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<document id="8D22D2EADF24258167572621AA2391B5" ID-CLB-Dataset="298695" ID-DOI="10.3897/evolsyst.8.124213" ID-GBIF-Dataset="07480afa-995b-4bfe-a1fe-799319db3f89" ID-ZooBank="8FFA1CD5-BAD6-455E-AF56-C13A25262059" ID-publisher-id="124213" URI-arpha="76887DB8-8714-522B-B010-D2066B4A73B6" URI-zoobank="http://zoobank.org/8FFA1CD5-BAD6-455E-AF56-C13A25262059" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1718879303986" checkinUser="pensoft" docAuthor="Jaume-Schinkel, Santiago" docDate="2024" docId="BBEA1DBADC8E549F805DC43A81F0CBA7" docLanguage="en" docName="Evolutionary Systematics 8(1): 145-154" docOrigin="Evolutionary Systematics 8 (1)" docSource="https://evolsyst.pensoft.net/article/124213/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Armillipora nawak Jaume-Schinkel 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="154" masterDocId="76887DB88714522BB010D2066B4A73B6" masterDocTitle="Further contributions to the taxonomy of Armillipora Quate (Diptera, Psychodidae) with the description of two additional new species" masterLastPageNumber="154" masterPageNumber="145" pageNumber="145" updateTime="1718903320353" updateUser="ExternalLinkService">
<mods:mods id="E4001B4FE6B715691F8D0D64FE256EAC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C518D018537EC9F581DEC9A241076EA5">
<mods:title id="AC2AAAF50FA10F6AB7937DC7392ED3B0">Further contributions to the taxonomy of Armillipora Quate (Diptera, Psychodidae) with the description of two additional new species</mods:title>
</mods:titleInfo>
<mods:name id="EA7B6483F2B22A4D2091B483E7A1FA97" type="personal">
<mods:role id="54E1A64061D1532490D71E79DDBC311A">
<mods:roleTerm id="F9969E2FBF31A8D951CB7908A1EFAED0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F03A4107A1922636F2F54F66178B30B9">Jaume-Schinkel, Santiago</mods:namePart>
<mods:nameIdentifier id="30825378427D0F03EBA76D5F42871BC4" type="ORCID">0000-0002-3502-9407</mods:nameIdentifier>
<mods:affiliation id="B8391A51E83C82A7EF78D6023A5897B6">Leibniz Institute for the Analysis of Biodiversity Change – Museum Koenig Bonn, Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:typeOfResource id="ED8D0089EECF036D7F8F33FDA14E0939">text</mods:typeOfResource>
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<mods:title id="0CB321AB844160121A38668AD0F4F070">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="E4628A35C242F8AE388E3F24A9AF1B0E">
<mods:date id="9EF2102C21F99B40C0073855F848452F">2024</mods:date>
<mods:detail id="9396076097B10FE0C2B68AB411E9BDF9" type="pubDate">
<mods:number id="3608BB60226EE08903DC2D49E3DA631B">2024-06-20</mods:number>
</mods:detail>
<mods:detail id="7B2D983540344AEE8893B503EE6E8C17" type="volume">
<mods:number id="15C4E6A077DFCF11BB424B02D2161571">8</mods:number>
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<treatment id="BBEA1DBADC8E549F805DC43A81F0CBA7" ID-DOI="http://doi.org/10.5281/zenodo.12186460" ID-GBIF-Taxon="233241200" ID-Zenodo-Dep="12186460" ID-arpha="BBEA1DBA-DC8E-549F-805D-C43A81F0CBA7" ID-zoobank="https://zoobank.org/33A151A9-81BF-41DF-A36C-949297C1DB78" LSID="urn:lsid:plazi:treatment:BBEA1DBADC8E549F805DC43A81F0CBA7" LSID-ZBK="urn:lsid:zoobank.org:act:33A151A9-81BF-41DF-A36C-949297C1DB78" httpUri="http://treatment.plazi.org/id/BBEA1DBADC8E549F805DC43A81F0CBA7">
<subSubSection id="A61222140C73CD798CA21D98B00FE8A0" type="nomenclature">
<paragraph id="408789CB7649D1BDE668991300E894A0">
<taxonomicName id="006E0AF45C28AAD361D5878C47D3287D" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nawak" status="sp. nov.">
<emphasis id="30697066648B5FC0F1EA2AF77FE24C96" italics="true">Armillipora nawak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69B2C70F6798E38E560E75293323C271" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="494D50F0C6C15851BF527B4F2DB28A20" type="description">
<paragraph id="538FF970F514547C35B35460A8007EE2">
<figureCitation id="E1E4BC86F194DFFBDD34C72C693B775E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the species of Armillipora Quate." httpUri="https://binary.pensoft.net/fig/1074830">Figs 1</figureCitation>
,
<figureCitation id="F86C6BFD8FBEF9BDB8A786D060E777F5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4</figureCitation>
,
<figureCitation id="7C4B643EF1563BE8DA3B11C79DE07AC0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVPAE" type="Differential diagnosis">
<paragraph id="8D3A4ECD839F704031E4B764B76A4EFB">
<heading id="859C5F35D6F46040E5BA94D5DA9AE13E" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="66FF6FFE3B0DC31F5E1754DA2D302B8E">
<taxonomicName id="A8C457A22C8CCC39C71C99580A4C9F29" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nawak" status="sp. nov.">
<emphasis id="914E1E8B7EC9CDDEEEA316D11C7AB42C" italics="true">Armillipora nawak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4B434C8D6F482681876821B0B13EF1D1" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="733EE24081F244920C791827B7651962" authorityName="Jezek, Obona &amp; Le Pont" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="suapiensis">
<emphasis id="66C78067D9F3AB9EE3E1900071427D91" italics="true">suapiensis</emphasis>
</taxonomicName>
group (gonostyli not fused). It can be easily differentiated from all the species in the group by the digitiform gonostyli with rounded apex (apex of gonostyli bifurcated in
<taxonomicName id="3E729E780714584F485A896AD730F074" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="53436D7ADF82B1F4F0682EA8BD6D6BC1" italics="true">A. bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="339A609E789300F0AE092658684BCDEA" rank="species">sp. nov.</taxonomicNameLabel>
, and apex of gonostyli pointed and sickle-shaped in
<taxonomicName id="3EC72868357633E304CF76810B356693" authorityName="Jezek, Obona &amp; Le Pont" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="suapiensis">
<emphasis id="AADC4BB9475651C208F73B2606523741" italics="true">A. suapiensis</emphasis>
</taxonomicName>
); the gonostyli are as long as the aedeagus (shorter than the aedeagus in
<taxonomicName id="9BC1375659FBA39B5B78D7C971AF7C03" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="F331EE49520E41438803E9009B61995E" italics="true">A. bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C97D6248DF7F7BDD0624E2BD5B166A" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="6874DEE9D9535150FDB86042BE420F7F" authorityName="Jezek, Obona &amp; Le Pont" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="suapiensis">
<emphasis id="2DFC9EDBF8DEA2235D2483758F7BCFA0" italics="true">A. suapiensis</emphasis>
</taxonomicName>
); the apex of the aedeagus with two digitiform sclerotized structures (apex of the aedeagus pointed without sclerotized structures in
<taxonomicName id="2F5AE2B4CAD73DAB0C05D86E2D8D90F7" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="2B65061D2CE01506F84356187AA99882" italics="true">A. bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39F84161D921947260D02226F9E7FAB1" rank="species">sp. nov.</taxonomicNameLabel>
, and in
<taxonomicName id="88774FDAA844306E70BDEE6E4B2F816D" authorityName="Jezek, Obona &amp; Le Pont" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="suapiensis">
<emphasis id="2BE48CA38530217F288075110D8D90C7" italics="true">A. suapiensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESSAE" type="etymology">
<paragraph id="EDB5C1FA31A087F3522941D46FE1ED52">
<heading id="7CDDF438B6F2F54568E6D8105B0EF6CB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1143F16C283BB8E6A0CFE739DD2FBE55">
The species epithet
<taxonomicName id="B420B9132B10BD436B67F63BF6AD7651" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nawak">
<emphasis id="BE223DBB9A0F563451091879FEED6781" italics="true">nawak</emphasis>
</taxonomicName>
derives from the Cabécar word: “
<emphasis id="141D95BA0B91EEBE72D2C34A5DA54BDB" italics="true">ña ̱ ́ wák</emphasis>
”, commonly used to refer to synanthropic psychodines. To be treated as a name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EETAE" type="material">
<paragraph id="B420143F16B33D0352E5235B76D80F4C">
<heading id="257734D2D26CCF1236332D46A0A52CF2" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="DA2A58399A48A80B9BE10F105A7F49EF">
<materialsCitation id="BE9A42BBB9071CB64A40698A718924E5" ID-GBIF-Occurrence="4895573304" collectingDate="2012-10-22" collectingDateMax="2012-11-01" collectingDateMin="2012-10-22" collectingMethod="Malaise Trap" collectionCode="LACM" collectorName="ZABDI project" country="Costa Rica" elevation="1600" latitude="10.0492" location="Zurqui de Moravia" longLatPrecision="7" longitude="-84.0037" municipality="Zurqui de Moravia" specimenCode="ZABDI- 186, INB 0004346462" specimenCount="1" specimenCount-male="1" stateProvince="San Jose Moravia" typeStatus="holotype">
<collectingCountry id="AD28783B47B93169FAC6D7D86E868276" name="Costa Rica">Costa Rica</collectingCountry>
–
<emphasis id="C16C8A8C3C33B72532DD856A3A43C435" bold="true">
<collectingRegion id="7ACCBA128778F467A3AA60A32A59A99A">San Jose Moravia</collectingRegion>
</emphasis>
•
<specimenCount id="AF07864C170AEDCD3544DE2A318C78F4" count="1" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="13FF2A3E3B262E4B94BE674BEDC2C914">Zurqui de Moravia</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-84.003700,10.049200]}">
-
<geoCoordinate id="77885FBD2B5B02BE60F13C7D769E6D47" degrees="10.0492" direction="north" orientation="latitude" precision="5" value="10.0492">10.0492 ° N</geoCoordinate>
, -
<geoCoordinate id="4385562754761BF81220DE5A0AE80A01" degrees="84.0037" direction="west" orientation="longitude" precision="5" value="-84.0037">84.0037 ° W</geoCoordinate>
</named-content>
; alt.
<quantity id="08872F4E1DB5B614A6468BFC002AA5CD" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="BA83E0A3BBCAFF302D9BC699960D7CFC" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 masl</elevation>
</quantity>
;
<date id="156FA3E3877C3188ED81D02C1440CB59" value="2012-10-22" valueMax="2012-11-01" valueMin="2012-10-22">
<collectingDate id="DABF8EAE1AD5BEB0D9A8C2A8E7249403" value="2012-10-22" valueMax="2012-11-01" valueMin="2012-10-22">22 Oct. – 01 Nov. 2012</collectingDate>
</date>
;
<collectorName id="A11B46A3238B08102D14806768E5F66F">ZABDI project</collectorName>
leg.;
<collectingMethod id="632FD21DF166D86AF2EC2274B0D1A296">Malaise Trap</collectingMethod>
<specimenCode id="B9D3DE10265490752C5A792E8994CD49">ZABDI- 186</specimenCode>
;
<abbrev id="ABBRID0ETTAE" xlink_title="Natural History Museum of Los Angeles County">
<collectionCode id="0EECBA5F4722BB85E8221B7EAE6CA52B" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</abbrev>
[
<specimenCode id="FF5213EB06F39EDBB35C84AAC19D039F" collectionCode="INB" country="Costa Rica" lsid="urn:lsid:biocol.org:col:14937" name="Instituto Nacional de Biodiversidad" type="Herbarium">INB 0004346462</specimenCode>
- INBIOCRI]
</materialsCitation>
.
<materialsCitation id="C9BD2B5F9BB05FF780E1A7D9603C490F" ID-GBIF-Occurrence="4895573305" collectingDate="2013-01-01" collectingDateMax="2013-01-07" collectingDateMin="2013-01-01" collectingMethod="Malaise Trap" collectionCode="LACM" collectorName="ZABDI project" country="Costa Rica" county="Coto Brus" elevation="1500" latitude="10.0492" location="Estacion Biologica Las Alturas" longLatPrecision="7" longitude="-84.0037" municipality="Las Tablas" specimenCode="ZABDI- 544, INB 0004415321" specimenCount="1" specimenCount-male="1" stateProvince="Punteras" typeStatus="paratype">
<emphasis id="AE17F310286AB3556DB69D29FAE66D02" bold="true">
<emphasis id="E484FB99DAFCB2D06D8DEE8693A2A410" bold="true" italics="true">
<typeStatus id="E6BE53B9E36DD8F2E5EB8881D8E4AA60" type="paratype">Paratype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="C6864D9201F3C196014A34CF955BDB66" name="Costa Rica">Costa Rica</collectingCountry>
–
<emphasis id="1FFDA72F0CBFAB873648CD3F604FCE9C" bold="true">
<collectingRegion id="B264C4DCBA292CF8431E8F2AF1E90DD5">Punteras</collectingRegion>
</emphasis>
•
<specimenCount id="74FFBDD5D950C7CD2D7040670D1B69DA" count="1" type="male">1 ♂</specimenCount>
;
<collectingCounty id="369A371C1C76E1C37429B3BE506E83C1">Coto Brus</collectingCounty>
,
<collectingMunicipality id="0C1F63BFC7ABB12F42550194671D3B6B">Las Tablas</collectingMunicipality>
,
<location id="A132941DBBCE612533EE06BAC177C16F" LSID="urn:lsid:plazi:treatment:BBEA1DBADC8E549F805DC43A81F0CBA7:A132941DBBCE612533EE06BAC177C16F" country="Costa Rica" county="Coto Brus" latitude="10.0492" longLatPrecision="7" longitude="-84.0037" municipality="Las Tablas" name="Estacion Biologica Las Alturas" stateProvince="Punteras">Estacion Biologica Las Alturas</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAUAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.832200,8.951900]}">
-
<geoCoordinate id="39D1737B9B5DE97F16CE9481FA10A1E8" degrees="8.9519" direction="north" orientation="latitude" precision="5" value="8.9519">8.9519 ° N</geoCoordinate>
, -
<geoCoordinate id="F8B06DBA38AC4587CF79F3CAB1734D0F" degrees="82.8322" direction="west" orientation="longitude" precision="5" value="-82.8322">82.8322 ° W</geoCoordinate>
</named-content>
;
<quantity id="5582244EDA461FBA6C0200B1168BA149" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<elevation id="C74B6EAE1D48C563752EAF778229D58D" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 masl</elevation>
</quantity>
;
<date id="52CB188F692E05FA806DB255020499AD" value="2013-01-01" valueMax="2013-01-07" valueMin="2013-01-01">
<collectingDate id="A6F800B7EB609E9225F70442F8C9F93A" value="2013-01-01" valueMax="2013-01-07" valueMin="2013-01-01">01 Jan. – 07 Jan. 2013</collectingDate>
</date>
;
<collectorName id="3DB2FD8C381D1913FD23F2828920EE4A">ZABDI project</collectorName>
leg.;
<collectingMethod id="5F2276065F7A31B7C2500871C6E2AB30">Malaise Trap</collectingMethod>
<specimenCode id="80EBDB17947A4EAE9C4B00475CC782AA">ZABDI- 544</specimenCode>
;
<abbrev id="ABBRID0EEUAE" xlink_title="Natural History Museum of Los Angeles County">
<collectionCode id="ECF6E8F0B55AF855229E9B59DEB8E46E" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</abbrev>
[
<specimenCode id="580691C699975CEC603EEF05D9EE225C" collectionCode="INB" country="Costa Rica" lsid="urn:lsid:biocol.org:col:14937" name="Instituto Nacional de Biodiversidad" type="Herbarium">INB 0004415321</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIUAE" type="description">
<paragraph id="7622E722BDD548AC7738E87E17E28F32">
<heading id="078B531D2030D430CDDCD6662E051EE3" reason="title">Description.</heading>
</paragraph>
<paragraph id="5B1241DC109718A638DE6646DEAA1C81">(N = 2). Wing length: 3.02, width: 1.09. Head length: 0.50, width: 0.60. Antennal segments: scape: 0.12); pedicel: 0.07; flagellomeres 1–4 0.17, flagellomeres 5–14 missing in examined material. Palpal segment 1: 0.10, palpal segments 2–4 missing in examined material.</paragraph>
<paragraph id="07BC1A6D2C79091B5680086555FB1F1D">
<emphasis id="0203FA6078DF6E073231F989B640BEC1" bold="true">Male.</emphasis>
Head (Fig.
<figureCitation id="EB677CCC88379F63BD89CF8C0FC716B4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 A</figureCitation>
). About 1.20 times wider than its length; eye bridge separated by about half facet’s diameter, with four rows of facets; interocular suture absent; the frontal patch of alveoli is divided. Antennal scape almost cylindrical, about 1.66 times the length of the spherical pedicel; flagellomeres fusiform and longer than the scape, with scattered alveoli in the surface; each flagellomere with two rows of alveoli; flagellomeres after the fourth absent in examined material; ascoids undistinguishable in examined material. First palpal segment cylindrical; labium without any strong sclerites; labella elongated and teardrop-shaped, with 5–6 scattered setae on the surface and with one apical spiniform tooth on each.
</paragraph>
<caption id="06BD0B9AA40A3F8DBDDB3AF13BA33424" ID-arpha="7AA76D65-D2AA-5C56-9A4D-D34CEA73AF90" httpUri="https://binary.pensoft.net/fig/1074833" startId="F4">
<paragraph id="3B952A2C2D1EEE3E5DE67013ED9EF330">
<label id="C09DEB8B2F21F0D84A4E5262D96BFC39">Figure 4.</label>
</paragraph>
<paragraph id="22B08130AD6A7B81101710BD4194E2EA">
<taxonomicName id="9EE8B601568CEC580C7D9474EB6628B0" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nawak" status="sp. nov.">
<emphasis id="2375A608114430D857482C145DF30185" italics="true">Armillipora nawak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AFA0D3B90A97D9CFC53588BBA4EE46E" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0D6D97DB712EE31ACBB586E3953EE79A" bold="true">A – E</emphasis>
♂, holotype (INB 0004346462).
<emphasis id="4316D7B46B03366986340E201844B579" bold="true">A.</emphasis>
Head;
<emphasis id="206CF6D57D0F43AACF47E0A9E7831731" bold="true">B.</emphasis>
Thorax;
<emphasis id="3FAB00A0953D9E902BAEFBD4356D327F" bold="true">C.</emphasis>
Wing;
<emphasis id="508A61179E6D548645AF8B004A71DEC5" bold="true">D.</emphasis>
Genitalia, ventral view;
<emphasis id="F6C4A86D94CA651A2B032B87C18CE0B4" bold="true">E.</emphasis>
Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (
<emphasis id="41E8427592D180060AB61FAB73EC84BF" bold="true">A</emphasis>
); 0.10 mm (
<emphasis id="3A90D55F9F32589A42E4CFA7B8427281" bold="true">B</emphasis>
); 0.50 mm (
<emphasis id="E913163826EC79F3F28C26E81A5BC81B" bold="true">C</emphasis>
); 0.10 mm (
<emphasis id="1E60E84ADEC8BB5313E0726E6A49A85C" bold="true">D – E</emphasis>
).
</paragraph>
</caption>
<caption id="FFB52D9927C67E243B7F4E13CEB843B1" ID-arpha="1BCD6E16-72E0-5432-B97F-4498BC45604B" httpUri="https://binary.pensoft.net/fig/1074834" startId="F5">
<paragraph id="2CDD6A828FF783F5589BE843E13C301B">
<label id="0C94E7F7180850EFF07BB553F9EB64D2">Figure 5.</label>
</paragraph>
<paragraph id="2B9E0781B04CD1A96050FAC85786968D">
<taxonomicName id="C5BCC29A0B1D568AE7B51D0B51330D78" authority="Jaume-Schinkel, 2024" authorityName="Jaume-Schinkel" authorityYear="2024" class="Insecta" family="Psychodidae" genus="Armillipora" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nawak" status="sp. nov.">
<emphasis id="DD132852A6F9D024060EC53D1C3D42AA" italics="true">Armillipora nawak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AF8C708FB9055AAA97BB4644936E40F" rank="species">sp. nov.</taxonomicNameLabel>
♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm.
</paragraph>
</caption>
<paragraph id="2825650E95BA762F204C3E8908DA95BE">
Thorax (Fig.
<figureCitation id="F78EAC91EEAB6CD8396ED278D1F3DEBF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 B</figureCitation>
). It does not present allurement organs, with a single patch of alveoli in the paraterguite and antepronotum; all coxae with a stripe of one to two rows of alveoli. Wing (Fig.
<figureCitation id="2B76842AAC298AB6F965C531BFB10EFF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 C</figureCitation>
) length about three times its width; wing membrane hyaline, with darkened spots in the apex of longitudinal veins (as seen in Fig.
<figureCitation id="A0E36085999CDA9EB96028F291ED2F92" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 C</figureCitation>
); Sc vein short ending at the origin of R
<subScript id="4926BE7835CD3C2B6821B61F3439FA41">1</subScript>
; radial fork basal to medial fork; R
<subScript id="871FC05D4B95FA748A0B96275D51EB68">2 + 3</subScript>
not joining R
<subScript id="406F9E3949537CFA64E9101C993697BF">4</subScript>
; R
<subScript id="BC4D2A5CDD72E041F05614C15E695297">5</subScript>
ending at the wing apex; CuA ending at wing margin.
</paragraph>
<paragraph id="A63873B867AA2836B27B2098C0273D9D">
Terminalia (Figs
<figureCitation id="7C6B280B49D26CC7B01BE338B34669D6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 D – E</figureCitation>
,
<figureCitation id="2BF588D4DC4F746B4C4C3220FDD0DBDA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
). Hypandrium on ventral view band-like, sclerotized and joining the base of the reduced gonocoxites; gonostyli not fused with gonocoxites and clearly divergent, digitiform with a rounded apex (as in Fig.
<figureCitation id="34212B14DE02C34EC9994FAA0B73D6AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 D – E</figureCitation>
,
<figureCitation id="3E5C05AA50F82FCCF3D62B532BBCCE7F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
), each gonostylus with two apical and one subapical alveoli; aedeagus on ventral view straight, basal 2 / 3 rds looking like a single shaft, while apical third as two distinct heavily sclerotized digitiform structures, the aedeagus ends beyond the apex of the gonostyli; ejaculatory apodeme slightly shorter than the aedeagus, broader on the base, and tapering towards its apex, in ventral view the basal margin is rounded, and the width of the basal margin is the same as the length of the ejaculatory apodeme; parameral sclerites seem absent; gonocoxal apodemes (Figs
<figureCitation id="1B1B523CC0DD4ACB07170CD4590E97D2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 D</figureCitation>
,
<figureCitation id="01DDBB6055D33958E9B5867E41AB7D64" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
) fitting in a concavity on the underside of the ejaculatory apodeme, and joining the reduced gonocoxites; epandrium rectangular (Fig.
<figureCitation id="F5D0CBFF4ABD2F3BABB1E8D7EE0BC4FA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
), about 3 times wider than its length; hypoproct tongue-shaped and covered in setulae, epiproct not observed; epandrial appendages (Figs
<figureCitation id="218BB025619448DF9EAE954E06D2FFA5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 D – E</figureCitation>
,
<figureCitation id="174034ECEB41CF04FACDDBC45986E74B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
) barely hemispherical, prolonged and tapering distally, covered with setae; epandrial appendages with one preapical, tip-folded and short tenacula (Fig.
<figureCitation id="C6F6DDF3A434FF097206E3A37EE10A72" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
) with a basal darkened patch of accessory tenacula, the accessory tenacula varies in length, from shorter to longer than the epandrial appendages (Figs
<figureCitation id="E777D6055557C331E58B941BC6D1A40E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Armillipora nawak sp. nov. A – E ♂, holotype (INB 0004346462). A. Head; B. Thorax; C. Wing; D. Genitalia, ventral view; E. Aedeagus, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn apo = gonocoxal apodeme; gns = gonostylus; ten = tenacula with folded tip. Scale bars: 0.25 mm (A); 0.10 mm (B); 0.50 mm (C); 0.10 mm (D – E)." httpUri="https://binary.pensoft.net/fig/1074833">4 D</figureCitation>
,
<figureCitation id="2CE0849EAF21C8EE46B19AC3B0368161" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Armillipora nawak sp. nov. ♂, holotype (INB 0004346462). Genitalia, ventral view. Abbreviations: acc ten = accessory tenacula; aed = aedeagus; eja = ejaculatory apodeme; ep ap = epandrial appendage; gn con = gonocoxal condyle; gns = gonostyli; hyp = hypandrium; ten = tenacula with folded tip. Scale bar: 0.10 mm." httpUri="https://binary.pensoft.net/fig/1074834">5</figureCitation>
).
</paragraph>
<paragraph id="9EF2056367F5FF73A2C225A0A3992D70">
<emphasis id="AAA2DCD0D52E5D7A2C5CC7C17FF5973A" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFZAE" type="distribution">
<paragraph id="584A886AFA1C65FC89566E86D5219BCE">
<heading id="BF0993BDA73C52C1829D40DCC08773C0" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="75CEFB600BA74B977D4E8A07C04F7C60">
The species is only known from the type locality in
<collectingCountry id="8D10C84E5050E06F07055EC4E828697A" name="Costa Rica">Costa Rica</collectingCountry>
(Fig.
<figureCitation id="BB2F645943247912D98257B1FF957B17" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the species of Armillipora Quate." httpUri="https://binary.pensoft.net/fig/1074830">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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