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<document id="E3E337F0368876700235A9B628C94614" ID-DOI="10.5281/zenodo.179195" ID-GBIF-Dataset="e0a7d367-be4e-4031-acad-42a17507fadf" ID-ISSN="1175-5326" ID-Zenodo-Dep="179195" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459941676950" checkinUser="plazi" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2007" docId="BB10DE286C3F2C6E4485FC3DFE6E2ED0" docLanguage="en" docName="zt01621p031.pdf" docOrigin="Zootaxa 1621" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hymenicoides Kemp 1917" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="4729A6506C3E2C6D4412FF97FF8C2C32" masterDocTitle="On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species" masterLastPageNumber="31" masterPageNumber="17" pageNumber="18" updateTime="1698230522269" updateUser="plazi">
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<mods:title id="3688D8354B31A6878C964CB4CE0C4400">On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species</mods:title>
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<mods:namePart id="75E54257CDB3046BD3C54C1B0E5FF315">Naruse, Tohru</mods:namePart>
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<mods:namePart id="0241820A13DB152AF9A10A001A3C1DBD">Ng, Peter K. L.</mods:namePart>
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<mods:date id="B273F5E3B5640EA18D9FCCC5A208B8F3">2007</mods:date>
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<mods:identifier id="AE4C434FCE9BF164E0F10BBCE5B34577" type="DOI">10.5281/zenodo.179195</mods:identifier>
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<treatment id="BB10DE286C3F2C6E4485FC3DFE6E2ED0" ID-DOI="http://doi.org/10.5281/zenodo.6251670" ID-GBIF-Taxon="119375995" ID-Zenodo-Dep="6251670" LSID="urn:lsid:plazi:treatment:BB10DE286C3F2C6E4485FC3DFE6E2ED0" httpUri="http://treatment.plazi.org/id/BB10DE286C3F2C6E4485FC3DFE6E2ED0" lastPageId="3" lastPageNumber="20" pageId="1" pageNumber="18">
<subSubSection id="7BA33CB56C3F2C6C4485FC3DFD1E281E" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="33066F3E6C3F2C6C4485FC3DFE522FF6" blockId="1.[151,478,882,964]" box="[151,478,938,964]" pageId="1" pageNumber="18">
<heading id="684ED8526C3F2C6C4485FC3DFE522FF6" bold="true" box="[151,478,938,964]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
<taxonomicName id="F4B914BD6C3F2C6C4485FC3DFE522FF6" ID-CoL="7PL39" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[151,478,938,964]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3F2C6C4485FC3DFE522FF6" bold="true" box="[151,478,938,964]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FC3DFECE2FF6" bold="true" box="[151,322,938,964]" italics="true" pageId="1" pageNumber="18">Hymenicoides</emphasis>
<bibRefCitation id="572812CF6C3F2C6C455BFC3DFE522FF6" author="Kemp" box="[329,478,938,964]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="33066F3E6C3F2C6C4485FC62FD1E281E" blockId="1.[151,1436,1013,1068]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FC62FECA283E" box="[151,326,1013,1036]" italics="true" pageId="1" pageNumber="18">Hymenicoides—</emphasis>
<bibRefCitation id="572812CF6C3F2C6C455EFC62FE58283E" author="Kemp" box="[332,468,1013,1036]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
: 267;
<bibRefCitation id="572812CF6C3F2C6C4607FC62FD17283E" author="Lucas" box="[533,667,1013,1036]" pageId="1" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas, 1980</bibRefCitation>
: 196;
<bibRefCitation id="572812CF6C3F2C6C46CEFC62FC35283E" author="Ng" box="[732,953,1013,1036]" pageId="1" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: 50 (part); Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C411BFC62FA14283E" author="Forges" box="[1289,1432,1013,1036]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
: 460 (part);
<bibRefCitation id="572812CF6C3F2C6C452DFB82FDD5281E" author="Guinot" box="[319,601,1045,1068]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard, 1998</bibRefCitation>
: 685.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6C4485FBCAFB5C2845" box="[151,1232,1117,1143]" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="33066F3E6C3F2C6C4485FBCAFB5C2845" blockId="1.[151,1437,1117,2023]" box="[151,1232,1117,1143]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FBCAFEBE2845" bold="true" box="[151,306,1117,1143]" pageId="1" pageNumber="18">
<typeStatus id="EC02D19C6C3F2C6C4485FBCAFF582845" box="[151,212,1117,1143]" pageId="1" pageNumber="18">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="F4B914BD6C3F2C6C4552FBCAFD542845" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[320,728,1117,1143]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C4552FBCAFDB12844" box="[320,573,1117,1142]" italics="true" pageId="1" pageNumber="18">Hymenicoides carteri</emphasis>
<bibRefCitation id="572812CF6C3F2C6C4657FBCAFD542845" author="Kemp" box="[581,728,1117,1143]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</taxonomicName>
, by monotypy; gender of genus masculine.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6C44D4FB12FE702A4D" pageId="1" pageNumber="18" type="diagnosis">
<paragraph id="33066F3E6C3F2C6C44D4FB12FE702A4D" blockId="1.[151,1437,1117,2023]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C44D4FB12FECA28AD" bold="true" box="[198,326,1157,1183]" pageId="1" pageNumber="18">Diagnosis.</emphasis>
Carapace oval, dorsal surface concave; grooves distinct; rostrum absent or very weak; antenna with base of basal article placed posterior-inner part of base of eye; eyes, antennae, antennules visible dorsally; third maxillipeds narrow, not covering more than three-quarters of mouth field when closed, merus rectangular, dactylus styliform, approximately twice length of propodus; male chelae relatively stout, manus with high, prominent, dorso-ventrally compressed tubercle on distal part of outer surface, tubercle absent in females. Female vulvae placed on imaginary line joining inner ends of sutures between sternites 5 and 6 on medially fused plate of thoracic sternum, vulva with basal mount. Male abdomen-pleotelson without fused segments, male pleotelson distinctly trilobed, inner surface thickened externally, forming socket for sternal button (
<figureCitation id="AB8273BB6C3F2C6C44E1FA52FEBB29ED" box="[243,311,1477,1503]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
a). G1 stout, bent outwards medially; distal part with prominent distal inner processes, swollen distal outer angle, long, dorsal bursiform projections. Female abdomen with 6 distinctly demarcated segments, boundary between first and second segments (
<taxonomicName id="F4B914BD6C3F2C6C470BF982FC032A1C" box="[793,911,1557,1582]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C470BF982FC032A1C" box="[793,911,1557,1582]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
) or between second and third segments (
<taxonomicName id="F4B914BD6C3F2C6C416DF982FF7E2A64" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C3F2C6C416DF982FF7E2A64" italics="true" pageId="1" pageNumber="18">H. robertsi</emphasis>
</taxonomicName>
) movable; long, biramous pleopods on second to fifth segments, developed from distal outer end of inner surface of each segment.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6F44D4F91AFBFF2DF8" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="discussion">
<paragraph id="33066F3E6C3F2C6C44D4F91AFDF62BA5" blockId="1.[151,1437,1117,2023]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C44D4F91AFEB42A95" bold="true" box="[198,312,1677,1703]" pageId="1" pageNumber="18">Remarks</emphasis>
. The distinctive trilobate pleotelson of
<taxonomicName id="F4B914BD6C3F2C6C4714F91AFCF62A94" box="[774,890,1677,1702]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C4714F91AFCF62A94" box="[774,890,1677,1702]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
has received some attention from taxonomists due to its unusual shape.
<bibRefCitation id="572812CF6C3F2C6C45D5F922FD2E2AFD" author="Lucas" box="[455,674,1717,1743]" pageId="1" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas (1980: 197)</bibRefCitation>
regarded that the lateral lobe of male pleotelson as “a cavity to receive the apice of the first pleopod”. Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C4746F94AFBBA2AC5" author="Forges" box="[852,1078,1757,1783]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges (1997: 470)</bibRefCitation>
suggested that the lateral lobe of the pleotelson may be derived from the inserted plate (see Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C4067F892FA992B2D" author="Forges" box="[1141,1301,1797,1823]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
:
<figureCitation id="AB8273BB6C3F2C6C4130F892FAE62B2D" box="[1314,1386,1797,1823]" captionStart="FIGURE 6" captionStartId="8.[151,255,1506,1530]" captionTargetBox="[151,1436,184,1485]" captionTargetId="figure@8.[151,1436,184,1485]" captionTargetPageId="8" captionText="FIGURE 6. Hymenicoides robertsi new species. a – d, right second to fifth pleopods; e, exopods. Paratype, ovigerous female (ZRC 2007.0113), 4.2 × 5.0 mm." httpUri="https://zenodo.org/record/179201/files/figure.png" pageId="1" pageNumber="18">Fig. 6</figureCitation>
, for mobile intercalated platelet of
<taxonomicName id="F4B914BD6C3F2C6C461BF8BAFD682B74" box="[521,740,1837,1862]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="01CDB32C6C3F2C6C461BF8BAFD682B74" box="[521,740,1837,1862]" italics="true" pageId="1" pageNumber="18">Odiomaris pilosus</emphasis>
</taxonomicName>
).
<bibRefCitation id="572812CF6C3F2C6C4712F8BAFB0A2B75" author="Guinot" box="[768,1158,1837,1863]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard (1998: 685)</bibRefCitation>
subsequently hypothesized that the abdominal socket, which is placed on the inner surface of the lateral lobe (or the intercalated platelet), is homologous with the uropod.
</paragraph>
<paragraph id="33066F3E6C3F2C6F44D4F832FBFF2DF8" blockId="1.[151,1437,1117,2023]" lastBlockId="2.[151,1436,152,458]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
Our examination of the
<typeStatus id="EC02D19C6C3F2C6C45F3F832FD9F2B8D" box="[481,531,1957,1983]" pageId="1" pageNumber="18">type</typeStatus>
material of
<taxonomicName id="F4B914BD6C3F2C6C46B1F832FC942B8C" box="[675,792,1957,1982]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C46B1F832FC942B8C" box="[675,792,1957,1982]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
and
<taxonomicName id="F4B914BD6C3F2C6C4740F832FC5E2B8C" box="[850,978,1957,1982]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3F2C6C4740F832FC5E2B8C" box="[850,978,1957,1982]" italics="true" pageId="1" pageNumber="18">H. robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3F2C6C47C8F832FBE62B8D" bold="true" box="[986,1130,1957,1983]" pageId="1" pageNumber="18">
<taxonomicNameLabel id="1AFE0E576C3F2C6C47C8F832FBE62B8D" box="[986,1130,1957,1983]" pageId="1" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
, shows that the inner surface of the lateral lobe of the male pleotelson is domed and that it is externally surrounded by a thickened rim, forming a semicircular cavity (
<figureCitation id="AB8273BB6C3C2C6F461AFF0FFDC12C80" box="[520,589,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
b). When the abdomen is closed, the cavity overlays the sternal button (
<figureCitation id="AB8273BB6C3C2C6F448DFF57FF6A2CE8" box="[159,230,192,218]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
a). This does not support the premise that the lateral lobe of the pleotelson is to accommodate the unusual tip of the G1 (
<figureCitation id="AB8273BB6C3C2C6F458EFF7FFE622D30" box="[412,494,232,258]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="2" pageNumber="19">Figs. 2</figureCitation>
a, b, 5c, d). In fact, even when the abdomen is open, the G1 can easily be accommodated within the sterno-abdominal cavity (
<figureCitation id="AB8273BB6C3C2C6F46BDFE87FD7E2D18" box="[687,754,272,298]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
a). The sternal buttons and the lateral cavities of the pleotelson function instead to “lock” the abdomen with the thorax. It should be noted that the G1 of
<taxonomicName id="F4B914BD6C3C2C6F4134FEAFFA102D63" box="[1318,1436,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3C2C6F4134FEAFFA102D63" box="[1318,1436,312,337]" italics="true" pageId="2" pageNumber="19">H. carteri</emphasis>
</taxonomicName>
closely resembles that of
<taxonomicName id="F4B914BD6C3C2C6F45DAFEF7FD732D4B" box="[456,767,352,377]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="xenomorpha">
<emphasis id="01CDB32C6C3C2C6F45DAFEF7FD732D4B" box="[456,767,352,377]" italics="true" pageId="2" pageNumber="19">Cancrocaeca xenomorpha</emphasis>
</taxonomicName>
(see also Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3C2C6F4062FEF7FA822D48" author="Forges" box="[1136,1294,352,378]" pageId="2" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
), and in the latter species, the male abdomen is more typical in shape, lacking the trilobite pleotelson, as well as sternal buttons, seen in
<taxonomicName id="F4B914BD6C3C2C6F4547FE27FE772DFB" box="[341,507,432,457]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3C2C6F4547FE27FE772DFB" box="[341,507,432,457]" italics="true" pageId="2" pageNumber="19">Hymenicoides</emphasis>
</taxonomicName>
(Naruse, T., Ng, P.K.L. &amp; Guinot, D., in manuscript).
</paragraph>
</subSubSection>
<caption id="67C63FB66C3C2C6F4485F8EAFB212BC7" httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19" targetBox="[268,1318,501,1901]" targetPageId="2">
<paragraph id="33066F3E6C3C2C6F4485F8EAFB212BC7" blockId="2.[151,1436,1917,2037]" pageId="2" pageNumber="19">
<emphasis id="01CDB32C6C3C2C6F4485F8EAFE952BA7" bold="true" box="[151,281,1917,1941]" pageId="2" pageNumber="19">FIGURE 1.</emphasis>
Male abdominal cavity and pleotelson of
<taxonomicName id="F4B914BD6C3C2C6F46CFF8E9FC5C2BA7" box="[733,976,1918,1941]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F46CFF8E9FC5C2BA7" box="[733,976,1918,1941]" italics="true" pageId="2" pageNumber="19">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F47C4F8EAFBD52BA7" bold="true" box="[982,1113,1917,1941]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F47C4F8EAFBD52BA7" box="[982,1113,1917,1941]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="F4B914BD6C3C2C6F4086F8E9FA802BA7" box="[1172,1292,1918,1941]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3C2C6F4086F8E9FA802BA7" box="[1172,1292,1918,1941]" italics="true" pageId="2" pageNumber="19">Limnopilos</emphasis>
</taxonomicName>
species. a, b,
<taxonomicName id="F4B914BD6C3C2C6F4485F809FE042B87" box="[151,392,1950,1973]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F4485F809FE042B87" box="[151,392,1950,1973]" italics="true" pageId="2" pageNumber="19">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F4583F80AFD992B87" bold="true" box="[401,533,1949,1973]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F4583F80AFD992B87" box="[401,533,1949,1973]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d,
<taxonomicName id="F4B914BD6C3C2C6F403FF809FA102B87" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[1069,1436,1950,1973]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C3C2C6F403FF809FB352B87" box="[1069,1209,1950,1973]" italics="true" pageId="2" pageNumber="19">L. naiyanetri</emphasis>
Chuang &amp; Ng, 1991
</taxonomicName>
(ZRC 1991.6521–6555, 4.9 × 5.3 mm); e, f,
<taxonomicName id="F4B914BD6C3C2C6F4666F829FC802BE7" box="[628,780,1982,2005]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F4666F829FC802BE7" box="[628,780,1982,2005]" italics="true" pageId="2" pageNumber="19">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F4706F82AFC1B2BE7" bold="true" box="[788,919,1981,2005]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F4706F82AFC1B2BE7" box="[788,919,1981,2005]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G1; b, d, f, inner surface of male pleotelson.
</paragraph>
</caption>
<subSubSection id="7BA33CB56C3D2C6E44D4FF0FFBCB2EA0" pageId="3" pageNumber="20" type="reference_group">
<paragraph id="33066F3E6C3D2C6E44D4FF0FFC552DC0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
<taxonomicName id="F4B914BD6C3D2C6E44D4FF0FFD442C80" ID-CoL="72CL4" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[198,712,152,178]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C3D2C6E44D4FF0FFE402C83" box="[198,460,152,177]" italics="true" pageId="3" pageNumber="20">Limnopilos naiyanetri</emphasis>
<bibRefCitation id="572812CF6C3D2C6E45C6FF0FFD442C80" author="Chuang" box="[468,712,152,178]" pageId="3" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
(
<typeStatus id="EC02D19C6C3D2C6E46CAFF0FFC802C80" box="[728,780,152,178]" pageId="3" pageNumber="20">type</typeStatus>
species of
<taxonomicName id="F4B914BD6C3D2C6E4786FF0FFA942C80" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[916,1304,152,178]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4786FF0FFB942C83" box="[916,1048,152,177]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
<bibRefCitation id="572812CF6C3D2C6E4033FF0FFA942C80" author="Chuang" box="[1057,1304,152,178]" pageId="3" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
),
<taxonomicName id="F4B914BD6C3D2C6E413CFF0FFE062CE8" ID-CoL="8WB9Z" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="microrhynchus" status="comb. nov.">
<emphasis id="01CDB32C6C3D2C6E413CFF0FFE8E2CEB" italics="true" pageId="3" pageNumber="20">L. microrhynchus</emphasis>
(Ng, 1995)
</taxonomicName>
<taxonomicNameLabel id="1AFE0E576C3D2C6E4580FF57FDED2CE8" box="[402,609,192,218]" pageId="3" pageNumber="20">new combination</taxonomicNameLabel>
, and
<taxonomicName id="F4B914BD6C3D2C6E468CFF57FCCB2CEB" ID-CoL="6QCKT" box="[670,839,192,217]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C3D2C6E468CFF57FCCB2CEB" box="[670,839,192,217]" italics="true" pageId="3" pageNumber="20">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3D2C6E475CFF28FC522CEB" bold="true" box="[846,990,191,217]" pageId="3" pageNumber="20">
<taxonomicNameLabel id="1AFE0E576C3D2C6E475CFF28FC522CEB" box="[846,990,191,217]" pageId="3" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
, all possess the sternal button and the pleotelson lateral cavity (
<figureCitation id="AB8273BB6C3D2C6E45ADFF7FFD8C2D30" box="[447,512,232,258]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
c–f). The lateral cavities of these three species, however, are relatively smaller and the rim is more distal in position on the cavity when compared with those of
<taxonomicName id="F4B914BD6C3D2C6E4071FE87FB542D1B" ID-CoL="3NJ2G" box="[1123,1240,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E4071FE87FB542D1B" box="[1123,1240,272,297]" italics="true" pageId="3" pageNumber="20">H. carteri</emphasis>
</taxonomicName>
and
<taxonomicName id="F4B914BD6C3D2C6E4106FE87FA182D1B" ID-CoL="3NJ2J" box="[1300,1428,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C3D2C6E4106FE87FA182D1B" box="[1300,1428,272,297]" italics="true" pageId="3" pageNumber="20">H. robertsi</emphasis>
</taxonomicName>
. This may be due to the different shape of the G1 of
<taxonomicName id="F4B914BD6C3D2C6E46ECFEAFFC282D63" ID-CoL="7PL39" box="[766,932,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E46ECFEAFFC282D63" box="[766,932,312,337]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
. The lateral cavity of
<taxonomicName id="F4B914BD6C3D2C6E40B9FEAFFADD2D63" ID-CoL="7PL39" box="[1195,1361,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E40B9FEAFFADD2D63" box="[1195,1361,312,337]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
needs to be thickened along the external margins to effectively engage the sternal button, three sides (anterior, inner and posterior) which are surrounded by the G
<quantity id="F441C2DB6C3D2C6E46A9FE1FFD6A2D93" box="[699,742,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="20" unit="in" value="1.0">
1
<emphasis id="01CDB32C6C3D2C6E46DDFE1FFD6A2D93" box="[719,742,392,417]" italics="true" pageId="3" pageNumber="20">in</emphasis>
</quantity>
situ (
<figureCitation id="AB8273BB6C3D2C6E4739FE1FFCE32D90" box="[811,879,392,418]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
a). However, the sternal buttons of
<taxonomicName id="F4B914BD6C3D2C6E410AFE1FFA102D93" ID-CoL="7PMTG" box="[1304,1436,392,417]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E410AFE1FFA102D93" box="[1304,1436,392,417]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
are placed more distantly from the distal outer angle of the G1 (
<figureCitation id="AB8273BB6C3D2C6E4769FE27FC312DF8" box="[891,957,432,458]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 – 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
c, e), and allow the lateral cavity to more easily lock onto the sternal button by way of the distally thickened rim.
</paragraph>
<paragraph id="33066F3E6C3D2C6E44D4FD97FBCB2EA0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
The G1s of the
<taxonomicName id="F4B914BD6C3D2C6E4594FD97FDA02E2B" ID-CoL="7PL39" box="[390,556,512,537]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4594FD97FDA02E2B" box="[390,556,512,537]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
species (
<figureCitation id="AB8273BB6C3D2C6E468EFD97FD692E28" box="[668,741,512,538]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
a, b, 5c, d) differ from those of
<taxonomicName id="F4B914BD6C3D2C6E4075FD97FB672E2B" ID-CoL="7PMTG" box="[1127,1259,512,537]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4075FD97FB672E2B" box="[1127,1259,512,537]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
in several key aspects: they have a proportionately stouter shaft (vs. moderately stout in
<taxonomicName id="F4B914BD6C3D2C6E47EEFDBFFB0C2E73" ID-CoL="7PMTG" box="[1020,1152,552,577]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E47EEFDBFFB0C2E73" box="[1020,1152,552,577]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
), more prominent distal inner processes (vs. moderately stout in
<taxonomicName id="F4B914BD6C3D2C6E467EFDC7FD7C2E5B" ID-CoL="7PMTG" box="[620,752,592,617]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E467EFDC7FD7C2E5B" box="[620,752,592,617]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
) and with the distal outer angle more swollen than in
<taxonomicName id="F4B914BD6C3D2C6E4176FDC7FF612EA3" ID-CoL="7PMTG" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4176FDC7FF612EA3" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
species (
<figureCitation id="AB8273BB6C3D2C6E454BFDEFFE122EA0" box="[345,414,632,658]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="3" pageNumber="20">Fig. 8</figureCitation>
c;
<bibRefCitation id="572812CF6C3D2C6E45A8FDEFFD212EA0" author="Ng" box="[442,685,632,658]" pageId="3" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: Fig. 21H; Ng, 1995: Fig. 14A, B).
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3D2C6E44D4FD37FE6E2ED0" pageId="3" pageNumber="20" type="description">
<paragraph id="33066F3E6C3D2C6E44D4FD37FE6E2ED0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
The marked differences in the chela, abdomen and G1 lead us to now conclude that
<taxonomicName id="F4B914BD6C3D2C6E408DFD37FAAF2E8B" box="[1183,1315,672,697]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E408DFD37FAAF2E8B" box="[1183,1315,672,697]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
should be resurrected as a valid genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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