E3D546" box="[159,237,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1547,1571]" captionTargetBox="[151,1435,179,1521]" captionTargetId="figure-18@2.[151,1435,179,1521]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus inopinatus Holthuis & Gottlieb, 1958: male from Kuwait, OUMNH.ZC.2020-01-001 [fcn KUW-011], A—dorsal view; B—lateral view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407194" httpUri="https://zenodo.org/record/4407194/files/figure.png" pageId="1" pageNumber="190">Figs. 1</figureCitation>
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,
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<figureCitation id="ACEB12A0FFF8FFBFFF43FD9C2309D546" box="[251,263,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="1" pageNumber="190">3</figureCitation>
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A–C)
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<treatmentCitation id="B5712834FFF8FFBFFF2FFD502085D584" author="Holthuis, L. B. & Gottlieb, E." box="[151,651,651,675]" page="42" pageId="1" pageNumber="190" year="1958">
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<emphasis id="06A4D237FFF8FFBFFF2FFD502354D584" box="[151,346,651,675]" italics="true" pageId="1" pageNumber="190">Alpheus inopinatus</emphasis>
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<bibRefCitation id="504173D4FFF8FFBFFED8FD502085D584" author="Holthuis, L. B. & Gottlieb, E." box="[352,651,651,675]" pageId="1" pageNumber="190" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb 1958: 42</bibRefCitation>
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, figs. 8, 9;
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<treatmentCitation id="B5712834FFF8FFBFFD41FD50264CD584" author="Lewinsohn, C. & Holthuis, L. B." box="[761,1090,651,675]" page="51" pageId="1" pageNumber="190" year="1964">
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<bibRefCitation id="504173D4FFF8FFBFFD41FD50264CD584" author="Lewinsohn, C. & Holthuis, L. B." box="[761,1090,651,675]" pageId="1" pageNumber="190" pagination="45 - 63" refId="ref6559" refString="Lewinsohn, C. & Holthuis, L. B. (1964) New records of decapod Crustacea from the Mediterranean coast of Israel and the eastern Mediterranean. Zoologische Mededelingen, 40, 45 - 63." type="journal article" year="1964">Lewinsohn & Holthuis 1964: 51</bibRefCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFBF4FD502757D584" author="Tirmizi, N. M. & Kazmi, M. A." box="[1100,1369,651,675]" page="99" pageId="1" pageNumber="190" year="1969">
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<bibRefCitation id="504173D4FFF8FFBFFBF4FD502757D584" author="Tirmizi, N. M. & Kazmi, M. A." box="[1100,1369,651,675]" pageId="1" pageNumber="190" pagination="99 - 100" refId="ref6661" refString="Tirmizi, N. M. & Kazmi, M. A. (1969) The occurrence of Alpheus inopinatus Holthuis & Gottlieb, 1958 in the Arabian Sea (Decapoda, Alpheidae). Crustaceana, 16, 99 - 100. https: // doi. org / 10.1163 / 156854068 X 00250" type="journal article" year="1969">Tirmizi & Kazmi 1969: 99</bibRefCitation>
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(379), fig. 1a;
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<treatmentCitation id="B5712834FFF8FFBFFEABFD712011D5E5" author="Karim, S. I. & Hafizur, R." box="[275,543,682,706]" page="26" pageId="1" pageNumber="190" year="1974">
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<bibRefCitation id="504173D4FFF8FFBFFEABFD712011D5E5" author="Karim, S. I. & Hafizur, R." box="[275,543,682,706]" pageId="1" pageNumber="190" pagination="25 - 28" refId="ref6366" refString="Karim, S. I. & Hafizur, R. (1974) On some Carides of Karachi coasts. Records of Zoological Survey, 6, 25 - 28, pls. 1 - 4." type="journal article" year="1974">Karim & Hafizur 1974: 26</bibRefCitation>
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</treatmentCitation>
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, pl. 4, figs. 1–4b;
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<treatmentCitation id="B5712834FFF8FFBFFD68FD7121E0D5E5" author="Banner, D. M. & Banner, A. H." box="[720,1006,682,706]" page="223" pageId="1" pageNumber="190" year="1978">
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<bibRefCitation id="504173D4FFF8FFBFFD68FD7121E0D5E5" author="Banner, D. M. & Banner, A. H." box="[720,1006,682,706]" pageId="1" pageNumber="190" pagination="215 - 257" refId="ref5338" refString="Banner, D. M. & Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner & Banner 1978: 223</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFC4FFD712704D5E5" author="Kazmi, M. A. & Kazmi, Q. B." box="[1015,1290,682,706]" page="153" pageId="1" pageNumber="190" year="1979">
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<bibRefCitation id="504173D4FFF8FFBFFC4FFD712704D5E5" author="Kazmi, M. A. & Kazmi, Q. B." box="[1015,1290,682,706]" pageId="1" pageNumber="190" pagination="151 - 157" refId="ref6404" refString="Kazmi, M. A. & Kazmi, Q. B. (1979) A check-list of marine caridean prawns of Pakistan. Biologia, 25, 151 - 157." type="journal article" year="1979">Kazmi & Kazmi 1979: 153</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFAABFD7122FED5C7" author="Barkati, S." page="355" pageId="1" pageNumber="190" year="1980">
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<bibRefCitation id="504173D4FFF8FFBFFAABFD7122FED5C7" author="Barkati, S." pageId="1" pageNumber="190" pagination="355 - 359" refId="ref5486" refString="Barkati, S. (1980) Sex ratio, fecundity and larval development of Alpheus inopinatus Holthuis and Gottlieb, 1958 (Crustacea, Decapoda, Caridea). Proceedings of the First Pakistan Congress of Zoology, B, 355 - 359." type="book chapter" year="1980">Barkati 1980: 355</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFF43FD1223B2D5C7" author="Kocatas, A." box="[251,444,712,737]" page="162" pageId="1" pageNumber="190" year="1981">
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<bibRefCitation id="504173D4FFF8FFBFFF43FD1223B2D5C7" author="Kocatas, A." box="[251,444,712,737]" pageId="1" pageNumber="190" pagination="161 - 162" refId="ref6437" refString="Kocatas, A. (1981) Liste preliminaires et repartition des crustaces decapodes des eaux turques. Rapports de la Commission Internationale de la Mer Mediterranee, 27, 161 - 162." type="journal article" year="1981">Kocataş 1981: 162</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFE7FFD1220E7D5C6" author="Banner, D. M. & Banner, A. H." box="[455,745,713,737]" page="241" pageId="1" pageNumber="190" year="1982">
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<bibRefCitation id="504173D4FFF8FFBFFE7FFD1220E7D5C6" author="Banner, D. M. & Banner, A. H." box="[455,745,713,737]" pageId="1" pageNumber="190" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982: 241</bibRefCitation>
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</treatmentCitation>
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, fig. 75;
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<treatmentCitation id="B5712834FFF8FFBFFCFCFD132676D5C6" author="Lewinsohn, C. & Galil, B." box="[836,1144,712,737]" page="208" pageId="1" pageNumber="190" year="1982">
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<bibRefCitation id="504173D4FFF8FFBFFCFCFD132676D5C6" author="Lewinsohn, C. & Galil, B." box="[836,1144,712,737]" pageId="1" pageNumber="190" pagination="207 - 210" refId="ref6514" refString="Lewinsohn, C. & Galil, B. (1982). Notes on species of Alpheus (Crustacea Decapoda) from the Mediterranean coast of Israel. Quaderni del Laboratorio di Tecnologia della Pesca, Ancona, 3, 207 - 210." type="journal article" year="1982">Lewinsohn & Galil 1982: 208</bibRefCitation>
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;
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<treatmentCitation id="B5712834FFF8FFBFFB39FD13276DD5C6" author="Afzal, M. & Javed, A. & Barkati, S." box="[1153,1379,712,737]" page="339" pageId="1" pageNumber="190" year="1986">
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<bibRefCitation id="504173D4FFF8FFBFFB39FD13276DD5C6" author="Afzal, M. & Javed, A. & Barkati, S." box="[1153,1379,712,737]" pageId="1" pageNumber="190" pagination="337 - 340" refId="ref4790" refString="Afzal, M., Javed, A. & Barkati, S. (1986) Checklist, synonymy and key to the genera and species of alpheid shrimps of Karachi coast. Pakistan Journal of Agriculture Research, 7, 337 - 340." type="journal article" year="1986">
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Afzal
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<emphasis id="06A4D237FFF8FFBFFB78FD1226FAD5C7" box="[1216,1268,712,736]" italics="true" pageId="1" pageNumber="190">et al.</emphasis>
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1986: 339
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;
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<treatmentCitation id="B5712834FFF8FFBFFAD6FD12232DD5D8" author="Jeng, M. S." page="13" pageId="1" pageNumber="190" year="1997">
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<bibRefCitation id="504173D4FFF8FFBFFAD6FD12232DD5D8" author="Jeng, M. S." pageId="1" pageNumber="190" pagination="1 - 66" refId="ref6296" refString="Jeng, M. S. (1997) Studies on the land and aquatic decapod crustacean fauna of the Kenting National Park (II). Conservation Research Report, 96, 1 - 66." type="journal article" year="1997">Jeng 1997: 13</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="B5712834FFF8FFBFFE96FD3C2019D5D8" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[302,535,743,767]" page="343" pageId="1" pageNumber="190" year="2020">
|
||
<bibRefCitation id="504173D4FFF8FFBFFE96FD3C2019D5D8" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[302,535,743,767]" pageId="1" pageNumber="190" pagination="337 - 354" refId="ref5531" refString="Cunha, A. M., Terossi, M., Mantelatto, F. M. & Almeida, A. O. (2020) Delimiting the snapping shrimp Alpheus lobidens De Haan, 1849 (Caridea: Alpheidae) based on morphological and molecular data. Zootaxa, 4718 (3), 337 - 354. https: // doi. org / 10.11646 / zootaxa. 4718.3.3" type="journal article" year="2020">
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFE96FD3C237ED5D8" box="[302,368,743,767]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="190">Cunha</collectingCountry>
|
||
<emphasis id="06A4D237FFF8FFBFFECEFD3323ADD5D8" box="[374,419,743,767]" italics="true" pageId="1" pageNumber="190">et al</emphasis>
|
||
. 2020: 343
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 351, fig 5C–D, 6C–D, 7C–D.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFF8FFBFFF2FFCDD2645D41A" blockId="1.[151,1437,651,859]" pageId="1" pageNumber="190">
|
||
<treatmentCitationGroup id="14C0290BFFF8FFBFFF2FFCDD2645D41A" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF2FFCDD2343D439" box="[151,333,774,798]" italics="true" pageId="1" pageNumber="190">
|
||
<taxonomicName id="F3D075A6FFF8FFBFFF2FFCDD2347D439" ID-CoL="84W8B" authorityName="sensu De Haan" authorityYear="1849" box="[151,329,774,798]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="lobidens">Alpheus lobidens</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
—
|
||
<treatmentCitation id="B5712834FFF8FFBFFECDFCDD2010D439" author="Jones & D. A" box="[373,542,774,798]" page="152" pageId="1" pageNumber="190" year="1986">
|
||
<bibRefCitation id="504173D4FFF8FFBFFECDFCDD2010D439" author="Jones & D. A" box="[373,542,774,798]" pageId="1" pageNumber="190" refId="ref6333" refString="Jones. D. A. (1986) A field guide to the sea shores of Kuwait and the Arabian Gulf. University of Kuwait, Kuwait City, 192 pp." type="book" year="1986">Jones 1986: 152</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 42 (
|
||
<emphasis id="06A4D237FFF8FFBFFDCDFCDD20B7D43A" box="[629,697,774,797]" italics="true" pageId="1" pageNumber="190">partim</emphasis>
|
||
?);
|
||
<treatmentCitation id="B5712834FFF8FFBFFD63FCDD218CD439" author="Geldiay, R." box="[731,898,774,798]" page="3" pageId="1" pageNumber="190" year="1969">
|
||
<bibRefCitation id="504173D4FFF8FFBFFD63FCDD218CD439" author="Geldiay, R." box="[731,898,774,798]" pageId="1" pageNumber="190" pagination="1 - 17" refId="ref6098" refString="Geldiay, R. 1969. A report on the collection of Natantia (Crustacea, Decapoda) along the coast of Turkey from the eastern Mediterranean to the vicinity of Izmir. Scientific Reports of the Faculty of Science Ege University, 74, 1 - 17." type="journal article" year="1969">Geldiay 1969: 3</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 5 (partim?);
|
||
<treatmentCitation id="B5712834FFF8FFBFFBB7FCDD2732D439" author="Kocatas, A. & Katagan, T." box="[1039,1340,774,798]" page="65" pageId="1" pageNumber="190" year="2003">
|
||
<bibRefCitation id="504173D4FFF8FFBFFBB7FCDD2732D439" author="Kocatas, A. & Katagan, T." box="[1039,1340,774,798]" pageId="1" pageNumber="190" pagination="63 - 74" refId="ref6471" refString="Kocatas, A. & Katagan, T. (2003) The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East, 29, 63 - 74. https: // doi. org / 10.1080 / 09397140.2003.10637971" type="journal article" year="2003">Kocataş & Kataĝan 2003: 65</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 70 (
|
||
<emphasis id="06A4D237FFF8FFBFFAC9FCDC22E9D41B" italics="true" pageId="1" pageNumber="190">partim</emphasis>
|
||
?);
|
||
<treatmentCitation id="B5712834FFF8FFBFFEB1FCFF2010D41B" ID-DOI="http://doi.org/10.5281/zenodo.5255792" ID-Zenodo-Dep="5255792" author="Dehghani, A. & Sari A. & Naderloo, R." box="[265,542,804,829]" httpUri="http://treatment.plazi.org/id/730587B4FF8A3C13FF06FA3EFB74FD8C" page="486" pageId="1" pageNumber="190" year="2019">
|
||
<bibRefCitation id="504173D4FFF8FFBFFEB1FCFF2010D41B" author="Dehghani, A. & Sari A. & Naderloo, R." box="[265,542,804,829]" pageId="1" pageNumber="190" pagination="479 - 504" refId="ref5860" refString="Dehghani, A., Sari A. & Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
|
||
Dehghani
|
||
<emphasis id="06A4D237FFF8FFBFFEC9FCFE23ABD41B" box="[369,421,804,828]" italics="true" pageId="1" pageNumber="190">et al.</emphasis>
|
||
2019a: 486
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 4A–D (
|
||
<emphasis id="06A4D237FFF8FFBFFD26FCFE20ECD41B" box="[670,738,805,828]" italics="true" pageId="1" pageNumber="190">partim</emphasis>
|
||
?) [not
|
||
<taxonomicName id="F3D075A6FFF8FFBFFC93FCFE264FD41A" ID-CoL="84W8B" authority="De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" box="[811,1089,804,829]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFF8FFBFFC93FCFE2193D41B" box="[811,925,804,828]" italics="true" pageId="1" pageNumber="190">A. lobidens</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFF8FFBFFF2FFC9821FAD47C" blockId="1.[151,1437,651,859]" box="[151,1012,835,859]" pageId="1" pageNumber="190">
|
||
<treatmentCitationGroup id="14C0290BFFF8FFBFFF2FFC9821FAD47C" box="[151,1012,835,859]" pageId="1" pageNumber="190">
|
||
<taxonomicName id="F3D075A6FFF8FFBFFF2FFC9822E7D47C" ID-CoL="84LB5" authorityName="Fabricius" authorityYear="1798" box="[151,233,835,859]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="190" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06A4D237FFF8FFBFFF2FFC9822E7D47C" box="[151,233,835,859]" italics="true" pageId="1" pageNumber="190">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
sp. —
|
||
<treatmentCitation id="B5712834FFF8FFBFFE8AFC9F2022D47C" author="Forest, J. & Guinot, D." box="[306,556,835,859]" page="6" pageId="1" pageNumber="190" year="1958">
|
||
<bibRefCitation id="504173D4FFF8FFBFFE8AFC9F2022D47C" author="Forest, J. & Guinot, D." box="[306,556,835,859]" pageId="1" pageNumber="190" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot 1958: 6</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs. 3–7 (see
|
||
<treatmentCitation id="B5712834FFF8FFBFFD7EFC9F21E7D47C" author="Banner, D. M. & Banner, A. H." box="[710,1001,835,859]" page="243" pageId="1" pageNumber="190" year="1982">
|
||
<bibRefCitation id="504173D4FFF8FFBFFD7EFC9F21E7D47C" author="Banner, D. M. & Banner, A. H." box="[710,1001,835,859]" pageId="1" pageNumber="190" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982: 243</bibRefCitation>
|
||
</treatmentCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CCA5DAEFFF8FFBFFF2FFC522723D2E7" pageId="1" pageNumber="190" type="materials_examined">
|
||
<paragraph id="346F0E25FFF8FFBFFF2FFC522109D35C" blockId="1.[151,1437,905,2012]" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF2FFC522373D484" bold="true" box="[151,381,905,931]" pageId="1" pageNumber="190">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="84B80478FFF8FFBFFE36FC52231CD4CB" ID-GBIF-Occurrence="3026647305" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & S. De Grave & Z. Sattari" country="Kuwait" latitude="29.354662" location="Shaab" longLatPrecision="1" longitude="48.029827" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFE36FC5223EBD484" box="[398,485,905,931]" name="Kuwait" pageId="1" pageNumber="190">Kuwait</collectingCountry>
|
||
:
|
||
<specimenCount id="22D6C5ACFFF8FFBFFE4AFC52204AD484" box="[498,580,905,931]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFDEAFC5220CAD484" box="[594,708,905,931]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFD70FC5220E0D484" box="[712,750,905,931]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-001,
|
||
<location id="310F58FEFFF8FFBFFC28FC5221D9D484" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFC28FC5221D9D484" box="[912,983,905,931]" country="Kuwait" latitude="29.354662" longLatPrecision="1" longitude="48.029827" name="Shaab" pageId="1" pageNumber="190">Shaab</location>
|
||
,
|
||
<geoCoordinate id="51E468E2FFF8FFBFFC5DFC52269CD484" box="[997,1170,905,931]" degrees="29" direction="north" minutes="21" orientation="latitude" pageId="1" pageNumber="190" precision="1" seconds="16.78" value="29.354662">29°21’16.78”N</geoCoordinate>
|
||
<geoCoordinate id="51E468E2FFF8FFBFFB24FC522746D484" box="[1180,1352,905,931]" degrees="48" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="190" precision="1" seconds="47.38" value="48.029827">48°01’47.38”E</geoCoordinate>
|
||
, mixed rocky-muddy intertidal, under rocks, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFDDCFC7520C2D4E0" box="[612,716,941,968]" pageId="1" pageNumber="190">A. Anker</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFD6EFC752173D4E0" box="[726,893,941,968]" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFC3FFC76261CD4E0" box="[903,1042,941,968]" pageId="1" pageNumber="190">S. De Grave</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFBA4FC752689D4E0" box="[1052,1159,941,968]" pageId="1" pageNumber="190">Z. Sattari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFB29FC762767D4E0" box="[1169,1385,941,968]" pageId="1" pageNumber="190" value="2019-09-25">
|
||
<collectingDate id="502AD10DFFF8FFBFFB29FC762767D4E0" box="[1169,1385,941,968]" pageId="1" pageNumber="190" value="2019-09-25">25 September 2019</collectingDate>
|
||
</date>
|
||
[fcn KUW-011
|
||
</materialsCitation>
|
||
];
|
||
<materialsCitation id="84B80478FFF8FFBFFE9DFC0A2796D4CB" ID-GBIF-Occurrence="3026647302" box="[293,1432,977,1004]" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & S. De Grave & Z. Sattari" country="Kuwait" latitude="29.354662" location="Shaab" longLatPrecision="1" longitude="48.029827" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFE9DFC0A237BD4CC" box="[293,373,977,1003]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFE3BFC0A23FDD4CC" box="[387,499,977,1003]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFE4DFC0A2014D4CC" box="[501,538,977,1003]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
. 2020-01-002, same collection data as for previous specimen [fcn KUW-030]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFF2FFC2E278FD337" ID-GBIF-Occurrence="3026647304" box="[151,1409,1013,1040]" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & S. De Grave & Z. Sattari" country="Kuwait" latitude="29.354662" location="Shaab" longLatPrecision="1" longitude="48.029827" pageId="1" pageNumber="190" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFF2FFC2E22FCD328" box="[151,242,1013,1039]" pageId="1" pageNumber="190" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="22D6C5ACFFF8FFBFFF46FC2E236AD328" box="[254,356,1013,1039]" pageId="1" pageNumber="190" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFEC8FC2E23ECD328" box="[368,482,1013,1039]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFE5DFC2E2005D328" box="[485,523,1013,1039]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-003, same collection data as for previous specimens [fcn KUW-034]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFA36FC2E213BD370" ID-GBIF-Occurrence="3026647301" collectingDate="2019-10-03" collectionCode="OUMNH, ZC" collectorName="M. Al-Kandari" country="Kuwait" latitude="29.39015" location="Failaka Island" longLatPrecision="1" longitude="48.398785" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFA36FC2E22DED314" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFF58FBC2235FD314" box="[224,337,1049,1075]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFEECFBC22374D314" box="[340,378,1049,1075]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-004,
|
||
<location id="310F58FEFFF8FFBFFDA6FBC220CDD314" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFDA6FBC220CDD314" box="[542,707,1049,1075]" country="Kuwait" latitude="29.39015" longLatPrecision="1" longitude="48.398785" name="Failaka Island" pageId="1" pageNumber="190">Failaka Island</location>
|
||
, eastern side,
|
||
<geoCoordinate id="51E468E2FFF8FFBFFCD4FBC221F3D314" box="[876,1021,1049,1075]" degrees="29" direction="north" minutes="23.409" orientation="latitude" pageId="1" pageNumber="190" precision="1" value="29.39015">29°23.409’N</geoCoordinate>
|
||
<geoCoordinate id="51E468E2FFF8FFBFFBB0FBC226ABD314" box="[1032,1189,1049,1075]" degrees="48" direction="east" minutes="23.927" orientation="longitude" pageId="1" pageNumber="190" precision="1" value="48.398785">48° 23.927’E</geoCoordinate>
|
||
, mixed rocky-muddy intertidal, under rocks, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFE68FBE52076D370" box="[464,632,1085,1112]" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFD3DFBE6213BD370" box="[645,821,1085,1111]" pageId="1" pageNumber="190" value="2019-10-03">
|
||
<collectingDate id="502AD10DFFF8FFBFFD3DFBE6213BD370" box="[645,821,1085,1111]" pageId="1" pageNumber="190" value="2019-10-03">3 October 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
[fcn KUW-077];
|
||
<materialsCitation id="84B80478FFF8FFBFFBB8FBE6210AD35C" ID-GBIF-Occurrence="3026647306" collectingDate="2019-12-14" collectionCode="KISR" collectorName="M. Al-Kandari" country="Kuwait" location="Al Julaia'a" municipality="Seashell Hotel" pageId="1" pageNumber="190" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFBB8FBE62641D370" box="[1024,1103,1085,1111]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="22D6C5ACFFF8FFBFFBE4FBE626CFD370" box="[1116,1217,1085,1111]" pageId="1" pageNumber="190" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFB75FBE6271ED370" box="[1229,1296,1085,1111]" pageId="1" pageNumber="190">KISR</collectionCode>
|
||
,
|
||
<location id="310F58FEFFF8FFBFFAA2FBE6279BD370" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFAA2FBE6279BD370" box="[1306,1429,1085,1111]" country="Kuwait" municipality="Seashell Hotel" name="Al Julaia'a" pageId="1" pageNumber="190">Al Julaia’a</location>
|
||
,
|
||
<collectingMunicipality id="D40B945FFFF8FFBFFF2FFBBA2333D35C" box="[151,317,1121,1147]" pageId="1" pageNumber="190">Seashell Hotel</collectingMunicipality>
|
||
, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFEC0FBB92011D35C" box="[376,543,1121,1148]" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFD93FBBA210AD35C" box="[555,772,1121,1147]" pageId="1" pageNumber="190" value="2019-12-14">
|
||
<collectingDate id="502AD10DFFF8FFBFFD93FBBA210AD35C" box="[555,772,1121,1147]" pageId="1" pageNumber="190" value="2019-12-14">14 December 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFF8FFBFFF7FFB5E2723D2E7" blockId="1.[151,1437,905,2012]" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF7FFB5E23C0D3B8" bold="true" box="[199,462,1157,1183]" pageId="1" pageNumber="190">Comparative material</emphasis>
|
||
.
|
||
<taxonomicName id="F3D075A6FFF8FFBFFE66FB5E2177D3B8" authority="De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" box="[478,889,1157,1184]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFF8FFBFFE66FB5E2038D3B8" box="[478,566,1157,1183]" italics="true" pageId="1" pageNumber="190">Alpheus</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFF8FFBFFDDDFB5E20CCD3B8" box="[613,706,1157,1183]" italics="true" pageId="1" pageNumber="190">lobidens</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
.
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFC3FFB5E21D2D3B8" box="[903,988,1157,1183]" name="Kuwait" pageId="1" pageNumber="190">Kuwait</collectingCountry>
|
||
:
|
||
<materialsCitation id="84B80478FFF8FFBFFC50FB5E20ECD3CF" ID-GBIF-Occurrence="3026647310" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="M. Al-Kandari & Z. Sattari" latitude="29.394266" location="Ras Ajuza" longLatPrecision="1" longitude="47.997334" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFC50FB5E2636D3B8" box="[1000,1080,1157,1183]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
, 1 ovigerous female,
|
||
<collectionCode id="52C196E0FFF8FFBFFA95FB5E2797D3B8" box="[1325,1433,1157,1183]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFF2FFB7222B3D3E3" box="[151,189,1193,1220]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-005,
|
||
<location id="310F58FEFFF8FFBFFEE4FB7123C1D3E4" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFEE4FB7123C1D3E4" box="[348,463,1193,1220]" latitude="29.394266" longLatPrecision="1" longitude="47.997334" name="Ras Ajuza" pageId="1" pageNumber="190">Ras Ajuza</location>
|
||
,
|
||
<geoCoordinate id="51E468E2FFF8FFBFFE62FB722067D3E3" box="[474,617,1193,1220]" degrees="29" direction="north" minutes="23.656" orientation="latitude" pageId="1" pageNumber="190" precision="1" value="29.394266">29°23.656’N</geoCoordinate>
|
||
<geoCoordinate id="51E468E2FFF8FFBFFDC8FB7220F0D3E3" box="[624,766,1193,1220]" degrees="47" direction="east" minutes="59.840" orientation="longitude" pageId="1" pageNumber="190" precision="1" value="47.997334">47°59.840’E</geoCoordinate>
|
||
, mixed rocky-muddy intertidal, under rocks, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFAA4FB7122CAD3C0" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFF6AFB152333D3C0" box="[210,317,1229,1256]" pageId="1" pageNumber="190">Z. Sattari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFEF2FB16202AD3C0" box="[330,548,1229,1256]" pageId="1" pageNumber="190" value="2019-09-25">
|
||
<collectingDate id="502AD10DFFF8FFBFFEF2FB16202AD3C0" box="[330,548,1229,1256]" pageId="1" pageNumber="190" value="2019-09-25">25 September 2019</collectingDate>
|
||
</date>
|
||
[fcn KUW-027]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFD57FB16205CD22B" ID-GBIF-Occurrence="3026647303" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="M. Al-Kandari & Z. Sattari" latitude="29.394266" location="Ras Ajuza" longLatPrecision="1" longitude="47.997334" pageId="1" pageNumber="190" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFD57FB16215DD3C0" box="[751,851,1229,1255]" pageId="1" pageNumber="190" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFCD9FB1621DED3C0" box="[865,976,1229,1255]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFC6BFB1621F6D3C0" box="[979,1016,1229,1255]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-006, same collection data as for previous specimens [fcn KUW-029]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFDE7FB2A2344D273" ID-GBIF-Occurrence="3026647311" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & S. De Grave & Z. Sattari" latitude="29.354662" location="Shaab" longLatPrecision="1" longitude="48.029827" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFDE7FB2A20A2D22C" box="[607,684,1265,1291]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFD00FB2A2129D22C" box="[696,807,1265,1291]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFC93FB2A215ED22C" box="[811,848,1265,1291]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-007,
|
||
<location id="310F58FEFFF8FFBFFC56FB2A263AD22C" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFC56FB2A263AD22C" box="[1006,1076,1265,1291]" latitude="29.354662" longLatPrecision="1" longitude="48.029827" name="Shaab" pageId="1" pageNumber="190">Shaab</location>
|
||
,
|
||
<geoCoordinate id="51E468E2FFF8FFBFFB87FB2A26E6D22C" box="[1087,1256,1265,1291]" degrees="29" direction="north" minutes="21" orientation="latitude" pageId="1" pageNumber="190" precision="1" seconds="16.78" value="29.354662">29°21’16.78”N</geoCoordinate>
|
||
<geoCoordinate id="51E468E2FFF8FFBFFB48FB2A2799D22C" box="[1264,1431,1265,1291]" degrees="48" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="190" precision="1" seconds="47.38" value="48.029827">48°01’47.38”E</geoCoordinate>
|
||
, mixed rocky-muddy intertidal, under rocks, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFD1DFACD2105D208" box="[677,779,1301,1328]" pageId="1" pageNumber="190">A. Anker</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFCADFACD21B6D208" box="[789,952,1301,1328]" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFC7BFACE2642D208" box="[963,1100,1301,1328]" pageId="1" pageNumber="190">S. De Grave</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFBEEFACD26B0D208" box="[1110,1214,1301,1328]" pageId="1" pageNumber="190">Z. Sattari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFB70FACE2792D208" box="[1224,1436,1301,1328]" pageId="1" pageNumber="190" value="2019-09-25">
|
||
<collectingDate id="502AD10DFFF8FFBFFB70FACE2792D208" box="[1224,1436,1301,1328]" pageId="1" pageNumber="190" value="2019-09-25">25 September 2019</collectingDate>
|
||
</date>
|
||
[fcn KUW-035]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFEEEFAE22370D25F" ID-GBIF-Occurrence="3026647308" collectingDate="2019-09-25" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & S. De Grave & Z. Sattari" latitude="29.354662" location="Shaab" longLatPrecision="1" longitude="48.029827" pageId="1" pageNumber="190" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFEEEFAE223ADD274" box="[342,419,1337,1363]" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
, 1 ovigerous female,
|
||
<collectionCode id="52C196E0FFF8FFBFFD29FAE2210ED274" box="[657,768,1337,1363]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFCBCFAE22127D274" box="[772,809,1337,1363]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-008, same collection data as for previous specimen [fcn KUW-036]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFE31FA86278CD2BB" ID-GBIF-Occurrence="3026647309" collectingDate="2019-10-07" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & Z. Sattari" latitude="29.3258" location="Sulaibikhat Bay" longLatPrecision="8" longitude="47.8225" pageId="1" pageNumber="190" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFE31FA8623E4D250" box="[393,490,1373,1399]" pageId="1" pageNumber="190" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFE4CFA86206DD250" box="[500,611,1373,1399]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFDDFFA862082D250" box="[615,652,1373,1399]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-009,
|
||
<location id="310F58FEFFF8FFBFFC90FA8621D6D25F" LSID="urn:lsid:plazi:treatment:BC79BF33FFF8FFB8FF2FFDF92332D2E6:310F58FEFFF8FFBFFC90FA8621D6D25F" box="[808,984,1373,1400]" latitude="29.3258" longLatPrecision="8" longitude="47.8225" name="Sulaibikhat Bay" pageId="1" pageNumber="190">Sulaibikhat Bay</location>
|
||
,
|
||
<geoCoordinate id="51E468E2FFF8FFBFFC5BFA86267FD25F" box="[995,1137,1373,1400]" degrees="29" direction="north" minutes="19.548" orientation="latitude" pageId="1" pageNumber="190" precision="1" value="29.3258">29°19.548’N</geoCoordinate>
|
||
<geoCoordinate id="51E468E2FFF8FFBFFBC0FA8626F6D25F" box="[1144,1272,1373,1400]" degrees="47" direction="east" minutes="49.35" orientation="longitude" pageId="1" pageNumber="190" precision="9" value="47.8225">47°49.35’E</geoCoordinate>
|
||
, intertidal mud flat, in burrows, suction (yabby) pump, leg.
|
||
<collectorName id="99256BF3FFF8FFBFFD38FA5920E9D2BC" box="[640,743,1409,1436]" pageId="1" pageNumber="190">A. Anker</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFD4BFA592196D2BC" box="[755,920,1409,1436]" pageId="1" pageNumber="190">M. Al-Kandari</collectorName>
|
||
,
|
||
<collectorName id="99256BF3FFF8FFBFFC1DFA592600D2BC" box="[933,1038,1409,1436]" pageId="1" pageNumber="190">Z. Sattari</collectorName>
|
||
,
|
||
<date id="406E28E5FFF8FFBFFBA2FA5926C9D2BC" box="[1050,1223,1409,1436]" pageId="1" pageNumber="190" value="2019-10-07">
|
||
<collectingDate id="502AD10DFFF8FFBFFBA2FA5926C9D2BC" box="[1050,1223,1409,1436]" pageId="1" pageNumber="190" value="2019-10-07">7 October 2019</collectingDate>
|
||
</date>
|
||
[fcn KUW-107]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="84B80478FFF8FFBFFA36FA5A2727D2E7" ID-GBIF-Occurrence="3026647307" collectingDate="2019-10-07" collectionCode="OUMNH, ZC" collectorName="A. Anker & M. Al-Kandari & Z. Sattari" latitude="29.3258" location="Sulaibikhat Bay" longLatPrecision="8" longitude="47.8225" pageId="1" pageNumber="190" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="22D6C5ACFFF8FFBFFA36FA5A22C1D298" pageId="1" pageNumber="190" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="22D6C5ACFFF8FFBFFF62FA7E2335D298" box="[218,315,1445,1471]" pageId="1" pageNumber="190" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="52C196E0FFF8FFBFFEFEFA7E23BBD298" box="[326,437,1445,1471]" pageId="1" pageNumber="190">OUMNH</collectionCode>
|
||
.
|
||
<collectionCode id="52C196E0FFF8FFBFFE01FA7E23D0D298" box="[441,478,1445,1471]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="190" type="University or college">ZC</collectionCode>
|
||
.2020-01-010, same collection data as for previous specimen [fcn KUW-108]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CCA5DAEFFF8FFBFFF7FFA122069D0FC" pageId="1" pageNumber="190" type="description">
|
||
<paragraph id="346F0E25FFF8FFBFFF7FFA12234FD120" blockId="1.[151,1437,905,2012]" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF7FFA122340D2C4" bold="true" box="[199,334,1481,1507]" pageId="1" pageNumber="190">Description</emphasis>
|
||
. See
|
||
<bibRefCitation id="504173D4FFF8FFBFFE32FA1220B5D2C3" author="Holthuis, L. B. & Gottlieb, E." box="[394,699,1481,1508]" pageId="1" pageNumber="190" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb (1958)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFF8FFBFFD7EFA1221D5D2C3" author="Tirmizi, N. M. & Kazmi, M. A." box="[710,987,1481,1508]" pageId="1" pageNumber="190" pagination="99 - 100" refId="ref6661" refString="Tirmizi, N. M. & Kazmi, M. A. (1969) The occurrence of Alpheus inopinatus Holthuis & Gottlieb, 1958 in the Arabian Sea (Decapoda, Alpheidae). Crustaceana, 16, 99 - 100. https: // doi. org / 10.1163 / 156854068 X 00250" type="journal article" year="1969">Tirmizi & Kazmi (1969)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFF8FFBFFC5FFA1226F7D2C4" author="Karim, S. I. & Hafizur, R." box="[999,1273,1481,1507]" pageId="1" pageNumber="190" pagination="25 - 28" refId="ref6366" refString="Karim, S. I. & Hafizur, R. (1974) On some Carides of Karachi coasts. Records of Zoological Survey, 6, 25 - 28, pls. 1 - 4." type="journal article" year="1974">Karim & Hafizur (1974)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="504173D4FFF8FFBFFA97FA112333D120" author="Banner, D. M. & Banner, A. H." pageId="1" pageNumber="190" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner (1982)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFF8FFBFFF7FF9CA271BD024" blockId="1.[151,1437,905,2012]" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF7FF9CA2379D10C" bold="true" box="[199,375,1553,1579]" pageId="1" pageNumber="190">Colour pattern</emphasis>
|
||
. Pale greenish or bluish grey to various tones of brown, often darker in females; pleon with three pairs of small dark spots distributed as following: one pair of small lateral spots on second pleuron, one pair of small dorsolateral spots on third pleuron (sometimes very faint and inconspicuous) and one pair of larger, more conspicuous lateral spots on fourth pleuron; mesial side of major and minor chelae greenish-brownish, with some olive or orange areas; dactylar plunger of major chela creamy white with some bluish tinge proximally; antennular and antennal flagella pale greenish-yellow; second pereiopods and walking legs (third to fifth pereiopods) pale reddish or pinkish with whitish articulations; tail fan brownish or greenish, with bluish area on uropodal exopod (
|
||
<figureCitation id="ACEB12A0FFF8FFBFFBD2F93226BBD024" box="[1130,1205,1769,1795]" captionStart="FIGURE 1" captionStartId="2.[151,250,1547,1571]" captionTargetBox="[151,1435,179,1521]" captionTargetId="figure-18@2.[151,1435,179,1521]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus inopinatus Holthuis & Gottlieb, 1958: male from Kuwait, OUMNH.ZC.2020-01-001 [fcn KUW-011], A—dorsal view; B—lateral view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407194" httpUri="https://zenodo.org/record/4407194/files/figure.png" pageId="1" pageNumber="190">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="ACEB12A0FFF8FFBFFB7AF93226C0D024" box="[1218,1230,1769,1795]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="1" pageNumber="190">3</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFF8FFBFFF7FF8D62069D0FC" blockId="1.[151,1437,905,2012]" pageId="1" pageNumber="190">
|
||
<emphasis id="06A4D237FFF8FFBFFF7FF8D623B2D000" bold="true" box="[199,444,1805,1831]" pageId="1" pageNumber="190">General distribution</emphasis>
|
||
. Possibly widely distributed in the Indo-West Pacific with records from the Red Sea,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFAD7F8D622DDD06C" name="Djibouti" pageId="1" pageNumber="190">Djibouti</collectingCountry>
|
||
, Arabian-Persian Gulf (
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFE65F8EA203AD06C" box="[477,564,1841,1867]" name="Kuwait" pageId="1" pageNumber="190">Kuwait</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFDF8F8E9207ED06B" box="[576,624,1842,1868]" name="Iran" pageId="1" pageNumber="190">Iran</collectingCountry>
|
||
, see Remarks),
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFC99F8EA2166D06C" box="[801,872,1841,1867]" name="Oman" pageId="1" pageNumber="190">Oman</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFCCCF8EA21D8D06C" box="[884,982,1841,1867]" name="Pakistan" pageId="1" pageNumber="190">Pakistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFC59F8E92666D06B" box="[993,1128,1842,1868]" name="Hong Kong" pageId="1" pageNumber="190">Hong Kong</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFBCBF8EA26C6D06C" box="[1139,1224,1841,1867]" name="Taiwan" pageId="1" pageNumber="190">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFB6AF8EA2737D06C" box="[1234,1337,1841,1867]" name="Australia" pageId="1" pageNumber="190">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFF8FFBFFAFFF8EA2330D048" author="Tirmizi, N. M. & Kazmi, M. A." pageId="1" pageNumber="190" pagination="99 - 100" refId="ref6661" refString="Tirmizi, N. M. & Kazmi, M. A. (1969) The occurrence of Alpheus inopinatus Holthuis & Gottlieb, 1958 in the Arabian Sea (Decapoda, Alpheidae). Crustaceana, 16, 99 - 100. https: // doi. org / 10.1163 / 156854068 X 00250" type="journal article" year="1969">Tirmizi & Kazmi 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFF8FFBFFEF1F88D2043D048" author="Banner, D. M. & Banner, A. H." box="[329,589,1877,1904]" pageId="1" pageNumber="190" pagination="215 - 257" refId="ref5338" refString="Banner, D. M. & Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner & Banner 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFF8FFBFFDEFF88E209FD048" author="Banner, D. M. & Banner, A. H." box="[599,657,1877,1903]" pageId="1" pageNumber="190" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFF8FFBFFD24F88E20D9D048" author="Banner, D. M. & Banner, A. H." box="[668,727,1877,1903]" pageId="1" pageNumber="190" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFF8FFBFFD5AF88D2158D048" author="Jeng, M. S." box="[738,854,1877,1904]" pageId="1" pageNumber="190" pagination="1 - 66" refId="ref6296" refString="Jeng, M. S. (1997) Studies on the land and aquatic decapod crustacean fauna of the Kenting National Park (II). Conservation Research Report, 96, 1 - 66." type="journal article" year="1997">Jeng 1997</bibRefCitation>
|
||
; present study); however, all records from outside of the western Indian Ocean require confirmation (see Remarks). Lessepsian migrant with a well-established population in the eastern Mediterranean Sea, with confirmed records from
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFC23F84621D7D090" box="[923,985,1949,1975]" name="Israel" pageId="1" pageNumber="190">Israel</collectingCountry>
|
||
and
|
||
<collectingCountry id="4CC74EB5FFF8FFBFFBA9F846266FD090" box="[1041,1121,1949,1975]" name="Turkey" pageId="1" pageNumber="190">Turkey</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFF8FFBFFBC8F8462799D09F" author="Holthuis, L. B. & Gottlieb, E." box="[1136,1431,1949,1976]" pageId="1" pageNumber="190" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFF8FFBFFF2FF819233DD0FC" author="Kocatas, A." box="[151,307,1985,2012]" pageId="1" pageNumber="190" pagination="161 - 162" refId="ref6437" refString="Kocatas, A. (1981) Liste preliminaires et repartition des crustaces decapodes des eaux turques. Rapports de la Commission Internationale de la Mer Mediterranee, 27, 161 - 162." type="journal article" year="1981">Kocataş 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFF8FFBFFE87F81A2054D0FC" author="Lewinsohn, C. & Galil, B." box="[319,602,1985,2011]" pageId="1" pageNumber="190" pagination="207 - 210" refId="ref6514" refString="Lewinsohn, C. & Galil, B. (1982). Notes on species of Alpheus (Crustacea Decapoda) from the Mediterranean coast of Israel. Quaderni del Laboratorio di Tecnologia della Pesca, Ancona, 3, 207 - 210." type="journal article" year="1982">Lewinsohn & Galil 1982</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="60AF5EADFFFBFFBCFF2FF9D020F6D160" ID-DOI="http://doi.org/10.5281/zenodo.4407194" ID-Zenodo-Dep="4407194" httpUri="https://zenodo.org/record/4407194/files/figure.png" pageId="2" pageNumber="191" startId="2.[151,250,1547,1571]" targetBox="[151,1435,179,1521]" targetPageId="2">
|
||
<paragraph id="346F0E25FFFBFFBCFF2FF9D020F6D160" blockId="2.[151,1436,1547,1607]" pageId="2" pageNumber="191">
|
||
<emphasis id="06A4D237FFFBFFBCFF2FF9D02301D104" bold="true" box="[151,271,1547,1571]" pageId="2" pageNumber="191">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="F3D075A6FFFBFFBCFEA6F9D0210FD104" authority="Holthuis & Gottlieb, 1958" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[286,769,1547,1571]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFBFFBCFEA6F9D023EAD104" box="[286,484,1547,1571]" italics="true" pageId="2" pageNumber="191">Alpheus inopinatus</emphasis>
|
||
<bibRefCitation id="504173D4FFFBFFBCFE54F9D0210FD104" author="Holthuis, L. B. & Gottlieb, E." box="[492,769,1547,1571]" pageId="2" pageNumber="191" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
: male from Kuwait, OUMNH.ZC.2020-01-001 [fcn KUW-011], A—dorsal view; B—lateral view. Photographs by A. Anker.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7CCA5DAEFFFBFFBCFF7FF9AE20F7D1FF" pageId="2" pageNumber="191" type="biology_ecology">
|
||
<paragraph id="346F0E25FFFBFFBCFF7FF9AE20F7D1FF" blockId="2.[151,1437,1653,2004]" pageId="2" pageNumber="191">
|
||
<emphasis id="06A4D237FFFBFFBCFF7FF9AE2328D1A8" bold="true" box="[199,294,1653,1679]" pageId="2" pageNumber="191">Ecology</emphasis>
|
||
. Largely intertidal, sometimes also in the shallow subtidal (to at least
|
||
<quantity id="F328A3C0FFFBFFBCFBFEF9AE267BD1B7" box="[1094,1141,1653,1680]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="191" unit="m" value="6.0">6 m</quantity>
|
||
), rocky and mixed sandyrocky shores, often heavily silted or with a significant mud component; typically on sand/mud under large rocks and coral rubble (
|
||
<bibRefCitation id="504173D4FFFBFFBCFE94F9662046D1F0" author="Lewinsohn, C. & Galil, B." box="[300,584,1725,1751]" pageId="2" pageNumber="191" pagination="207 - 210" refId="ref6514" refString="Lewinsohn, C. & Galil, B. (1982). Notes on species of Alpheus (Crustacea Decapoda) from the Mediterranean coast of Israel. Quaderni del Laboratorio di Tecnologia della Pesca, Ancona, 3, 207 - 210." type="journal article" year="1982">Lewinsohn & Galil 1982</bibRefCitation>
|
||
; present study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7CCA5DAEFFFBFFB8FF7FF93A2332D2E6" lastPageId="6" lastPageNumber="195" pageId="2" pageNumber="191" type="discussion">
|
||
<paragraph id="346F0E25FFFBFFBCFF7FF93A2325D0F3" blockId="2.[151,1437,1653,2004]" pageId="2" pageNumber="191">
|
||
<emphasis id="06A4D237FFFBFFBCFF7FF93A233AD1DC" bold="true" box="[199,308,1761,1787]" pageId="2" pageNumber="191">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="F3D075A6FFFBFFBCFEFDF93A2011D1DC" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[325,543,1761,1787]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFBFFBCFEFDF93A2011D1DC" box="[325,543,1761,1787]" italics="true" pageId="2" pageNumber="191">Alpheus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="F3D075A6FFFBFFBCFD6DF9392600D1DC" authority="De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" box="[725,1038,1761,1788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFBFFBCFD6DF9392158D1DC" box="[725,854,1761,1788]" italics="true" pageId="2" pageNumber="191">A. lobidens</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
species complex, together with
|
||
<taxonomicName id="F3D075A6FFFBFFBCFA3CF93923BAD038" authority="Heller, 1862" authorityName="sensu Heller" authorityYear="1862" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFBFFBCFA3CF939232DD038" italics="true" pageId="2" pageNumber="191">A. crassimanus</emphasis>
|
||
<bibRefCitation id="504173D4FFFBFFBCFE92F8DE23BAD038" author="Heller, C." box="[298,436,1797,1823]" pageId="2" pageNumber="191" pagination="519 - 528" refId="ref6196" refString="Heller, C. (1862) Neue Crustaceen gesammelt wahrend der Weltumsegelung der k. k. Fregatte Novara. Zweiter vorlaufiger Beri- cht. Verhandlungen der Koeniglich-Kaiserlichen Zoologisch-Botanischen Gesellschaft, Wien, 28, 519 - 528." type="journal article" year="1862">Heller, 1862</bibRefCitation>
|
||
</taxonomicName>
|
||
(currently considered to be a synonym of
|
||
<taxonomicName id="F3D075A6FFFBFFBCFC34F8DD2604D038" authorityName="sensu De Haan" authorityYear="1849" box="[908,1034,1797,1824]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFBFFBCFC34F8DD2604D038" box="[908,1034,1797,1824]" italics="true" pageId="2" pageNumber="191">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="F3D075A6FFFBFFBCFB99F8DD2323D064" authority="Banner & Banner, 1974" authorityName="Banner & Banner" authorityYear="1974" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="polynesicus">
|
||
<emphasis id="06A4D237FFFBFFBCFB99F8DD2727D038" box="[1057,1321,1797,1824]" italics="true" pageId="2" pageNumber="191">A. lobidens polynesicus</emphasis>
|
||
<bibRefCitation id="504173D4FFFBFFBCFA97F8DD2323D064" author="Banner, A. H. & Banner, D. M." pageId="2" pageNumber="191" pagination="423 - 437" refId="ref5281" refString="Banner, A. H. & Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner & Banner, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
(originally described as
|
||
<taxonomicName id="F3D075A6FFFBFFBCFDF9F8F12133D064" box="[577,829,1833,1860]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="polynesica">
|
||
<emphasis id="06A4D237FFFBFFBCFDF9F8F12133D064" box="[577,829,1833,1860]" italics="true" pageId="2" pageNumber="191">A. lobidens polynesica</emphasis>
|
||
</taxonomicName>
|
||
), and the recently described
|
||
<taxonomicName id="F3D075A6FFFBFFBCFBC7F8F123ECD040" authority="Dehghani, Sari & Naderloo, 2019" authorityName="Dehghani, Sari & Naderloo" authorityYear="2019" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="mohammadpouri">
|
||
<emphasis id="06A4D237FFFBFFBCFBC7F8F12753D064" box="[1151,1373,1833,1860]" italics="true" pageId="2" pageNumber="191">A. mohammadpouri</emphasis>
|
||
Dehghani, Sari & Naderloo, 2019
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="504173D4FFFBFFBCFE48F89620F7D040" author="Banner, A. H. & Banner, D. M." box="[496,761,1869,1896]" pageId="2" pageNumber="191" pagination="423 - 437" refId="ref5281" refString="Banner, A. H. & Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner & Banner 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFBFFBCFCBCF8962131D040" author="Banner, D. M. & Banner, A. H." box="[772,831,1869,1895]" pageId="2" pageNumber="191" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFFBFFBCFCF2F8962648D04F" author="Dehghani, A. & Sari, A. & Naderloo, R." box="[842,1094,1869,1896]" pageId="2" pageNumber="191" pagination="911 - 920" refId="ref5931" refString="Dehghani, A., Sari, A. & Naderloo, R. (2019 b) Three new species of narrowly endemic snapping shrimp, genus Alpheus (Decapoda: Caridea: Alpheidae) from the Persian Gulf. Journal of the Marine Biological Association of the United Kingdom, 99, 911 - 920. https: // doi. org / 10.1017 / S 0025315418000693" type="journal article" year="2019">
|
||
Dehghani
|
||
<emphasis id="06A4D237FFFBFFBCFC06F89521F8D040" box="[958,1014,1869,1895]" italics="true" pageId="2" pageNumber="191">et al.</emphasis>
|
||
2019b
|
||
</bibRefCitation>
|
||
), together with a large number of undescribed species. The complicated taxonomic situation of
|
||
<taxonomicName id="F3D075A6FFFBFFBCFCD6F8A921E3D0AC" authorityName="sensu De Haan" authorityYear="1849" box="[878,1005,1905,1932]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFBFFBCFCD6F8A921E3D0AC" box="[878,1005,1905,1932]" italics="true" pageId="2" pageNumber="191">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
and the presence of several cryptic or pseudocryptic species in this species complex make identifications of specimens extremely difficult (
|
||
<bibRefCitation id="504173D4FFFBFFBCFAB6F84E232ED0F3" author="Anker, A. & De Grave, S." pageId="2" pageNumber="191" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="60AF5EADFFFAFFBDFF2FF9C02086D17F" ID-DOI="http://doi.org/10.5281/zenodo.4407198" ID-Zenodo-Dep="4407198" httpUri="https://zenodo.org/record/4407198/files/figure.png" pageId="3" pageNumber="192" startId="3.[151,250,1563,1587]" targetBox="[151,1435,181,1540]" targetPageId="3">
|
||
<paragraph id="346F0E25FFFAFFBDFF2FF9C02086D17F" blockId="3.[151,1437,1563,1624]" pageId="3" pageNumber="192">
|
||
<emphasis id="06A4D237FFFAFFBDFF2FF9C02301D113" bold="true" box="[151,271,1563,1588]" pageId="3" pageNumber="192">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="F3D075A6FFFAFFBDFEA5F9C7209BD113" authority="De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" box="[285,661,1564,1588]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFAFFBDFEA5F9C72361D113" box="[285,367,1564,1588]" italics="true" pageId="3" pageNumber="192">Alpheus</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFAFFBDFE20F9C723E0D113" box="[408,494,1564,1588]" italics="true" pageId="3" pageNumber="192">lobidens</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
: male from Kuwait, OUMNH.ZC.2020-01-005 [fcn KUW-027], A—dorsal view; B—lateral view. Photographs by A. Anker.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="346F0E25FFFAFFBDFF7FF95D21CCD0C2" blockId="3.[151,1437,1670,2021]" pageId="3" pageNumber="192">
|
||
<taxonomicName id="F3D075A6FFFAFFBDFF7FF95D23AED187" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[199,416,1670,1696]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFAFFBDFF7FF95D23AED187" box="[199,416,1670,1696]" italics="true" pageId="3" pageNumber="192">Alpheus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
was originally described based on material from the Mediterranean coast of
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFAABF95D275AD187" box="[1299,1364,1670,1696]" name="Israel" pageId="3" pageNumber="192">Israel</collectingCountry>
|
||
, as an “immigrant from the Red Sea” (
|
||
<bibRefCitation id="504173D4FFFAFFBDFDBBF9712122D1E2" author="Holthuis, L. B. & Gottlieb, E." box="[515,812,1706,1733]" pageId="3" pageNumber="192" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb 1958</bibRefCitation>
|
||
). This species is morphologically very close to another species of the
|
||
<taxonomicName id="F3D075A6FFFAFFBDFE80F91523B9D1CF" authorityName="sensu De Haan" authorityYear="1849" box="[312,439,1742,1768]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFAFFBDFE80F91523B9D1CF" box="[312,439,1742,1768]" italics="true" pageId="3" pageNumber="192">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex, which is present in the Gulf and possibly throughout the western and northern Indian Ocean (including the Red Sea), and identified either as
|
||
<taxonomicName id="F3D075A6FFFAFFBDFCEFF92921D8D02B" authorityName="sensu De Haan" authorityYear="1849" box="[855,982,1778,1804]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFAFFBDFCEFF92921D8D02B" box="[855,982,1778,1804]" italics="true" pageId="3" pageNumber="192">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="EB6BB087FFFAFFBDFC5DF9292619D02B" box="[997,1047,1778,1804]" pageId="3" pageNumber="192">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFB39F92926CDD02B" box="[1153,1219,1778,1804]" name="Japan" pageId="3" pageNumber="192">Japan</collectingCountry>
|
||
) or
|
||
<taxonomicName id="F3D075A6FFFAFFBDFB49F9292792D02B" authorityName="sensu Heller" authorityYear="1862" box="[1265,1436,1778,1804]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFAFFBDFB49F9292792D02B" box="[1265,1436,1778,1804]" italics="true" pageId="3" pageNumber="192">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="EB6BB087FFFAFFBDFF27F8CD22DFD017" box="[159,209,1814,1840]" pageId="3" pageNumber="192">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFE84F8CD23FFD017" box="[316,497,1814,1840]" name="India" pageId="3" pageNumber="192">Nicobar Islands</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFE45F8CD2035D016" box="[509,571,1814,1841]" name="India" pageId="3" pageNumber="192">India</collectingCountry>
|
||
). Like
|
||
<taxonomicName id="F3D075A6FFFAFFBDFD32F8CD212ED017" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[650,800,1814,1840]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFAFFBDFD32F8CD212ED017" box="[650,800,1814,1840]" italics="true" pageId="3" pageNumber="192">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
, this species also penetrated into the Mediterranean Sea through the
|
||
<collectingRegion id="F614C0C7FFFAFFBDFEA5F8E1235DD073" box="[285,339,1850,1876]" country="Egypt" name="As Suways" pageId="3" pageNumber="192">Suez</collectingRegion>
|
||
Canal, spreading along the North African coasts from
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFC17F8E121FCD073" box="[943,1010,1850,1876]" name="Egypt" pageId="3" pageNumber="192">Egypt</collectingCountry>
|
||
westwards to
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFB28F8E126E8D073" box="[1168,1254,1850,1876]" name="Tunisia" pageId="3" pageNumber="192">Tunisia</collectingCountry>
|
||
, and northwards to
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFF0DF88522FED05F" box="[181,240,1886,1912]" name="Syria" pageId="3" pageNumber="192">Syria</collectingCountry>
|
||
and
|
||
<collectingCountry id="4CC74EB5FFFAFFBDFE91F8852377D05F" box="[297,377,1886,1912]" name="Turkey" pageId="3" pageNumber="192">Turkey</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFFAFFBDFE31F885205BD05F" author="Hasan, H. & Zeini, A. & Noel, P. Y." box="[393,597,1886,1912]" pageId="3" pageNumber="192" pagination="513 - 536" refId="ref6147" refString="Hasan, H., Zeini, A. & Noel, P. Y. (2008) The marine decapod Crustacea of the area of Lattakia, Syria. Crustaceana, 81, 513 - 536. https: // doi. org / 10.1163 / 156854008784092175" type="journal article" year="2008">
|
||
Hasan
|
||
<emphasis id="06A4D237FFFAFFBDFE60F884201CD05F" box="[472,530,1886,1912]" italics="true" pageId="3" pageNumber="192">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFFAFFBDFDDAF8852129D05E" author="Bakir, K. & Ilkyaz, A. T. & Aydin, C. & Turkmen, G." box="[610,807,1886,1913]" pageId="3" pageNumber="192" pagination="651 - 656" refId="ref5216" refString="Bakir, K., Ilkyaz, A. T., Aydin, C. & Turkmen, G. (2015) The presence of Alpheus lobidens De Haan, 1849 (Decapoda, Alpheidae) on the Turkish Aegean Sea coast. Crustaceana, 88, 651 - 656. https: // doi. org / 10.1163 / 15685403 - 00003434" type="journal article" year="2015">
|
||
Bakir
|
||
<emphasis id="06A4D237FFFAFFBDFD10F88420ECD05F" box="[680,738,1886,1912]" italics="true" pageId="3" pageNumber="192">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Older Mediterranean records of this species are under the name
|
||
<taxonomicName id="F3D075A6FFFAFFBDFEBBF85923A0D0BB" authorityName="sensu Heller" authorityYear="1862" box="[259,430,1922,1948]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFAFFBDFEBBF85923A0D0BB" box="[259,430,1922,1948]" italics="true" pageId="3" pageNumber="192">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
(e.g., Balss 1936;
|
||
<bibRefCitation id="504173D4FFFAFFBDFDC6F8592179D0BA" author="Forest, J. & Guinot, D." box="[638,887,1922,1949]" pageId="3" pageNumber="192" pagination="24 - 43" refId="ref6023" refString="Forest, J. & Guinot, D. (1956) Sur une collection de crustaces decapodes et stomatopodes des mers tunisiennes. Bulletin de la Station Oceanographique Salammbo, 53, 24 - 43." type="journal article" year="1956">Forest & Guinot 1956</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFAFFBDFC3AF85921B3D0BA" author="Forest, J. & Guinot, D." box="[898,957,1922,1949]" pageId="3" pageNumber="192" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFFAFFBDFC70F85926E7D0BA" author="Holthuis, L. B. & Gottlieb, E." box="[968,1257,1922,1949]" pageId="3" pageNumber="192" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFFAFFBDFB4DF859279ED0BA" author="Geldiay, R." box="[1269,1424,1922,1949]" pageId="3" pageNumber="192" pagination="1 - 17" refId="ref6098" refString="Geldiay, R. 1969. A report on the collection of Natantia (Crustacea, Decapoda) along the coast of Turkey from the eastern Mediterranean to the vicinity of Izmir. Scientific Reports of the Faculty of Science Ege University, 74, 1 - 17." type="journal article" year="1969">Geldiay 1969</bibRefCitation>
|
||
), which was later placed in the synonymy of
|
||
<taxonomicName id="F3D075A6FFFAFFBDFDC1F87D20F8D0E7" authorityName="sensu De Haan" authorityYear="1849" box="[633,758,1958,1984]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFAFFBDFDC1F87D20F8D0E7" box="[633,758,1958,1984]" italics="true" pageId="3" pageNumber="192">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(see discussion in
|
||
<bibRefCitation id="504173D4FFFAFFBDFC7FF87D26C2D0E7" author="Banner, A. H. & Banner, D. M." box="[967,1228,1958,1984]" pageId="3" pageNumber="192" pagination="423 - 437" refId="ref5281" refString="Banner, A. H. & Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner & Banner 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFAFFBDFB6FF87D271FD0E7" author="Banner, D. M. & Banner, A. H." box="[1239,1297,1958,1984]" pageId="3" pageNumber="192" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFAFFBDFAA4F87D2756D0E7" author="Banner, D. M. & Banner, A. H." box="[1308,1368,1958,1984]" pageId="3" pageNumber="192" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
), with more recent records under the name
|
||
<taxonomicName id="F3D075A6FFFAFFBDFD88F81120A0D0C3" authorityName="sensu De Haan" authorityYear="1849" box="[560,686,1994,2020]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFAFFBDFD88F81120A0D0C3" box="[560,686,1994,2020]" italics="true" pageId="3" pageNumber="192">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="504173D4FFFAFFBDFD4DF81121B8D0C2" author="Bakir, K. & Ilkyaz, A. T. & Aydin, C. & Turkmen, G." box="[757,950,1994,2021]" pageId="3" pageNumber="192" pagination="651 - 656" refId="ref5216" refString="Bakir, K., Ilkyaz, A. T., Aydin, C. & Turkmen, G. (2015) The presence of Alpheus lobidens De Haan, 1849 (Decapoda, Alpheidae) on the Turkish Aegean Sea coast. Crustaceana, 88, 651 - 656. https: // doi. org / 10.1163 / 15685403 - 00003434" type="journal article" year="2015">
|
||
Bakir
|
||
<emphasis id="06A4D237FFFAFFBDFC82F810217CD0C3" box="[826,882,1994,2020]" italics="true" pageId="3" pageNumber="192">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="60AF5EADFFFDFFBAFF2FFA5626DFD112" ID-DOI="http://doi.org/10.5281/zenodo.4407200" ID-Zenodo-Dep="4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="4" pageNumber="193" startId="4.[151,250,1421,1445]" targetBox="[151,1435,181,1397]" targetPageId="4">
|
||
<paragraph id="346F0E25FFFDFFBAFF2FFA5626DFD112" blockId="4.[151,1437,1421,1589]" pageId="4" pageNumber="193">
|
||
<emphasis id="06A4D237FFFDFFBAFF2FFA562303D282" bold="true" box="[151,269,1421,1445]" pageId="4" pageNumber="193">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="F3D075A6FFFDFFBAFEA2FA5620FED282" authority="Holthuis & Gottlieb, 1958" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[282,752,1421,1445]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFDFFBAFEA2FA5623D0D282" box="[282,478,1421,1445]" italics="true" pageId="4" pageNumber="193">Alpheus inopinatus</emphasis>
|
||
<bibRefCitation id="504173D4FFFDFFBAFE5CFA5620FED282" author="Holthuis, L. B. & Gottlieb, E." box="[484,752,1421,1445]" pageId="4" pageNumber="193" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
(A–C) and
|
||
<taxonomicName id="F3D075A6FFFDFFBAFCD1FA5626D7D282" authority="De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" box="[873,1241,1421,1445]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFCD1FA5621B5D282" box="[873,955,1421,1445]" italics="true" pageId="4" pageNumber="193">Alpheus</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFDFFBAFC5BFA562637D282" box="[995,1081,1421,1445]" italics="true" pageId="4" pageNumber="193">lobidens</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
(D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—ma-jor chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="346F0E25FFFDFFBAFF7FF9BF23A7D0BA" blockId="4.[151,1437,1635,2022]" pageId="4" pageNumber="193">
|
||
In the present study, we tentatively continue to accept the synonymy of
|
||
<taxonomicName id="F3D075A6FFFDFFBAFC40F9BF26AAD15A" authorityName="sensu Heller" authorityYear="1862" box="[1016,1188,1636,1662]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFDFFBAFC40F9BF26AAD15A" box="[1016,1188,1636,1662]" italics="true" pageId="4" pageNumber="193">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="F3D075A6FFFDFFBAFB5EF9BF2768D15A" authorityName="sensu De Haan" authorityYear="1849" box="[1254,1382,1635,1662]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFB5EF9BF2768D15A" box="[1254,1382,1635,1662]" italics="true" pageId="4" pageNumber="193">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
proposed by
|
||
<bibRefCitation id="504173D4FFFDFFBAFEB9F953201ED186" author="Banner, A. H. & Banner, D. M." box="[257,528,1671,1698]" pageId="4" pageNumber="193" pagination="423 - 437" refId="ref5281" refString="Banner, A. H. & Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: Additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner & Banner (1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFDFFBAFDA2F95C2058D186" author="Banner, D. M. & Banner, A. H." box="[538,598,1671,1697]" pageId="4" pageNumber="193" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
), who compared the Japanese specimens they identified as
|
||
<taxonomicName id="F3D075A6FFFDFFBAFB5FF953276AD186" authorityName="sensu De Haan" authorityYear="1849" box="[1255,1380,1671,1698]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFB5FF953276AD186" box="[1255,1380,1671,1698]" italics="true" pageId="4" pageNumber="193">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
with the material from other areas of the Indo-West Pacific. However, we feel that a thorough morphological and molecular study is needed to establish the true identity of this taxon in light of the fact that their view of
|
||
<taxonomicName id="F3D075A6FFFDFFBAFAB8F90B2770D1CE" authorityName="sensu De Haan" authorityYear="1849" box="[1280,1406,1743,1770]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFAB8F90B2770D1CE" box="[1280,1406,1743,1770]" italics="true" pageId="4" pageNumber="193">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
as a single widespread species has now been refuted (see also
|
||
<bibRefCitation id="504173D4FFFDFFBAFC9FF92821FFD02A" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[807,1009,1779,1805]" pageId="4" pageNumber="193" pagination="337 - 354" refId="ref5531" refString="Cunha, A. M., Terossi, M., Mantelatto, F. M. & Almeida, A. O. (2020) Delimiting the snapping shrimp Alpheus lobidens De Haan, 1849 (Caridea: Alpheidae) based on morphological and molecular data. Zootaxa, 4718 (3), 337 - 354. https: // doi. org / 10.11646 / zootaxa. 4718.3.3" type="journal article" year="2020">
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFC9FF928217ED02A" box="[807,880,1779,1805]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="193">Cunha</collectingCountry>
|
||
<emphasis id="06A4D237FFFDFFBAFCCFF92F21A0D02A" box="[887,942,1779,1805]" italics="true" pageId="4" pageNumber="193">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
). The colour pattern of
|
||
<taxonomicName id="F3D075A6FFFDFFBAFB4AF92F2792D02A" authorityName="sensu Heller" authorityYear="1862" box="[1266,1436,1780,1806]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFDFFBAFB4AF92F2792D02A" box="[1266,1436,1780,1806]" italics="true" pageId="4" pageNumber="193">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
is currently unknown, which is especially relevant in light of growing evidence of the presence of several cryptic or pseudocryptic species within the
|
||
<taxonomicName id="F3D075A6FFFDFFBAFD8CF8E720BAD072" authorityName="sensu De Haan" authorityYear="1849" box="[564,692,1851,1878]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFD8CF8E720BAD072" box="[564,692,1851,1878]" italics="true" pageId="4" pageNumber="193">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex distinguishable mainly by colour patterns, for instance, in
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFF0CF8842328D05E" box="[180,294,1887,1913]" name="Singapore" pageId="4" pageNumber="193">Singapore</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFFDFFBAFE8DF884205CD05D" author="Anker, A. & De Grave, S." box="[309,594,1887,1914]" pageId="4" pageNumber="193" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
),
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFDDDF88420B9D05E" box="[613,695,1887,1913]" name="Taiwan" pageId="4" pageNumber="193">Taiwan</collectingCountry>
|
||
(A. Anker, pers. obs.),
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFC78F8BB260ED05D" box="[960,1024,1888,1914]" name="Japan" pageId="4" pageNumber="193">Japan</collectingCountry>
|
||
(K. Nomura, pers. comm.), and
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFAD7F8BB2792D05D" box="[1391,1436,1888,1914]" name="Iran" pageId="4" pageNumber="193">Iran</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFFDFFBAFF27F8582393D0BA" author="Dehghani, A. & Sari A. & Naderloo, R." box="[159,413,1923,1949]" pageId="4" pageNumber="193" pagination="479 - 504" refId="ref5860" refString="Dehghani, A., Sari A. & Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
|
||
Dehghani
|
||
<emphasis id="06A4D237FFFDFFBAFEACF85F2343D0BA" box="[276,333,1923,1949]" italics="true" pageId="4" pageNumber="193">et al.</emphasis>
|
||
2019a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFDFFBBFF7FF8732054D6AD" blockId="4.[151,1437,1635,2022]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193">
|
||
<bibRefCitation id="504173D4FFFDFFBAFF7FF87323D9D0E5" author="Forest, J. & Guinot, D." box="[199,471,1959,1986]" pageId="4" pageNumber="193" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot (1958)</bibRefCitation>
|
||
were aware of the presence of two morphologically distinct species from the
|
||
<taxonomicName id="F3D075A6FFFDFFBAFAE4F87322EED0C2" authorityName="sensu De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFDFFBAFAE4F87322EED0C2" italics="true" pageId="4" pageNumber="193">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex on the Mediterranean coast of
|
||
<collectingCountry id="4CC74EB5FFFDFFBAFD10F81020E7D0C2" box="[680,745,1995,2021]" name="Israel" pageId="4" pageNumber="193">Israel</collectingCountry>
|
||
, viz.
|
||
<taxonomicName id="F3D075A6FFFDFFBAFC9EF81721B5D0C2" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[806,955,1996,2022]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFDFFBAFC9EF81721B5D0C2" box="[806,955,1996,2022]" italics="true" pageId="4" pageNumber="193">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
(reported as
|
||
<taxonomicName id="F3D075A6FFFDFFBAFBE8F81026A4D0C2" authorityName="Fabricius" authorityYear="1798" box="[1104,1194,1995,2021]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06A4D237FFFDFFBAFBE8F81026A4D0C2" box="[1104,1194,1995,2021]" italics="true" pageId="4" pageNumber="193">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
sp., as it was already being studied by Holthuis & Gottlieb) and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD3CFF4C210AD796" authorityName="sensu De Haan" authorityYear="1849" box="[644,772,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFD3CFF4C210AD796" box="[644,772,151,177]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(reported by them as
|
||
<taxonomicName id="F3D075A6FFFCFFBBFBBAFF4C26A1D797" authorityName="sensu Heller" authorityYear="1862" box="[1026,1199,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFCFFBBFBBAFF4C26A1D797" box="[1026,1199,151,177]" italics="true" pageId="5" pageNumber="194">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
). These authors also examined several specimens in Henri Coutière’s alpheid collection in the Muséum National d’Histoire Naturelle in Paris (MNHN), which morphologically corresponded well to
|
||
<taxonomicName id="F3D075A6FFFCFFBBFCCDFF042602D7DF" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[885,1036,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFCCDFF042602D7DF" box="[885,1036,223,249]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
, viz. two specimens from the Gulf of
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFF0FFED822F5D63A" box="[183,251,259,285]" name="Oman" pageId="5" pageNumber="194">Oman</collectingCountry>
|
||
and a mixed lot from
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFE45FED82053D63A" box="[509,605,259,285]" name="Djibouti" pageId="5" pageNumber="194">Djibouti</collectingCountry>
|
||
, containing both
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC9BFED821B4D63B" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[803,954,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFC9BFED821B4D63B" box="[803,954,259,285]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC4BFED8267DD63A" authorityName="sensu De Haan" authorityYear="1849" box="[1011,1139,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFC4BFED8267DD63A" box="[1011,1139,259,285]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
. Interestingly,
|
||
<bibRefCitation id="504173D4FFFCFFBBFAA5FED8234AD666" author="Holthuis, L. B. & Gottlieb, E." pageId="5" pageNumber="194" pagination="1 - 126" refId="ref6236" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb (1958)</bibRefCitation>
|
||
contrasted
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE7DFEFC2054D667" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[453,602,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFE7DFEFC2054D667" box="[453,602,295,321]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD22FEFC2116D666" authorityName="sensu De Haan" authorityYear="1849" box="[666,792,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFD22FEFC2116D666" box="[666,792,295,321]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
only superficially, comparing it with three more distantly related species of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFEDEFE9023CED642" authorityName="Fabricius" authorityYear="1798" box="[358,448,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06A4D237FFFCFFBBFEDEFE9023CED642" box="[358,448,331,357]" italics="true" pageId="5" pageNumber="194">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<bibRefCitation id="504173D4FFFCFFBBFD8DFE90214AD642" author="Forest, J. & Guinot, D." box="[565,836,330,357]" pageId="5" pageNumber="194" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot (1958)</bibRefCitation>
|
||
provided a full account of the morphological differences between the two taxa (see below).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFCFFBBFF7FFE48278BD5D6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="194">
|
||
<bibRefCitation id="504173D4FFFCFFBBFF7FFE4823D4D68A" author="Banner, D. M. & Banner, A. H." box="[199,474,403,429]" pageId="5" pageNumber="194" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner & Banner (1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFCFFBBFE5EFE482011D68A" author="Banner, D. M. & Banner, A. H." box="[486,543,403,429]" pageId="5" pageNumber="194" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
) questioned the separation of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFCCDFE482605D68B" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[885,1035,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFCCDFE482605D68B" box="[885,1035,403,429]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="F3D075A6FFFCFFBBFBE9FE4826DED68A" authorityName="sensu De Haan" authorityYear="1849" box="[1105,1232,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFBE9FE4826DED68A" box="[1105,1232,403,429]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
and synonymised the former species. The Hawaiian subspecies,
|
||
<emphasis id="06A4D237FFFCFFBBFD1CFE6C21BBD6F6" box="[676,949,439,465]" italics="true" pageId="5" pageNumber="194">
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD1CFE6C21BED6F6" authorityName="Banner & Banner" authorityYear="1974" box="[676,944,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="polynesicus">A. lobidens polynesicus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
was also placed in the synonymy of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFAE5FE6C22EED6D2" authorityName="sensu De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFAE5FE6C22EED6D2" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
, despite some rather striking colour pattern differences (
|
||
<bibRefCitation id="504173D4FFFCFFBBFCD3FE002672D6D2" author="Banner, D. M. & Banner, A. H." box="[875,1148,475,501]" pageId="5" pageNumber="194" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner & Banner 1981</bibRefCitation>
|
||
). However, at least some of the synonymies proposed by
|
||
<bibRefCitation id="504173D4FFFCFFBBFE45FE242100D53E" author="Banner, D. M. & Banner, A. H." box="[509,782,511,537]" pageId="5" pageNumber="194" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner & Banner (1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFCFFBBFCA2FE24215DD53E" author="Banner, D. M. & Banner, A. H." box="[794,851,511,537]" pageId="5" pageNumber="194" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
) in the
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC11FE242626D53E" authorityName="sensu De Haan" authorityYear="1849" box="[937,1064,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFC11FE242626D53E" box="[937,1064,511,537]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex were premature (
|
||
<bibRefCitation id="504173D4FFFCFFBBFAECFE2422DCD51A" author="Anker, A." pageId="5" pageNumber="194" pagination="57 - 72" refId="ref4944" refString="Anker, A. (2001) Two new species of snapping shrimps from the Indo-Pacific, with remarks on colour patterns and sibling species in Alpheidae (Crustacea: Caridea). Raffles Bulletin of Zoology, 49, 57 - 72." type="journal article" year="2001">Anker 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="504173D4FFFCFFBBFF63FDF823FED519" author="Anker, A. & De Grave, S." box="[219,496,547,574]" pageId="5" pageNumber="194" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
). New material collected throughout the Indo-West Pacific clearly shows that several features of the colour pattern, such as the presence or absence of dark transverse bands on the pleon, the presence or absence of dark spots on the sides of the pleon, the markings on the mesial side of the major and minor chelae and the colour of the dactylar plunger, may provide additional differentiating characters between the species of the
|
||
<taxonomicName id="F3D075A6FFFCFFBBFAE7FD5422EED5EA" authorityName="sensu De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFAE7FD5422EED5EA" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<bibRefCitation id="504173D4FFFCFFBBFEEDFD682063D5EA" author="Anker, A. & De Grave, S." box="[341,621,691,717]" pageId="5" pageNumber="194" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
; A. Anker, pers. obs., see also
|
||
<figureCitation id="ACEB12A0FFFCFFBBFC04FD6821F1D5EA" box="[956,1023,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="5" pageNumber="194">Fig. 3</figureCitation>
|
||
).
|
||
<bibRefCitation id="504173D4FFFCFFBBFBABFD682744D5EA" author="De Grave, S. & Fransen, C. H. J. M." box="[1043,1354,691,717]" pageId="5" pageNumber="194" pagination="195 - 589" refId="ref5671" refString="De Grave, S. & Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85, 195 - 589." type="journal article" year="2011">De Grave & Fransen (2011)</bibRefCitation>
|
||
treated
|
||
<taxonomicName id="F3D075A6FFFCFFBBFF2FFD0C2322D5D7" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[151,300,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFF2FFD0C2322D5D7" box="[151,300,727,753]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
as a valid species, but continued to list
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD51FD0C21FFD5D6" authorityName="Banner & Banner" authorityYear="1974" box="[745,1009,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="polynesicus">
|
||
<emphasis id="06A4D237FFFCFFBBFD51FD0C21FFD5D6" box="[745,1009,727,753]" italics="true" pageId="5" pageNumber="194">A. lobidens polynesicus</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFAB8FD0C2770D5D6" authorityName="sensu De Haan" authorityYear="1849" box="[1280,1406,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFAB8FD0C2770D5D6" box="[1280,1406,727,753]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFCFFBBFF7FFD20269AD312" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="194">
|
||
The present material confirms that
|
||
<taxonomicName id="F3D075A6FFFCFFBBFDE6FD2020F8D433" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[606,758,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFDE6FD2020F8D433" box="[606,758,763,789]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
is clearly distinct from the west Indian Ocean taxon often recorded as
|
||
<taxonomicName id="F3D075A6FFFCFFBBFEA7FCC42390D41E" authorityName="sensu De Haan" authorityYear="1849" box="[287,414,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFEA7FCC42390D41E" box="[287,414,799,825]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
and thus represents a valid species (see also
|
||
<bibRefCitation id="504173D4FFFCFFBBFC22FCC4266CD41E" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[922,1122,799,825]" pageId="5" pageNumber="194" pagination="337 - 354" refId="ref5531" refString="Cunha, A. M., Terossi, M., Mantelatto, F. M. & Almeida, A. O. (2020) Delimiting the snapping shrimp Alpheus lobidens De Haan, 1849 (Caridea: Alpheidae) based on morphological and molecular data. Zootaxa, 4718 (3), 337 - 354. https: // doi. org / 10.11646 / zootaxa. 4718.3.3" type="journal article" year="2020">
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFC22FCC421EDD41E" box="[922,995,799,825]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="5" pageNumber="194">Cunha</collectingCountry>
|
||
<emphasis id="06A4D237FFFCFFBBFC52FCC4262DD41E" box="[1002,1059,799,825]" italics="true" pageId="5" pageNumber="194">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
for molecular comparison). The common and widespread taxon from the western Indian Ocean (e.g., Red Sea, Gulf,
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFB1BFC982708D479" box="[1187,1286,835,862]" name="Pakistan" pageId="5" pageNumber="194">Pakistan</collectingCountry>
|
||
), which penetrated into the Mediterranean Sea most likely does not represent
|
||
<taxonomicName id="F3D075A6FFFCFFBBFCC8FCBC26E5D4A6" authority="sensu De Haan (1849)" authorityName="sensu De Haan" authorityYear="1849" box="[880,1259,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFCC8FCBC263FD4A7" box="[880,1073,871,897]" italics="true" pageId="5" pageNumber="194">A. lobidens sensu</emphasis>
|
||
De Haan (1849)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFA96FCBC2760D4A6" box="[1326,1390,871,897]" name="Japan" pageId="5" pageNumber="194">Japan</collectingCountry>
|
||
and is herein referred to
|
||
<taxonomicName id="F3D075A6FFFCFFBBFEC7FC50202AD482" authorityName="sensu De Haan" authorityYear="1849" box="[383,548,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFEC7FC502396D482" box="[383,408,907,933]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFE7DFC50202AD482" box="[453,548,907,933]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
(see Comparative material). Whether this taxon is identical to
|
||
<taxonomicName id="F3D075A6FFFCFFBBFB48FC50237DD4ED" authority="sensu Heller (1862)" authorityName="sensu Heller" authorityYear="1862" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFCFFBBFB48FC5022DBD4EF" italics="true" pageId="5" pageNumber="194">A. crassimanus sensu</emphasis>
|
||
<bibRefCitation id="504173D4FFFCFFBBFF64FC74237DD4ED" author="Heller, C." box="[220,371,943,970]" pageId="5" pageNumber="194" pagination="519 - 528" refId="ref6196" refString="Heller, C. (1862) Neue Crustaceen gesammelt wahrend der Weltumsegelung der k. k. Fregatte Novara. Zweiter vorlaufiger Beri- cht. Verhandlungen der Koeniglich-Kaiserlichen Zoologisch-Botanischen Gesellschaft, Wien, 28, 519 - 528." type="journal article" year="1862">Heller (1862)</bibRefCitation>
|
||
</taxonomicName>
|
||
remains to be shown.
|
||
<taxonomicName id="F3D075A6FFFCFFBBFDD5FC74214DD4EF" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[621,835,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFDD5FC74214DD4EF" box="[621,835,943,969]" italics="true" pageId="5" pageNumber="194">Alpheus inopinatus</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="F3D075A6FFFCFFBBFBFEFC7426E9D4EE" authorityName="sensu De Haan" authorityYear="1849" box="[1094,1255,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFBFEFC742650D4EE" box="[1094,1118,943,969]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFB30FC7426E9D4EE" box="[1160,1255,943,969]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
by at least three morphological characters, in addition to a small but consistent difference in their colour patterns. Both species can also be separated morphologically from
|
||
<taxonomicName id="F3D075A6FFFCFFBBFDE6FC2C2166D336" authorityName="Banner & Banner" authorityYear="1974" box="[606,872,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="polynesicus">
|
||
<emphasis id="06A4D237FFFCFFBBFDE6FC2C2166D336" box="[606,872,1015,1041]" italics="true" pageId="5" pageNumber="194">A. lobidens polynesicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC27FC2C2670D336" authorityName="Dehghani, Sari & Naderloo" authorityYear="2019" box="[927,1150,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="mohammadpouri">
|
||
<emphasis id="06A4D237FFFCFFBBFC27FC2C2670D336" box="[927,1150,1015,1041]" italics="true" pageId="5" pageNumber="194">A. mohammadpouri</emphasis>
|
||
</taxonomicName>
|
||
. The taxonomic status of these taxa will be discussed elsewhere, as part of a larger study of the
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC1FFBC0262BD312" authorityName="sensu De Haan" authorityYear="1849" box="[935,1061,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFC1FFBC0262BD312" box="[935,1061,1051,1077]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFCFFBBFF7FFBE423B1D2E5" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="194">
|
||
Many of the differentiating characters between
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD54FBE4218AD37F" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[748,900,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFD54FBE4218AD37F" box="[748,900,1087,1113]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC79FBE42666D37E" authorityName="sensu De Haan" authorityYear="1849" box="[961,1128,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFC79FBE421D7D37E" box="[961,985,1087,1113]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFBB1FBE42666D37E" box="[1033,1128,1087,1113]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
listed by
|
||
<bibRefCitation id="504173D4FFFCFFBBFB67FBE422ECD35A" author="Forest, J. & Guinot, D." pageId="5" pageNumber="194" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot (1958)</bibRefCitation>
|
||
were confirmed in the present material. The most important character separating the two species is the absence of a sharp distomesial tooth on the meri of the major and minor cheliped in the former species. In some specimens of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFF44FB70239DD3E3" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[252,403,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFF44FB70239DD3E3" box="[252,403,1195,1221]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
, the distomesial margin terminates in a subacute angle, sometimes protruding as a minute tooth, which is not the same structure as a much stronger acute tooth, situated slightly below the distomesial angle, in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFF0CFB28235BD22A" authorityName="sensu De Haan" authorityYear="1849" box="[180,341,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFF0CFB2822C2D22A" box="[180,204,1267,1293]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFF4EFB28235BD22A" box="[246,341,1267,1293]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="504173D4FFFCFFBBFE30FB28208CD22A" author="Forest, J. & Guinot, D." box="[392,642,1266,1293]" pageId="5" pageNumber="194" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot 1958</bibRefCitation>
|
||
: figs. 1, 3, 4). The male minor cheliped has a much stouter merus and a stouter and more sculptured chela in
|
||
<emphasis id="06A4D237FFFCFFBBFD80FACC20DAD217" box="[568,724,1303,1329]" italics="true" pageId="5" pageNumber="194">
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD80FACC20DED217" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[568,720,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">A. inopinatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
compared to the noticeably slenderer merus and chela of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFAD8FACC22F8D272" authorityName="sensu De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFAD8FACC2776D216" box="[1376,1400,1303,1329]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFF2FFAE022F8D272" box="[151,246,1339,1365]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="504173D4FFFCFFBBFE92FAE0202BD272" author="Forest, J. & Guinot, D." box="[298,549,1338,1365]" pageId="5" pageNumber="194" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot 1958</bibRefCitation>
|
||
: figs. 2, 5). The third and fourth pereiopods are also stouter in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFB59FAE02779D273" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[1249,1399,1339,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFB59FAE02779D273" box="[1249,1399,1339,1365]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
, as expressed by the more inflated merus, with a length-width ratio of about 3.0 (cf.
|
||
<bibRefCitation id="504173D4FFFCFFBBFB91FA842728D25E" author="Forest, J. & Guinot, D." box="[1065,1318,1374,1401]" pageId="5" pageNumber="194" pagination="4 - 16" refId="ref6060" refString="Forest, J. & Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest & Guinot 1958</bibRefCitation>
|
||
: fig. 7)
|
||
<emphasis id="06A4D237FFFCFFBBFAC6FA842792D25F" box="[1406,1436,1375,1400]" italics="true" pageId="5" pageNumber="194">vs.</emphasis>
|
||
more than 4.0 in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFEE1FA5823F3D2BA" authorityName="sensu De Haan" authorityYear="1849" box="[345,509,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFEE1FA58237CD2BA" box="[345,370,1411,1437]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFE26FA5823F3D2BA" box="[414,509,1411,1437]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFDFBFA58209BD2BA" box="[579,661,1411,1437]" name="Kuwait" pageId="5" pageNumber="194">Kuwait</collectingCountry>
|
||
or the specimens from
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFC18FA582609D2BA" box="[928,1031,1411,1437]" name="Australia" pageId="5" pageNumber="194">Australia</collectingCountry>
|
||
identified as
|
||
<taxonomicName id="F3D075A6FFFCFFBBFB19FA58272ED2BA" authorityName="sensu De Haan" authorityYear="1849" box="[1185,1312,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFB19FA58272ED2BA" box="[1185,1312,1411,1437]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="504173D4FFFCFFBBFAF3FA58235FD2E6" author="Banner, D. M. & Banner, A. H." pageId="5" pageNumber="194" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner (1982</bibRefCitation>
|
||
: fig. 78f).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFCFFBBFF7FFA1021B5D06A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="194">
|
||
In life,
|
||
<taxonomicName id="F3D075A6FFFCFFBBFEA4FA1023BDD2C3" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[284,435,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFEA4FA1023BDD2C3" box="[284,435,1483,1509]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE55FA10209DD2C2" authorityName="sensu De Haan" authorityYear="1849" box="[493,659,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFE55FA102008D2C2" box="[493,518,1483,1509]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFD8CFA10209DD2C2" box="[564,659,1483,1509]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
can be separated by their colour patterns, although the difference is rather subtle. The pleon of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE7AFA342059D12F" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[450,599,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFE7AFA342059D12F" box="[450,599,1519,1545]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
presents three pairs of small dark spots: one pair of lateral spots on the second pleuron, one pair of very small dorsolateral spots on the third pleuron and one pair of more conspicuous lateral spots on the fourth pleuron (
|
||
<figureCitation id="ACEB12A0FFFCFFBBFE6BF9EC201AD176" box="[467,532,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1547,1571]" captionTargetBox="[151,1435,179,1521]" captionTargetId="figure-18@2.[151,1435,179,1521]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus inopinatus Holthuis & Gottlieb, 1958: male from Kuwait, OUMNH.ZC.2020-01-001 [fcn KUW-011], A—dorsal view; B—lateral view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407194" httpUri="https://zenodo.org/record/4407194/files/figure.png" pageId="5" pageNumber="194">Fig. 1</figureCitation>
|
||
); all these spots are absent in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFCDCF9EC260BD176" authorityName="sensu De Haan" authorityYear="1849" box="[868,1029,1591,1617]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFCDCF9EC2172D176" box="[868,892,1591,1617]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFC1EF9EC260BD176" box="[934,1029,1591,1617]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="ACEB12A0FFFCFFBBFBACF9EC2657D176" box="[1044,1113,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1563,1587]" captionTargetBox="[151,1435,181,1540]" captionTargetId="figure-19@3.[151,1435,181,1540]" captionTargetPageId="3" captionText="FIGURE 2. Alpheus cf. lobidens De Haan, 1849: male from Kuwait, OUMNH.ZC.2020-01-005 [fcn KUW-027], A—dorsal view; B—lateral view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407198" httpUri="https://zenodo.org/record/4407198/files/figure.png" pageId="5" pageNumber="194">Fig. 2</figureCitation>
|
||
). The dactylar plunger of the major chela of both species is similar in colour, whitish with creamy or yellow tinge, although in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFB6FF9802762D153" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[1239,1388,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFB6FF9802762D153" box="[1239,1388,1627,1653]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
, the base of the plunger has a bluish tinge (
|
||
<figureCitation id="ACEB12A0FFFCFFBBFDECF9A420A2D1BE" box="[596,684,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="5" pageNumber="194">Fig. 3C</figureCitation>
|
||
), whilst in
|
||
<taxonomicName id="F3D075A6FFFCFFBBFC97F9A421DAD1BE" authorityName="sensu De Haan" authorityYear="1849" box="[815,980,1663,1689]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFC97F9A42149D1BE" box="[815,839,1663,1689]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFCCDF9A421DAD1BE" box="[885,980,1663,1689]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
, the anterior surface has a more intense pale orange colour (
|
||
<figureCitation id="ACEB12A0FFFCFFBBFEC2F97823DED19A" box="[378,464,1699,1725]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="5" pageNumber="194">Fig. 3F</figureCitation>
|
||
). The Iranian material of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD4BF978217CD19A" authorityName="sensu De Haan" authorityYear="1849" box="[755,882,1699,1725]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFD4BF978217CD19A" box="[755,882,1699,1725]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
listed by
|
||
<bibRefCitation id="504173D4FFFCFFBBFC5AF97826E1D19A" author="Dehghani, A. & Sari A. & Naderloo, R." box="[994,1263,1699,1725]" pageId="5" pageNumber="194" pagination="479 - 504" refId="ref5860" refString="Dehghani, A., Sari A. & Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
|
||
Dehghani
|
||
<emphasis id="06A4D237FFFCFFBBFBEFF978269ED19A" box="[1111,1168,1699,1725]" italics="true" pageId="5" pageNumber="194">et al.</emphasis>
|
||
(2019a)
|
||
</bibRefCitation>
|
||
likely contains both
|
||
<taxonomicName id="F3D075A6FFFCFFBBFF69F91C2369D1C7" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[209,359,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFF69F91C2369D1C7" box="[209,359,1735,1761]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE18F91C204AD1C6" authorityName="sensu De Haan" authorityYear="1849" box="[416,580,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFE18F91C23B6D1C6" box="[416,440,1735,1761]" italics="true" pageId="5" pageNumber="194">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFCFFBBFE5DF91C204AD1C6" box="[485,580,1735,1761]" italics="true" pageId="5" pageNumber="194">lobidens</emphasis>
|
||
</taxonomicName>
|
||
, based on an analysis of the colour patterns of the four illustrated individuals. Generally, many records under the name “
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD1FF9302129D022" authorityName="sensu De Haan" authorityYear="1849" box="[679,807,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFD1FF9302129D022" box="[679,807,1771,1797]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
” from the Gulf,
|
||
<collectingRegion id="F614C0C7FFFCFFBBFC50F9302646D022" box="[1000,1096,1771,1797]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="194">Red Sea</collectingRegion>
|
||
, Mediterranean Sea and other localities may refer to
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE2EF8D42022D00F" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[406,556,1807,1833]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFE2EF8D42022D00F" box="[406,556,1807,1833]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
as the two species have been commonly confused in the past, especially after the synonymisation of
|
||
<taxonomicName id="F3D075A6FFFCFFBBFE2FF8E82022D06B" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[407,556,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFCFFBBFE2FF8E82022D06B" box="[407,556,1843,1869]" italics="true" pageId="5" pageNumber="194">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="504173D4FFFCFFBBFDEEF8E82168D06A" author="Banner, D. M. & Banner, A. H." box="[598,870,1843,1869]" pageId="5" pageNumber="194" pagination="1 - 99" refId="ref5379" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner & Banner (1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="504173D4FFFCFFBBFCCAF8E821A0D06A" author="Banner, D. M. & Banner, A. H." box="[882,942,1843,1869]" pageId="5" pageNumber="194" pagination="1 - 357" refId="ref5421" refString="Banner, D. M. & Banner, A. H. (1982) The Alpheid Shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFCFFB8FF7FF88C2311D6D1" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,1473]" lastPageId="6" lastPageNumber="195" pageId="5" pageNumber="194">
|
||
<bibRefCitation id="504173D4FFFCFFBBFF7FF88C23F6D055" author="Anker, A. & De Grave, S." box="[199,504,1879,1906]" pageId="5" pageNumber="194" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave (2016)</bibRefCitation>
|
||
, while reporting
|
||
<taxonomicName id="F3D075A6FFFCFFBBFD03F88C2134D056" authorityName="sensu De Haan" authorityYear="1849" box="[699,826,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFD03F88C2134D056" box="[699,826,1879,1905]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<emphasis id="06A4D237FFFCFFBBFCD4F88C2669D056" box="[876,1127,1879,1905]" italics="true" pageId="5" pageNumber="194">
|
||
<taxonomicName id="F3D075A6FFFCFFBBFCD4F88C263CD057" box="[876,1074,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="195" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="sensu">
|
||
A. lobidens
|
||
<taxonomicNameLabel id="1D976F4CFFFCFFBBFC4CF88C263CD057" box="[1012,1074,1879,1904]" pageId="5" pageNumber="194" sensu="lato">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
) from
|
||
<collectingCountry id="4CC74EB5FFFCFFBBFB00F88C2722D056" box="[1208,1324,1879,1905]" name="Singapore" pageId="5" pageNumber="194">Singapore</collectingCountry>
|
||
, provided colour photographs of three different colour morphs, possibly three different species, of the
|
||
<taxonomicName id="F3D075A6FFFCFFBBFB79F8A0274CD0B2" authorityName="sensu De Haan" authorityYear="1849" box="[1217,1346,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFCFFBBFB79F8A0274CD0B2" box="[1217,1346,1915,1941]" italics="true" pageId="5" pageNumber="194">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
species complex. They can be briefly described as (1) pleon non-banded, with two conspicuous lateral black spots on the second and fourth pleura and two small medially fused black spots on the dorsal surface of the fifth pleonite (
|
||
<bibRefCitation id="504173D4FFFCFFB8FAECF818236CD796" author="Anker, A. & De Grave, S." lastPageId="6" lastPageNumber="195" pageId="5" pageNumber="194" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
: fig. 16A, E); (2) pleon banded, with two small dorsolateral black spots on the third pleuron, two very large and conspicuous black lateral spots on the fourth pleuron and two smaller black lateral patches on the fifth pleuron (
|
||
<emphasis id="06A4D237FFFFFFB8FF44FF04233CD7DE" box="[252,306,223,249]" italics="true" pageId="6" pageNumber="195">idem</emphasis>
|
||
.: fig. 16C, D); and (3) pleon non-banded, without spots (
|
||
<emphasis id="06A4D237FFFFFFB8FC16FF0421EAD7DE" box="[942,996,223,249]" italics="true" pageId="6" pageNumber="195">idem</emphasis>
|
||
.: fig. 17). The arrangement of the spots of the Singaporean specimens appears to be different from that of
|
||
<taxonomicName id="F3D075A6FFFFFFB8FCCEFED82605D63B" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[886,1035,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFFFFB8FCCEFED82605D63B" box="[886,1035,259,285]" italics="true" pageId="6" pageNumber="195">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the dactylar plunger of the major chela of the two black-spotted colour morphs from
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FCFCFEFC21B8D666" box="[836,950,295,321]" name="Singapore" pageId="6" pageNumber="195">Singapore</collectingCountry>
|
||
is bright orange, which makes them clearly different from both
|
||
<taxonomicName id="F3D075A6FFFFFFB8FECEFE902002D643" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[374,524,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFFFFB8FECEFE902002D643" box="[374,524,331,357]" italics="true" pageId="6" pageNumber="195">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
and the specimens herein identified as
|
||
<taxonomicName id="F3D075A6FFFFFFB8FC72FE902663D642" authorityName="sensu De Haan" authorityYear="1849" box="[970,1133,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FC72FE9021ECD642" box="[970,994,331,357]" italics="true" pageId="6" pageNumber="195">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFFFFB8FBB6FE902663D642" box="[1038,1133,331,357]" italics="true" pageId="6" pageNumber="195">lobidens</emphasis>
|
||
</taxonomicName>
|
||
, both with whitish-creamy plungers (
|
||
<figureCitation id="ACEB12A0FFFFFFB8FEB0FEB42371D6AE" box="[264,383,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1421,1445]" captionTargetBox="[151,1435,181,1397]" captionTargetId="figure-18@4.[151,1435,181,1397]" captionTargetPageId="4" captionText="FIGURE 3. Alpheus inopinatus Holthuis & Gottlieb, 1958 (A–C) and Alpheus cf. lobidens De Haan, 1849 (D–F), male chelipeds, all specimens from Kuwait, A—major chela of male, OUMNH.ZC. 2020-01-003 [fcn KUW-034]; B—minor chela of the same male; C—major chela of male, OUMNH.ZC.2020-01-001 [fcn KUW-011], fingers opened to show the plunger; D—major chela of male, OUMNH.ZC. 2020-01-008 [fcn KUW-036]; E—minor chela of the same male; F—major chela of male, OUMNH.ZC.2020-01-005 [fcn KUW-027], fingers opened to show the plunger. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4407200" httpUri="https://zenodo.org/record/4407200/files/figure.png" pageId="6" pageNumber="195">Fig. 3C, F</figureCitation>
|
||
). Therefore, all material identified as
|
||
<taxonomicName id="F3D075A6FFFFFFB8FC91FEB421B1D6AF" authorityName="Holthuis & Gottlieb" authorityYear="1958" box="[809,959,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="inopinatus">
|
||
<emphasis id="06A4D237FFFFFFB8FC91FEB421B1D6AF" box="[809,959,367,393]" italics="true" pageId="6" pageNumber="195">A. inopinatus</emphasis>
|
||
</taxonomicName>
|
||
from outside of the western Indian Ocean (
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FF27FE482327D68A" box="[159,297,403,429]" name="Hong Kong" pageId="6" pageNumber="195">Hong Kong</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FE8DFE482384D68A" box="[309,394,403,429]" name="Taiwan" pageId="6" pageNumber="195">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FE2EFE4823F1D689" box="[406,511,403,430]" name="Australia" pageId="6" pageNumber="195">Australia</collectingCountry>
|
||
) needs to be carefully re-examined. These observations also contribute to the remaining uncertainty about the taxonomic status of
|
||
<taxonomicName id="F3D075A6FFFFFFB8FD69FE6C2172D6F7" authorityName="sensu Heller" authorityYear="1862" box="[721,892,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="crassimanus">
|
||
<emphasis id="06A4D237FFFFFFB8FD69FE6C2172D6F7" box="[721,892,439,465]" italics="true" pageId="6" pageNumber="195">A. crassimanus</emphasis>
|
||
</taxonomicName>
|
||
, currently regarded as a synonym of
|
||
<taxonomicName id="F3D075A6FFFFFFB8FAA6FE6C2792D6F6" authorityName="sensu De Haan" authorityYear="1849" box="[1310,1436,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FAA6FE6C2792D6F6" box="[1310,1436,439,465]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(see above).
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFFFFB8FF7FFE242634D3E2" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="195">
|
||
Recently,
|
||
<bibRefCitation id="504173D4FFFFFFB8FE80FE242018D53E" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[312,534,511,537]" pageId="6" pageNumber="195" pagination="337 - 354" refId="ref5531" refString="Cunha, A. M., Terossi, M., Mantelatto, F. M. & Almeida, A. O. (2020) Delimiting the snapping shrimp Alpheus lobidens De Haan, 1849 (Caridea: Alpheidae) based on morphological and molecular data. Zootaxa, 4718 (3), 337 - 354. https: // doi. org / 10.11646 / zootaxa. 4718.3.3" type="journal article" year="2020">
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FE80FE24238FD53E" box="[312,385,511,537]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="195">Cunha</collectingCountry>
|
||
<emphasis id="06A4D237FFFFFFB8FE32FE2423CDD53E" box="[394,451,511,537]" italics="true" pageId="6" pageNumber="195">et al.</emphasis>
|
||
(2020)
|
||
</bibRefCitation>
|
||
redescribed
|
||
<taxonomicName id="F3D075A6FFFFFFB8FD10FE242129D53E" authorityName="sensu De Haan" authorityYear="1849" box="[680,807,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FD10FE242129D53E" box="[680,807,511,537]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="F614C0C7FFFFFFB8FCD6FE2421D8D53E" box="[878,982,511,537]" country="Japan" name="Nagasaki" pageId="6" pageNumber="195">Nagasaki</collectingRegion>
|
||
on the basis of what they considered to be topotypic specimens, collected from the vicinity of
|
||
<collectingRegion id="F614C0C7FFFFFFB8FD4FFDF8216DD51A" box="[759,867,547,573]" country="Japan" name="Nagasaki" pageId="6" pageNumber="195">Nagasaki</collectingRegion>
|
||
. However, the type locality of
|
||
<taxonomicName id="F3D075A6FFFFFFB8FB00FDF82738D51A" authorityName="sensu De Haan" authorityYear="1849" box="[1208,1334,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FB00FDF82738D51A" box="[1208,1334,547,573]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FAEDFDF82796D51A" box="[1365,1432,547,573]" name="Japan" pageId="6" pageNumber="195">Japan</collectingCountry>
|
||
, i.e., the entire Japanese archipelago and not specifically
|
||
<collectingRegion id="F614C0C7FFFFFFB8FCAAFD9C2174D546" box="[786,890,583,609]" country="Japan" name="Nagasaki" pageId="6" pageNumber="195">Nagasaki</collectingRegion>
|
||
(
|
||
<bibRefCitation id="504173D4FFFFFFB8FC32FD9C26FCD546" author="De Grave, S. & Fransen, C. H. J. M." box="[906,1266,582,609]" pageId="6" pageNumber="195" pagination="195 - 589" refId="ref5671" refString="De Grave, S. & Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85, 195 - 589." type="journal article" year="2011">De Grave & Fransen 2011: 385</bibRefCitation>
|
||
). Only six species described in
|
||
<bibRefCitation id="504173D4FFFFFFB8FED8FDB02001D5A2" author="De Haan, W." box="[352,527,618,645]" pageId="6" pageNumber="195" pagination="1 - 243" refId="ref5723" refString="De Haan, W. (1833 - 1850) Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam, jussuet auspicilis superiorum, qui summum in India Batava Imperium tenent, suspecto, annis 1823 - 1830 collegit, notis, observationibus et dumbrationibus illustravit. Fasc. 1 - 8. Apud Auctorem, Lugduni Batavorum, xvii + xxxi + 244 pp., 74 pls. [pp. i-xvii + i-xxxi + 1 - 244, pls. 1 - 55 + A-Q + 1 + 2. pp. i-xxi + vii-xvii + ix-xvi + 1 - 243, pls. 1 - 55 + A-Q, Circ. pl. 2]" type="book chapter" year="1833">De Haan (1833</bibRefCitation>
|
||
–1850) actually have a type locality that is geographically more restricted than merely
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FF55FD542323D58E" box="[237,301,655,681]" name="Japan" pageId="6" pageNumber="195">Japan</collectingCountry>
|
||
(see Yamagushi & Baba 1993), so it cannot be assumed that the
|
||
<typeStatus id="EB6BB087FFFFFFB8FBB6FD54267ED58E" box="[1038,1136,655,681]" pageId="6" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="F3D075A6FFFFFFB8FB2FFD542718D58E" authorityName="sensu De Haan" authorityYear="1849" box="[1175,1302,655,681]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FB2FFD542718D58E" box="[1175,1302,655,681]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
was indeed collected in the
|
||
<collectingRegion id="F614C0C7FFFFFFB8FEF1FD6823BFD5EA" box="[329,433,691,717]" country="Japan" name="Nagasaki" pageId="6" pageNumber="195">Nagasaki</collectingRegion>
|
||
area. Furthermore, it is well known that the botanical and zoological collections gathered by Von Siebold and Bürger, a part of which was studied by De Haan, came from all over
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FB37FD0C26DCD5D6" box="[1167,1234,727,753]" name="Japan" pageId="6" pageNumber="195">Japan</collectingCountry>
|
||
. For example, the type series of the giant spider crab
|
||
<taxonomicName id="F3D075A6FFFFFFB8FD9DFD2021CED431" authority="(Temminck)" baseAuthorityName="Temminck" baseAuthorityYear="1836" box="[549,960,763,790]" class="Malacostraca" family="Inachidae" genus="Macrocheira" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="kaempferi">
|
||
<emphasis id="06A4D237FFFFFFB8FD9DFD202120D432" box="[549,814,763,789]" italics="true" pageId="6" pageNumber="195">Macrocheira kaempferi</emphasis>
|
||
(Temminck)
|
||
</taxonomicName>
|
||
was collected in Suruga Province, further east, whereas their collection also contained cold-water decapods from much further north, e.g.,
|
||
<taxonomicName id="F3D075A6FFFFFFB8FB5AFCC4235DD47A" authority="Von Siebold" authorityName="Von Siebold" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="japonicus">
|
||
<emphasis id="06A4D237FFFFFFB8FB5AFCC422B2D47B" italics="true" pageId="6" pageNumber="195">Lithodes japonicus</emphasis>
|
||
Von Siebold
|
||
</taxonomicName>
|
||
, now known as
|
||
<taxonomicName id="F3D075A6FFFFFFB8FDA9FC9821CCD479" authority="(Tilesius)" baseAuthorityName="Tilesius" baseAuthorityYear="1815" box="[529,962,835,862]" class="Malacostraca" family="Lithodidae" genus="Paralithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="camtschaticus">
|
||
<emphasis id="06A4D237FFFFFFB8FDA9FC982145D47A" box="[529,843,835,861]" italics="true" pageId="6" pageNumber="195">Paralithodes camtschaticus</emphasis>
|
||
(Tilesius)
|
||
</taxonomicName>
|
||
. The authors simply “…assumed that the topotypes from Shimabara,
|
||
<collectingRegion id="F614C0C7FFFFFFB8FE68FCBC2035D4A6" box="[464,571,871,897]" country="Japan" name="Nagasaki" pageId="6" pageNumber="195">Nagasaki</collectingRegion>
|
||
,
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FDFFFCBC2089D4A6" box="[583,647,871,897]" name="Japan" pageId="6" pageNumber="195">Japan</collectingCountry>
|
||
(RMNH PEG 25134) correspond to
|
||
<emphasis id="06A4D237FFFFFFB8FB9FFCBC2735D4A7" box="[1063,1339,871,897]" italics="true" pageId="6" pageNumber="195">
|
||
<taxonomicName id="F3D075A6FFFFFFB8FB9FFCBC26E5D4A7" box="[1063,1259,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="sensu">
|
||
A. lobidens
|
||
<taxonomicNameLabel id="1D976F4CFFFFFFB8FB15FCBC26E5D4A7" box="[1197,1259,871,896]" pageId="6" pageNumber="195" sensu="stricto">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
stricto
|
||
</emphasis>
|
||
, i.e., are conspecifics of the damaged
|
||
<typeStatus id="EB6BB087FFFFFFB8FE65FC502049D482" box="[477,583,907,933]" pageId="6" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
…”, without a direct comparison to the
|
||
<typeStatus id="EB6BB087FFFFFFB8FC46FC50266AD482" box="[1022,1124,907,933]" pageId="6" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
. Although it is possible that
|
||
<bibRefCitation id="504173D4FFFFFFB8FF2FFC74237FD4EE" author="Cunha, A. M. & Terossi, M. & Mantelatto, F. M. & Almeida, A. O." box="[151,369,943,969]" pageId="6" pageNumber="195" pagination="337 - 354" refId="ref5531" refString="Cunha, A. M., Terossi, M., Mantelatto, F. M. & Almeida, A. O. (2020) Delimiting the snapping shrimp Alpheus lobidens De Haan, 1849 (Caridea: Alpheidae) based on morphological and molecular data. Zootaxa, 4718 (3), 337 - 354. https: // doi. org / 10.11646 / zootaxa. 4718.3.3" type="journal article" year="2020">
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FF2FFC7422EED4EE" box="[151,224,943,969]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="195">Cunha</collectingCountry>
|
||
<emphasis id="06A4D237FFFFFFB8FF5FFC74232ED4EE" box="[231,288,943,969]" italics="true" pageId="6" pageNumber="195">et al.</emphasis>
|
||
(2020)
|
||
</bibRefCitation>
|
||
did redescribe what in future could be considered as true
|
||
<taxonomicName id="F3D075A6FFFFFFB8FC41FC742676D4EE" authorityName="sensu De Haan" authorityYear="1849" box="[1017,1144,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FC41FC742676D4EE" box="[1017,1144,943,969]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
(i.e.
|
||
<taxonomicName id="F3D075A6FFFFFFB8FB09FC742310D4CA" authority="sensu De Haan, 1849" authorityName="sensu De Haan" authorityYear="1849" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FB09FC74277AD4EF" box="[1201,1396,943,969]" italics="true" pageId="6" pageNumber="195">A. lobidens sensu</emphasis>
|
||
De Haan, 1849
|
||
</taxonomicName>
|
||
), it should be noted that numerous species of the
|
||
<taxonomicName id="F3D075A6FFFFFFB8FCEBFC0821DCD4CA" authorityName="sensu De Haan" authorityYear="1849" box="[851,978,979,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FCEBFC0821DCD4CA" box="[851,978,979,1005]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex occur in
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FB10FC0826E5D4CA" box="[1192,1259,979,1005]" name="Japan" pageId="6" pageNumber="195">Japan</collectingCountry>
|
||
, an observation based primarily on the diversity of colour patterns (K. Nomura, pers. comm.). As no colour pattern was documented for
|
||
<taxonomicName id="F3D075A6FFFFFFB8FF07FBC0238FD313" box="[191,385,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="subSpecies" species="lobidens" subSpecies="sensu">
|
||
<emphasis id="06A4D237FFFFFFB8FF07FBC0238FD313" box="[191,385,1051,1077]" italics="true" pageId="6" pageNumber="195">A. lobidens sensu</emphasis>
|
||
</taxonomicName>
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FE30FBC023DFD312" box="[392,465,1051,1077]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="195">Cunha</collectingCountry>
|
||
et al. (20202), any further comparison with other taxa is hindered, as it is obvious now that colour patterns, along with molecular analyses and morphology, will be crucial to resolve this challenging species complex. For example, in
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FE49FBB8206DD35A" box="[497,611,1123,1149]" name="Singapore" pageId="6" pageNumber="195">Singapore</collectingCountry>
|
||
(
|
||
<bibRefCitation id="504173D4FFFFFFB8FDC9FBB82182D359" author="Anker, A. & De Grave, S." box="[625,908,1123,1150]" pageId="6" pageNumber="195" pagination="343 - 454" refId="ref5031" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker & De Grave 2016</bibRefCitation>
|
||
),
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FC18FBB821FCD35A" box="[928,1010,1123,1149]" name="Kuwait" pageId="6" pageNumber="195">Kuwait</collectingCountry>
|
||
(present study) and some localities in eastern
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FF53FB5C235CD386" box="[235,338,1159,1185]" name="Australia" pageId="6" pageNumber="195">Australia</collectingCountry>
|
||
(A. Anker, pers. obs.), two or more species of the
|
||
<taxonomicName id="F3D075A6FFFFFFB8FCC7FB5C21F2D386" authorityName="sensu De Haan" authorityYear="1849" box="[895,1020,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FCC7FB5C21F2D386" box="[895,1020,1159,1185]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
complex occur syntopically, with the easiest separation between them being consistent differences in the colour patterns.
|
||
</paragraph>
|
||
<paragraph id="346F0E25FFFFFFB8FF7FFB142332D2E6" blockId="6.[151,1437,151,1473]" pageId="6" pageNumber="195">
|
||
<bibRefCitation id="504173D4FFFFFFB8FF7FFB1423DFD3CE" author="Al-Maliky, T. H. Y. & Al-Khafaji, K. K. & Khalaf, T. A." box="[199,465,1231,1257]" pageId="6" pageNumber="195" pagination="47 - 53" refId="ref4888" refString="Al-Maliky, T. H. Y., Al-Khafaji, K. K. & Khalaf, T. A. (2017) New records of the snapping shrimp Alpheus edwardsii (Audouin, 1826) (Crustacea: Alpheoidea) in Basrah, Iraq. Arthropods, 6, 47 - 53." type="journal article" year="2017">
|
||
Al-Maliky
|
||
<emphasis id="06A4D237FFFFFFB8FEFDFB142381D3CE" box="[325,399,1231,1257]" italics="true" pageId="6" pageNumber="195">et al. (</emphasis>
|
||
2017)
|
||
</bibRefCitation>
|
||
reported and illustrated
|
||
<taxonomicName id="F3D075A6FFFFFFB8FD5EFB142676D3CD" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[742,1144,1231,1258]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="edwardsii">
|
||
<emphasis id="06A4D237FFFFFFB8FD5EFB1421BBD3CE" box="[742,949,1231,1257]" italics="true" pageId="6" pageNumber="195">Alpheus edwardsii</emphasis>
|
||
(
|
||
<bibRefCitation id="504173D4FFFFFFB8FC7DFB14267ED3CD" author="Audouin, V." box="[965,1136,1231,1258]" pageId="6" pageNumber="195" pagination="77 - 98" refId="ref5123" refString="Audouin, V. (1826) Explication sommaire des planches de crustaces de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, Membre de l'Institut; offrant un expos des characteres naturels des genres avec la distinction des especes. Animaux invertebres. In: Description de l'Egypte ou receuil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise, publie par les ordres de sa Majeste l'Empereur Napoleon le Grand. Imprimerie Imperiale, Paris, pp. 77 - 98." type="book chapter" year="1826">Audouin, 1826</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Basrah area in
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FAD0FB142797D3CE" box="[1384,1433,1231,1257]" name="Iraq" pageId="6" pageNumber="195">Iraq</collectingCountry>
|
||
. Although their photographs of preserved specimens are in very low resolution, the dorsal shoulder of the major chela is not overhanging the adjacent transverse groove (
|
||
<emphasis id="06A4D237FFFFFFB8FCAFFACC2143D216" box="[791,845,1303,1329]" italics="true" pageId="6" pageNumber="195">idem</emphasis>
|
||
: figs. 2, 3A), whilst the merus of the major cheliped seems to be armed with a strong distomesial tooth (
|
||
<emphasis id="06A4D237FFFFFFB8FD77FAE0210BD272" box="[719,773,1339,1365]" italics="true" pageId="6" pageNumber="195">idem</emphasis>
|
||
: fig. 3B, D). Therefore, the Iraqi material can be tentatively referred to
|
||
<taxonomicName id="F3D075A6FFFFFFB8FEAEFA8423B4D25E" authorityName="sensu De Haan" authorityYear="1849" box="[278,442,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FEAEFA842320D25E" box="[278,302,1375,1401]" italics="true" pageId="6" pageNumber="195">A.</emphasis>
|
||
cf.
|
||
<emphasis id="06A4D237FFFFFFB8FEE3FA8423B4D25E" box="[347,442,1375,1401]" italics="true" pageId="6" pageNumber="195">lobidens</emphasis>
|
||
</taxonomicName>
|
||
, as suggested by Al-Khandari
|
||
<emphasis id="06A4D237FFFFFFB8FCAFFA84215FD25E" box="[791,849,1375,1401]" italics="true" pageId="6" pageNumber="195">et al.</emphasis>
|
||
(2020: p. 254; mind a small error therein, read correctly: “
|
||
<bibRefCitation id="504173D4FFFFFFB8FF49FA58200FD2BA" author="Al-Maliky, T. H. Y. & Al-Khafaji, K. K. & Khalaf, T. A." box="[241,513,1411,1437]" pageId="6" pageNumber="195" pagination="47 - 53" refId="ref4888" refString="Al-Maliky, T. H. Y., Al-Khafaji, K. K. & Khalaf, T. A. (2017) New records of the snapping shrimp Alpheus edwardsii (Audouin, 1826) (Crustacea: Alpheoidea) in Basrah, Iraq. Arthropods, 6, 47 - 53." type="journal article" year="2017">
|
||
Al-Maliky
|
||
<emphasis id="06A4D237FFFFFFB8FECAFA5823A2D2BA" box="[370,428,1411,1437]" italics="true" pageId="6" pageNumber="195">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
, who erroneously reported
|
||
<taxonomicName id="F3D075A6FFFFFFB8FC81FA5821B6D2BA" authorityName="sensu De Haan" authorityYear="1849" box="[825,952,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="lobidens">
|
||
<emphasis id="06A4D237FFFFFFB8FC81FA5821B6D2BA" box="[825,952,1411,1437]" italics="true" pageId="6" pageNumber="195">A. lobidens</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="F3D075A6FFFFFFB8FC58FA582660D2BA" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[992,1134,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="195" phylum="Arthropoda" rank="species" species="edwardsii">
|
||
<emphasis id="06A4D237FFFFFFB8FC58FA582660D2BA" box="[992,1134,1411,1437]" italics="true" pageId="6" pageNumber="195">A. edwardsii</emphasis>
|
||
</taxonomicName>
|
||
from Shatt-Al-Basrah and Faw in
|
||
<collectingCountry id="4CC74EB5FFFFFFB8FF53FA7C232FD2E6" box="[235,289,1447,1473]" name="Iraq" pageId="6" pageNumber="195">Iraq</collectingCountry>
|
||
.”).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |