307 lines
42 KiB
XML
307 lines
42 KiB
XML
<document id="0819E0C9961D0D919C1B6A618E959496" ID-CLB-Dataset="25169" ID-DOI="10.11646/zootaxa.4688.2.8" ID-GBIF-Dataset="22249957-a026-4e29-8a3d-cb31812ed2a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3516228" ID-ZooBank="DF78B9EF-FEEA-4D42-9D5B-FCE58039C277" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571755596709" checkinUser="plazi" docAuthor="Polizei, Thiago T. S. & Barclay, Maxwell V. L." docDate="2019" docId="BB7C7021FFC8FF99FF56FBF5C9225DAF" docLanguage="en" docName="zootaxa.4688.2.8.pdf.imf" docOrigin="Zootaxa 4688 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pilielmis shepardi Polizei & Barclay 2019, sp. nov." docType="treatment" docUuid="90DCFE7C-4089-4A52-A0B6-67BB24E334E2" docUuidSource="ZooBank" docVersion="7" lastPageNumber="287" masterDocId="47450859FFCAFF9CFFC1FFD3CD1C5E08" masterDocTitle="First records of the genera Neocylloepus and Pilielmis (Coleoptera: Elmidae Elminae) from Venezuela, with the description of Pilielmis shepardi sp. nov." masterLastPageNumber="288" masterPageNumber="282" pageNumber="284" updateTime="1698760549112" updateUser="ExternalLinkService">
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<mods:mods id="1441AF697085C6A563060481150CDA9C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B36CDE6419AD40ADC34373660C0B226B">
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<mods:title id="E2A628B5B06095E4470DB82CAA4E6805">First records of the genera Neocylloepus and Pilielmis (Coleoptera: Elmidae Elminae) from Venezuela, with the description of Pilielmis shepardi sp. nov.</mods:title>
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</mods:titleInfo>
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<mods:name id="B7F3C5CBA77879D107F7BCA6D658CD1B" type="personal">
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<mods:role id="4807E25AAC98C2AB785001E3F6E5537C">
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<mods:roleTerm id="8D5F43D1C34B7B03A4636DFDC64EBFB3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="010E08AC6D70133B079BBE87666BD3C0">Polizei, Thiago T. S.</mods:namePart>
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</mods:name>
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<mods:name id="86DCB05A2D27DCCAD6B3A16A59BD87C1" type="personal">
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<mods:role id="2790956247999355CD823A0815E7E4F8">
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<mods:roleTerm id="D583CCF573515A0A2406E4E99AC3AD8C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="20CBEF3608113DE64BBD605FD89A09E0">Barclay, Maxwell V. L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="51A5A3BE5946FFB97B741CCCB74E0A72">text</mods:typeOfResource>
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<mods:relatedItem id="D8F5EE491F80A314DD165B883F78EB92" type="host">
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<mods:titleInfo id="67E71F7F38B201836627EF94BDD1AE64">
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<mods:title id="4928FE4791FE6BAD00EFE782B85CC8FD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C732361EC156344BB189245934316590">
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<mods:date id="849C5756B052DB00CF56E55119E7E6FA">2019</mods:date>
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<mods:detail id="AC7ABE46BCDC6F9240869CAA7DE22F23" type="pubDate">
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<mods:number id="CA3286BEDFABB00FE2B6F607311F7203">2019-10-22</mods:number>
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</mods:detail>
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<mods:detail id="E9544957BBAB56C4F0E1D2FC4638363A" type="volume">
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<mods:number id="575AC061F4C30734441226D125B58295">4688</mods:number>
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</mods:detail>
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<mods:detail id="02649A9DCFA34B63C1A0069363EE1DED" type="issue">
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<mods:number id="F949B1F0E77B1D1D26DEF702CA0A3530">2</mods:number>
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</mods:detail>
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<mods:start id="9986D0E2E3E81433815F412D2BDF06FE">282</mods:start>
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<mods:end id="E060B6A53028ACC6BCAE88DE610349E5">288</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="42C45C3AFDDC1D63CEB0D037FAC3E81B">journal article</mods:classification>
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<mods:identifier id="914D80262002272BB81F2AC14B6F78DB" type="CLB-Dataset">25169</mods:identifier>
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<mods:identifier id="5B198B040BDAA54A874CA974992AB487" type="DOI">10.11646/zootaxa.4688.2.8</mods:identifier>
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<mods:identifier id="2A18589F12CA5CD8DA9996B988D0CBEC" type="GBIF-Dataset">22249957-a026-4e29-8a3d-cb31812ed2a8</mods:identifier>
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<mods:identifier id="1082A09B6E7F66C61D3254C8ED3F04B2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A70ECF54197773B9BD57E6172DE1B706" type="Zenodo-Dep">3516228</mods:identifier>
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<mods:identifier id="34C2DFB8C96583EF889FABCB466A86BA" type="ZooBank">DF78B9EF-FEEA-4D42-9D5B-FCE58039C277</mods:identifier>
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</mods:mods>
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<treatment id="BB7C7021FFC8FF99FF56FBF5C9225DAF" ID-GBIF-Taxon="159811125" LSID="urn:lsid:zoobank.org:act:90DCFE7C-4089-4A52-A0B6-67BB24E334E2" httpUri="http://treatment.plazi.org/id/BB7C7021FFC8FF99FF56FBF5C9225DAF" lastPageId="5" lastPageNumber="287" pageId="2" pageNumber="284">
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<subSubSection id="7BCF92BCFFC8FF9EFF56FBF5CCCD5A48" box="[151,465,1062,1088]" pageId="2" pageNumber="284" type="nomenclature">
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<paragraph id="336AC137FFC8FF9EFF56FBF5CCCD5A48" blockId="2.[151,970,1062,1160]" box="[151,465,1062,1088]" pageId="2" pageNumber="284">
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<taxonomicName id="F4D5BAB4FFC8FF9EFF56FBF5CC715A48" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[151,365,1062,1088]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="284" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
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<emphasis id="01A11D25FFC8FF9EFF56FBF5CC715A48" box="[151,365,1062,1088]" italics="true" pageId="2" pageNumber="284">Pilielmis shepardi</emphasis>
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</taxonomicName>
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<emphasis id="01A11D25FFC8FF9EFEB5FBF5CCCD5A48" bold="true" box="[372,465,1062,1088]" pageId="2" pageNumber="284">
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<taxonomicNameLabel id="1A92A05EFFC8FF9EFEB5FBF5CCCD5A48" box="[372,465,1062,1088]" pageId="2" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="7BCF92BCFFC8FF9EFF56FB9ACC095A6C" box="[151,277,1097,1124]" pageId="2" pageNumber="284" type="description">
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<paragraph id="336AC137FFC8FF9EFF56FB9ACC095A6C" blockId="2.[151,970,1062,1160]" box="[151,277,1097,1124]" pageId="2" pageNumber="284">
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(
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<figureCitation id="ABEEDDB2FFC8FF9EFF5EFB9ACC105A6C" box="[159,268,1097,1124]" captionStart-0="FIGURES 3–5" captionStart-1="FIGURES 6–10" captionStartId-0="3.[151,264,1500,1525]" captionStartId-1="4.[151,264,1956,1981]" captionTargetBox-0="[216,1371,345,1475]" captionTargetBox-1="[260,1320,357,1922]" captionTargetId-0="figure@3.[216,1372,345,1475]" captionTargetId-1="figure@4.[254,1333,345,1931]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." captionText-1="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.3516232" figureDoi-1="http://doi.org/10.5281/zenodo.3516234" httpUri-0="https://zenodo.org/record/3516232/files/figure.png" httpUri-1="https://zenodo.org/record/3516234/files/figure.png" pageId="2" pageNumber="284">Figs 3-10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="7BCF92BCFFC8FF9EFF56FBBECED65A80" box="[151,970,1133,1160]" pageId="2" pageNumber="284" type="nomenclature">
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<paragraph id="336AC137FFC8FF9EFF56FBBECED65A80" blockId="2.[151,970,1062,1160]" box="[151,970,1133,1160]" pageId="2" pageNumber="284">
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<uri id="4744CD35FFC8FF9EFF56FBBECED65A80" box="[151,970,1133,1160]" pageId="2" pageNumber="284">
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urn:lsid:zoobank.org:act:
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<uuid id="4773FBE2FFC8FF9EFE68FBBECED65A80" box="[425,970,1133,1160]" pageId="2" pageNumber="284">90DCFE7C-4089-4A52-A0B6-67BB24E334E2</uuid>
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</uri>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="7BCF92BCFFC8FF9EFF56FB65CE245B68" pageId="2" pageNumber="284" type="materials_examined">
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<paragraph id="336AC137FFC8FF9EFF56FB65CFA65AC7" blockId="2.[151,1437,1205,2024]" box="[151,698,1205,1232]" pageId="2" pageNumber="284">
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<emphasis id="01A11D25FFC8FF9EFF56FB65CC2A5AD8" bold="true" box="[151,310,1205,1232]" pageId="2" pageNumber="284">
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<typeStatus id="EC6E7F95FFC8FF9EFF56FB65CDCE5AD8" box="[151,210,1206,1232]" pageId="2" pageNumber="284">Type</typeStatus>
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locality.
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</emphasis>
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<location id="360A97ECFFC8FF9EFEFDFB66CC8F5AC7" LSID="urn:lsid:plazi:treatment:BB7C7021FFC8FF99FF56FBF5C9225DAF:360A97ECFFC8FF9EFEFDFB66CC8F5AC7" box="[316,403,1205,1231]" country="Venezuela" name="Icabaru" pageId="2" pageNumber="284" stateProvince="Bolivar">Icabarú</location>
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,
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<collectingRegion id="F1110FD5FFC8FF9EFE5FFB66CF285AD8" box="[414,564,1205,1232]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="284">Bolivar State</collectingRegion>
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,
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<collectingCountry id="4BC281A7FFC8FF9EFDFEFB66CFAA5AC7" box="[575,694,1205,1231]" name="Venezuela" pageId="2" pageNumber="284">Venezuela</collectingCountry>
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.
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</paragraph>
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<paragraph id="336AC137FFC8FF9EFF06FB09CE515B10" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="284">
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<emphasis id="01A11D25FFC8FF9EFF06FB09CC695AFC" bold="true" box="[199,373,1241,1268]" pageId="2" pageNumber="284">Type material.</emphasis>
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<materialsCitation id="83BDCB6AFFC8FF9EFEBDFB0ACE515B10" ID-GBIF-Occurrence="2428542130" collectingDate="1987-07-04" collectorName="M. A. Ivie" country="Venezuela" location="Sta. Elena de Uairen" municipality="Icabaru" pageId="2" pageNumber="284" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
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<typeStatus id="EC6E7F95FFC8FF9EFEBDFB0ACCFB5AFB" box="[380,487,1241,1267]" pageId="2" pageNumber="284" type="holotype">Holotype</typeStatus>
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, male, with label data:
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<collectingCountry id="4BC281A7FFC8FF9EFD2AFB09CE8A5AFC" box="[747,918,1242,1268]" name="Venezuela" pageId="2" pageNumber="284">VENEZUELA</collectingCountry>
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:
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<collectingRegion id="F1110FD5FFC8FF9EFC61FB0ACEE85AFB" box="[928,1012,1241,1267]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="284">Bolivar</collectingRegion>
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/ nr.
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<collectingMunicipality id="D30E5B4DFFC8FF9EFBEFFB0AC9995AFB" box="[1070,1157,1241,1267]" pageId="2" pageNumber="284">Icabarú</collectingMunicipality>
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, ahia lm SW /
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<location id="360A97ECFFC8FF9EFAEAFB09CC195B1F" LSID="urn:lsid:plazi:treatment:BB7C7021FFC8FF99FF56FBF5C9225DAF:360A97ECFFC8FF9EFAEAFB09CC195B1F" country="Venezuela" municipality="Icabaru" name="Sta. Elena de Uairen" pageId="2" pageNumber="284" stateProvince="Bolivar">Sta. Elena de Uairen</location>
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/
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<date id="476BE7F7FFC8FF9EFEDAFB2DCCDC5B10" box="[283,448,1278,1304]" pageId="2" pageNumber="284" value="1987-07-04">
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||
<collectingDate id="572F1E1FFFC8FF9EFEDAFB2DCCDC5B10" box="[283,448,1278,1304]" pageId="2" pageNumber="284" value="1987-07-04">
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||
04 JULY
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<quantity id="F42D6CD2FFC8FF9EFE44FB2DCCDC5B10" box="[389,448,1278,1304]" metricMagnitude="6" metricUnit="m" metricValue="1.98749" pageId="2" pageNumber="284" unit="m" value="1987490.0">1987</quantity>
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</collectingDate>
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</date>
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, 490m / at light,
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<collectorName id="9E20A4E1FFC8FF9EFDBCFB2ECFF85B1F" box="[637,740,1277,1303]" pageId="2" pageNumber="284">M.A.Ivie</collectorName>
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(MIZA).
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</materialsCitation>
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||
</paragraph>
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<paragraph id="336AC137FFC8FF9EFF06FAF1CE245B68" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="284">
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<materialsCitation id="83BDCB6AFFC8FF9EFF06FAF1CE285B68" ID-GBIF-Occurrence="2428542128" collectingDate="1987-07-04" collectorName="M. A. Ivie" country="Venezuela" location="Sta. Elena de Uairen" municipality="Icabaru" pageId="2" pageNumber="284" specimenCount="34" stateProvince="Bolivar" typeStatus="paratype">
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<typeStatus id="EC6E7F95FFC8FF9EFF06FAF1CC245B34" box="[199,312,1314,1340]" pageId="2" pageNumber="284" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="25D30ABEFFC8FF9EFE84FAF1CCFF5B33" box="[325,483,1313,1340]" pageId="2" pageNumber="284" type="generic">34 specimens</specimenCount>
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, same data as holotype (6 to be deposited in MIZA,
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<quantity id="F42D6CD2FFC8FF9EFB86FAF2C9685B33" box="[1095,1140,1313,1339]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="284" unit="in" value="6.0">6 in</quantity>
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MZSP,
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<quantity id="F42D6CD2FFC8FF9EFB15FAF2C81D5B33" box="[1236,1281,1313,1339]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="284" unit="in" value="6.0">6 in</quantity>
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BMNH,
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<quantity id="F42D6CD2FFC8FF9EFAAEFAF2C8805B33" box="[1391,1436,1313,1339]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="284" unit="in" value="6.0">6 in</quantity>
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MAIC,
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<quantity id="F42D6CD2FFC8FF9EFF2FFA95CC3B5B57" box="[238,295,1349,1376]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="284" unit="in" value="10.0">10 in</quantity>
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USNM); (MZSP 31140-31142; 42273-42275)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="7BCF92BCFFC8FF9EFF06FABAC96D5853" pageId="2" pageNumber="284" type="diagnosis">
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<paragraph id="336AC137FFC8FF9EFF06FABAC96D5853" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="284">
|
||
<emphasis id="01A11D25FFC8FF9EFF06FABACC5E5B8C" bold="true" box="[199,322,1385,1412]" pageId="2" pageNumber="284">Diagnosis.</emphasis>
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Body subparallel. Head surface covered with micropunctures; antennomeres uniformly coloured. Pronotum moderately wider than long, with a deep and broad longitudinal impression on disc. Elytra 1.5 times longer than wide, one sublateral carina on interval VI, extending 3/4 of elytral length; apex and humeral angle yellowish in colour. Epipleura covered by tomentum. Hypomerum with a belt of tomentum reaching anterior and lateral margins. Disc of the first abdominal ventrite without tomentum. Aedeagus with the phallobase approximately as long as wide; parameres reaching ¾ of penis length; penis 1.5x the length of phallobase, ca 3.6x as long as wide, lateral margin subpararell, lateral margin near the apex with sensorial pores, apex sharp.
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</paragraph>
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</subSubSection>
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<subSubSection id="7BCF92BCFFC8FF99FF06F9B6CFD25F83" lastPageId="5" lastPageNumber="287" pageId="2" pageNumber="284" type="description">
|
||
<paragraph id="336AC137FFC8FF9EFF06F9B6CCA258AC" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="284">
|
||
<emphasis id="01A11D25FFC8FF9EFF06F9B6CC455888" bold="true" box="[199,345,1637,1664]" pageId="2" pageNumber="284">Description.</emphasis>
|
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Male: General colour light brownish, elytral apex and humeral angles yellowish. Total length
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<quantity id="F42D6CD2FFC8FF9EFAB8F9B5CDD958AC" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="284" unit="mm" value="2.1">2.1 mm</quantity>
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, greatest width
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<quantity id="F42D6CD2FFC8FF9EFEB7F959CCA658AC" box="[374,442,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="284" unit="mm" value="1.0">1 mm</quantity>
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.
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</paragraph>
|
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<paragraph id="336AC137FFC8FF9FFF06F97EC8025F4B" blockId="2.[151,1437,1205,2024]" lastBlockId="3.[151,1436,153,323]" lastPageId="3" lastPageNumber="285" pageId="2" pageNumber="284">
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Head: Head capsule partially retractable; without impressions; surface with micropunctures separated from each other by their diameter, each with one seta. Eyes black; laterally protruding (
|
||
<figureCitation id="ABEEDDB2FFC8FF9EFBE3F902C99458E4" box="[1058,1160,1745,1772]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="2" pageNumber="284">Figs. 3-5</figureCitation>
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). Antennae filiform with 11 antennomeres; uniformly coloured. Clypeus broad and concave;
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<date id="476BE7F7FFC8FF9EFC53F925CEB15918" box="[914,941,1782,1808]" pageId="2" pageNumber="284">4x</date>
|
||
as wide as long; surface punctured. Frontoclypeal suture straight. Labrum (
|
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<figureCitation id="ABEEDDB2FFC8FF9EFDC4F8CACF54593C" box="[517,584,1817,1844]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1956,1981]" captionTargetBox="[260,1320,357,1922]" captionTargetId="figure@4.[254,1333,345,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="2" pageNumber="284">Fig. 6</figureCitation>
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) with lateral margins rounded; surface without setae, excepted by short row of spines on disc near the anterior margin, and antero-lateral margin with long setae. Mandible (
|
||
<figureCitation id="ABEEDDB2FFC8FF9EFB0BF8EEC8125950" box="[1226,1294,1853,1880]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1956,1981]" captionTargetBox="[260,1320,357,1922]" captionTargetId="figure@4.[254,1333,345,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="2" pageNumber="284">Fig. 7</figureCitation>
|
||
) with lateral lobe strongly pronounced; three apical teeth moderately sharp; prostheca feebly sclerotized with four bilobed lobes; lateral margin with short setae. Maxilla (
|
||
<figureCitation id="ABEEDDB2FFC8FF9EFD9EF856CFBD59A8" box="[607,673,1925,1952]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1956,1981]" captionTargetBox="[260,1320,357,1922]" captionTargetId="figure@4.[254,1333,345,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="2" pageNumber="284">Fig. 8</figureCitation>
|
||
) with cardo subtriangular; stipes large, palpifer elongate; palpi with four palpomeres, without setae; galea palpiform and 2-articulated, with long thin setae at apex and five long setae laterointernally; lacinia subrectangular,
|
||
<date id="476BE7F7FFC8FF9EFD98F81DCF6959E0" box="[601,629,1998,2024]" pageId="2" pageNumber="284">2x</date>
|
||
as long as wide, with three tufts of very thin and long setae anteriorly and long, thin setae anteriorly and externally. Labium (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFCC2FF4ACE5A5EBB" box="[771,838,153,179]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1956,1981]" captionTargetBox="[260,1320,357,1922]" captionTargetId="figure@4.[254,1333,345,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="3" pageNumber="285">Fig. 9</figureCitation>
|
||
) with mentum subrectangular; lateral margins feebly sinuous, with four long setae on each side, anterolateral angles sharp; palpiger present; palpi with three palpomeres, palpomeres 1 and 2 transverse; apical palpomere larger than basal two combined, with sensorial pores and two short setae on lateral margin; ligula wider than long, anterior margin straight, anterior angles rounded and produced laterally with short spines, ventral surface with two transverse rows of spines on disc. Gena with tomentum.
|
||
</paragraph>
|
||
<caption id="67AA91BFFFC9FF9FFF56FA0FCF255811" ID-DOI="http://doi.org/10.5281/zenodo.3516232" ID-Zenodo-Dep="3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285" startId="3.[151,264,1500,1525]" targetBox="[216,1371,345,1475]" targetPageId="3">
|
||
<paragraph id="336AC137FFC9FF9FFF56FA0FCF255811" blockId="3.[151,1436,1500,1561]" pageId="3" pageNumber="285">
|
||
<emphasis id="01A11D25FFC9FF9FFF56FA0FCC205BFD" bold="true" box="[151,316,1500,1525]" pageId="3" pageNumber="285">FIGURES 3–5.</emphasis>
|
||
Habitus
|
||
<taxonomicName id="F4D5BAB4FFC9FF9FFE5AFA0ECF495BFD" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[411,597,1501,1525]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="01A11D25FFC9FF9FFE5AFA0ECF495BFD" box="[411,597,1501,1525]" italics="true" pageId="3" pageNumber="285">Pilielmis shepardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01A11D25FFC9FF9FFD9DFA0ECFB35BFD" bold="true" box="[604,687,1501,1525]" pageId="3" pageNumber="285">
|
||
<taxonomicNameLabel id="1A92A05EFFC9FF9FFD9DFA0ECFB35BFD" box="[604,687,1501,1525]" pageId="3" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="336AC137FFC9FF98FF06F994C9EF5F4B" blockId="3.[151,1437,1607,2030]" lastBlockId="4.[151,1437,153,323]" lastPageId="4" lastPageNumber="286" pageId="3" pageNumber="285">
|
||
Thorax: Pronotum moderately wider than long (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFD23F994CE3B5869" box="[738,807,1607,1633]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Fig. 3</figureCitation>
|
||
). Anterior margin and anterior angles moderately acute; lateral margins convex with granules; posterior angles acute; posterior margin smooth with two prescutellar fovea. Pronotal surface covered with golden setae on disc and, with micropunctures separated by their own diameter; posterior half alutaceous. Sublateral carinae complete and sinuous, reaching the anterior and posterior margin (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFAAAF960CDD958F9" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Figs 3, 5</figureCitation>
|
||
). Longitudinal impression on disc deep and broad. Elytra (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFCA7F904CECF58F9" box="[870,979,1750,1777]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Figs. 3, 5</figureCitation>
|
||
) 1.5x longer than wide, anterior margin straight; lateral margin slightly arcuate and apex truncate; apex yellowish in colour. Median suture elevated. Humeral angle elevated yellowish in colour. One short subcarinal basal on interval IV and one sublateral carina on interval VI, extending 3/4 of elytral length; surface with puncture rows separated by double rows of setae on each interval. Scutellum cordiform. Epipleura covered with tomentum (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFC48F8B4C93D5989" box="[905,1057,1894,1921]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Figs. 4 and 5</figureCitation>
|
||
). Hind wings macropterous. Prosternum as long as wide, with lateral portion covered by tomentum (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFC53F858C93959AD" box="[914,1061,1930,1957]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Figs. 4 and 5</figureCitation>
|
||
). Hypomeron
|
||
<date id="476BE7F7FFC9FF9FFB06F858C9FF59AD" box="[1223,1251,1931,1957]" pageId="3" pageNumber="285">3x</date>
|
||
as long as wide, with a belt of tomentum reaching the anterior and lateral margins (
|
||
<figureCitation id="ABEEDDB2FFC9FF9FFCB9F87CC91F59C1" box="[888,1027,1966,1993]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="3" pageNumber="285">Figs 4 and 5</figureCitation>
|
||
). Prosternal process as long as wide; posterior margin elevated; surface with micropunctures; apex truncate. Mesoventrite with slightly broad depression to receive prosternal process; surface similar to prosternum. Metaventrite broad; lateral part covered by tomentum, disc alutaceous, with a longitudinal line, surface with few short setae. Legs except the tarsi covered with tomentum. Coxae and trochanter rounded. Femora narrowed at distal margin,
|
||
<date id="476BE7F7FFCEFF98FC56FF32CEAF5EF3" box="[919,947,225,251]" pageId="4" pageNumber="286">4x</date>
|
||
as long as wide. Tibiae elongate and thin, approximately
|
||
<date id="476BE7F7FFCEFF98FE81FED6CC405F17" box="[320,348,261,287]" pageId="4" pageNumber="286">6x</date>
|
||
as long as wide; a row of spines on distal half. Tarsus elongate, apical tarsomere longer than the basal four combined, with long setae on apex. Claws large, thin with a tooth near the base (
|
||
<figureCitation id="ABEEDDB2FFCEFF98FB52FEFAC9FA5F4B" box="[1171,1254,296,323]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="4" pageNumber="286">Fig. 5a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="67AA91BFFFCEFF98FF56F877C91859E9" ID-DOI="http://doi.org/10.5281/zenodo.3516234" ID-Zenodo-Dep="3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="4" pageNumber="286" startId="4.[151,264,1956,1981]" targetBox="[260,1320,357,1922]" targetPageId="4">
|
||
<paragraph id="336AC137FFCEFF98FF56F877C91859E9" blockId="4.[151,1436,1956,2017]" pageId="4" pageNumber="286">
|
||
<emphasis id="01A11D25FFCEFF98FF56F877CC5A59B5" bold="true" box="[151,326,1956,1981]" pageId="4" pageNumber="286">FIGURES 6–10.</emphasis>
|
||
<taxonomicName id="F4D5BAB4FFCEFF98FE8AF876CF1F59B5" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[331,515,1957,1981]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="01A11D25FFCEFF98FE8AF876CF1F59B5" box="[331,515,1957,1981]" italics="true" pageId="4" pageNumber="286">Pilielmis shepardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01A11D25FFCEFF98FDC9F876CF7959B5" bold="true" box="[520,613,1956,1981]" pageId="4" pageNumber="286">
|
||
<taxonomicNameLabel id="1A92A05EFFCEFF98FDC9F876CF4B59B5" box="[520,599,1957,1981]" pageId="4" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
</emphasis>
|
||
6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="336AC137FFCFFF99FF06FF4AC9465EDF" blockId="5.[151,1437,153,936]" pageId="5" pageNumber="287">
|
||
Abdomen: Five visible ventrites (
|
||
<figureCitation id="ABEEDDB2FFCFFF99FD83FF4ACF9B5EBB" box="[578,647,153,179]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="5" pageNumber="287">Fig. 4</figureCitation>
|
||
), with margins moderately arcuate; surface covered by tomentum and micropunctures. Ventrite I with disc lacking tomentum. Ventrite V with apex rounded.
|
||
</paragraph>
|
||
<paragraph id="336AC137FFCFFF99FF06FF32C9755F6F" blockId="5.[151,1437,153,936]" pageId="5" pageNumber="287">
|
||
Genitalia: Total length
|
||
<quantity id="F42D6CD2FFCFFF99FE12FF32CF205EF3" box="[467,572,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="287" unit="mm" value="0.37">0.37 mm</quantity>
|
||
, greatest width
|
||
<quantity id="F42D6CD2FFCFFF99FD37FF32CE415EF3" box="[758,861,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="287" unit="mm" value="0.13">0.13 mm</quantity>
|
||
(
|
||
<figureCitation id="ABEEDDB2FFCFFF99FCAEFF32CEDA5EF3" box="[879,966,225,251]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1956,1981]" captionTargetBox="[260,1320,357,1922]" captionTargetId="figure@4.[254,1333,345,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Pilielmis shepardi sp. nov. (6) Labrum, ventral view. (7) Right mandible, ventral view. (8) Left maxilla, dorsal view. (9) Labium, ventral view. (10) Male genitalia, ventral view. Scale bars = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.3516234" httpUri="https://zenodo.org/record/3516234/files/figure.png" pageId="5" pageNumber="287">Fig. 10</figureCitation>
|
||
). Phallobase as long as wide. Parameres reaching ¾ of penis length, wider at base than at apex, rounded apically. Penis 1.5x the length of phallobase, ca 3.6x, basolateral apophyses twice as long as wide; lateral margin moderately sinuous and concave, apically with sensorial pores, apex slightly sharp; fibula approximately the length of the penis; corona present.
|
||
</paragraph>
|
||
<paragraph id="336AC137FFCFFF99FF06FEA2CFD25F83" blockId="5.[151,1437,153,936]" box="[199,718,369,395]" pageId="5" pageNumber="287">Female: External morphology similar to male.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BCF92BCFFCFFF99FF06FE46CF265FDB" pageId="5" pageNumber="287" type="etymology">
|
||
<paragraph id="336AC137FFCFFF99FF06FE46CF265FDB" blockId="5.[151,1437,153,936]" pageId="5" pageNumber="287">
|
||
<emphasis id="01A11D25FFCFFF99FF06FE46CC525FA7" bold="true" box="[199,334,405,431]" pageId="5" pageNumber="287">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FDFDFE46CF815FA7" authorityName="Polizei & Barclay" authorityYear="2019" box="[572,669,405,431]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="shepardi">
|
||
<emphasis id="01A11D25FFCFFF99FDFDFE46CF815FA7" box="[572,669,405,431]" italics="true" pageId="5" pageNumber="287">shepardi</emphasis>
|
||
</taxonomicName>
|
||
is a tribute to Dr. William Shepard in honour of his many contributions to the study of the riffle beetles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BCF92BCFFCFFF99FF06FE0FC9225DAF" pageId="5" pageNumber="287" type="discussion">
|
||
<paragraph id="336AC137FFCFFF99FF06FE0FC9EC5CFB" blockId="5.[151,1437,153,936]" pageId="5" pageNumber="287">
|
||
<emphasis id="01A11D25FFCFFF99FF06FE0FCC535FFF" bold="true" box="[199,335,476,503]" pageId="5" pageNumber="287">Comments.</emphasis>
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FE97FE0ECF3E5FFF" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[342,546,477,503]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="01A11D25FFCFFF99FE97FE0ECF3E5FFF" box="[342,546,477,503]" italics="true" pageId="5" pageNumber="287">Pilielmis shepardi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1A92A05EFFCFFF99FDE8FE0ECF9C5FFF" box="[553,640,477,503]" pageId="5" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be easily differentiated from all other congeners by the colour pattern and the morphology of the male genitalia. The new species resembles
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FC0EFDD1C9D35C13" authority="Hinton, 1971" authorityName="Hinton" authorityYear="1971" box="[975,1231,513,539]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="apama">
|
||
<emphasis id="01A11D25FFCFFF99FC0EFDD1C92A5C13" box="[975,1078,514,539]" italics="true" pageId="5" pageNumber="287">P. apama</emphasis>
|
||
<bibRefCitation id="5744BCC6FFCFFF99FBFDFDD2C9D35C13" author="Hinton, H. E." box="[1084,1231,513,539]" pageId="5" pageNumber="287" pagination="161 - 166" refId="ref3673" refString="Hinton, H. E. (1971) Pilielmis, a new genus of Elmidae (Coleoptera). Entomologist's Monthly Magazine, 107, 161 - 166." type="journal article" year="1971">
|
||
Hinton,
|
||
<quantity id="F42D6CD2FFCFFF99FB56FDD2C9D35C13" box="[1175,1231,513,539]" metricMagnitude="1" metricUnit="m" metricValue="5.00634" pageId="5" pageNumber="287" unit="in" value="1971.0">1971</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in body shape and size, rounded trochanters and the first abdominal ventrite with the disc lacking tomentum, but can be differentiated because in
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FED0FD99CC645C6B" authorityName="Hinton" authorityYear="1971" box="[273,376,586,611]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="apama">
|
||
<emphasis id="01A11D25FFCFFF99FED0FD99CC645C6B" box="[273,376,586,611]" italics="true" pageId="5" pageNumber="287">P. apama</emphasis>
|
||
</taxonomicName>
|
||
the antennae have the two first antennomeres distinctly paler than the remainder, and the colour pattern of the body is dark brown to blackish. In the male genitalia,
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FC4FFDBDCEEA5C8F" authorityName="Hinton" authorityYear="1971" box="[910,1014,622,647]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="apama">
|
||
<emphasis id="01A11D25FFCFFF99FC4FFDBDCEEA5C8F" box="[910,1014,622,647]" italics="true" pageId="5" pageNumber="287">P. apama</emphasis>
|
||
</taxonomicName>
|
||
has the phallobase approximately the same length as the penis, and the penis is wider basally than apically with apex rounded.
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FBB0FD41C9F05CA3" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[1137,1260,657,683]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="01A11D25FFCFFF99FBB0FD41C9F05CA3" box="[1137,1260,657,683]" italics="true" pageId="5" pageNumber="287">P. shepardi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1A92A05EFFCFFF99FB32FD42C8555CA3" box="[1267,1353,657,683]" pageId="5" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has the antennae uniformly coloured, the body pattern colour brownish to yellow. The phallobase is half of the penis length, and the penis has the lateral margin moderately sinuous and concave, and the apex slightly pointed.
|
||
</paragraph>
|
||
<paragraph id="336AC137FFCFFF99FF06FD2EC9225DAF" blockId="5.[151,1437,153,936]" pageId="5" pageNumber="287">
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FF06FD2ECC375D1F" authorityName="Hinton" authorityYear="1971" box="[199,299,765,791]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01A11D25FFCFFF99FF06FD2ECC375D1F" box="[199,299,765,791]" italics="true" pageId="5" pageNumber="287">Pilielmis</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FE33FD2ECF8B5D1F" authorityName="Hinton" authorityYear="1940" box="[498,663,765,791]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01A11D25FFCFFF99FE33FD2ECF8B5D1F" box="[498,663,765,791]" italics="true" pageId="5" pageNumber="287">Hexacylloepus</emphasis>
|
||
</taxonomicName>
|
||
but can be easily differentiated by the presence of only one sublateral carina on the elytra, and the claws bearing a tooth near the base (
|
||
<figureCitation id="ABEEDDB2FFCFFF99FCB3FCF2CED95D33" box="[882,965,800,827]" captionStart="FIGURES 3–5" captionStartId="3.[151,264,1500,1525]" captionTargetBox="[216,1371,345,1475]" captionTargetId="figure@3.[216,1372,345,1475]" captionTargetPageId="3" captionText="FIGURES 3–5. Habitus Pilielmis shepardi sp. nov. male, holotype. (3) Dorsal view. (4) Ventral view. (5) Lateral view. (5a) Hind tarsi and claw. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.3516232" httpUri="https://zenodo.org/record/3516232/files/figure.png" pageId="5" pageNumber="287">Fig. 5a</figureCitation>
|
||
).
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FC18FCF2C9625D33" authorityName="Hinton" authorityYear="1940" box="[985,1150,801,827]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01A11D25FFCFFF99FC18FCF2C9625D33" box="[985,1150,801,827]" italics="true" pageId="5" pageNumber="287">Hexacylloepus</emphasis>
|
||
</taxonomicName>
|
||
has two sublateral carina on the elytra and claws lack a tooth. The
|
||
<typeStatus id="EC6E7F95FFCFFF99FDA5FC96CF885D57" box="[612,660,837,863]" pageId="5" pageNumber="287">type</typeStatus>
|
||
material of the new species was housed in USNM as
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FB31FC96C8895D57" authorityName="Hinton" authorityYear="1940" box="[1264,1429,837,863]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01A11D25FFCFFF99FB31FC96C8895D57" box="[1264,1429,837,863]" italics="true" pageId="5" pageNumber="287">Hexacylloepus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FF56FCBACC7E5D8B" authority="Polizei & Barclay, 2019" authorityName="Polizei & Barclay" authorityYear="2019" box="[151,354,873,899]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="01A11D25FFCFFF99FF56FCBACC7E5D8B" box="[151,354,873,899]" italics="true" pageId="5" pageNumber="287">Pilielmis shepardi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1A92A05EFFCFFF99FEA9FCBACCA25D8B" box="[360,446,873,899]" pageId="5" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
represents the first record of the genus
|
||
<taxonomicName id="F4D5BAB4FFCFFF99FCB2FCBACECB5D8B" authorityName="Hinton" authorityYear="1971" box="[883,983,873,899]" class="Insecta" family="Elmidae" genus="Pilielmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01A11D25FFCFFF99FCB2FCBACECB5D8B" box="[883,983,873,899]" italics="true" pageId="5" pageNumber="287">Pilielmis</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<collectingCountry id="4BC281A7FFCFFF99FBC5FCBAC9675D8B" box="[1028,1147,873,899]" name="Venezuela" pageId="5" pageNumber="287">Venezuela</collectingCountry>
|
||
. Until now this genus was recorded only from
|
||
<collectingCountry id="4BC281A7FFCFFF99FEB7FC5ECCA65DAF" box="[374,442,909,935]" name="Brazil" pageId="5" pageNumber="287">Brazil</collectingCountry>
|
||
(North – Amazon region),
|
||
<collectingCountry id="4BC281A7FFCFFF99FD2DFC5ECE465DAF" box="[748,858,909,935]" name="Colombia" pageId="5" pageNumber="287">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="4BC281A7FFCFFF99FC51FC5EC9265DAF" box="[912,1082,909,935]" name="French Guiana" pageId="5" pageNumber="287">French Guiana</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |