1208 lines
150 KiB
XML
1208 lines
150 KiB
XML
<document id="20AD95AE4E74829BC18AF4275B22EF64" ID-DOI="10.11646/zootaxa.5570.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734603" ID-ZooBank="BBC01CC8-D0A1-4032-9D2D-4F18BFD5761E" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1737736144597" checkinUser="plazi" docAuthor="Le, Manh Van, Nguyen, Luan Thanh, Murphy, Robert W., Tran, Tuyet-Dzung Thi, Nguyen, Sang Ngoc & Che, Jing" docDate="2025" docId="BB6B87E6FFBF7560FF3AFA881BCFBBD3" docLanguage="en" docName="zootaxa.5570.1.2.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnonectes splendissimus Le, Nguyen, Murphy, Tran, Nguyen & Che, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="71" masterDocId="4752FF9EFFB8756EFFADFFFC1A47BE12" masterDocTitle="A new fanged frog of the genus Limnonectes Fitzinger, 1843 (Anura: Dicroglossidae) from Langbian Plateau, southern Vietnam" masterLastPageNumber="80" masterPageNumber="57" pageNumber="64" updateTime="1738057485176" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D26519E1D72C021F82198D0A89359911">
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<mods:title id="1009A3493F011D2DF96B0553815A9077">A new fanged frog of the genus Limnonectes Fitzinger, 1843 (Anura: Dicroglossidae) from Langbian Plateau, southern Vietnam</mods:title>
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<mods:name id="03837CBF9CD78B07F8F42C5A1220BC67" type="personal">
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<mods:roleTerm id="9AD721EBF3849F6930A52C8E1BB2742E">Author</mods:roleTerm>
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<mods:namePart id="4658012FEEE26B3574DE07C2E9FD8CC2">Le, Manh Van</mods:namePart>
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<mods:affiliation id="D788A48A38A8F4953F64C315909D9D8A">Key Laboratory of Genetic Evolution and Animal Models, and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China & Kunming College of Life Science, University of Chinese Academy of Sciences, Kunming 650204, China & Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Dist. 3, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:name id="58CCCACD83450F96BCCE2B5E5F990036" type="personal">
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<mods:roleTerm id="AD0F7BF397FBF6CA24C22A5A3B28C1EC">Author</mods:roleTerm>
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<mods:namePart id="C5679C3E798FBC382F70BDAADEA8E117">Nguyen, Luan Thanh</mods:namePart>
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<mods:affiliation id="ADBB397DF985E8D7B5878B5090597EDA">Asian Turtle Program of Indo-Myanmar Conservation, No. 1806, C 14 Bac Ha Building, To Huu St., Nam Tu Liem Dist., Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="CE3D3649647FEE985B642372C0800925">Murphy, Robert W.</mods:namePart>
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<mods:affiliation id="CD094E8A84D66A4435ECC408A23941D9">Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, Toronto, Ontario M 5 S 2 C 6, Canada</mods:affiliation>
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</mods:name>
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<mods:namePart id="9A12FF7C862BEB949AB5311F140EE10F">Tran, Tuyet-Dzung Thi</mods:namePart>
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<mods:affiliation id="19E4892EB273889F85C5DB42B870D228">Asian Turtle Program of Indo-Myanmar Conservation, No. 1806, C 14 Bac Ha Building, To Huu St., Nam Tu Liem Dist., Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier id="996041C97F90EE657C9DC31453E8260F" type="email">tuyetdung26@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9622F71679039FB140D9A1E8AE5479D6">Nguyen, Sang Ngoc</mods:namePart>
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<mods:affiliation id="0FC90B1FA9F1AAFB2C10EA7A5B5749DC">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan St., Dist. 3, Ho Chi Minh City, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="12DE70509379925955C63EABDABF470B">Che, Jing</mods:namePart>
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<mods:affiliation id="81ACD3AE7D73075347D509078F7634C0">Key Laboratory of Genetic Evolution and Animal Models, and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China & Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
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<mods:title id="FA664BE204FCD3DEA90FFF0BA283797D">Zootaxa</mods:title>
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<mods:part id="93C1BDCBDAF65B64EF0E765E106EAA28">
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<mods:date id="02D8B6BE66BBFC67AB9ABFAECC235996">2025</mods:date>
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<mods:number id="45C7249F7E7DD4FCC5820AE9D1AC870C">2025-01-16</mods:number>
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<mods:number id="46F06988EA2CDAB3E1DE480678DEC7EF">5570</mods:number>
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<mods:url id="DB11B5A6E239CFBDF727A907D407F4AE">https://doi.org/10.11646/zootaxa.5570.1.2</mods:url>
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<mods:classification id="9D31F878F4511D49293A3817A52A88C9">journal article</mods:classification>
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<mods:identifier id="9D465B87527FABC76D0E4B3E3A2BAAFB" type="ISSN">1175-5326</mods:identifier>
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<emphasis id="01B6EAE2FFBF7569FF3AFA881BA6BB9D" bold="true" box="[151,481,1396,1423]" italics="true" pageId="7" pageNumber="64">Limnonectes splendissimus</emphasis>
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</emphasis>
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<subSubSection id="7BD8657BFFBF7569FF3AFA411809B92D" pageId="7" pageNumber="64" type="materials_examined">
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<typeStatus id="EC798852FFBF7569FF3AFA411B41BBC5" box="[151,262,1469,1495]" pageId="7" pageNumber="64" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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, an
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(
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&
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<figureCitation id="ABF92A75FFBF7569FD74FA4118AEBBC5" box="[729,745,1469,1495]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-17@9.[151,1436,181,1501]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Limnonectes splendissimus sp. nov. in life. A. Dorsal surface; B. Ventral surface; C. Nuptial pad on finger I; scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.14734611" httpUri="https://zenodo.org/record/14734611/files/figure.png" pageId="7" pageNumber="64">4</figureCitation>
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), collected from
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,
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<collectingMunicipality id="D319AC8AFFBF7569FB0FFA401F1BBBC4" box="[1186,1372,1468,1494]" pageId="7" pageNumber="64">Di Linh District</collectingMunicipality>
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,
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<collectingRegion id="F106F812FFBF7569FAC4FA411B02BBE8" country="Vietnam" name="Lam Dong" pageId="7" pageNumber="64">Lam Dong Province</collectingRegion>
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,
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<collectingCountry id="4BD57660FFBF7569FEFFFA1C1BF3BBE8" box="[338,436,1504,1530]" name="Vietnam" pageId="7" pageNumber="64">Vietnam</collectingCountry>
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; coordinates
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11
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27’11.6”N
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,
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108
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05’32.3”E
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;
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elevation
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<quantity id="F43A9B15FFBF7569FBB3FA1C1E32BBE9" box="[1054,1141,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="7" pageNumber="64" unit="m" value="1369.0">1369 m</quantity>
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a.s.l.
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</elevation>
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by
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<collectorName id="9E375326FFBF7569FB4CFA1C1F2CBBE9" box="[1249,1387,1504,1531]" pageId="7" pageNumber="64">Manh V. Le</collectorName>
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and
|
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<collectorName id="9E375326FFBF7569FF3AF9F91B1EB80D" box="[151,345,1541,1567]" pageId="7" pageNumber="64">Tuan M. Nguyen</collectorName>
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, on
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<date id="477C1030FFBF7569FE2AF9F91822B80D" box="[391,613,1540,1567]" pageId="7" pageNumber="64" value="2023-09-22">
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<collectingDate id="5738E9D8FFBF7569FE2AF9F91822B80D" box="[391,613,1540,1567]" pageId="7" pageNumber="64" value="2023-09-22">22 September 2023</collectingDate>
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</date>
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.
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<paragraph id="337D36F0FFBF7569FF6AF9D51809B92D" blockId="7.[151,1437,1468,2035]" pageId="7" pageNumber="64">
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<materialsCitation id="83AA3CADFFBF7569FF6AF9D51FDBB874" collectingDate="2023-09-22" collectionCode="ITBCZ" collectorName="Manh V. Le & Tuan M. Nguyen" country="Vietnam" elevation="1369" latitude="11.453222" location="Gung Re Commune" longLatPrecision="1" longitude="108.09231" municipality="Di Linh District" pageId="7" pageNumber="64" specimenCode="ITBCZ 8675, 8676, ITBCZ 8677, ITBCZ 8678" specimenCount="14" specimenCount-adult="3" specimenCount-female="1" specimenCount-male="2" specimenCount-subadult="1" stateProvince="Lam Dong Province" typeStatus="paratype">
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<emphasis id="01B6EAE2FFBF7569FF6AF9D51B01B851" bold="true" box="[199,326,1577,1603]" pageId="7" pageNumber="64">
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<typeStatus id="EC798852FFBF7569FF6AF9D51B06B851" box="[199,321,1577,1603]" pageId="7" pageNumber="64" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCount id="25C4FD79FFBF7569FEE3F9D51854B850" box="[334,531,1576,1603]" count="7" pageId="7" pageNumber="64" type="generic">Seven specimens</specimenCount>
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:
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,
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<specimenCode id="63649E8BFFBF7569FD10F9D418B2B850" box="[701,757,1576,1602]" collectionCode="ITBCZ" pageId="7" pageNumber="64">8676</specimenCode>
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(
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<specimenCount id="25C4FD79FFBF7569FCA9F9D51928B850" box="[772,879,1576,1603]" count="2" pageId="7" pageNumber="64" type="adult">two adult</specimenCount>
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<specimenCount id="25C4FD79FFBF7569FCDAF9D419FCB850" box="[887,955,1576,1602]" pageId="7" pageNumber="64" type="male">males</specimenCount>
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,
|
||
<figureCitation id="ABF92A75FFBF7569FC6AF9D41E05B851" box="[967,1090,1576,1603]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="7" pageNumber="64">Fig. 5A–D</figureCitation>
|
||
),
|
||
<specimenCode id="63649E8BFFBF7569FBFAF9D51EA0B850" box="[1111,1255,1576,1603]" pageId="7" pageNumber="64">ITBCZ 8677</specimenCode>
|
||
(
|
||
<specimenCount id="25C4FD79FFBF7569FB5BF9D41F10B850" box="[1270,1367,1576,1602]" pageId="7" pageNumber="64" type="subadult">subadult</specimenCount>
|
||
<specimenCount id="25C4FD79FFBF7569FAF2F9D41FDFB850" box="[1375,1432,1576,1602]" pageId="7" pageNumber="64" type="male">male</specimenCount>
|
||
,
|
||
<figureCitation id="ABF92A75FFBF7569FF3AF9B01B4EB875" box="[151,265,1612,1639]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="7" pageNumber="64">Fig. 5E, F</figureCitation>
|
||
),
|
||
<specimenCode id="63649E8BFFBF7569FEB0F9B11BEBB874" box="[285,428,1612,1639]" pageId="7" pageNumber="64">ITBCZ 8678</specimenCode>
|
||
(
|
||
<specimenCount id="25C4FD79FFBF7569FE16F9B01BB2B874" box="[443,501,1612,1638]" pageId="7" pageNumber="64" type="adult">adult</specimenCount>
|
||
<specimenCount id="25C4FD79FFBF7569FE56F9B0180BB874" box="[507,588,1612,1638]" pageId="7" pageNumber="64" type="female">female</specimenCount>
|
||
), collected by the same collectors, on the same date and site as the holotype;
|
||
</materialsCitation>
|
||
<materialsCitation id="83AA3CADFFBF7569FF3AF98D1826B8C1" collectingDate="2023-09-23" collectionCode="ITBCZ" collectorName="Manh V. Le & Tuan M. Nguyen" country="Vietnam" county="Duc Trong District" elevation="1611" latitude="11.827611" location="Voi Mountain" longLatPrecision="1" longitude="108.40811" municipality="Hiep Thanh Commune" pageId="7" pageNumber="64" specimenCode="ITBCZ 8679" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Lam Dong Province">
|
||
<specimenCode id="63649E8BFFBF7569FF3AF98D1B6FB898" box="[151,296,1648,1675]" pageId="7" pageNumber="64">ITBCZ 8679</specimenCode>
|
||
(
|
||
<specimenCount id="25C4FD79FFBF7569FE9AF98C1B36B898" box="[311,369,1648,1674]" pageId="7" pageNumber="64" type="adult">adult</specimenCount>
|
||
<specimenCount id="25C4FD79FFBF7569FED5F98C1B80B898" box="[376,455,1648,1674]" pageId="7" pageNumber="64" type="female">female</specimenCount>
|
||
,
|
||
<figureCitation id="ABF92A75FFBF7569FE7EF98C180AB899" box="[467,589,1648,1675]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="7" pageNumber="64">Fig. 5G, H</figureCitation>
|
||
) collected from
|
||
<location id="361D602BFFBF7569FCAAF98C19E2B898" LSID="urn:lsid:plazi:treatment:BB6B87E6FFBF7560FF3AFA881BCFBBD3:361D602BFFBF7569FCAAF98C19E2B898" box="[775,933,1648,1674]" country="Vietnam" county="Duc Trong District" latitude="11.827611" longLatPrecision="1" longitude="108.40811" municipality="Hiep Thanh Commune" name="Voi Mountain" pageId="7" pageNumber="64" stateProvince="Lam Dong Province">Voi Mountain</location>
|
||
,
|
||
<collectingMunicipality id="D319AC8AFFBF7569FC1DF98C1EF2B899" box="[944,1205,1648,1675]" pageId="7" pageNumber="64">Hiep Thanh Commune</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="DA1C4E7CFFBF7569FB6DF98D1FDFB898" box="[1216,1432,1648,1675]" pageId="7" pageNumber="64">Duc Trong District</collectingCounty>
|
||
,
|
||
<collectingRegion id="F106F812FFBF7569FF3AF9691B3DB8BC" box="[151,378,1684,1711]" country="Vietnam" name="Lam Dong" pageId="7" pageNumber="64">Lam Dong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="4BD57660FFBF7569FE29F9681BA1B8BC" box="[388,486,1684,1710]" name="Vietnam" pageId="7" pageNumber="64">Vietnam</collectingCountry>
|
||
; coordinates
|
||
<geoCoordinate id="56F65037FFBF7569FDD5F9691953B8BD" box="[632,788,1685,1711]" degrees="11" direction="north" minutes="49" orientation="latitude" pageId="7" pageNumber="64" precision="1" seconds="39.4" value="11.827611">
|
||
11
|
||
<superScript id="C4B79BB8FFBF7569FD3EF96918DCB8B1" attach="none" box="[659,667,1685,1699]" fontSize="6" pageId="7" pageNumber="64">o</superScript>
|
||
49’39.4”N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="56F65037FFBF7569FCB2F969198FB8BD" box="[799,968,1685,1711]" degrees="108" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="64" precision="1" seconds="29.2" value="108.40811">
|
||
108
|
||
<superScript id="C4B79BB8FFBF7569FCE4F9691916B8B1" attach="none" box="[841,849,1685,1699]" fontSize="6" pageId="7" pageNumber="64">o</superScript>
|
||
24’29.2”E
|
||
</geoCoordinate>
|
||
;
|
||
<elevation id="B8EFD1C3FFBF7569FC7FF9681ED5B8BD" box="[978,1170,1684,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="7" pageNumber="64" unit="m" value="1611.0">
|
||
elevation
|
||
<quantity id="F43A9B15FFBF7569FB92F9681ED5B8BD" box="[1087,1170,1684,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="7" pageNumber="64" unit="m" value="1611.0">1611 m</quantity>
|
||
</elevation>
|
||
a.s.l by
|
||
<collectorName id="9E375326FFBF7569FB41F9681F29B8BD" box="[1260,1390,1684,1711]" pageId="7" pageNumber="64">Manh V. Le</collectorName>
|
||
and
|
||
<collectorName id="9E375326FFBF7569FF3AF9451B11B8C1" box="[151,342,1721,1747]" pageId="7" pageNumber="64">Tuan M. Nguyen</collectorName>
|
||
, on
|
||
<date id="477C1030FFBF7569FE2EF945181AB8C1" box="[387,605,1720,1747]" pageId="7" pageNumber="64" value="2023-09-23">
|
||
<collectingDate id="5738E9D8FFBF7569FE2EF945181AB8C1" box="[387,605,1720,1747]" pageId="7" pageNumber="64" value="2023-09-23">23 September 2023</collectingDate>
|
||
</date>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="83AA3CADFFBF7569FDC5F9451E70B8C1" box="[616,1079,1720,1747]" collectionCode="ITBCZ" pageId="7" pageNumber="64" specimenCode="ITBCZ 3626" specimenCount="2" specimenCount-female="1" specimenCount-subadult="1">
|
||
<specimenCode id="63649E8BFFBF7569FDC5F94518B1B8C0" box="[616,758,1720,1747]" pageId="7" pageNumber="64">ITBCZ 3626</specimenCode>
|
||
(
|
||
<specimenCount id="25C4FD79FFBF7569FCA9F9441922B8C0" box="[772,869,1720,1746]" pageId="7" pageNumber="64" type="subadult">subadult</specimenCount>
|
||
<specimenCount id="25C4FD79FFBF7569FCC7F94419FEB8C0" box="[874,953,1720,1746]" pageId="7" pageNumber="64" type="female">female</specimenCount>
|
||
,
|
||
<figureCitation id="ABF92A75FFBF7569FC6EF9441E68B8C1" box="[963,1071,1720,1747]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="7" pageNumber="64">Fig. 5I, K</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="83AA3CADFFBF7569FB90F9441809B92D" collectingDate="2023-02-25" collectionCode="ITBCZ" collectorName="Luan T. Nguyen & Manh V. Le & Tuan M. Nguyen & Dzung T. T. Tran" country="Vietnam" county="Duc Trong District" elevation="1488" latitude="11.827611" location="Voi Mountain" longLatPrecision="1" longitude="108.40811" municipality="Hiep Thanh Commune" pageId="7" pageNumber="64" specimenCode="ITBCZ 3627" specimenCount="1" specimenCount-juvenile="1" stateProvince="Lam Dong Province">
|
||
and
|
||
<specimenCode id="63649E8BFFBF7569FBC6F9451EBDB8C0" box="[1131,1274,1720,1747]" pageId="7" pageNumber="64">ITBCZ 3627</specimenCode>
|
||
(
|
||
<specimenCount id="25C4FD79FFBF7569FAAAF9441F21B8C0" box="[1287,1382,1720,1746]" pageId="7" pageNumber="64" type="juvenile">juvenile</specimenCount>
|
||
,
|
||
<figureCitation id="ABF92A75FFBF7569FADDF9441AA7B8E5" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="7" pageNumber="64">Fig. 5L, M</figureCitation>
|
||
) collected from
|
||
<location id="361D602BFFBF7569FE30F920187CB8E4" LSID="urn:lsid:plazi:treatment:BB6B87E6FFBF7560FF3AFA881BCFBBD3:361D602BFFBF7569FE30F920187CB8E4" box="[413,571,1756,1782]" country="Vietnam" county="Duc Trong District" latitude="11.827611" longLatPrecision="1" longitude="108.40811" municipality="Hiep Thanh Commune" name="Voi Mountain" pageId="7" pageNumber="64" stateProvince="Lam Dong Province">Voi Mountain</location>
|
||
,
|
||
<collectingMunicipality id="D319AC8AFFBF7569FDE5F9201909B8E5" box="[584,846,1756,1783]" pageId="7" pageNumber="64">Hiep Thanh Commune</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="DA1C4E7CFFBF7569FCF6F9211E73B8E4" box="[859,1076,1756,1783]" pageId="7" pageNumber="64">Duc Trong District</collectingCounty>
|
||
,
|
||
<collectingRegion id="F106F812FFBF7569FBECF9211F6EB8E4" box="[1089,1321,1756,1783]" country="Vietnam" name="Lam Dong" pageId="7" pageNumber="64">Lam Dong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="4BD57660FFBF7569FA9BF9201FDFB8E4" box="[1334,1432,1756,1782]" name="Vietnam" pageId="7" pageNumber="64">Vietnam</collectingCountry>
|
||
; coordinates
|
||
<geoCoordinate id="56F65037FFBF7569FEB3F8FD1BFDB909" box="[286,442,1793,1819]" degrees="11" direction="north" minutes="49" orientation="latitude" pageId="7" pageNumber="64" precision="1" seconds="39.4" value="11.827611">
|
||
11
|
||
<superScript id="C4B79BB8FFBF7569FE95F8FD1B07B91D" attach="none" box="[312,320,1793,1807]" fontSize="6" pageId="7" pageNumber="64">o</superScript>
|
||
49’39.4”N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="56F65037FFBF7569FE69F8FD182AB909" box="[452,621,1793,1819]" degrees="108" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="64" precision="1" seconds="29.2" value="108.40811">
|
||
108
|
||
<superScript id="C4B79BB8FFBF7569FE43F8FD1BB1B91D" attach="none" box="[494,502,1793,1807]" fontSize="6" pageId="7" pageNumber="64">o</superScript>
|
||
24’29.2”E
|
||
</geoCoordinate>
|
||
;
|
||
<elevation id="B8EFD1C3FFBF7569FDDAF8FC1971B909" box="[631,822,1792,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.488" pageId="7" pageNumber="64" unit="m" value="1488.0">
|
||
elevation
|
||
<quantity id="F43A9B15FFBF7569FD4EF8FD1971B909" box="[739,822,1793,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.488" pageId="7" pageNumber="64" unit="m" value="1488.0">1488 m</quantity>
|
||
</elevation>
|
||
a.s.l by
|
||
<collectorName id="9E375326FFBF7569FC22F8FD1E04B909" box="[911,1091,1793,1819]" pageId="7" pageNumber="64">Luan T. Nguyen</collectorName>
|
||
,
|
||
<collectorName id="9E375326FFBF7569FBE0F8FC1E97B909" box="[1101,1232,1792,1819]" pageId="7" pageNumber="64">Manh V. Le</collectorName>
|
||
,
|
||
<collectorName id="9E375326FFBF7569FB77F8FD1FDFB909" box="[1242,1432,1793,1819]" pageId="7" pageNumber="64">Tuan M. Nguyen</collectorName>
|
||
,
|
||
<collectorName id="9E375326FFBF7569FF3AF8D91B15B92D" box="[151,338,1829,1855]" pageId="7" pageNumber="64">Dzung T.T. Tran</collectorName>
|
||
, on
|
||
<date id="477C1030FFBF7569FE2DF8D8180CB92D" box="[384,587,1828,1855]" pageId="7" pageNumber="64" value="2023-02-25">
|
||
<collectingDate id="5738E9D8FFBF7569FE2DF8D8180CB92D" box="[384,587,1828,1855]" pageId="7" pageNumber="64" value="2023-02-25">25 February 2023</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BD8657BFFBF7566FF6AF8B41B12BF9B" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64" type="diagnosis">
|
||
<paragraph id="337D36F0FFBF7566FF6AF8B41B12BF9B" blockId="7.[151,1437,1468,2035]" lastBlockId="8.[151,1437,151,393]" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64">
|
||
<emphasis id="01B6EAE2FFBF7569FF6AF8B41B05B970" bold="true" box="[199,322,1864,1890]" pageId="7" pageNumber="64">Diagnosis.</emphasis>
|
||
The new species belongs to the genus
|
||
<taxonomicName id="F4C24D73FFBF7569FD55F8B519C1B970" authorityName="Fitzinger" authorityYear="1843" box="[760,902,1865,1890]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="genus">
|
||
<emphasis id="01B6EAE2FFBF7569FD55F8B519C1B970" box="[760,902,1865,1890]" italics="true" pageId="7" pageNumber="64">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
based on both molecular analyses (
|
||
<bibRefCitation id="57534B01FFBF7569FABBF8B41A96B995" author="Portik, D. M. & Streicher, J. W. & Wiens, J. J." pageId="7" pageNumber="64" refId="ref15896" refString="Portik, D. M., Streicher, J. W. & Wiens, J. J. (2023) Frog phylogeny: a time-calibrated, species-level tree based on hundreds of loci and 5,242 species. Molecular phylogenetics and evolution, 188, 107907. https: // doi. org / 10.1016 / j. ympev. 2023.107907" type="journal volume" year="2023">
|
||
Portik
|
||
<emphasis id="01B6EAE2FFBF7569FAC9F8B51FDAB970" box="[1380,1437,1864,1890]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
|
||
2023
|
||
</bibRefCitation>
|
||
) and morphological assessment because it has fang-like odontoid processes on its lower jaw, and with males being larger than females (
|
||
<bibRefCitation id="57534B01FFBF7569FE6FF86D18ECB9B9" author="Emerson, S. B. & Inger, R. & Iskandar, D." box="[450,683,1936,1963]" pageId="7" pageNumber="64" pagination="131 - 142" refId="ref13042" refString="Emerson, S. B., Inger, R. & Iskandar, D. (2000) Molecular systematics and biogeography of the fanged frogs of Southeast Asia. Molecular Phylogenetics and Evolution, 16, 131 - 142. https: // doi. org / 10.1006 / mpev. 2000.0778" type="journal article" year="2000">
|
||
Emerson
|
||
<emphasis id="01B6EAE2FFBF7569FD82F86D1820B9B8" box="[559,615,1936,1962]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="F4C24D73FFBF7569FD13F86D19A9B9B8" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[702,1006,1936,1962]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFBF7569FD13F86D19A9B9B8" box="[702,1006,1936,1962]" italics="true" pageId="7" pageNumber="64">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFBF7569FC58F86D1E08B9B9" bold="true" box="[1013,1103,1937,1963]" pageId="7" pageNumber="64">
|
||
<taxonomicNameLabel id="1A855799FFBF7569FC58F86D1E08B9B9" box="[1013,1103,1937,1963]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all of its congeners by a combination of the following morphological characters: large body size (SVL 104.1–107.0 mm in adult males,
|
||
<quantity id="F43A9B15FFBF7569FEECF8251BA3B9E1" box="[321,484,2009,2035]" metricMagnitude="-2" metricUnit="m" metricValue="9.245" metricValueMax="9.77" metricValueMin="8.72" pageId="7" pageNumber="64" unit="mm" value="92.45" valueMax="97.7" valueMin="87.2">87.2–97.7 mm</quantity>
|
||
in adult females); head length longer than wide (HW/HL = 0.91±
|
||
<quantity id="F43A9B15FFBF7569FB6CF8251F48B9E0" box="[1217,1295,2008,2035]" metricMagnitude="-4" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="64" unit="in" value="0.01">0.01 in</quantity>
|
||
adult males, 0.87±
|
||
<quantity id="F43A9B15FFB07566FF7AFF6B1B60BEA3" box="[215,295,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="65" unit="in" value="0.08">0.08 in</quantity>
|
||
adult females); males with increased head height and a nuptial pad on Finger I; tympanum distinct; eye diameter about three-fifths of tympanum-eye distance in adult males (ED/TED = 0.62±0.13) while longer than in adult females (ED/TED = 1.40±0.02); presence of vomerine teeth; odontoid developed (OH 8.0±
|
||
<quantity id="F43A9B15FFB07566FB41FF231F05BEEB" box="[1260,1346,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="65" unit="mm" value="0.6">0.6 mm</quantity>
|
||
in adult males, 4.5±
|
||
<quantity id="F43A9B15FFB07566FEB7FEFE1B36BF0F" box="[282,369,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="65" unit="mm" value="0.9">0.9 mm</quantity>
|
||
in adult females); finger length relative III>I>II>IV, no webbing between fingers; relative toe lengths IV>III>V>II>I; toes webbed to distal of terminal phalanx, formula I0-0II0-0III0-0IV0-0V; throat pattern of juveniles distinct with black and white spots that cover both the throat and the anterior half of venter, posterior half of venter yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="67BD6678FFB07566FF3AFBD119CBBA9F" ID-DOI="http://doi.org/10.5281/zenodo.14734609" ID-Zenodo-Dep="14734609" httpUri="https://zenodo.org/record/14734609/files/figure.png" pageId="8" pageNumber="65" startId="8.[151,250,1069,1093]" targetBox="[158,1427,427,1035]" targetPageId="8" targetType="figure">
|
||
<paragraph id="337D36F0FFB07566FF3AFBD119CBBA9F" blockId="8.[151,1436,1069,1165]" pageId="8" pageNumber="65">
|
||
<emphasis id="01B6EAE2FFB07566FF3AFBD11B54BA57" bold="true" box="[151,275,1069,1093]" pageId="8" pageNumber="65">FIGURE 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="F4C24D73FFB07566FE31FBD218F7BA57" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[412,688,1069,1093]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB07566FE31FBD218F7BA57" box="[412,688,1069,1093]" italics="true" pageId="8" pageNumber="65">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB07566FD1BFBD21940BA54" bold="true" box="[694,775,1070,1094]" pageId="8" pageNumber="65">
|
||
<taxonomicNameLabel id="1A855799FFB07566FD1BFBD21940BA54" box="[694,775,1070,1094]" pageId="8" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in preservative. A. Dorsal surface; B. Ventral surface; C. Pattern of posterior surface of thighs; D. Pattern of throat surface; E. Vomerine teeth and choanae; F. Odontoid; G. Ventral surface of hands; H. Ventral surface of foot. Scale bars 5 mm. Photos by Manh V. Le.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7BD8657BFFB07563FF6AFB231824BFE7" lastPageId="13" lastPageNumber="70" pageId="8" pageNumber="65" type="description">
|
||
<paragraph id="337D36F0FFB07566FF6AFB23191CB8B8" blockId="8.[151,1437,1247,1958]" pageId="8" pageNumber="65">
|
||
<emphasis id="01B6EAE2FFB07566FF6AFB231BAABAE8" bold="true" box="[199,493,1247,1274]" pageId="8" pageNumber="65">
|
||
Description of
|
||
<typeStatus id="EC798852FFB07566FED2FB1C1BAFBAE8" box="[383,488,1248,1274]" pageId="8" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male, large body size, SVL
|
||
<quantity id="F43A9B15FFB07566FC26FB1C1E43BAE8" box="[907,1028,1248,1275]" metricMagnitude="-1" metricUnit="m" metricValue="1.054" pageId="8" pageNumber="65" unit="mm" value="105.4">105.4 mm</quantity>
|
||
; habitus robust with enlarged head (HL/SVL = 0.48); head length longer than wide (HW/HL = 0.91); snout round anteriorly in dorsal view, projecting beyond lower jaw; rostral length greater than eye diameter (RL
|
||
<quantity id="F43A9B15FFB07566FCD5FAD419A7BB50" box="[888,992,1320,1347]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="8" pageNumber="65" unit="mm" value="15.3">15.3 mm</quantity>
|
||
, ED
|
||
<quantity id="F43A9B15FFB07566FBB0FAD41E3EBB50" box="[1053,1145,1320,1346]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="65" unit="mm" value="8.8">8.8 mm</quantity>
|
||
); nostril lateral, elongate, ovoid, closer to tip of snout than to eye (NS
|
||
<quantity id="F43A9B15FFB07566FD04FAB01942BB74" box="[681,773,1356,1382]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="8" pageNumber="65" unit="mm" value="6.1">6.1 mm</quantity>
|
||
, EN
|
||
<quantity id="F43A9B15FFB07566FCEEFAB019E6BB74" box="[835,929,1356,1382]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="65" unit="mm" value="9.4">9.4 mm</quantity>
|
||
); interorbital distance wider than internarial distance and upper eyelid width (IOD
|
||
<quantity id="F43A9B15FFB07566FDE3FA8C18F2BB98" box="[590,693,1392,1418]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="65" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, IND
|
||
<quantity id="F43A9B15FFB07566FD51FA8C1911BB98" box="[764,854,1392,1418]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="65" unit="mm" value="8.2">8.2 mm</quantity>
|
||
, UEW
|
||
<quantity id="F43A9B15FFB07566FC06FA8C1E41BB98" box="[939,1030,1392,1418]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="65" unit="mm" value="6.4">6.4 mm</quantity>
|
||
); canthus rounded; loreals concave; eye diameter approximately equal tympanum diameter (ED/TD = 1.05), eye closer to nostril than tympanum (EN
|
||
<quantity id="F43A9B15FFB07566FF3AFA441AB4BBC0" box="[151,243,1464,1490]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="8" pageNumber="65" unit="mm" value="9.4">9.4 mm</quantity>
|
||
, TED
|
||
<quantity id="F43A9B15FFB07566FEEFFA441BEABBC0" box="[322,429,1464,1490]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="8" pageNumber="65" unit="mm" value="14.2">14.2 mm</quantity>
|
||
); tympanum distinct, slightly elongate, distance between tympanum to eye longer than eye diameter and tympanum diameter (TED
|
||
<quantity id="F43A9B15FFB07566FD3EFA2018BCBBE4" box="[659,763,1500,1526]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="8" pageNumber="65" unit="mm" value="14.2">14.2 mm</quantity>
|
||
, ED
|
||
<quantity id="F43A9B15FFB07566FC98FA2019C9BBE4" box="[821,910,1500,1526]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="8" pageNumber="65" unit="mm" value="8.8">8.8 mm</quantity>
|
||
, TD
|
||
<quantity id="F43A9B15FFB07566FC65FA201E64BBE4" box="[968,1059,1500,1526]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="65" unit="mm" value="8.4">8.4 mm</quantity>
|
||
). Hypertrophied jaw musculature (adductors) visible on dorsum; supratympanic fold robust, extending from eye to angle of jaw; vomerine teeth on oblique ridges, deflected towards the inside of the choanae, the distance between vomerine teeth rows shorter than distance from vomerine and the choanae. Choanae oval, perpendicular to longitudinal axis of body. Odontoid robust at base tapering to pointed tips, length almost twice depth of mandible. Symphysial knob at mandibular symphysis. Tongue notched deep posterior, about one-fourth of the tongue.
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB07566FF6AF94F1BB7B9B4" blockId="8.[151,1437,1247,1958]" pageId="8" pageNumber="65">
|
||
<emphasis id="01B6EAE2FFB07566FF6AF94F1B00B8DF" bold="true" box="[199,327,1715,1741]" pageId="8" pageNumber="65">Forelimbs.</emphasis>
|
||
Arms medium in length, upper arm length (UAL) 21.0 mm, forearm length (FAL)
|
||
<quantity id="F43A9B15FFB07566FB55F9481F27B8DC" box="[1272,1376,1716,1742]" metricMagnitude="-2" metricUnit="m" metricValue="4.36" pageId="8" pageNumber="65" unit="mm" value="43.6">43.6 mm</quantity>
|
||
. Tips of finger rounded not expanded into discs with fingers pad elevated; finger length relative III>I>II>IV, no webbing between fingers; fringe on sides of finger I absent, distinct between fingers II–III, indistinct between fingers III–IV; subarticular tubercles prominent, round, formula 1, 1, 1, 1 for finger I, II, III, IV respectively; thenar tubercle at base of finger I large (TTL
|
||
<quantity id="F43A9B15FFB07566FE22F8B81BAEB94C" box="[399,489,1860,1887]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="65" unit="mm" value="5.1">5.1 mm</quantity>
|
||
), oval, distinct elevated forward; inner metacarpal tubercle at base of finger III (IMC
|
||
<quantity id="F43A9B15FFB07566FF3AF8941AB6B990" box="[151,241,1896,1922]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="65" unit="mm" value="4.3">4.3 mm</quantity>
|
||
), slightly rounded; outer metacarpal tubercle small (OMT
|
||
<quantity id="F43A9B15FFB07566FCD3F894199FB990" box="[894,984,1896,1922]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="8" pageNumber="65" unit="mm" value="3.9">3.9 mm</quantity>
|
||
), at base of finger IV, elongated; distinct nuptial pad present on finger I.
|
||
</paragraph>
|
||
<caption id="67BD6678FFB17567FF3AFA081BEFB822" ID-DOI="http://doi.org/10.5281/zenodo.14734611" ID-Zenodo-Dep="14734611" httpUri="https://zenodo.org/record/14734611/files/figure.png" pageId="9" pageNumber="66" startId="9.[151,250,1524,1548]" targetBox="[151,1436,181,1501]" targetPageId="9" targetType="figure">
|
||
<paragraph id="337D36F0FFB17567FF3AFA081BEFB822" blockId="9.[151,1436,1524,1585]" pageId="9" pageNumber="66">
|
||
<emphasis id="01B6EAE2FFB17567FF3AFA081B53B81E" bold="true" box="[151,276,1524,1548]" pageId="9" pageNumber="66">FIGURE 4.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="F4C24D73FFB17567FE33FA0918F5B81F" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[414,690,1525,1549]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB17567FE33FA0918F5B81F" box="[414,690,1525,1549]" italics="true" pageId="9" pageNumber="66">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB17567FD14FA09194CB81F" bold="true" box="[697,779,1525,1549]" pageId="9" pageNumber="66">
|
||
<taxonomicNameLabel id="1A855799FFB17567FD14FA09194CB81F" box="[697,779,1525,1549]" pageId="9" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life. A. Dorsal surface; B. Ventral surface; C. Nuptial pad on finger I; scale bars 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="337D36F0FFB17567FF6AF97F1BF3B98B" blockId="9.[151,1437,1667,1945]" pageId="9" pageNumber="66">
|
||
<emphasis id="01B6EAE2FFB17567FF6AF97F1B0BB88F" bold="true" box="[199,332,1667,1693]" pageId="9" pageNumber="66">Hindlimbs.</emphasis>
|
||
Robust and long (total length
|
||
<quantity id="F43A9B15FFB17567FD36F97F1957B88F" box="[667,784,1667,1693]" metricMagnitude="-1" metricUnit="m" metricValue="1.814" pageId="9" pageNumber="66" unit="mm" value="181.4">181.4 mm</quantity>
|
||
). Tibia length longer than thigh length (FeL
|
||
<quantity id="F43A9B15FFB17567FB55F97F1F19B88F" box="[1272,1374,1667,1694]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="9" pageNumber="66" unit="mm" value="52.6">52.6 mm</quantity>
|
||
; TBL
|
||
<quantity id="F43A9B15FFB17567FF3AF95B1ABDB8D3" box="[151,250,1703,1730]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" pageId="9" pageNumber="66" unit="mm" value="55.5">55.5 mm</quantity>
|
||
) and approximately 2.9 times of tibia wide. Tips of toes rounded not expanded into discs, no circummarginal groove, toe pads elevated; relative toe lengths IV>III>V>II>I; toes webbed to distal of terminal phalanx, formula I0-0II0-0III0-0IV0-0V; movable flap of skin distinct on postaxial side toe V from base of toe pad to the metatarsus; movable flap of skin on preaxial side of toe I from base of toe pad to the inner metatarsal tubercle continuing as distinct fold on distal half of tarsus; subarticular tubercles prominent, oval, elevated, formula 1, 1, 2, 3, 2 for toe I, II, III, IV, V, respectively; inner metatarsal tubercle elongated (IMT
|
||
<quantity id="F43A9B15FFB17567FC1EF8A71E4CB967" box="[947,1035,1883,1909]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="66" unit="mm" value="6.2">6.2 mm</quantity>
|
||
) with elevated postaxial border; no outer metatarsal tubercle.
|
||
</paragraph>
|
||
<caption id="67BD6678FFB27564FF3AFA6B1898B80E" ID-DOI="http://doi.org/10.5281/zenodo.14734613" ID-Zenodo-Dep="14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="10" pageNumber="67" startId="10.[151,250,1431,1455]" targetBox="[151,1436,181,1407]" targetPageId="10" targetType="figure">
|
||
<paragraph id="337D36F0FFB27564FF3AFA6B1898B80E" blockId="10.[151,1437,1431,1564]" pageId="10" pageNumber="67">
|
||
<emphasis id="01B6EAE2FFB27564FF3AFA6B1B55BBBD" bold="true" box="[151,274,1431,1455]" pageId="10" pageNumber="67">FIGURE 5.</emphasis>
|
||
Color variations in life of
|
||
<taxonomicName id="F4C24D73FFB27564FDB0FA641977BBBD" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[541,816,1431,1455]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB27564FDB0FA641977BBBD" box="[541,816,1431,1455]" italics="true" pageId="10" pageNumber="67">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB27564FC98FA6B19C2BBBD" bold="true" box="[821,901,1431,1455]" pageId="10" pageNumber="67">
|
||
<taxonomicNameLabel id="1A855799FFB27564FC98FA6B19C2BBBD" box="[821,901,1431,1455]" pageId="10" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="337D36F0FFB27564FF6AF99119EDB905" blockId="10.[151,1437,1645,2032]" pageId="10" pageNumber="67">
|
||
<emphasis id="01B6EAE2FFB27564FF6AF9911B42B895" bold="true" box="[199,261,1645,1671]" pageId="10" pageNumber="67">Skin.</emphasis>
|
||
Body smooth in general; dorsal surface of head almost smooth, upper of eyelid with some tubercles; supratympanic fold distinct, extending from eye to angle of jaw; dorsolateral fold absent; dorsal surface of flanks rough scabrous with larger tubercles; dorsal surface of forelimbs smooth; dorsal surface hindlimbs smooth with distinctly small tubercles on half of shank and foot region; throat, chest, belly and ventral surface of thighs smooth, numerous white distinctly small tubercles focus on pre-cloacal region.
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB27565FF6AF8DD1EEEBF9B" blockId="10.[151,1437,1645,2032]" lastBlockId="11.[151,1437,151,537]" lastPageId="11" lastPageNumber="68" pageId="10" pageNumber="67">
|
||
<emphasis id="01B6EAE2FFB27564FF6AF8DD1BE4B929" bold="true" box="[199,419,1825,1851]" pageId="10" pageNumber="67">Coloration in life.</emphasis>
|
||
Dorsal surface dark reddish brown with black and interrupted mottling patterns; pupil rounded, black, and reflects orangy-red under flash photography (
|
||
<figureCitation id="ABF92A75FFB27564FC2CF8B9199BB972" box="[897,988,1861,1888]" captionStart="FIGURE 3" captionStartId="8.[151,250,1069,1093]" captionTargetBox="[158,1427,427,1035]" captionTargetId="figure-127@8.[151,1436,417,1046]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Limnonectes splendissimus sp. nov. in preservative. A. Dorsal surface; B. Ventral surface; C. Pattern of posterior surface of thighs; D. Pattern of throat surface; E. Vomerine teeth and choanae; F. Odontoid; G. Ventral surface of hands; H. Ventral surface of foot. Scale bars 5 mm. Photos by Manh V. Le." figureDoi="http://doi.org/10.5281/zenodo.14734609" httpUri="https://zenodo.org/record/14734609/files/figure.png" pageId="10" pageNumber="67">Fig. 3A</figureCitation>
|
||
); light orange interorbital bar, bordered posteriorly by black; iris with mixed between light orangish red and light greyish spots; tympanums particolored with upper black and lower light reddish brown; rostrum reddish brown; upper lip with bars that extend all the way to behind the eye and the orangy-red section is just beneath the tympanum area; lower lip with black and white barring and orange spots; flank surfaces reddish brown with some black spots, orangish red tubercles, and two yellow spots near the armpit and groin; dorsal and anterior surfaces of limbs reddish brown and orangish red, and interrupted with fainted black crossbars; dorsal surface of fingers and toes orangish red with black spots; posterior surface of forelimbs black with light spots; posterior surfaces of thighs pattern mixed between yellow or light yellow with black spots; throat surface pattern grey and white, outer margin mixed with orange, white and black barring, and black spots near base on forelimbs; middle part of ventral surface white, light golden on latter half, with yellow outer margins; ventral surface of forelimbs white; ventral surface of hindlimbs reddish brown or brown with some grey lines, and outer margin yellow; ventral surface of forefoots and hindfoots black (
|
||
<figureCitation id="ABF92A75FFB37565FBFAFE931EDCBF9B" box="[1111,1179,367,393]" captionStart="FIGURE 3" captionStartId="8.[151,250,1069,1093]" captionTargetBox="[158,1427,427,1035]" captionTargetId="figure-127@8.[151,1436,417,1046]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Limnonectes splendissimus sp. nov. in preservative. A. Dorsal surface; B. Ventral surface; C. Pattern of posterior surface of thighs; D. Pattern of throat surface; E. Vomerine teeth and choanae; F. Odontoid; G. Ventral surface of hands; H. Ventral surface of foot. Scale bars 5 mm. Photos by Manh V. Le." figureDoi="http://doi.org/10.5281/zenodo.14734609" httpUri="https://zenodo.org/record/14734609/files/figure.png" pageId="11" pageNumber="68">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB37565FF6AFE6E1809BC0B" blockId="11.[151,1437,151,537]" pageId="11" pageNumber="68">
|
||
<emphasis id="01B6EAE2FFB37565FF6AFE6E184DBFBE" bold="true" box="[199,522,402,428]" pageId="11" pageNumber="68">Coloration in preservation.</emphasis>
|
||
Dorsum, rostrum, and limbs surfaces faded to brown; surfaces of post-dorsal part, shank, and foot with white small tubercles remain distinct; posterior surfaces of thighs faded to dark grey and some bright brindles; throat surface pattern faded to grey and cream, outer margin faded to cream with black barring; ventral surface of limbs cream (
|
||
<figureCitation id="ABF92A75FFB37565FE56FE031807BC0B" box="[507,576,511,537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-17@9.[151,1436,181,1501]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Limnonectes splendissimus sp. nov. in life. A. Dorsal surface; B. Ventral surface; C. Nuptial pad on finger I; scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.14734611" httpUri="https://zenodo.org/record/14734611/files/figure.png" pageId="11" pageNumber="68">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="67BD6678FFB37565FF3AF9071988B94E" ID-DOI="http://doi.org/10.5281/zenodo.14734615" ID-Zenodo-Dep="14734615" httpUri="https://zenodo.org/record/14734615/files/figure.png" pageId="11" pageNumber="68" startId="11.[151,250,1787,1811]" targetBox="[158,1430,571,1751]" targetPageId="11" targetType="figure">
|
||
<paragraph id="337D36F0FFB37565FF3AF9071988B94E" blockId="11.[151,1437,1787,1884]" pageId="11" pageNumber="68">
|
||
<emphasis id="01B6EAE2FFB37565FF3AF9071B51B901" bold="true" box="[151,278,1787,1811]" pageId="11" pageNumber="68">FIGURE 6.</emphasis>
|
||
Color variation of paratypes in preservative of
|
||
<taxonomicName id="F4C24D73FFB37565FCBCF9001E6FB906" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[785,1064,1788,1812]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB37565FCBCF9001E6FB906" box="[785,1064,1788,1812]" italics="true" pageId="11" pageNumber="68">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB37565FB9CF9001EC2B906" bold="true" box="[1073,1157,1788,1812]" pageId="11" pageNumber="68">
|
||
<taxonomicNameLabel id="1A855799FFB37565FB9CF9001EC2B906" box="[1073,1157,1788,1812]" pageId="11" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8678, adult female; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 8677, subadult male; L&M. ITBCZ 3626, subadult male; N&O. ITBCZ 2627, juvenile.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="67BD6678FFB47562FF3AFAE71F1FB8F6" ID-DOI="http://doi.org/10.5281/zenodo.14734617" ID-Zenodo-Dep="14734617" httpUri="https://zenodo.org/record/14734617/files/figure.png" pageId="12" pageNumber="69" startId="12.[151,250,1307,1331]" targetBox="[151,1436,181,1284]" targetPageId="12" targetType="figure">
|
||
<paragraph id="337D36F0FFB47562FF3AFAE71F1FB8F6" blockId="12.[151,1437,1307,1764]" pageId="12" pageNumber="69">
|
||
<emphasis id="01B6EAE2FFB47562FF3AFAE71B54BB26" bold="true" box="[151,275,1307,1332]" pageId="12" pageNumber="69">FIGURE 7.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="F4C24D73FFB47562FE48FAE018F4BB26" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[485,691,1308,1332]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="poilani">
|
||
<emphasis id="01B6EAE2FFB47562FE48FAE018F4BB26" box="[485,691,1308,1332]" italics="true" pageId="12" pageNumber="69">Limnonectes poilani</emphasis>
|
||
</taxonomicName>
|
||
group, emphasizing the necessity of further study on those populations of
|
||
<taxonomicName id="F4C24D73FFB47562FF3AFABC1B5CBB4A" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[151,283,1344,1368]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" isUncertain="true" kingdom="Animalia" order="Anura" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="poilani">
|
||
<emphasis id="01B6EAE2FFB47562FF3AFABC1AEBBB45" box="[151,172,1344,1367]" italics="true" pageId="12" pageNumber="69">L.</emphasis>
|
||
cf.
|
||
<emphasis id="01B6EAE2FFB47562FF7EFABC1B5CBB4A" box="[211,283,1344,1368]" italics="true" pageId="12" pageNumber="69">poilani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F4C24D73FFB47562FEE0FABC1851BB4A" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[333,534,1344,1368]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" isUncertain="true" kingdom="Animalia" order="Anura" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="splendissimus">
|
||
<emphasis id="01B6EAE2FFB47562FEE0FABC1B25BB45" box="[333,354,1344,1367]" italics="true" pageId="12" pageNumber="69">L.</emphasis>
|
||
cf.
|
||
<emphasis id="01B6EAE2FFB47562FE24FABC1851BB4A" box="[393,534,1344,1368]" italics="true" pageId="12" pageNumber="69">splendissimus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="01B6EAE2FFB47562FD8EFAC31838BB45" bold="true" box="[547,639,1343,1367]" pageId="12" pageNumber="69">Vietnam</emphasis>
|
||
:
|
||
<emphasis id="01B6EAE2FFB47562FD20FAC318E7BB45" bold="true" box="[653,672,1343,1367]" pageId="12" pageNumber="69">1.</emphasis>
|
||
Minh Hoa, Quang Binh Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FC4DFABC1EEDBB4A" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[992,1194,1344,1368]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FB94FABC1E2ABB4A" box="[1081,1133,1344,1368]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FB11FAC31E88BB45" bold="true" box="[1212,1231,1343,1367]" pageId="12" pageNumber="69">2.</emphasis>
|
||
Huong Hoa, Quang Tri Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FEADFA981BCBBB6E" author="Bourret, R." box="[256,396,1380,1404]" pageId="12" pageNumber="69" refId="ref12571" refString="Bourret, R. (1942) Les batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi, 547 pp." type="book" year="1942">Bourret 1942</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="57534B01FFB47562FE3AFA981827BB6E" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[407,608,1380,1404]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FE42FA981863BB6E" box="[495,548,1380,1404]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
; this study);
|
||
<emphasis id="01B6EAE2FFB47562FD4BFA9F18BFBB69" bold="true" box="[742,760,1379,1403]" pageId="12" pageNumber="69">3.</emphasis>
|
||
A Luoi, Thua Thien Hue Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FBEAFA981F55BB6E" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[1095,1298,1380,1404]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FB0DFA981E92BB6E" box="[1184,1237,1380,1404]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FA88FA9F1F7FBB69" bold="true" box="[1317,1336,1379,1403]" pageId="12" pageNumber="69">4.</emphasis>
|
||
Phu Loc, Thua Thien Hue Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FE2EFA74180CBBB2" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[387,587,1416,1440]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FE76FA741849BBB2" box="[475,526,1416,1440]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FDF1FA741829BBB2" bold="true" box="[604,622,1416,1440]" pageId="12" pageNumber="69">5.</emphasis>
|
||
Son Tra, Da Nang City (this study);
|
||
<emphasis id="01B6EAE2FFB47562FC4FFA7B19B3BB8D" bold="true" box="[994,1012,1415,1439]" pageId="12" pageNumber="69">6.</emphasis>
|
||
Tay Giang, Quang Nam Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FA9FFA741AA9BBD6" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FA27FA741AF5BBD6" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FF52FA501B55BBD6" bold="true" box="[255,274,1452,1476]" pageId="12" pageNumber="69">7.</emphasis>
|
||
Nam Giang, Quang Nam Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FDF6FA501966BBD6" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[603,801,1452,1476]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FD1FFA5018A2BBD6" box="[690,741,1452,1476]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FC9FFA571902BBD1" bold="true" box="[818,837,1451,1475]" pageId="12" pageNumber="69">8.</emphasis>
|
||
Nam Tra My, Quang Nam Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FB34FA501F05BBD6" author="Frost, D. R. & Grant, T. & Faivovich, J. & Bain, R. & Haas, A. & Haddad, C. F. B. & de Sa, R. O. & Channing, A. & Wilkinson, M. & Donnellan, S. C. & Raxworthy, C. & Campbell, J. A. & Blotto, B. L. & Moler, P. & Drewes, R. C. & Nussbaum, R. A. & Lynch, J. D. & Green, D. M. & Wheeler, W. C." box="[1177,1346,1452,1476]" pageId="12" pageNumber="69" pagination="1 - 291" refId="ref13418" refString="Frost, D. R., Grant, T., Faivovich, J., Bain, R., Haas, A., Haddad, C. F. B., de Sa, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. & Wheeler, W. C. (2006) The Amphibian Tree of Life. Bulletin of the American Museum of Natural History, 297, 1 - 291. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 297 [0001: TATOL] 2.0. CO; 2" type="journal article" year="2006">
|
||
Frost
|
||
<emphasis id="01B6EAE2FFB47562FB78FA501F40BBD6" box="[1237,1287,1452,1476]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="57534B01FFB47562FAE1FA501B4CBBFA" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FF3AFA2C1A8BBBFA" box="[151,204,1488,1512]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FEB3FA331B76BBF5" bold="true" box="[286,305,1487,1511]" pageId="12" pageNumber="69">9.</emphasis>
|
||
Son Ha, Quang Ngai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FDF3FA2C194DBBFA" author="Tran, D. T. A. & Le, Q. K. & Le, K. V. & Vu, T. N. & Nguyen, T. Q. & Bohme, W. & Ziegler, T." box="[606,778,1488,1512]" pageId="12" pageNumber="69" pagination="111 - 119" refId="ref16828" refString="Tran, D. T. A., Le, Q. K., Le, K. V., Vu, T. N., Nguyen, T. Q., Bohme, W. & Ziegler, T. (2010) First and preliminary frog records (Amphibia: Anura) from Quang Ngai Province, Vietnam. Herpetology Notes, 3, 111 - 119." type="journal article" year="2010">
|
||
Tran
|
||
<emphasis id="01B6EAE2FFB47562FD3AFA2C188BBBFA" box="[663,716,1488,1512]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FCB3FA33197ABBF5" bold="true" box="[798,829,1487,1511]" pageId="12" pageNumber="69">10.</emphasis>
|
||
Ngoc Linh, Kon Tum Prov. (Orlov 2004);
|
||
<emphasis id="01B6EAE2FFB47562FAADFA331F58BBF5" bold="true" box="[1280,1311,1487,1511]" pageId="12" pageNumber="69">11.</emphasis>
|
||
Kon Plong, Kon Tum Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FE92FA081840B81E" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[319,519,1524,1548]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FE3AFA081B8DB81E" box="[407,458,1524,1548]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FDB5FA0F187FB819" bold="true" box="[536,568,1523,1547]" pageId="12" pageNumber="69">12.</emphasis>
|
||
Chu Mom Ray, Kon Tum (
|
||
<bibRefCitation id="57534B01FFB47562FCE1FA081E7DB81E" author="Jestrzemski, D. & Schutz, S. & Nguyen, T. Q. & Ziegler, T." box="[844,1082,1524,1548]" pageId="12" pageNumber="69" pagination="88 - 110" refId="ref14185" refString="Jestrzemski, D., Schutz, S., Nguyen, T. Q. & Ziegler, T. (2013) A survey of amphibians and reptiles in Chu Mom Ray National Park, Vietnam, with implications for herpetofaunal conservation. Asian Journal of Conservation Biology, 2, 88 - 110." type="journal article" year="2013">
|
||
Jestrzemski
|
||
<emphasis id="01B6EAE2FFB47562FC67FA0819BAB81E" box="[970,1021,1524,1548]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FBE6FA0F1E2CB819" bold="true" box="[1099,1131,1523,1547]" pageId="12" pageNumber="69">13.</emphasis>
|
||
Kon Chu Rang, Gia Lai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FF33F9E41B20B822" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[158,359,1560,1584]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FF5BF9E41B6DB822" box="[246,298,1560,1584]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FED5F9EB1BDFB83D" bold="true" box="[376,408,1559,1583]" pageId="12" pageNumber="69">14.</emphasis>
|
||
Tram Lap, Gia Lai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FD08F9E41914B83D" author="Chen, L. Q. & Murphy, R. W. & Lathrop, A. & Ngo, A. & Orlov, N. L. & Ho, C. T. & Somorjai, I. L. M." box="[677,851,1559,1584]" pageId="12" pageNumber="69" pagination="231 - 243" refId="ref12728" refString="Chen, L. Q., Murphy, R. W., Lathrop, A., Ngo, A., Orlov, N. L., Ho, C. T. & Somorjai, I. L. M. (2005) Taxonomy chaos in Asian ranid frogs: an initial phylogenetic resolution. Herpetological Journal, 15, 231 - 243." type="journal article" year="2005">
|
||
Chen
|
||
<emphasis id="01B6EAE2FFB47562FD4FF9E41951B822" box="[738,790,1560,1584]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FCC9F9EB19C3B83D" bold="true" box="[868,900,1559,1583]" pageId="12" pageNumber="69">15.</emphasis>
|
||
K’Bang, Gia Lai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FBD0F9E41F02B822" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[1149,1349,1560,1584]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FB78F9E41F4EB822" box="[1237,1289,1560,1584]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FAFAF9EB1F30B83D" bold="true" box="[1367,1399,1559,1583]" pageId="12" pageNumber="69">16.</emphasis>
|
||
An Khe, Gia Lai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FEC8F9C01812B846" author="Emerson, S. B. & Ward, R." box="[357,597,1596,1620]" pageId="12" pageNumber="69" pagination="537 - 553" refId="ref12984" refString="Emerson, S. B. & Ward, R. (1998) Male secondary sexual characteristics, sexual selection, and molecular divergence in fanged ranid frogs of Southeast Asia. Zoological Journal of the Linnean Society, 122, 537 - 553. https: // doi. org / 10.1006 / zjls. 1996.0099" type="journal article" year="1998">Emerson & Ward 1998</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FDC5F9C718CFB841" bold="true" box="[616,648,1595,1619]" pageId="12" pageNumber="69">17.</emphasis>
|
||
Krong Pa, Gia Lai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FC39F9C01E19B846" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[916,1118,1596,1620]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FC40F9C01E66B846" box="[1005,1057,1596,1620]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FBDDF9C71ED7B841" bold="true" box="[1136,1168,1595,1619]" pageId="12" pageNumber="69">18.</emphasis>
|
||
York Don, Dak Lak Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FF33F99C1B20B86A" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[158,359,1632,1656]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FF5AF99C1B6DB86A" box="[247,298,1632,1656]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FED4F9A31BDEB865" bold="true" box="[377,409,1631,1655]" pageId="12" pageNumber="69">19.</emphasis>
|
||
Chu Yang Sin, Dak Lak Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FD77F99C19E3B86A" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[730,932,1632,1656]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FC9EF99C1920B86A" box="[819,871,1632,1656]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FC18F9A31992B865" bold="true" box="[949,981,1631,1655]" pageId="12" pageNumber="69">20.</emphasis>
|
||
Lac Duong, Lam Dong Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FABCF99C1A89B88E" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FAC4F99C1FDAB86A" box="[1385,1437,1632,1656]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FF4CF97F1B46B889" bold="true" box="[225,257,1667,1691]" pageId="12" pageNumber="69">21.</emphasis>
|
||
Duc Trong, Lam Dong Prov. (this study);
|
||
<emphasis id="01B6EAE2FFB47562FD14F97F189FB889" bold="true" box="[697,728,1667,1691]" pageId="12" pageNumber="69">22.</emphasis>
|
||
Di Linh, Lam Dong Prov. (this study);
|
||
<emphasis id="01B6EAE2FFB47562FBDEF97F1ED4B889" bold="true" box="[1139,1171,1667,1691]" pageId="12" pageNumber="69">23.</emphasis>
|
||
Da Teh, Lam Dong Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FF33F9541B20B8D2" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[158,359,1704,1728]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FF5BF9541B6DB8D2" box="[246,298,1704,1728]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FED5F95B1BDFB8AD" bold="true" box="[376,408,1703,1727]" pageId="12" pageNumber="69">24.</emphasis>
|
||
Cat Tien, Dong Nai Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FD03F9541930B8D2" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[686,887,1704,1728]" pageId="12" pageNumber="69" refId="ref15218" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="01B6EAE2FFB47562FCAAF954197DB8D2" box="[775,826,1704,1728]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
<emphasis id="01B6EAE2FFB47562FC25F95B19BEB8AD" bold="true" box="[904,1017,1703,1727]" pageId="12" pageNumber="69">Cambodia</emphasis>
|
||
:
|
||
<emphasis id="01B6EAE2FFB47562FBABF95B1E61B8AD" bold="true" box="[1030,1062,1703,1727]" pageId="12" pageNumber="69">25.</emphasis>
|
||
O’Rang, Mondulkiri Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FAE8F9541AA8B8F6" author="Stuart, B. L. & Sok, K. & Neang, T." pageId="12" pageNumber="69" pagination="129 - 155" refId="ref16574" refString="Stuart, B. L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology. Singapore, 54, 129 - 155." type="journal article" year="2006">
|
||
Stuart
|
||
<emphasis id="01B6EAE2FFB47562FA27F9541AF5B8F6" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="01B6EAE2FFB47562FEADF9371B67B8F1" bold="true" box="[256,288,1739,1763]" pageId="12" pageNumber="69">26.</emphasis>
|
||
Pichrada, Mondulkiri Prov (
|
||
<bibRefCitation id="57534B01FFB47562FDEFF93018B0B8F6" author="Stuart, B. L. & Sok, K. & Neang, T." box="[578,759,1740,1764]" pageId="12" pageNumber="69" pagination="129 - 155" refId="ref16574" refString="Stuart, B. L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology. Singapore, 54, 129 - 155." type="journal article" year="2006">
|
||
Stuart
|
||
<emphasis id="01B6EAE2FFB47562FD2AF93018FDB8F6" box="[647,698,1740,1764]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
).
|
||
<emphasis id="01B6EAE2FFB47562FCAAF930197CB8F6" bold="true" box="[775,827,1740,1764]" pageId="12" pageNumber="69">Laos</emphasis>
|
||
:
|
||
<emphasis id="01B6EAE2FFB47562FCE4F937192EB8F1" bold="true" box="[841,873,1739,1763]" pageId="12" pageNumber="69">27.</emphasis>
|
||
Dakcheung, Xekong Prov. (
|
||
<bibRefCitation id="57534B01FFB47562FB2AF9301F0BB8F6" author="Brakels, P. & Maury, N. & Nguyen, T. V." box="[1159,1356,1740,1764]" pageId="12" pageNumber="69" refId="ref12593" refString="Brakels, P., Maury, N. & Nguyen, T. V. (2021) Geographic distribution: Limnonectes poilani. Herpetological Review, 52, 792." type="journal volume" year="2021">
|
||
Brakels
|
||
<emphasis id="01B6EAE2FFB47562FB71F9301F57B8F6" box="[1244,1296,1740,1764]" italics="true" pageId="12" pageNumber="69">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="337D36F0FFB47563FF6AF8CA18D1BF0F" blockId="12.[151,1437,1846,2016]" lastBlockId="13.[151,1436,150,825]" lastPageId="13" lastPageNumber="70" pageId="12" pageNumber="69">
|
||
<emphasis id="01B6EAE2FFB47562FF6AF8CA1B78B942" bold="true" box="[199,319,1846,1872]" pageId="12" pageNumber="69">Variation.</emphasis>
|
||
Variations in measurements and morphological features of the
|
||
<typeStatus id="EC798852FFB47562FBBCF8CA1E06B942" box="[1041,1089,1846,1872]" pageId="12" pageNumber="69">type</typeStatus>
|
||
series are provided in
|
||
<tableCitation id="7E40034BFFB47562FAE5F8CA1FDBB943" box="[1352,1436,1846,1873]" captionStart="TABLE 5" captionStartId="16.[151,239,962,986]" captionTargetPageId="16" captionText="TABLE 5. Morphological characters of thetype series of Limnonectes splendissimus sp. nov., measurements in millimeters (abbreviations see Material and methods)." httpUri="http://table.plazi.org/id/67BD6678FFA8757EFF3AFC3E181FBDEC" pageId="12" pageNumber="69" tableUuid="67BD6678FFA8757EFF3AFC3E181FBDEC">Table 5</tableCitation>
|
||
and color variations in life and preservation follow in
|
||
<figureCitation id="ABF92A75FFB47562FCA8F8A61914B967" box="[773,851,1882,1909]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="12" pageNumber="69">Figs. 5</figureCitation>
|
||
&
|
||
<figureCitation id="ABF92A75FFB47562FCD6F8A619CBB966" box="[891,908,1882,1908]" captionStart="FIGURE 6" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[158,1430,571,1751]" captionTargetId="figure-200@11.[151,1436,561,1764]" captionTargetPageId="11" captionText="FIGURE 6. Color variation of paratypes in preservative of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8678, adult female; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 8677, subadult male; L&M. ITBCZ 3626, subadult male; N&O. ITBCZ 2627, juvenile." figureDoi="http://doi.org/10.5281/zenodo.14734615" httpUri="https://zenodo.org/record/14734615/files/figure.png" pageId="12" pageNumber="69">6</figureCitation>
|
||
. Body color in life varies between individuals within the
|
||
<typeStatus id="EC798852FFB47562FEBDF8821B07B98A" box="[272,320,1918,1944]" pageId="12" pageNumber="69">type</typeStatus>
|
||
series (
|
||
<figureCitation id="ABF92A75FFB47562FE3AF88218CAB98A" box="[407,653,1918,1945]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="12" pageNumber="69">Fig. 5A, C, E, G, I, L</figureCitation>
|
||
). Juvenile ITBCZ 2627 has a very distinct belly with black and white pattern that covers both the throat and half of ventral, a half ventral latter yellow (
|
||
<figureCitation id="ABF92A75FFB47562FB8EF85E1EE6B9AE" box="[1059,1185,1954,1981]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="12" pageNumber="69">Fig. 5L, M</figureCitation>
|
||
). In subadult male and female ITBCZ 8677 and ITBCZ 3626, throat surface pattern faded to grey and white, while the ventral surface is generally yellow (
|
||
<figureCitation id="ABF92A75FFB57563FECEFF6B1B9EBEA3" box="[355,473,150,177]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="13" pageNumber="70">Fig. 5F, K</figureCitation>
|
||
). In adult males and female ITBCZ 8675, 8676 and 8679, the ventral surfaces change from yellow to ochre, yellowish-brown or beige (
|
||
<figureCitation id="ABF92A75FFB57563FD16FF471911BEC7" box="[699,854,186,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="13" pageNumber="70">Fig. 5B, D, H</figureCitation>
|
||
). Body color in preservation changed from brown to grey (
|
||
<figureCitation id="ABF92A75FFB57563FF7AFF231BA1BEEB" box="[215,486,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[158,1430,571,1751]" captionTargetId="figure-200@11.[151,1436,561,1764]" captionTargetPageId="11" captionText="FIGURE 6. Color variation of paratypes in preservative of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8678, adult female; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 8677, subadult male; L&M. ITBCZ 3626, subadult male; N&O. ITBCZ 2627, juvenile." figureDoi="http://doi.org/10.5281/zenodo.14734615" httpUri="https://zenodo.org/record/14734615/files/figure.png" pageId="13" pageNumber="70">Fig. 6A, C, E, G, I, L, N</figureCitation>
|
||
). Throat surface pattern in juvenile and subadults is distinct (
|
||
<figureCitation id="ABF92A75FFB57563FB25FF231F61BEEB" box="[1160,1318,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[158,1430,571,1751]" captionTargetId="figure-200@11.[151,1436,561,1764]" captionTargetPageId="11" captionText="FIGURE 6. Color variation of paratypes in preservative of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8678, adult female; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 8677, subadult male; L&M. ITBCZ 3626, subadult male; N&O. ITBCZ 2627, juvenile." figureDoi="http://doi.org/10.5281/zenodo.14734615" httpUri="https://zenodo.org/record/14734615/files/figure.png" pageId="13" pageNumber="70">Fig. 6K, M, O</figureCitation>
|
||
) but faded in adult males and females (
|
||
<figureCitation id="ABF92A75FFB57563FE7CFEFF18CDBF0F" box="[465,650,259,285]" captionStart="FIGURE 6" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[158,1430,571,1751]" captionTargetId="figure-200@11.[151,1436,561,1764]" captionTargetPageId="11" captionText="FIGURE 6. Color variation of paratypes in preservative of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8678, adult female; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 8677, subadult male; L&M. ITBCZ 3626, subadult male; N&O. ITBCZ 2627, juvenile." figureDoi="http://doi.org/10.5281/zenodo.14734615" httpUri="https://zenodo.org/record/14734615/files/figure.png" pageId="13" pageNumber="70">Fig. 6B, D, F, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB57563FF6AFEDA1824BFE7" blockId="13.[151,1436,150,825]" pageId="13" pageNumber="70">
|
||
<emphasis id="01B6EAE2FFB57563FF6AFEDA1BF1BF52" bold="true" box="[199,438,294,320]" pageId="13" pageNumber="70">Sexual dimorphism.</emphasis>
|
||
Adult males have larger body sizes than females (SVL 104.1–107.0 mm in adult males vs.
|
||
<quantity id="F43A9B15FFB57563FF3AFEB71B7EBF77" box="[151,313,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="9.245" metricValueMax="9.77" metricValueMin="8.72" pageId="13" pageNumber="70" unit="mm" value="92.45" valueMax="97.7" valueMin="87.2">87.2–97.7 mm</quantity>
|
||
in adult females); an increased head height; a nuptial pad on finger I present (vs. absent); tympanum-eye distance greater than the eye diameter in adult males and shorter in females (ED/TED = 0.62±
|
||
<quantity id="F43A9B15FFB57563FAA6FE931F1CBF9B" box="[1291,1371,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.302" pageId="13" pageNumber="70" unit="in" value="0.13">0.13 in</quantity>
|
||
adult males vs. 1.40±
|
||
<quantity id="F43A9B15FFB57563FEEAFE6F1BE9BFBF" box="[327,430,403,429]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="70" unit="mm" value="0.02">0.02 mm</quantity>
|
||
in adult females); odontoid height tends to be more prominent (OH 8.0±
|
||
<quantity id="F43A9B15FFB57563FB45FE6F1F78BFBF" box="[1256,1343,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="70" unit="mm" value="0.6">0.6 mm</quantity>
|
||
in adult males vs. 4.5±
|
||
<quantity id="F43A9B15FFB57563FE92FE4A1BDFBFC3" box="[319,408,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="70" unit="mm" value="0.9">0.9 mm</quantity>
|
||
in adult females). Juveniles, subadults, and adults represent differences in throat surface pattern and ventral surface color (
|
||
<figureCitation id="ABF92A75FFB57563FDBCFE271812BFE7" box="[529,597,474,501]" captionStart="FIGURE 5" captionStartId="10.[151,250,1431,1455]" captionTargetBox="[151,1436,181,1407]" captionTargetId="figure-23@10.[151,1436,181,1407]" captionTargetPageId="10" captionText="FIGURE 5. Color variations in life of Limnonectes splendissimus sp. nov. A&B. ITBCZ 8675, adult male; C&D. ITBCZ 8676, adult male; E&F. ITBCZ 8677, subadult male; G&H. ITBCZ 8679, adult female; I&K. ITBCZ 3626, subadult male; L&M. ITBCZ 2627, juvenile; N–P. juvenile, no voucher; Q. Subadult, no voucher, in-situ photo. A–H photos by Manh V. Le; I–P photos by Luan T. Nguyen; Q photo by Tuan M. Nguyen." figureDoi="http://doi.org/10.5281/zenodo.14734613" httpUri="https://zenodo.org/record/14734613/files/figure.png" pageId="13" pageNumber="70">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BD8657BFFB57563FF6AFE031889BC73" pageId="13" pageNumber="70" type="etymology">
|
||
<paragraph id="337D36F0FFB57563FF6AFE031889BC73" blockId="13.[151,1436,150,825]" pageId="13" pageNumber="70">
|
||
<emphasis id="01B6EAE2FFB57563FF6AFE031B09BC0B" bold="true" box="[199,334,511,537]" pageId="13" pageNumber="70">Etymology.</emphasis>
|
||
Name from Latin
|
||
<taxonomicName id="F4C24D73FFB57563FD81FE03189FBC0B" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[556,728,511,537]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="splendissimus">“splendissimus</taxonomicName>
|
||
” refers to the splendid coloration on the underside of young individuals. We recommend “Splendid Fanged Frog” and “Ếch nhẽo đốm” as the common English name and Vietnamese name of the new species, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BD8657BFFB57563FF6AFD971B58BD2B" pageId="13" pageNumber="70" type="biology_ecology">
|
||
<paragraph id="337D36F0FFB57563FF6AFD971B58BD2B" blockId="13.[151,1436,150,825]" pageId="13" pageNumber="70">
|
||
<emphasis id="01B6EAE2FFB57563FF6AFD971825BC96" bold="true" box="[199,610,618,645]" pageId="13" pageNumber="70">Natural history and Distribution.</emphasis>
|
||
The new species is currently known from type locality in
|
||
<collectingRegion id="F106F812FFB57563FAB2FD971ABABCBB" country="Vietnam" name="Lam Dong" pageId="13" pageNumber="70">Lam Dong Province</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="4BD57660FFB57563FEC0FD731B8BBCBB" box="[365,460,655,681]" name="Vietnam" pageId="13" pageNumber="70">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="ABF92A75FFB57563FE74FD73185BBCBB" box="[473,540,655,681]" captionStart="FIGURE 7" captionStartId="12.[151,250,1307,1331]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-23@12.[151,1436,181,1284]" captionTargetPageId="12" captionText="FIGURE 7. Distribution map of Limnonectes poilani group, emphasizing the necessity of further study on those populations of L. cf. poilani and L. cf. splendissimus. Vietnam: 1. Minh Hoa, Quang Binh Prov. (Nguyen et al. 2009); 2. Huong Hoa, Quang Tri Prov. (Bourret 1942; Nguyen et al. 2009; this study); 3. A Luoi, Thua Thien Hue Prov. (Nguyen et al. 2009); 4. Phu Loc, Thua Thien Hue Prov. (Nguyen et al. 2009); 5. Son Tra, Da Nang City (this study); 6. Tay Giang, Quang Nam Prov. (Nguyen et al. 2009); 7. Nam Giang, Quang Nam Prov. (Nguyen et al. 2009); 8. Nam Tra My, Quang Nam Prov. (Frost et al. 2006; Nguyen et al. 2009); 9. Son Ha, Quang Ngai Prov. (Tran et al. 2010); 10. Ngoc Linh, Kon Tum Prov. (Orlov 2004); 11. Kon Plong, Kon Tum Prov. (Nguyen et al. 2009); 12. Chu Mom Ray, Kon Tum (Jestrzemski et al. 2013); 13. Kon Chu Rang, Gia Lai Prov. (Nguyen et al. 2009); 14. Tram Lap, Gia Lai Prov. (Chen et al. 2005); 15. K’Bang, Gia Lai Prov. (Nguyen et al. 2009); 16. An Khe, Gia Lai Prov. (Emerson & Ward 1998); 17. Krong Pa, Gia Lai Prov. (Nguyen et al. 2009); 18. York Don, Dak Lak Prov. (Nguyen et al. 2009); 19. Chu Yang Sin, Dak Lak Prov. (Nguyen et al. 2009); 20. Lac Duong, Lam Dong Prov. (Nguyen et al. 2009); 21. Duc Trong, Lam Dong Prov. (this study); 22. Di Linh, Lam Dong Prov. (this study); 23. Da Teh, Lam Dong Prov. (Nguyen et al. 2009); 24. Cat Tien, Dong Nai Prov. (Nguyen et al. 2009). Cambodia: 25. O’Rang, Mondulkiri Prov. (Stuart et al. 2006); 26. Pichrada, Mondulkiri Prov (Stuart et al. 2006). Laos: 27. Dakcheung, Xekong Prov. (Brakels et al. 2021)." figureDoi="http://doi.org/10.5281/zenodo.14734617" httpUri="https://zenodo.org/record/14734617/files/figure.png" pageId="13" pageNumber="70">Fig. 7</figureCitation>
|
||
). All specimens were discovered between 20:00 and 24:00 in evergreen forests at elevations exceeding
|
||
<quantity id="F43A9B15FFB57563FE2AFD4F1B9CBCDF" box="[391,475,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="70" unit="m" value="1300.0">1300 m</quantity>
|
||
a.s.l. The
|
||
<typeStatus id="EC798852FFB57563FDE3FD4F18F7BCDF" box="[590,688,691,717]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="EC798852FFB57563FD48FD4F1917BCDF" box="[741,848,691,717]" pageId="13" pageNumber="70" type="paratype">paratypes</typeStatus>
|
||
<specimenCode id="63649E8BFFB57563FCFAFD4F1E57BCDF" box="[855,1040,690,717]" collectionCode="ITBCZ" pageId="13" pageNumber="70">ITBCZ 8675–78</specimenCode>
|
||
were obtained from a small swamp after rainfall (
|
||
<figureCitation id="ABF92A75FFB57563FE9DFD2B1BCEBCE3" box="[304,393,727,753]" captionStart="FIGURE 8" captionStartId="13.[151,250,1730,1754]" captionTargetBox="[151,1436,849,1707]" captionTargetId="figure-313@13.[151,1436,849,1707]" captionTargetPageId="13" captionText="FIGURE 8. Microhabitats of Limnonectes splendissimus sp. nov. from Lam Dong Province, Vietnam. A. the small swamp; B. the larger swamp; C. the stream in the evergreen forest; D. the water hole along the stream." figureDoi="http://doi.org/10.5281/zenodo.14734619" httpUri="https://zenodo.org/record/14734619/files/figure.png" pageId="13" pageNumber="70">Fig. 8A</figureCitation>
|
||
).
|
||
<typeStatus id="EC798852FFB57563FE31FD2B1BB9BCE3" box="[412,510,727,753]" pageId="13" pageNumber="70" type="paratype">Paratype</typeStatus>
|
||
<specimenCode id="63649E8BFFB57563FDA9FD2B18D4BCE3" box="[516,659,727,753]" collectionCode="ITBCZ" pageId="13" pageNumber="70">ITBCZ 8679</specimenCode>
|
||
was collected from a larger swamp (
|
||
<figureCitation id="ABF92A75FFB57563FB84FD2B1EC7BCE3" box="[1065,1152,727,753]" captionStart="FIGURE 8" captionStartId="13.[151,250,1730,1754]" captionTargetBox="[151,1436,849,1707]" captionTargetId="figure-313@13.[151,1436,849,1707]" captionTargetPageId="13" captionText="FIGURE 8. Microhabitats of Limnonectes splendissimus sp. nov. from Lam Dong Province, Vietnam. A. the small swamp; B. the larger swamp; C. the stream in the evergreen forest; D. the water hole along the stream." figureDoi="http://doi.org/10.5281/zenodo.14734619" httpUri="https://zenodo.org/record/14734619/files/figure.png" pageId="13" pageNumber="70">Fig. 8B</figureCitation>
|
||
).
|
||
<typeStatus id="EC798852FFB57563FB3EFD2B1F47BCE3" box="[1171,1280,727,753]" pageId="13" pageNumber="70" type="paratype">Paratypes</typeStatus>
|
||
<specimenCode id="63649E8BFFB57563FAABFD2B1FDFBCE3" box="[1286,1432,727,753]" collectionCode="ITBCZ" pageId="13" pageNumber="70">ITBCZ 2627</specimenCode>
|
||
,
|
||
<specimenCode id="63649E8BFFB57563FF3AFD071A88BD07" box="[151,207,763,789]" collectionCode="ITBCZ" pageId="13" pageNumber="70">3626</specimenCode>
|
||
were retrieved from a water hole along the stream.
|
||
<specimenCount id="25C4FD79FFB57563FCA4FD071921BD07" box="[777,870,763,789]" pageId="13" pageNumber="70" type="female">Females</specimenCount>
|
||
carry
|
||
<specimenCount id="25C4FD79FFB57563FC01FD071998BD07" box="[940,991,763,789]" pageId="13" pageNumber="70" type="egg">eggs</specimenCount>
|
||
in
|
||
<collectingMunicipality id="D319AC8AFFB57563FBACFD071E3CBD07" box="[1025,1147,763,789]" pageId="13" pageNumber="70">September</collectingMunicipality>
|
||
, and
|
||
<specimenCount id="25C4FD79FFB57563FB1EFD071F5FBD07" box="[1203,1304,763,789]" pageId="13" pageNumber="70" type="juvenile">juveniles</specimenCount>
|
||
were found in
|
||
<location id="361D602BFFB57563FF19FCE31B5CBD2B" LSID="urn:lsid:plazi:treatment:BB6B87E6FFBF7560FF3AFA881BCFBBD3:361D602BFFB57563FF19FCE31B5CBD2B" box="[180,283,799,825]" municipality="September" name="February" pageId="13" pageNumber="70">February</location>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="67BD6678FFB57563FF3AF93E1E74B8EF" ID-DOI="http://doi.org/10.5281/zenodo.14734619" ID-Zenodo-Dep="14734619" httpUri="https://zenodo.org/record/14734619/files/figure.png" pageId="13" pageNumber="70" startId="13.[151,250,1730,1754]" targetBox="[151,1436,849,1707]" targetPageId="13" targetType="figure">
|
||
<paragraph id="337D36F0FFB57563FF3AF93E1E74B8EF" blockId="13.[151,1436,1730,1790]" pageId="13" pageNumber="70">
|
||
<emphasis id="01B6EAE2FFB57563FF3AF93E1B54B8C8" bold="true" box="[151,275,1730,1754]" pageId="13" pageNumber="70">FIGURE 8.</emphasis>
|
||
Microhabitats of
|
||
<taxonomicName id="F4C24D73FFB57563FE64F93F189BB8C8" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[457,732,1730,1754]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB57563FE64F93F189BB8C8" box="[457,732,1730,1754]" italics="true" pageId="13" pageNumber="70">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB57563FD4EF93E1974B8C8" bold="true" box="[739,819,1730,1754]" pageId="13" pageNumber="70">
|
||
<taxonomicNameLabel id="1A855799FFB57563FD4EF93E1974B8C8" box="[739,819,1730,1754]" pageId="13" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Lam Dong Province, Vietnam. A. the small swamp; B. the larger swamp; C. the stream in the evergreen forest; D. the water hole along the stream.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7BD8657BFFB57560FF6AF8AC1BCFBBD3" lastPageId="14" lastPageNumber="71" pageId="13" pageNumber="70" type="diagnosis">
|
||
<paragraph id="337D36F0FFB57560FF6AF8AC1B3ABEC7" blockId="13.[151,1437,1872,2043]" lastBlockId="14.[151,1437,151,1473]" lastPageId="14" lastPageNumber="71" pageId="13" pageNumber="70">
|
||
<emphasis id="01B6EAE2FFB57563FF6AF8AC1B2AB978" bold="true" box="[199,365,1872,1898]" pageId="13" pageNumber="70">Comparisons.</emphasis>
|
||
<taxonomicName id="F4C24D73FFB57563FEDFF8AD18E7B978" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[370,672,1872,1898]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB57563FEDFF8AD18E7B978" box="[370,672,1872,1898]" italics="true" pageId="13" pageNumber="70">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB57563FD08F8AC18BBB978" bold="true" box="[677,764,1872,1898]" pageId="13" pageNumber="70">
|
||
<taxonomicNameLabel id="1A855799FFB57563FD08F8AC18BBB978" box="[677,764,1872,1898]" pageId="13" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a visible tympanum, and therefore it can be distinguished from the following species that do not have a visible tympanum:
|
||
<taxonomicName id="F4C24D73FFB57563FCCCF8891EE9B99C" authority="(Ye et al. 2007)" baseAuthorityName="Ye" baseAuthorityYear="2007" box="[865,1198,1908,1934]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="bannaensis">
|
||
<emphasis id="01B6EAE2FFB57563FCCCF88919BCB99C" box="[865,1019,1908,1934]" italics="true" pageId="13" pageNumber="70">L. bannaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FBA5F8881EE1B99C" author="Ye, C. Y. & Fei, L. & Xie, F. & Jiang, J. P." box="[1032,1190,1908,1934]" pageId="13" pageNumber="70" pagination="545 - 550" refId="ref16938" refString="Ye, C. Y., Fei, L., Xie, F. & Jiang, J. P. (2007) A new Ranidae species from China - Limnonectes bannaensis (Ranidae: Anura). Zoological Research. Kunming, 28, 545 - 550." type="journal article" year="2007">
|
||
Ye
|
||
<emphasis id="01B6EAE2FFB57563FB80F8891E23B99C" box="[1069,1124,1908,1934]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB57563FB15F8891FDBB99C" authority="Dehling" authorityName="Dehling" authorityYear="2014" box="[1208,1436,1908,1934]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="deinodon">
|
||
<emphasis id="01B6EAE2FFB57563FB15F8891F7BB99C" box="[1208,1340,1908,1934]" italics="true" pageId="13" pageNumber="70">L. deinodon</emphasis>
|
||
Dehling
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FF32F8641B07B9A0" author="Dehling, J. M." box="[159,320,1944,1970]" pageId="13" pageNumber="70" pagination="17 - 30" refId="ref12895" refString="Dehling, J. M. (2014) Eine neue Fangzahnfroschart der Gattung Limnonectes (Anura: Dicroglossidae) vom Gunung Lawit, Malaiische Halbinsel. Sauria, 36, 17 - 30." type="journal article" year="2014">Dehling 2014</bibRefCitation>
|
||
);
|
||
<taxonomicName id="F4C24D73FFB57563FEF9F8651884B9A0" authority="(Stuart et al. 2020)" baseAuthorityName="Stuart" baseAuthorityYear="2020" box="[340,707,1944,1970]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="fastigatus">
|
||
<emphasis id="01B6EAE2FFB57563FEF9F8651BA7B9A0" box="[340,480,1944,1970]" italics="true" pageId="13" pageNumber="70">L. fastigatus</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FE5DF86418FCB9A0" author="Stuart, B. L. & Schoen, S. N. & Nelson, E. E. M. & Maher, H. & Neang, T. & Rowley, J. J. L. & McLeod, D. S." box="[496,699,1944,1970]" pageId="13" pageNumber="70" pagination="451 - 473" refId="ref16482" refString="Stuart, B. L., Schoen, S. N., Nelson, E. E. M., Maher, H., Neang, T., Rowley, J. J. L. & McLeod, D. S. (2020) A new fanged frog in the Limnonectes kuhlii complex (Anura: Dicroglossidae) from northeastern Cambodia. Zootaxa, 4894 (3), 451 - 473. https: // doi. org / 10.11646 / zootaxa. 4894.3.11" type="journal article" year="2020">
|
||
Stuart
|
||
<emphasis id="01B6EAE2FFB57563FD90F8651831B9A0" box="[573,630,1944,1970]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB57563FD62F8651E3CB9A0" authority="McLeod, Kelly & Barley" authorityName="McLeod, Kelly & Barley" authorityYear="2012" box="[719,1147,1944,1970]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="isanensis">
|
||
<emphasis id="01B6EAE2FFB57563FD62F8651912B9A0" box="[719,853,1945,1970]" italics="true" pageId="13" pageNumber="70">L. isanensis</emphasis>
|
||
McLeod, Kelly & Barley
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FB27F8641F36B9A0" author="McLeod, D. S. & Kelly, J. K. & Barley, A. J." box="[1162,1393,1944,1970]" pageId="13" pageNumber="70" pagination="261 - 274" refId="ref14950" refString="McLeod, D. S., Kelly, J. K. & Barley, A. J. (2012) " Same-same but different ": another new species of the Limnonectes kuhlii complex from Thailand (Anura: Dicroglossidae). Russian Journal of Herpetology, 19, 261 - 274." type="journal article" year="2012">
|
||
McLeod
|
||
<emphasis id="01B6EAE2FFB57563FB5EF8651F6BB9A0" box="[1267,1324,1944,1970]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="F4C24D73FFB57563FA2BF86518CFB9C4" authority="Matsui, Panha, Khonsue & Kuraishi" authorityName="Matsui, Panha, Khonsue & Kuraishi" authorityYear="2010" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="jarujini">
|
||
<emphasis id="01B6EAE2FFB57563FA2BF8651AACB9C4" italics="true" pageId="13" pageNumber="70">L. jarujini</emphasis>
|
||
Matsui, Panha, Khonsue & Kuraishi
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FD3AF840192DB9C4" author="Matsui, M. & Panha, S. & Khonsue, W. & Kuraishi, N." box="[663,874,1980,2006]" pageId="13" pageNumber="70" pagination="1 - 22" refId="ref14836" refString="Matsui, M., Panha, S., Khonsue, W. & Kuraishi, N. (2010) Two new species of the " kuhlii " complex of the genus Limnonectes from Thailand (Anura: Dicroglossidae). Zootaxa, 2615 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2615.1.1" type="journal article" year="2010">
|
||
Matsui
|
||
<emphasis id="01B6EAE2FFB57563FD43F8411960B9C4" box="[750,807,1980,2006]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="F4C24D73FFB57563FCD3F8411E93B9C4" authority="(Pham et al. 2018)" baseAuthorityName="Pham" baseAuthorityYear="2018" box="[894,1236,1980,2006]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="kiziriani">
|
||
<emphasis id="01B6EAE2FFB57563FCD3F84119BFB9C4" box="[894,1016,1980,2006]" italics="true" pageId="13" pageNumber="70">L. kiziriani</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FBAAF8401E8BB9C4" author="Pham, C. T. & Le, M. D. & Ngo, T. H. & Ziegler, T. & Nguyen, Q. T." box="[1031,1228,1980,2006]" pageId="13" pageNumber="70" pagination="115 - 130" refId="ref15581" refString="Pham, C. T., Le, M. D., Ngo, T. H., Ziegler, T. & Nguyen, Q. T. (2018) A new species of Limnonectes (Amphibia: Anura: Dicroglossidae) from Vietnam. Zootaxa, 4508 (1), 115 - 130. https: // doi. org / 10.11646 / zootaxa. 4508.1.7" type="journal article" year="2018">
|
||
Pham
|
||
<emphasis id="01B6EAE2FFB57563FBFDF8411ECFB9C4" box="[1104,1160,1980,2006]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB57563FB72F8411BA8B9E8" authority="(Boulenger) (Boulenger 1917)" authorityName="Boulenger" authorityYear="1917" baseAuthorityName="Boulenger" baseAuthorityYear="1917" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="macrognathus">
|
||
<emphasis id="01B6EAE2FFB57563FB72F8411FDBB9C4" box="[1247,1436,1980,2006]" italics="true" pageId="13" pageNumber="70">L. macrognathus</emphasis>
|
||
(Boulenger) (
|
||
<bibRefCitation id="57534B01FFB57563FE80F81C1BA1B9E8" author="Boulenger, G. A." box="[301,486,2016,2042]" pageId="13" pageNumber="70" pagination="413 - 418" refId="ref12470" refString="Boulenger, G. A. (1917) Descriptions of new frogs of the genus Rana. Annals and Magazine of Natural History, 20, 413 - 418. https: // doi. org / 10.1080 / 00222931709487029" type="journal article" year="1917">Boulenger 1917</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB57563FE56F81D1957B9E8" authority="McLeod" authorityName="McLeod" authorityYear="2008" box="[507,784,2016,2042]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="megastomias">
|
||
<emphasis id="01B6EAE2FFB57563FE56F81D18EDB9E8" box="[507,682,2017,2042]" italics="true" pageId="13" pageNumber="70">L. megastomias</emphasis>
|
||
McLeod
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FCB2F81C1983B9E8" author="McLeod, D. S." box="[799,964,2016,2042]" pageId="13" pageNumber="70" pagination="26 - 46" refId="ref14902" refString="McLeod, D. S. (2008) A new species of big-headed, fanged dicroglossine frog (Genus Limnonectes) from Thailand. Zootaxa, 1807 (1), 26 - 46. https: // doi. org / 10.11646 / zootaxa. 1807.1.2" type="journal article" year="2008">McLeod 2008</bibRefCitation>
|
||
);
|
||
<taxonomicName id="F4C24D73FFB57563FC7AF81D1F3DB9E9" authority="(McLeod et al. 2015)" baseAuthorityName="McLeod" baseAuthorityYear="2015" box="[983,1402,2016,2043]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="nguyenorum">
|
||
<emphasis id="01B6EAE2FFB57563FC7AF81D1EC7B9E8" box="[983,1152,2017,2042]" italics="true" pageId="13" pageNumber="70">L. nguyenorum</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB57563FB22F81C1F34B9E9" author="McLeod, D. S. & Kurlbaum, S. & Hoang, N. V." box="[1167,1395,2016,2043]" pageId="13" pageNumber="70" pagination="201 - 214" refId="ref15006" refString="McLeod, D. S., Kurlbaum, S. & Hoang, N. V. (2015) More of the same: a diminutive new species of the Limnonectes kuhlii complex from northern Vietnam (Anura: Dicroglossidae). Zootaxa, 3947 (2), 201 - 214. https: // doi. org / 10.11646 / zootaxa. 3947.2.4" type="journal article" year="2015">
|
||
McLeod
|
||
<emphasis id="01B6EAE2FFB57563FB5BF81D1F68B9E8" box="[1270,1327,2016,2042]" italics="true" pageId="13" pageNumber="70">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB57560FA2BF81D1846BEA3" authority="(Pham et al. 2020)" baseAuthorityName="Pham" baseAuthorityYear="2020" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" lastPageId="14" lastPageNumber="71" order="Anura" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="phuyenensis">
|
||
<emphasis id="01B6EAE2FFB57560FA2BF81D1B67BEA3" italics="true" lastPageId="14" lastPageNumber="71" pageId="13" pageNumber="70">L. phuyenensis</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB67560FE9DFF6B1BBEBEA3" author="Pham, C. T. & Do, D. T. & Le, D. M. & Ngo, H. T. & Nguyen, T. N. & Ziegler, T. & Nguyen, Q. T." box="[304,505,151,177]" pageId="14" pageNumber="71" pagination="387 - 402" refId="ref15494" refString="Pham, C. T., Do, D. T., Le, D. M., Ngo, H. T., Nguyen, T. N., Ziegler, T. & Nguyen, Q. T. (2020) A new species of Limnonectes (Amphibia: Anura: Dicroglossidae) from Vietnam. Zootaxa, 4894 (3), 387 - 402. https: // doi. org / 10.11646 / zootaxa. 4894.3.5" type="journal article" year="2020">
|
||
Pham
|
||
<emphasis id="01B6EAE2FFB67560FED7FF6B1BF3BEA3" box="[378,436,151,177]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="F4C24D73FFB67560FDA3FF6B1E53BEA3" authority="Matsui, Panha, Khonsue & Kuraishi" authorityName="Matsui, Panha, Khonsue & Kuraishi" authorityYear="2010" box="[526,1044,151,177]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="taylor">
|
||
<emphasis id="01B6EAE2FFB67560FDA3FF6B1837BEA3" box="[526,624,151,177]" italics="true" pageId="14" pageNumber="71">L. taylor</emphasis>
|
||
Matsui, Panha, Khonsue & Kuraishi
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="57534B01FFB67560FB89FF6B1EBCBEA3" author="Matsui, M. & Panha, S. & Khonsue, W. & Kuraishi, N." box="[1060,1275,151,177]" pageId="14" pageNumber="71" pagination="1 - 22" refId="ref14836" refString="Matsui, M., Panha, S., Khonsue, W. & Kuraishi, N. (2010) Two new species of the " kuhlii " complex of the genus Limnonectes from Thailand (Anura: Dicroglossidae). Zootaxa, 2615 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2615.1.1" type="journal article" year="2010">
|
||
Matsui
|
||
<emphasis id="01B6EAE2FFB67560FBD1FF6B1EE8BEA3" box="[1148,1199,151,177]" italics="true" pageId="14" pageNumber="71">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
); and
|
||
<taxonomicName id="F4C24D73FFB67560FAEDFF6B1B3EBEC7" authority="(Matsui et al. 2014)" baseAuthorityName="Matsui" baseAuthorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="utara">
|
||
<emphasis id="01B6EAE2FFB67560FAEDFF6B1FDABEA2" box="[1344,1437,151,176]" italics="true" pageId="14" pageNumber="71">L. utara</emphasis>
|
||
(
|
||
<bibRefCitation id="57534B01FFB67560FF32FF471B36BEC7" author="Matsui, M. & Belabut, D. M. & Ahmad, N." box="[159,369,187,213]" pageId="14" pageNumber="71" pagination="75 - 93" refId="ref14626" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014) Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">
|
||
Matsui
|
||
<emphasis id="01B6EAE2FFB67560FF58FF471B69BEC7" box="[245,302,187,213]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB67560FF6AFF231EF2BD4F" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="71">
|
||
Among species having a visible tympanum,
|
||
<taxonomicName id="F4C24D73FFB67560FD15FF231935BEEB" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[696,882,223,249]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB67560FD15FF231935BEEB" box="[696,882,223,249]" italics="true" pageId="14" pageNumber="71">L. splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB67560FCD5FF231995BEEB" bold="true" box="[888,978,223,249]" pageId="14" pageNumber="71">
|
||
<taxonomicNameLabel id="1A855799FFB67560FCD5FF231995BEEB" box="[888,978,223,249]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a larger body size (with SVL 104.1– 107.0 mm in males and
|
||
<quantity id="F43A9B15FFB67560FE32FEFF1806BF0F" box="[415,577,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="9.245" metricValueMax="9.77" metricValueMin="8.72" pageId="14" pageNumber="71" unit="mm" value="92.45" valueMax="97.7" valueMin="87.2">87.2–97.7 mm</quantity>
|
||
in females) than the following species:
|
||
<taxonomicName id="F4C24D73FFB67560FC55FEFF196FBF53" authority="Phimmachak, Sivongxay, Seateun, Yodthong, Rujirawan, Neang, Aowphol & Stuart" authorityName="Phimmachak, Sivongxay, Seateun, Yodthong, Rujirawan, Neang, Aowphol & Stuart" authorityYear="2018" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="coffeatus">
|
||
<emphasis id="01B6EAE2FFB67560FC55FEFF1E3DBF0F" box="[1016,1146,259,285]" italics="true" pageId="14" pageNumber="71">L. coffeatus</emphasis>
|
||
Phimmachak, Sivongxay, Seateun, Yodthong, Rujirawan, Neang, Aowphol & Stuart
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FCDAFEDB199BBF53" box="[887,988,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="14" pageNumber="71" unit="mm" value="37.9">37.9 mm</quantity>
|
||
in male and
|
||
<quantity id="F43A9B15FFB67560FBDFFEDB1F50BF53" box="[1138,1303,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="4.01" metricValueMax="4.21" metricValueMin="3.81" pageId="14" pageNumber="71" unit="mm" value="40.1" valueMax="42.1" valueMin="38.1">38.1–42.1 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FF3AFEB71BECBF77" author="Phimmachak, S. & Sivongxay, N. & Seateun, S. & Yodthong, S. & Rujirawan, A. & Neang, T. & Aowphol, A. & Stuart, B. L." box="[151,427,331,357]" pageId="14" pageNumber="71" pagination="325 - 340" refId="ref15817" refString="Phimmachak, S., Sivongxay, N., Seateun, S., Yodthong, S., Rujirawan, A., Neang, T., Aowphol, A. & Stuart, B. L. (2018) A new Limnonectes (Anura: Dicroglossidae) from southern Laos. Zootaxa, 4375 (3), 325 - 340. https: // doi. org / 10.11646 / zootaxa. 4375.3.2" type="journal article" year="2018">
|
||
Phimmachak
|
||
<emphasis id="01B6EAE2FFB67560FE9DFEB71B2FBF77" box="[304,360,331,357]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="F4C24D73FFB67560FE10FEB7187EBF77" baseAuthorityName="Smith" baseAuthorityYear="1922" box="[445,569,331,357]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="dabanus">
|
||
<emphasis id="01B6EAE2FFB67560FE10FEB7187EBF77" box="[445,569,331,357]" italics="true" pageId="14" pageNumber="71">L. dabanus</emphasis>
|
||
</taxonomicName>
|
||
(SVL 52.0–55.0 mm in males and 39.0 mm in female;
|
||
<bibRefCitation id="57534B01FFB67560FB33FEB71F64BF77" author="Smith, M. A." box="[1182,1315,331,357]" pageId="14" pageNumber="71" pagination="215 - 229" refId="ref16404" refString="Smith, M. A. (1922) The frogs allied to Rana doriae. Journal of the Natural History Society of Siam, 4, 215 - 229." type="journal article" year="1922">Smith 1922</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FA9BFEB71B58BF9B" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="01B6EAE2FFB67560FA9BFEB71FDBBF77" box="[1334,1436,331,357]" italics="true" pageId="14" pageNumber="71">L. doriae</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
(SVL 41.4–55.0 mm in males and
|
||
<quantity id="F43A9B15FFB67560FD1DFE921912BF9B" box="[688,853,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="4.484999999999999" metricValueMax="5.029999999999999" metricValueMin="3.94" pageId="14" pageNumber="71" unit="mm" value="44.849999999999994" valueMax="50.3" valueMin="39.4">39.4–50.3 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FC4CFE931E93BF9B" author="Yodthong, S. & Rujirawan, A. & Stuart, B. L. & Aowphol, A." box="[993,1236,367,393]" pageId="14" pageNumber="71" pagination="1 - 28" refId="ref16990" refString="Yodthong, S., Rujirawan, A., Stuart, B. L. & Aowphol, A. (2021) A new Limnonectes (Anura: Dicroglossidae) from southern Thailand. Animals. Basel, Switzerland, 11, 1 - 28. https: // doi. org / 10.3390 / ani 11020566" type="journal article" year="2021">
|
||
Yodthong
|
||
<emphasis id="01B6EAE2FFB67560FBFBFE931ED7BF9B" box="[1110,1168,367,393]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FB45FE931FDBBF9B" baseAuthorityName="Andersson" baseAuthorityYear="1916" box="[1256,1436,367,393]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="gyldenstolpei">
|
||
<emphasis id="01B6EAE2FFB67560FB45FE931FDBBF9B" box="[1256,1436,367,393]" italics="true" pageId="14" pageNumber="71">L. gyldenstolpei</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FF77FE6E1B3ABFBF" box="[218,381,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.975" metricValueMax="6.84" metricValueMin="5.11" pageId="14" pageNumber="71" unit="mm" value="59.75" valueMax="68.4" valueMin="51.1">51.1–68.4 mm</quantity>
|
||
in males and 23.0 in female;
|
||
<bibRefCitation id="57534B01FFB67560FD6DFE6F193EBFBF" author="Andersson, L. G." box="[704,889,403,429]" pageId="14" pageNumber="71" pagination="13 - 17" refId="ref12258" refString="Andersson, L. G. (1916) Zoological results of the Swedish zoological expeditions to Siam 1911 - 1912 and 1914. 3. Batrachians. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 55, 13 - 17." type="journal article" year="1916">Andersson 1916</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="57534B01FFB67560FC2FFE6F1E2EBFBF" author="Aowphol, A. & Rujirawan, A. & Taksintum, W. & Chuaynkern, Y. & Stuart, B. L." box="[898,1129,402,429]" pageId="14" pageNumber="71" pagination="258 - 270" refId="ref12297" refString="Aowphol, A., Rujirawan, A., Taksintum, W., Chuaynkern, Y. & Stuart, B. L. (2015) A new caruncle-bearing Limnonectes (Anura: Dicroglossidae) from northeastern Thailand. Zootaxa, 3956 (2), 258 - 270. https: // doi. org / 10.11646 / zootaxa. 3956.2.6" type="journal article" year="2015">
|
||
Aowphol
|
||
<emphasis id="01B6EAE2FFB67560FC5DFE6F1E6FBFBF" box="[1008,1064,403,429]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FBD1FE6F1FDABFBF" authority="(Stoliczka)" baseAuthorityName="Stoliczka" baseAuthorityYear="1870" box="[1148,1437,403,429]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="hascheanus">
|
||
<emphasis id="01B6EAE2FFB67560FBD1FE6F1F5BBFBF" box="[1148,1308,403,429]" italics="true" pageId="14" pageNumber="71">L. hascheanus</emphasis>
|
||
(Stoliczka)
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FF76FE4A1B39BFC3" box="[219,382,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" metricValueMax="2.54" metricValueMin="1.88" pageId="14" pageNumber="71" unit="mm" value="22.1" valueMax="25.4" valueMin="18.8">18.8–25.4 mm</quantity>
|
||
in males and 20.5–25.0 mm in females;
|
||
<bibRefCitation id="57534B01FFB67560FCEEFE4B1E60BFC3" author="Inger, R. F. & Stuart, B. L." box="[835,1063,439,465]" pageId="14" pageNumber="71" pagination="51 - 68" refId="ref14140" refString="Inger, R. F. & Stuart, B. L. (2010) Systematics of Limnonectes (Taylorana) Dubois. Current Herpetology. Kyoto, 29, 51 - 68. https: // doi. org / 10.3105 / 018.029.0201" type="journal article" year="2010">Inger & Stuart 2010</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FB97FE4B1F1EBFC3" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1929" box="[1082,1369,439,465]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="khammonensis">
|
||
<emphasis id="01B6EAE2FFB67560FB97FE4B1EBBBFC3" box="[1082,1276,439,465]" italics="true" pageId="14" pageNumber="71">L. khammonensis</emphasis>
|
||
(Smith)
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FF3AFE261ABABFE7" box="[151,253,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="14" pageNumber="71" unit="mm" value="37.5">37.5 mm</quantity>
|
||
in female;
|
||
<bibRefCitation id="57534B01FFB67560FE2CFE27184FBFE7" author="Smith, M. A." box="[385,520,475,501]" pageId="14" pageNumber="71" pagination="294 - 297" refId="ref16435" refString="Smith, M. A. (1929) Descriptions of a new skink from Christmas Island and a new frog from Annam. Annals and Magazine of Natural History, 3, 294 - 297. https: // doi. org / 10.1080 / 00222932908672972" type="journal article" year="1929">Smith 1929</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FDB3FE271883BFE7" box="[542,708,475,501]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="kohchangae">
|
||
<emphasis id="01B6EAE2FFB67560FDB3FE271883BFE7" box="[542,708,475,501]" italics="true" pageId="14" pageNumber="71">L. kohchangae</emphasis>
|
||
</taxonomicName>
|
||
(SVL 40.0–42.0 mm in males and 39.0–42.0 in females;
|
||
<bibRefCitation id="57534B01FFB67560FAF4FE271A94BC0B" author="Smith, M. A." pageId="14" pageNumber="71" pagination="215 - 229" refId="ref16404" refString="Smith, M. A. (1922) The frogs allied to Rana doriae. Journal of the Natural History Society of Siam, 4, 215 - 229." type="journal article" year="1922">Smith 1922</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FF45FE031E48BC0B" authority="Aowphol, Rujirawan, Taksinum, Chuaynkern & Stuart" authorityName="Aowphol, Rujirawan, Taksinum, Chuaynkern & Stuart" authorityYear="2015" box="[232,1039,511,537]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="lauhachindai">
|
||
<emphasis id="01B6EAE2FFB67560FF45FE031BDBBC0B" box="[232,412,511,537]" italics="true" pageId="14" pageNumber="71">L. lauhachindai</emphasis>
|
||
Aowphol, Rujirawan, Taksinum, Chuaynkern & Stuart
|
||
</taxonomicName>
|
||
(SVL 30.5–42.0 mm in males and
|
||
<quantity id="F43A9B15FFB67560FF3AFDDF1B7DBC2F" box="[151,314,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" metricValueMax="3.79" metricValueMin="3.29" pageId="14" pageNumber="71" unit="mm" value="35.4" valueMax="37.9" valueMin="32.9">32.9–37.9 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FE6CFDDF18EDBC2F" author="Aowphol, A. & Rujirawan, A. & Taksintum, W. & Chuaynkern, Y. & Stuart, B. L." box="[449,682,546,573]" pageId="14" pageNumber="71" pagination="258 - 270" refId="ref12297" refString="Aowphol, A., Rujirawan, A., Taksintum, W., Chuaynkern, Y. & Stuart, B. L. (2015) A new caruncle-bearing Limnonectes (Anura: Dicroglossidae) from northeastern Thailand. Zootaxa, 3956 (2), 258 - 270. https: // doi. org / 10.11646 / zootaxa. 3956.2.6" type="journal article" year="2015">
|
||
Aowphol
|
||
<emphasis id="01B6EAE2FFB67560FD9DFDDF1826BC2F" box="[560,609,547,573]" italics="true" pageId="14" pageNumber="71">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FD10FDDF197DBC2F" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[701,826,547,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="limborgi">
|
||
<emphasis id="01B6EAE2FFB67560FD10FDDF197DBC2F" box="[701,826,547,573]" italics="true" pageId="14" pageNumber="71">L. limborgi</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FC28FDDF19AEBC2F" box="[901,1001,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="14" pageNumber="71" unit="mm" value="31.2">31.2 mm</quantity>
|
||
in male and
|
||
<quantity id="F43A9B15FFB67560FBD5FDDF1F5CBC2F" box="[1144,1307,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.7399999999999998" metricValueMin="2.69" pageId="14" pageNumber="71" unit="mm" value="27.15" valueMax="27.4" valueMin="26.9">26.9–27.4 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FF3AFDBB1B2CBC73" author="Kohler, G. & Zwitzers, B. & Than, N. L. & Gupta, D. K. & Janke, A. & Pauls, S. U. & Thammachoti, P." box="[151,363,583,609]" pageId="14" pageNumber="71" pagination="1 - 35" refId="ref14419" refString="Kohler, G., Zwitzers, B., Than, N. L., Gupta, D. K., Janke, A., Pauls, S. U. & Thammachoti, P. (2021) Bioacoustics reveal hidden diversity in frogs: Two new species of the genus Limnonectes from Myanmar (Amphibia, Anura, Dicroglossidae). Diversity, 13, 1 - 35. https: // doi. org / 10.3390 / d 13090399" type="journal article" year="2021">
|
||
Köhler
|
||
<emphasis id="01B6EAE2FFB67560FF40FDBB1B60BC73" box="[237,295,583,609]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FED2FDBB18DDBC73" authority="(Inger)" baseAuthorityName="Inger" baseAuthorityYear="1966" box="[383,666,583,609]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="paramacrodon">
|
||
<emphasis id="01B6EAE2FFB67560FED2FDBB1804BC73" box="[383,579,583,609]" italics="true" pageId="14" pageNumber="71">L. paramacrodon</emphasis>
|
||
(Inger)
|
||
</taxonomicName>
|
||
(SVL up to
|
||
<quantity id="F43A9B15FFB67560FC87FDBA193DBC73" box="[810,890,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="71" unit="mm" value="75.0">75 mm</quantity>
|
||
in males;
|
||
<bibRefCitation id="57534B01FFB67560FC5CFDBB1E36BC73" author="Inger, R. F." box="[1009,1137,583,609]" pageId="14" pageNumber="71" pagination="1 - 402" refId="ref14055" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana, Zoology, 52, 1 - 402. https: // doi. org / 10.5962 / bhl. title. 3147" type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FB2BFDBB1FDABC73" authority="(Stoliczka)" baseAuthorityName="Stoliczka" baseAuthorityYear="1873" box="[1158,1437,583,609]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="plicatellus">
|
||
<emphasis id="01B6EAE2FFB67560FB2BFDBB1F5EBC73" box="[1158,1305,583,609]" italics="true" pageId="14" pageNumber="71">L. plicatellus</emphasis>
|
||
(Stoliczka)
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FF73FD971B77BC97" box="[222,304,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="71" unit="mm" value="36.0">36 mm</quantity>
|
||
in male and
|
||
<quantity id="F43A9B15FFB67560FE66FD97185ABC97" box="[459,541,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="71" unit="mm" value="34.0">34 mm</quantity>
|
||
in female;
|
||
<bibRefCitation id="57534B01FFB67560FD09FD971918BC97" author="Boulenger, G. A." box="[676,863,619,645]" pageId="14" pageNumber="71" pagination="1 - 226" refId="ref12511" refString="Boulenger, G. A. (1920). A monograph of the South Asian, Papuan, Melanesian and Australian frogs of the genus Rana. Records of the Indian Museum, 20, 1 - 226. https: // doi. org / 10.26515 / rzsi / v 20 / i 1 / 1920 / 163533" type="journal article" year="1920">Boulenger 1920</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FCD8FD971AB8BCBB" authority="Yodthong, Rujirawan, Stuart & Aowphol" authorityName="Yodthong, Rujirawan, Stuart & Aowphol" authorityYear="2021" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="pseudodoriae">
|
||
<emphasis id="01B6EAE2FFB67560FCD8FD971E6ABC97" box="[885,1069,619,645]" italics="true" pageId="14" pageNumber="71">L. pseudodoriae</emphasis>
|
||
Yodthong, Rujirawan, Stuart & Aowphol
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FEE3FD731BB5BCBB" box="[334,498,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="4.54" metricValueMax="4.82" metricValueMin="4.26" pageId="14" pageNumber="71" unit="mm" value="45.4" valueMax="48.2" valueMin="42.6">42.6–48.2 mm</quantity>
|
||
in males and 36.0–
|
||
<quantity id="F43A9B15FFB67560FD7EFD73197FBCBB" box="[723,824,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="4.11" pageId="14" pageNumber="71" unit="mm" value="41.1">41.1 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FC68FD731EFFBCBB" author="Yodthong, S. & Rujirawan, A. & Stuart, B. L. & Aowphol, A." box="[965,1208,655,681]" pageId="14" pageNumber="71" pagination="1 - 28" refId="ref16990" refString="Yodthong, S., Rujirawan, A., Stuart, B. L. & Aowphol, A. (2021) A new Limnonectes (Anura: Dicroglossidae) from southern Thailand. Animals. Basel, Switzerland, 11, 1 - 28. https: // doi. org / 10.3390 / ani 11020566" type="journal article" year="2021">
|
||
Yodthong
|
||
<emphasis id="01B6EAE2FFB67560FB97FD731E33BCBB" box="[1082,1140,655,681]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="F4C24D73FFB67560FB60FD731FDBBCBB" authorityName="Pham, Le, Nguyen, Ziegler, Wu & Nguyen" authorityYear="2017" box="[1229,1436,655,681]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="quangninhensis">
|
||
<emphasis id="01B6EAE2FFB67560FB60FD731FDBBCBB" box="[1229,1436,655,681]" italics="true" pageId="14" pageNumber="71">L. quangninhensis</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="F43A9B15FFB67560FF4DFD4E1BCFBCDF" box="[224,392,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.89" metricValueMin="5.01" pageId="14" pageNumber="71" unit="mm" value="59.5" valueMax="68.9" valueMin="50.1">50.1–68.9 mm</quantity>
|
||
in males and 45.5–63.0 mm in females;
|
||
<bibRefCitation id="57534B01FFB67560FCDBFD4F1E00BCDF" author="Pham, C. T. & Le, M. D. & Nguyen, T. T. & Ziegler, T. & Wu, J. Z. & Nguyen, T. Q." box="[886,1095,691,717]" pageId="14" pageNumber="71" pagination="545 - 558" refId="ref15654" refString="Pham, C. T., Le, M. D., Nguyen, T. T., Ziegler, T., Wu, J. Z. & Nguyen, T. Q. (2017) A new species of Limnonectes (Amphibia: Anura: Dicroglossidae) from Vietnam. Zootaxa, 4269 (4), 545 - 558. https: // doi. org / 10.11646 / zootaxa. 4269.4.8" type="journal article" year="2017">
|
||
Pham
|
||
<emphasis id="01B6EAE2FFB67560FC6FFD4F19B8BCDF" box="[962,1023,691,717]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
); and
|
||
<taxonomicName id="F4C24D73FFB67560FB3EFD4F19D4BCE3" authority="Phimmachak, Richards, Sivongsay, Seateun, Chuaykern, Makchai, Som & Stuart" authorityName="Phimmachak, Richards, Sivongsay, Seateun, Chuaykern, Makchai, Som & Stuart" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="savan">
|
||
<emphasis id="01B6EAE2FFB67560FB3EFD4F1EB0BCDE" box="[1171,1271,691,716]" italics="true" pageId="14" pageNumber="71">L. savan</emphasis>
|
||
Phimmachak, Richards, Sivongsay, Seateun, Chuaykern, Makchai, Som & Stuart
|
||
</taxonomicName>
|
||
(SVL 39.0–
|
||
<quantity id="F43A9B15FFB67560FB8FFD2A1ECFBCE3" box="[1058,1160,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="5.62" pageId="14" pageNumber="71" unit="mm" value="56.2">56.2 mm</quantity>
|
||
in males and
|
||
<quantity id="F43A9B15FFB67560FA81FD2A1A84BD07" metricMagnitude="-2" metricUnit="m" metricValue="4.755" metricValueMax="5.62" metricValueMin="3.8899999999999997" pageId="14" pageNumber="71" unit="mm" value="47.55" valueMax="56.2" valueMin="38.9">38.9–56.2 mm</quantity>
|
||
in females;
|
||
<bibRefCitation id="57534B01FFB67560FEE3FD071822BD07" author="Phimmachak, S. & Richards, S. J. & Sivongxay, N. & Seateun, S. & Chuaynkern, Y. & Makchai, S. & Som, E. H. & Stuart, B. L." box="[334,613,763,789]" pageId="14" pageNumber="71" pagination="133 - 156" refId="ref15734" refString="Phimmachak, S., Richards, S. J., Sivongxay, N., Seateun, S., Chuaynkern, Y., Makchai, S., Som, E. H. & Stuart, B. L. (2019) A new caruncle-bearing fanged frog (Limnonectes, Dicroglossidae) from Laos and Thailand. ZooKeys, 846, 133 - 156. https: // doi. org / 10.3897 / zookeys. 846.33200" type="journal article" year="2019">
|
||
Phimmachak
|
||
<emphasis id="01B6EAE2FFB67560FE45FD07185DBD07" box="[488,538,763,789]" italics="true" pageId="14" pageNumber="71">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
). For the large-body size group,
|
||
<taxonomicName id="F4C24D73FFB67560FC79FD071EC9BD07" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[980,1166,763,789]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB67560FC79FD071EC9BD07" box="[980,1166,763,789]" italics="true" pageId="14" pageNumber="71">L. splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB67560FB3BFD071EB7BD07" bold="true" box="[1174,1264,763,789]" pageId="14" pageNumber="71">
|
||
<taxonomicNameLabel id="1A855799FFB67560FB3BFD071EB7BD07" box="[1174,1264,763,789]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="F4C24D73FFB67560FA2BFD071B2ABD2B" authority="(Kiew)" baseAuthorityName="Kiew" baseAuthorityYear="1984" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="malesianus">
|
||
<emphasis id="01B6EAE2FFB67560FA2BFD071B52BD2B" italics="true" pageId="14" pageNumber="71">L. malesianus</emphasis>
|
||
(Kiew)
|
||
</taxonomicName>
|
||
by having a smaller body size in males, with SVL up to 107.0 mm (vs. SVL up to
|
||
<quantity id="F43A9B15FFB67560FA8DFCE21F38BD2B" box="[1312,1407,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="71" unit="mm" value="150.0">150 mm</quantity>
|
||
in males;
|
||
<bibRefCitation id="57534B01FFB67560FF4AFCBF1BD4BD4F" author="Das, I. & Jankowski, A. & Makmor, M. I. B. & Haas, A." box="[231,403,835,861]" pageId="14" pageNumber="71" pagination="141 - 174" refId="ref12832" refString="Das, I., Jankowski, A., Makmor, M. I. B. & Haas, A. (2007) Species diversity, elevational distribution and reproductive modes in an amphibian community at the Matang Range, Sarawak (Borneo). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 104, 141 - 174." type="journal article" year="2007">
|
||
Das
|
||
<emphasis id="01B6EAE2FFB67560FEB7FCBF1B15BD4F" box="[282,338,835,861]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
) and by having toes fully webbed (vs. toes ¾ webbed;
|
||
<bibRefCitation id="57534B01FFB67560FC56FCBF1EEDBD4F" author="Das, I. & Jankowski, A. & Makmor, M. I. B. & Haas, A." box="[1019,1194,835,861]" pageId="14" pageNumber="71" pagination="141 - 174" refId="ref12832" refString="Das, I., Jankowski, A., Makmor, M. I. B. & Haas, A. (2007) Species diversity, elevational distribution and reproductive modes in an amphibian community at the Matang Range, Sarawak (Borneo). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 104, 141 - 174." type="journal article" year="2007">
|
||
Das
|
||
<emphasis id="01B6EAE2FFB67560FB83FCBF1E18BD4F" box="[1070,1119,835,861]" italics="true" pageId="14" pageNumber="71">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB67560FF6AFC9B1BCABA27" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="71">
|
||
<taxonomicName id="F4C24D73FFB67560FF6AFC9B1BBCBD93" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[199,507,871,897]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB67560FF6AFC9B1BBCBD93" box="[199,507,871,897]" italics="true" pageId="14" pageNumber="71">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB67560FDA9FC9B1827BD93" bold="true" box="[516,608,871,897]" pageId="14" pageNumber="71">
|
||
<taxonomicNameLabel id="1A855799FFB67560FDA9FC9B1827BD93" box="[516,608,871,897]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically similar to
|
||
<taxonomicName id="F4C24D73FFB67560FC6CFC9B1EF9BD93" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1920" box="[961,1214,871,897]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="blythii">
|
||
<emphasis id="01B6EAE2FFB67560FC6CFC9B1E6DBD93" box="[961,1066,871,897]" italics="true" pageId="14" pageNumber="71">L. blythii</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
. However, the new species can be distinguished from
|
||
<taxonomicName id="F4C24D73FFB67560FDB6FC7718C6BDB7" baseAuthorityName="Boulenger" baseAuthorityYear="1920" box="[539,641,907,933]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="blythii">
|
||
<emphasis id="01B6EAE2FFB67560FDB6FC7718C6BDB7" box="[539,641,907,933]" italics="true" pageId="14" pageNumber="71">L. blythii</emphasis>
|
||
</taxonomicName>
|
||
by having smaller body size, with SVL 104.1–107.0 mm in males and
|
||
<quantity id="F43A9B15FFB67560FF3AFC531B7ABDDB" box="[151,317,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="9.245" metricValueMax="9.77" metricValueMin="8.72" pageId="14" pageNumber="71" unit="mm" value="92.45" valueMax="97.7" valueMin="87.2">87.2–97.7 mm</quantity>
|
||
in females (vs. larger body size of both sexes, SVL from 110.0 mm to nearly 260.0 mm;
|
||
<bibRefCitation id="57534B01FFB67560FAFEFC531A94BDFF" author="Taylor, E. H." pageId="14" pageNumber="71" pagination="265 - 599" refId="ref16787" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 43, 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor 1962</bibRefCitation>
|
||
); by males having a nuptial pad on first finger (vs. absent;
|
||
<bibRefCitation id="57534B01FFB67560FCCEFC2F19A8BDFF" author="Taylor, E. H." box="[867,1007,979,1005]" pageId="14" pageNumber="71" pagination="265 - 599" refId="ref16787" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 43, 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor 1962</bibRefCitation>
|
||
); and by having a tympanum diameter approximately eye diameter with ratio TD/ED 0.95±
|
||
<quantity id="F43A9B15FFB67560FD49FC0A1974BA03" box="[740,819,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="71" unit="in" value="0.07">0.07 in</quantity>
|
||
males and 0.76±
|
||
<quantity id="F43A9B15FFB67560FC5FFC0B1E06BA03" box="[1010,1089,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="71" unit="in" value="0.02">0.02 in</quantity>
|
||
females (vs. greatest ratio TD/ ED 0.6;
|
||
<bibRefCitation id="57534B01FFB67560FF58FBE71BC7BA27" author="Taylor, E. H." box="[245,384,1051,1077]" pageId="14" pageNumber="71" pagination="265 - 599" refId="ref16787" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. University of Kansas Science Bulletin, 43, 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor 1962</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="337D36F0FFB67560FF6AFBC31BCFBBD3" blockId="14.[151,1437,151,1473]" pageId="14" pageNumber="71">
|
||
<taxonomicName id="F4C24D73FFB67560FF6AFBC31BB0BA4B" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[199,503,1087,1113]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB67560FF6AFBC31BB0BA4B" box="[199,503,1087,1113]" italics="true" pageId="14" pageNumber="71">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB67560FE53FBC31810BA4B" bold="true" box="[510,599,1087,1113]" pageId="14" pageNumber="71">
|
||
<taxonomicNameLabel id="1A855799FFB67560FE53FBC31810BA4B" box="[510,599,1087,1113]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most closely to related to
|
||
<taxonomicName id="F4C24D73FFB67560FC3AFBC31E44BA4B" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[919,1027,1087,1113]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="poilani">
|
||
<emphasis id="01B6EAE2FFB67560FC3AFBC31E44BA4B" box="[919,1027,1087,1113]" italics="true" pageId="14" pageNumber="71">L. poilani</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="ABF92A75FFB67560FBBEFBC31E10BA4B" box="[1043,1111,1087,1113]" captionStart="FIGURE 9" captionStartId="17.[151,250,1879,1903]" captionTargetBox="[151,1436,737,1856]" captionTargetId="figure-165@17.[151,1436,733,1856]" captionTargetPageId="17" captionText="FIGURE 9. Color variation in life (A–M) and in preservative (N) of Limnonectes poilani from Bac Huong Hoa Nature Reserve, Quang Tri Province, Central Vietnam.A–E. ITBCZ 4439, adult male; F&G. no voucher, subadult; H&I. ITBCZ 4440, subadult male; K&L. subadult male, ITBCZ 4412; M&N. ITBCZ 8331, juvenile. Scale bars 5 mm.A–L photos by Luan T. Nguyen; M–N photos by Manh V. Le." figureDoi="http://doi.org/10.5281/zenodo.14734621" httpUri="https://zenodo.org/record/14734621/files/figure.png" pageId="14" pageNumber="71">Fig. 9</figureCitation>
|
||
). Measurements between the new species and
|
||
<taxonomicName id="F4C24D73FFB67560FEFBFB9F1B84BA6F" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[342,451,1123,1149]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="poilani">
|
||
<emphasis id="01B6EAE2FFB67560FEFBFB9F1B84BA6F" box="[342,451,1123,1149]" italics="true" pageId="14" pageNumber="71">L. poilani</emphasis>
|
||
</taxonomicName>
|
||
are provided in
|
||
<tableCitation id="7E40034BFFB67560FDD0FB9F1894BA6F" box="[637,723,1123,1149]" captionStart="TABLE 4" captionStartId="14.[151,239,1520,1544]" captionTargetPageId="14" captionText="TABLE 4. Statistical compared of Limnoectes splendissimus sp. nov. and L. poilani (SD = standard deviation; for other abbreviations see Material and methods). Bold figures indicate p <0.05." httpUri="http://table.plazi.org/id/67BD6678FFB67560FF3AFA0C19DCB83E" pageId="14" pageNumber="71" tableUuid="67BD6678FFB67560FF3AFA0C19DCB83E">Table 4</tableCitation>
|
||
.
|
||
<taxonomicName id="F4C24D73FFB67560FD72FB9F1E57BA6F" authorityName="Le & Nguyen & Murphy & Tran & Nguyen & Che" authorityYear="2025" box="[735,1040,1123,1149]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="splendissimus" status="sp. nov.">
|
||
<emphasis id="01B6EAE2FFB67560FD72FB9F1E57BA6F" box="[735,1040,1123,1149]" italics="true" pageId="14" pageNumber="71">Limnonectes splendissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="01B6EAE2FFB67560FBBAFB9F1E36BA6F" bold="true" box="[1047,1137,1123,1149]" pageId="14" pageNumber="71">
|
||
<taxonomicNameLabel id="1A855799FFB67560FBBAFB9F1E36BA6F" box="[1047,1137,1123,1149]" pageId="14" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="F4C24D73FFB67560FF3AFB7B1B43BAB3" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[151,260,1159,1185]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="poilani">
|
||
<emphasis id="01B6EAE2FFB67560FF3AFB7B1B43BAB3" box="[151,260,1159,1185]" italics="true" pageId="14" pageNumber="71">L. poilani</emphasis>
|
||
</taxonomicName>
|
||
by having larger body size, with SVL 104.1–107.0 mm in males and
|
||
<quantity id="F43A9B15FFB67560FBBEFB7B1EF1BAB3" box="[1043,1206,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="9.245" metricValueMax="9.77" metricValueMin="8.72" pageId="14" pageNumber="71" unit="mm" value="92.45" valueMax="97.7" valueMin="87.2">87.2–97.7 mm</quantity>
|
||
in females (vs. SVL
|
||
<quantity id="F43A9B15FFB67560FF3AFB571B7EBAD7" box="[151,313,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="8.669999999999998" metricValueMax="8.93" metricValueMin="8.41" pageId="14" pageNumber="71" unit="mm" value="86.69999999999999" valueMax="89.3" valueMin="84.1">84.1–89.3 mm</quantity>
|
||
in males and
|
||
<quantity id="F43A9B15FFB67560FE7DFB561835BAD7" box="[464,626,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="6.87" metricValueMax="6.99" metricValueMin="6.75" pageId="14" pageNumber="71" unit="mm" value="68.7" valueMax="69.9" valueMin="67.5">67.5–69.9 mm</quantity>
|
||
in females; SVL 93.0 mm in male [
|
||
<bibRefCitation id="57534B01FFB67560FC55FB571ED3BAD7" author="Bourret, R." box="[1016,1172,1195,1221]" pageId="14" pageNumber="71" refId="ref12571" refString="Bourret, R. (1942) Les batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi, 547 pp." type="book" year="1942">Bourret 1942</bibRefCitation>
|
||
]; SVL
|
||
<quantity id="F43A9B15FFB67560FB72FB561FC6BAD7" box="[1247,1409,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="7.58" metricValueMax="9.309999999999999" metricValueMin="5.85" pageId="14" pageNumber="71" unit="mm" value="75.8" valueMax="93.1" valueMin="58.5">58.5–93.1 mm</quantity>
|
||
in males and
|
||
<quantity id="F43A9B15FFB67560FEA0FB321BE8BAFB" box="[269,431,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="6.665" metricValueMax="7.959999999999999" metricValueMin="5.37" pageId="14" pageNumber="71" unit="mm" value="66.65" valueMax="79.6" valueMin="53.7">53.7–79.6 mm</quantity>
|
||
in females [
|
||
<bibRefCitation id="57534B01FFB67560FD9EFB3318B5BAFB" author="Inger, R. F. & Orlov, N. L. & Darevsky, I. S." box="[563,754,1231,1257]" pageId="14" pageNumber="71" pagination="1 - 46" refId="ref14098" refString="Inger, R. F., Orlov, N. L. & Darevsky, I. S. (1999) Frogs of Vietnam: A report on new collections. Fieldiana Zoology, 92, 1 - 46." type="journal article" year="1999">
|
||
Inger
|
||
<emphasis id="01B6EAE2FFB67560FDDAFB3318E9BAFB" box="[631,686,1231,1257]" italics="true" pageId="14" pageNumber="71">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
]); by having tympanum diameter approximately eye diameter with ratio TD/ED 0.95±
|
||
<quantity id="F43A9B15FFB67560FE0FFB0E1BA8BB1F" box="[418,495,1266,1293]" metricMagnitude="-3" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="71" unit="in" value="0.07">0.07 in</quantity>
|
||
males and 0.76±
|
||
<quantity id="F43A9B15FFB67560FD01FB0F18BDBB1F" box="[684,762,1267,1293]" metricMagnitude="-4" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="71" unit="in" value="0.02">0.02 in</quantity>
|
||
females (vs. ratio TD/ED 0.64±
|
||
<quantity id="F43A9B15FFB67560FBCDFB0F1EE9BB1F" box="[1120,1198,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="71" unit="in" value="0.07">0.07 in</quantity>
|
||
males and 0.66±
|
||
<quantity id="F43A9B15FFB67560FAC6FB0F1AEABB23" metricMagnitude="-4" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="71" unit="in" value="0.03">0.03 in</quantity>
|
||
females); by having smaller ratio between eye diameter and tympanum-eye, with ratio ED/TED 0.62±
|
||
<quantity id="F43A9B15FFB67560FAE1FAEB1FDBBB23" box="[1356,1436,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="3.302" pageId="14" pageNumber="71" unit="in" value="0.13">0.13 in</quantity>
|
||
adult males and 1.40±
|
||
<quantity id="F43A9B15FFB67560FE21FAC71B9CBB47" box="[396,475,1339,1365]" metricMagnitude="-4" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="71" unit="in" value="0.02">0.02 in</quantity>
|
||
adult females (vs. ED/TED 0.85±
|
||
<quantity id="F43A9B15FFB67560FCF5FAC619E2BB47" box="[856,933,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.7939999999999996" pageId="14" pageNumber="71" unit="in" value="0.11">0.11 in</quantity>
|
||
adult males and 1.77±
|
||
<quantity id="F43A9B15FFB67560FB0DFAC71EA8BB47" box="[1184,1263,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.826" pageId="14" pageNumber="71" unit="in" value="0.19">0.19 in</quantity>
|
||
adult females); and by juveniles having throat surface pattern very distinct with black and white spots that covers both the throat and anterior half of venter, and latter half of ventral yellow (vs. dark spots only in throat and ventral yellow-white, in juveniles;
|
||
<figureCitation id="ABF92A75FFB67560FE85FA5B1B3DBBD3" box="[296,378,1447,1473]" captionStart="FIGURE 10" captionStartId="18.[151,250,971,995]" captionTargetBox="[157,1433,189,944]" captionTargetId="figure-23@18.[151,1436,181,947]" captionTargetPageId="18" captionText="FIGURE 10. Differentiation of ventral surface between Limnonectes splendissimus sp. nov. (A–D) and L. poilani (E–H)." figureDoi="http://doi.org/10.5281/zenodo.14734624" httpUri="https://zenodo.org/record/14734624/files/figure.png" pageId="14" pageNumber="71">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |