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<document id="5E5D19355C7707FB02723E7747E6DAA6" ID-CLB-Dataset="53759" ID-DOI="10.6620/ZS.2018.57-18" ID-GBIF-Dataset="cff4ab26-6fdc-462b-aaa0-b8e8f87afd71" ID-PMC="PMC6624643" ID-PubMed="31966258" ID-Zenodo-Dep="8064335" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="tatiana" checkinTime="1687182456079" checkinUser="valdenar" docAuthor="Yoshida 1, Takahiro, Karner, Michael &amp; Hirowatari, Toshiya" docDate="2018" docId="BA4924401246FFFEFF1FF9DFFE6FA836" docLanguage="en" docName="ZoolStud.57.18.1-18.pdf" docOrigin="Zoological studies 57 (18)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Psammoecus harmandi , Grouvelle 1912" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="46705C38124FFFF5FFA6FFC2FF9FA960" masterDocTitle="A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="10" updateTime="1716449326147" updateUser="tatiana">
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<mods:title id="92DE247CFF6F609848CBB16AFE635D21">A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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<mods:namePart id="C9007DB61A16BF61952C75DB3E4090AC">Yoshida 1, Takahiro</mods:namePart>
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<mods:namePart id="6EAE983226014F7273D0C39EF9851696">Karner, Michael</mods:namePart>
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<treatment id="BA4924401246FFFEFF1FF9DFFE6FA836" ID-DOI="http://doi.org/10.5281/zenodo.8064345" ID-GBIF-Taxon="209705814" ID-Zenodo-Dep="8064345" LSID="urn:lsid:plazi:treatment:BA4924401246FFFEFF1FF9DFFE6FA836" httpUri="http://treatment.plazi.org/id/BA4924401246FFFEFF1FF9DFFE6FA836" lastPageId="11" lastPageNumber="11" pageId="9" pageNumber="10">
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<heading id="6917223A1246FFFCFF1FF9DFFD2FAF56" bold="true" box="[185,688,1564,1590]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="F5E0EED51246FFFCFF1FF9DFFD2FAF56" authority="Grouvelle, 1912" authorityName=", Grouvelle" authorityYear="1912" box="[185,688,1564,1590]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441246FFFCFF1FF9DFFD2FAF56" bold="true" box="[185,688,1564,1590]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFF1FF9DFFE7BAF56" bold="true" box="[185,484,1565,1590]" italics="true" pageId="9" pageNumber="10">Psammoecus harmandi</emphasis>
<bibRefCitation id="5671E8A71246FFFCFE4DF9DEFD2FAF56" author="Grouvelle A." box="[491,688,1564,1590]" pageId="9" pageNumber="10" pagination="81 - 94" refId="ref12816" refString="Grouvelle A. 1912. Psammoecus nouveaux du Musee de Leide. Notes from the Leyden Museum 34 (2): 81 - 94." type="journal article" year="1912">Grouvelle, 1912</bibRefCitation>
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<paragraph id="325F95561246FFFCFE96F9FFFDA6AF36" blockId="9.[185,688,1564,1622]" box="[304,569,1596,1622]" pageId="9" pageNumber="10">
(
<figureCitation id="AADB89D31246FFFCFE91F9FFFE00AF36" box="[311,415,1596,1622]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="9" pageNumber="10">Figs. 1C</figureCitation>
,
<figureCitation id="AADB89D31246FFFCFE0DF9FFFE26AF36" box="[427,441,1597,1622]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
D-F and 7)
</paragraph>
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<treatmentCitation id="B341B3471246FFFCFF2EF9BAFDCDAFEC" author="Grouvelle A." box="[136,594,1655,1676]" page="413" pageId="9" pageNumber="10" year="1912">
<taxonomicName id="F5E0EED51246FFFCFF2EF9BAFDCDAFEC" authority="Grouvelle, 1912: 413" authorityName=", Grouvelle" authorityPageNumber="413" authorityYear="1912" box="[136,594,1655,1676]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441246FFFCFF2EF9BAFEEDAFEC" box="[136,370,1656,1676]" italics="true" pageId="9" pageNumber="10">Psammoecus harmandi</emphasis>
<bibRefCitation id="5671E8A71246FFFCFEDAF9B5FDCDAFEC" author="Grouvelle A." box="[380,594,1655,1676]" pageId="9" pageNumber="10" pagination="81 - 94" refId="ref12816" refString="Grouvelle A. 1912. Psammoecus nouveaux du Musee de Leide. Notes from the Leyden Museum 34 (2): 81 - 94." type="journal article" year="1912">Grouvelle, 1912: 413</bibRefCitation>
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.
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locality:
<collectingCountry id="4AF7D5C61246FFFCFF1BF950FF74AFC6" box="[189,235,1682,1702]" name="India" pageId="9" pageNumber="10">India</collectingCountry>
(Darjeeling). -
<treatmentCitation id="B341B3471246FFFCFEDBF950FE65AFC6" author="Pal TK" box="[381,506,1682,1702]" page="19" pageId="9" pageNumber="10" year="1985">
<bibRefCitation id="5671E8A71246FFFCFEDBF950FE65AFC6" author="Pal TK" box="[381,506,1682,1702]" pageId="9" pageNumber="10" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal 1985: 19</bibRefCitation>
</treatmentCitation>
, figs 5, 32. -
<treatmentCitation id="B341B3471246FFFCFDDCF950FEF1AFA1" author="Sen Gupta T &amp; Pal TK" page="170" pageId="9" pageNumber="10" year="1996">
<bibRefCitation id="5671E8A71246FFFCFDDCF950FEF1AFA1" author="Sen Gupta T &amp; Pal TK" pageId="9" pageNumber="10" refId="ref13823" refString="Sen Gupta T, Pal TK. 1996. Fauna of India and the Adjacent Countries. Calvicornia [sic]: Coleoptera. Family Silvanidae. Zoological Survey of India, Calcutta." type="book" year="1996">Sen Gupta and Pal 1996: 170</bibRefCitation>
</treatmentCitation>
, figs 242, 269.
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</paragraph>
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<treatmentCitationGroup id="12F0B2781246FFFCFF2EF90AFDDEAF96" pageId="9" pageNumber="10">
<treatmentCitation id="B341B3471246FFFCFF2EF90AFD36AFBC" LSID="urn:lsid:zoobank.org:act:1E58DE65-FC6D-4E40-B55E-A952B311A454" author="Yoshida T &amp; Hirowatari T." box="[136,681,1736,1756]" httpUri="http://treatment.plazi.org/id/497581866A8D1AB91E01FDEF89691858" page="413" pageId="9" pageNumber="10" year="2014">
<taxonomicName id="F5E0EED51246FFFCFF2EF90AFD36AFBC" authority="Yoshida and Hirowatari, 2014: 413" authorityName="Yoshida and Hirowatari" authorityPageNumber="413" authorityYear="2014" box="[136,681,1736,1756]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="009449441246FFFCFF2EF90AFECDAFBC" box="[136,338,1736,1756]" italics="true" pageId="9" pageNumber="10">Psammoecus boreas</emphasis>
<bibRefCitation id="5671E8A71246FFFCFEFFF90AFD36AFBC" author="Yoshida T &amp; Hirowatari T." box="[345,681,1736,1756]" pageId="9" pageNumber="10" pagination="15 - 45" refId="ref13985" refString="Yoshida T, Hirowatari T. 2014. A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae). ZooKeys 403: 15 - 45. doi: 10.3897 / zookeys. 403.7145." type="journal article" year="2014">Yoshida and Hirowatari, 2014: 413</bibRefCitation>
</taxonomicName>
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.
<typeStatus id="ED5B2BF41246FFFCFD15F90AFD7EAFBC" box="[691,737,1736,1756]" pageId="9" pageNumber="10">Type</typeStatus>
locality:
<collectingCountry id="4AF7D5C61246FFFCFEACF920FED8AF96" box="[266,327,1762,1782]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
;
<typeStatus id="ED5B2BF41246FFFCFEF6F920FEE1AF96" box="[336,382,1762,1782]" pageId="9" pageNumber="10">Type</typeStatus>
examined. syn. nov.
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="12F0B2781246FFFCFF2EF93FFD5AAE71" box="[136,709,1789,1809]" pageId="9" pageNumber="10">
<treatmentCitation id="B341B3471246FFFCFF2EF93FFDB4AE71" author="Nakane T." box="[136,555,1789,1809]" page="195" pageId="9" pageNumber="10" year="1963">
<taxonomicName id="F5E0EED51246FFFCFF2EF93FFDB4AE71" authority=": Nakane 1963: 195" authorityName="Nakane" authorityPageNumber="195" authorityYear="1963" box="[136,555,1789,1809]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="009449441246FFFCFF2EF93FFEF2AE71" box="[136,365,1789,1809]" italics="true" pageId="9" pageNumber="10">Psammoecus triguttatus</emphasis>
:
<bibRefCitation id="5671E8A71246FFFCFEDFF93FFDB4AE71" author="Nakane T." box="[377,555,1789,1809]" pageId="9" pageNumber="10" pagination="2" refId="ref13568" refString="Nakane T. 1963. Silvanidae. In: Nakane T, Ohbayashi K, Nomura S, Kurosawa Y (eds) Iconographia Insectorum Japonicorum Colore naturali edita. Vol. 2 (Coleoptera), Hokuryukan, Tokyo. (in Japanese)" type="book chapter" year="1963">Nakane 1963: 195</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig.
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pl. 98.
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</paragraph>
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<taxonomicName id="F5E0EED51246FFFCFF2EF8DAFED6AE4C" box="[136,329,1816,1836]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="009449441246FFFCFF2EF8DAFE93AE4C" box="[136,268,1816,1836]" italics="true" pageId="9" pageNumber="10">Psammoecus</emphasis>
sp. 3
</taxonomicName>
,
<treatmentCitation id="B341B3471246FFFCFEF3F8DAFE68AE4C" author="Hirano Y." box="[341,503,1816,1836]" page="63" pageId="9" pageNumber="10" year="2009">
<bibRefCitation id="5671E8A71246FFFCFEF3F8DAFE68AE4C" author="Hirano Y." box="[341,503,1816,1836]" pageId="9" pageNumber="10" pagination="57 - 83" refId="ref13024" refString="Hirano Y. 2009. Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite). Kanagawa-chuho 168: 57 - 83. (in Japanese)" type="journal article" year="2009">Hirano 2009: 63</bibRefCitation>
</treatmentCitation>
, 66, 67, fig. 8. -
<treatmentCitation id="B341B3471246FFFCFD07F8DAFE8CAE26" author="Hirano Y." page="12" pageId="9" pageNumber="10" year="2010">
<bibRefCitation id="5671E8A71246FFFCFD07F8DAFE8CAE26" author="Hirano Y." pageId="9" pageNumber="10" refId="ref13051" refString="Hirano Y. 2010. Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae. Roppon-Ashi Entomological Books, Tokyo. (in Japanese, with English title)" type="book" year="2010">Hirano 2010: 12</bibRefCitation>
</treatmentCitation>
, 16.
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</paragraph>
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<paragraph id="325F95561246FFFCFF63F8AFFBB0AD96" blockId="9.[136,737,1901,1959]" lastBlockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFF63F8AFFE72AEE7" box="[197,493,1901,1927]" italics="true" pageId="9" pageNumber="10">S p e c i m e n s e x a m i n e d</emphasis>
: J A PA N:
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a l e, Yoshin, Tanzawa, Kanagawa Prefecture,
<date id="465EB3961246FFFCFD3BF84CFCF1A876" box="[669,878,253,1959]" pageId="9" pageNumber="10" value="1989-05-26">26-V-1989</date>
, Y. Hirano leg. (EUMJ; formerly
<typeStatus id="ED5B2BF41246FFFCFB5BFF3FFAFBA876" box="[1277,1380,253,278]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
of
<taxonomicName id="F5E0EED51246FFFCFC8BFEDFFBA7A856" authorityName="Yoshida and Hirowatari" authorityYear="2014" box="[813,1080,285,310]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="009449441246FFFCFC8BFEDFFBA7A856" box="[813,1080,285,310]" italics="true" pageId="9" pageNumber="10">Psammoecus boreas</emphasis>
</taxonomicName>
).
<collectingCountry id="4AF7D5C61246FFFCFBF3FEDFFB5CA856" box="[1109,1219,285,310]" name="Taiwan" pageId="9" pageNumber="10">TAIWAN</collectingCountry>
: [
<collectingRegion id="F0245BB41246FFFCFB7EFEDFFAE0A857" box="[1240,1407,285,311]" country="Taiwan" name="Yilan" pageId="9" pageNumber="10">Yilan County</collectingRegion>
]
<specimenCount id="24E65EDF1246FFFCFC8BFEFFFC1CA836" box="[813,899,317,342]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
&amp;
<specimenCount id="24E65EDF1246FFFCFC0DFEFFFC71A836" box="[939,1006,317,342]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, Taipingshan, Datong Township, alt.
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,
<date id="465EB3961246FFFCFC6BFE9FFBD6A816" box="[973,1097,349,374]" pageId="9" pageNumber="10" value="2004-04-01">1-IV-2004</date>
, T. Kurihara leg. (EUMJ). [
<collectingRegion id="F0245BB41246FFFCFC94FEBFFC70A8F7" box="[818,1007,381,407]" country="Taiwan" name="Hualien" pageId="9" pageNumber="10">Hualien County</collectingRegion>
]
<specimenCount id="24E65EDF1246FFFCFC5BFEBFFBA3A8F6" box="[1021,1084,381,406]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, Piluhsi For. Rec. Area,
<date id="465EB3961246FFFCFAF9FEBFFC19A8D6" pageId="9" pageNumber="10" value="1999-05-22">22- V-1999</date>
,
<quantity id="F51838B31246FFFCFC34FE5FFC6EA8D6" box="[914,1009,413,438]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="9" pageNumber="10" unit="m" value="2150.0">2150 m</quantity>
, S. E. Halbert &amp; C. W. L. O’Brien leg., Night Beating (FSCA). [
<collectingRegion id="F0245BB41246FFFCFBD9FE7FFAA9A8B7" box="[1151,1334,445,471]" country="Taiwan" name="Nantou" pageId="9" pageNumber="10">Nantou County</collectingRegion>
]
<specimenCount id="24E65EDF1246FFFCFAE5FE7FFA1DA8B6" box="[1347,1410,445,470]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, Pi Lu Chieh, alt.
<quantity id="F51838B31246FFFCFC5DFE1FFBC5A896" box="[1019,1114,477,502]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="10" unit="m" value="2400.0">2400 m</quantity>
,
<date id="465EB3961246FFFCFBCEFE1FFB7DA896" box="[1128,1250,477,502]" pageId="9" pageNumber="10" value="1986-09-06">6-IX-1986</date>
, K. Baba leg. (EUMJ);
<specimenCount id="24E65EDF1246FFFCFC3CFE3FFC43AB76" box="[922,988,509,534]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, Gaofeng,
<date id="465EB3961246FFFCFBC2FE3FFB70AB76" box="[1124,1263,509,534]" pageId="9" pageNumber="10" value="2009-05-07" valueMax="2009-05-08" valueMin="2009-05-07">7-8-V-2009</date>
, J. Aoki leg. (ELKU); 4 exs, Wushe, Ren-ai Township,
<date id="465EB3961246FFFCFA9AFDDFFCF3AB36" pageId="9" pageNumber="10" value="1983-03-16">16-III-1983</date>
, H. &amp; M. Townes leg. (FSCA); 2 exs, Meifeng, Ren-ai Township,
<date id="465EB3961246FFFCFBA5FD9FFBECAB16" box="[1027,1139,605,630]" pageId="9" pageNumber="10" value="1983-05-03">3-V-1983</date>
, Henry K. Townes leg., Insect Flight Trap. (FSCA);
<specimenCount id="24E65EDF1246FFFCFB21FDBFFB70ABF6" box="[1159,1263,637,662]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
,
<specimenCount id="24E65EDF1246FFFCFB5BFDBFFAF6ABF6" box="[1277,1385,637,662]" pageId="9" pageNumber="10" type="female">1 female</specimenCount>
&amp; 16 exs, Tun-yuan-trail, Jing Ying Village, Ren-ai Township,
<date id="465EB3961246FFFCFC14FD7FFBA7ABB6" box="[946,1080,701,726]" pageId="9" pageNumber="10" value="2014-07-05">5-VII-2014</date>
, T. Yoshida leg. (ELKU);
<specimenCount id="24E65EDF1246FFFCFAD0FD7FFCF8AB96" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
&amp; 4 exs, same locality,
<date id="465EB3961246FFFCFB26FD1FFA9EAB96" box="[1152,1281,733,758]" pageId="9" pageNumber="10" value="2014-07-06">6-VII-2014</date>
, T. Yoshida leg. (ELKU);
<specimenCount id="24E65EDF1246FFFCFC77FD3FFB8CAA76" box="[977,1043,765,790]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
,
<geoCoordinate id="57D4F3911246FFFCFB84FD3FFB5CAA76" box="[1058,1219,765,790]" degrees="24" direction="north" minutes="02" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="53.0" value="24.048056">N24°02'53.0&quot;</geoCoordinate>
<geoCoordinate id="57D4F3911246FFFCFB68FD3FFA1DAA76" box="[1230,1410,765,790]" degrees="121" direction="east" minutes="12" orientation="longitude" pageId="9" pageNumber="10" precision="1" seconds="55.5" value="121.215416">E121°12'55.5&quot;</geoCoordinate>
, Ren-ai Township,
<date id="465EB3961246FFFCFBADFCDFFB08AA56" box="[1035,1175,797,822]" pageId="9" pageNumber="10" value="2008-08-06">6-VIII-2008</date>
, Barclay, Mendel &amp; Ewers leg. (BMNH; studied by MK). [
<collectingRegion id="F0245BB41246FFFCFB7EFCFFFA1EAA37" box="[1240,1409,829,855]" country="Taiwan" name="Chiayi" pageId="9" pageNumber="10">Chiayi County</collectingRegion>
]
<specimenCount id="24E65EDF1246FFFCFC8BFC9FFCF2AA16" box="[813,877,861,886]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, Fenchihu, Zhuqi Township,
<date id="465EB3961246FFFCFB6AFC9FFAC1AA16" box="[1228,1374,861,886]" pageId="9" pageNumber="10" value="1966-07-10">10-VII-1966</date>
, H. Kamiya leg. (KUM); 5 exs, same locality,
<date id="465EB3961246FFFCFA95FCBFFCF2AAD6" pageId="9" pageNumber="10" value="1966-07-16">16-VII-1966</date>
, H. Kamiya leg. (KUM); 2 exs, same locality,
<date id="465EB3961246FFFCFC8BFC7FFC20AAB6" box="[813,959,957,982]" pageId="9" pageNumber="10" value="1966-07-23">23-VII-1966</date>
, H. Kamiya leg. (KUM); 7 exs, same locality,
<date id="465EB3961246FFFCFC2BFC1FFB82AA96" box="[909,1053,989,1014]" pageId="9" pageNumber="10" value="1966-07-24">24-VII-1966</date>
, H. Kamiya leg. (KUM);
<specimenCount id="24E65EDF1246FFFCFAE4FC1FFA1DAA96" box="[1346,1410,989,1014]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, same locality,
<date id="465EB3961246FFFCFC7EFC3FFBF7AD76" box="[984,1128,1021,1046]" pageId="9" pageNumber="10" value="1966-07-27">27-VII-1966</date>
, H. Kamiya leg. (KUM);
<specimenCount id="24E65EDF1246FFFCFC8BFBDFFCF1AD56" box="[813,878,1053,1078]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
, same locality,
<date id="465EB3961246FFFCFB88FBDFFB5FAD56" box="[1070,1216,1053,1078]" pageId="9" pageNumber="10" value="1966-07-28">28-VII-1966</date>
, H. Kamiya leg. (KUM). [
<collectingRegion id="F0245BB41246FFFCFC3AFBFFFBF3AD37" box="[924,1132,1085,1111]" country="Taiwan" name="Taitung" pageId="9" pageNumber="10">Taitung County</collectingRegion>
] 5 exs, Taimali-shan (Mt.), Jinfeng Township, alt. ca.
<quantity id="F51838B31246FFFCFB11FB9FFA87AD16" box="[1207,1304,1117,1142]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="10" unit="m" value="1300.0">1300 m</quantity>
,
<geoCoordinate id="57D4F3911246FFFCFA80FB9FFA1AAD16" box="[1318,1413,1117,1142]" degrees="22" direction="north" minutes="38" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="22.633333">22°38'N</geoCoordinate>
<geoCoordinate id="57D4F3911246FFFCFC8BFBBFFC00ADF6" box="[813,927,1149,1174]" degrees="120" direction="east" minutes="57" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="120.95">120°57'E</geoCoordinate>
,
<date id="465EB3961246FFFCFC0BFBBFFBA6ADF6" box="[941,1081,1149,1174]" pageId="9" pageNumber="10" value="2015-06-23">23-VI-2015</date>
, J. Yamasako leg. (ELKU). [
<collectingRegion id="F0245BB41246FFFCFC94FB5FFC6FADD7" box="[818,1008,1181,1207]" country="Taiwan" name="Kaohsiung" pageId="9" pageNumber="10">Kaohsiung City</collectingRegion>
]
<specimenCount id="24E65EDF1246FFFCFBA6FB5FFBC7ADD6" box="[1024,1112,1181,1206]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, vic. Fujieda (= Tengzhi) nr.
<collectingRegion id="F0245BB41246FFFCFCF7FB7FFC25ADB6" box="[849,954,1213,1238]" country="Taiwan" name="Taoyuan" pageId="9" pageNumber="10">Taoyuan</collectingRegion>
,
<collectingRegion id="F0245BB41246FFFCFC65FB7FFB1BADB6" box="[963,1156,1213,1238]" country="Taiwan" name="Taoyuan" pageId="9" pageNumber="10">Taoyuan District</collectingRegion>
, at light,
<date id="465EB3961246FFFCFB4DFB7FFAFDADB6" box="[1259,1378,1213,1238]" pageId="9" pageNumber="10" value="1976-04-06">6-IV-1976</date>
, K. Ushijima leg. (EUMJ).
</paragraph>
</subSubSection>
<subSubSection id="7AFAC6DD1246FFFCFCCFFB3FFAFAAF76" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="325F95561246FFFCFCCFFB3FFAFAAF76" blockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFCCFFB3FFC6DAC76" box="[873,1010,1277,1302]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
: This species resembles
<taxonomicName id="F5E0EED51246FFFCFACCFB3FFC5EAC56" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="009449441246FFFCFACCFB3FFAE3AC76" box="[1386,1404,1277,1302]" italics="true" pageId="9" pageNumber="10">P</emphasis>
.
<emphasis id="009449441246FFFCFC8BFADFFC5EAC56" box="[813,961,1309,1334]" italics="true" pageId="9" pageNumber="10">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F1246FFFCFC6AFADFFBAFAC56" box="[972,1072,1309,1334]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
but is easily distinguished by the absence of a row of long erect setae along each lateral elytral margin, the weak anterior angles of pronotum and the cone-shaped and stout parameres (see
<bibRefCitation id="5671E8A71246FFFCFBF5FA5FFCF0ACB6" author="Yoshida T &amp; Hirowatari T." pageId="9" pageNumber="10" pagination="15 - 45" refId="ref13985" refString="Yoshida T, Hirowatari T. 2014. A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae). ZooKeys 403: 15 - 45. doi: 10.3897 / zookeys. 403.7145." type="journal article" year="2014">Yoshida and Hirowatari 2014</bibRefCitation>
). This species also resembles
<taxonomicName id="F5E0EED51246FFFCFB73FA7FFA19ACB6" authorityName="Motschulsky" authorityYear="1858" box="[1237,1414,1469,1494]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="009449441246FFFCFB73FA7FFB78ACB6" box="[1237,1255,1469,1494]" italics="true" pageId="9" pageNumber="10">P</emphasis>
.
<emphasis id="009449441246FFFCFB50FA7FFA19ACB6" box="[1270,1414,1469,1494]" italics="true" pageId="9" pageNumber="10">trimaculatus</emphasis>
</taxonomicName>
and other closely related species, but it differs from them by the shorter lateral teeth of its pronotum.
</paragraph>
</subSubSection>
<subSubSection id="7AFAC6DD1246FFFCFCCFF9DFFB1BAE16" pageId="9" pageNumber="10" type="description">
<paragraph id="325F95561246FFFCFCCFF9DFFB1BAE16" blockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFCCFF9DFFC6CAF56" box="[873,1011,1565,1590]" italics="true" pageId="9" pageNumber="10">Coloration</emphasis>
: (
<figureCitation id="AADB89D31246FFFCFBB4F9DFFB18AF57" box="[1042,1159,1565,1591]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="9" pageNumber="10">Figs. 1C</figureCitation>
and
<figureCitation id="AADB89D31246FFFCFB74F9DFFB78AF56" box="[1234,1255,1565,1590]" captionStart="Fig" captionStartId="10.[174,208,1925,1945]" captionTargetBox="[273,1353,681,1898]" captionTargetId="figure-201@10.[273,1353,681,1898]" captionTargetPageId="10" captionText="Fig. 7. Habitus of Psammoecus harmandi Grouvelle in dorsal (A, C and D) and in ventral (B). (C) lighter colored specimen (D) darker colored specimen. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064351" httpUri="https://zenodo.org/record/8064351/files/figure.png" pageId="9" pageNumber="10">7</figureCitation>
). Head and pronotum yellowish-brown. Elytra sometimes a little lighter colored, with variable black maculae near posterior 1/3; round maculae around middle of each elytron, another round one near posterior 1/3 close to elytral suture (
<figureCitation id="AADB89D31246FFFCFB31F97FFA9FAFB7" box="[1175,1280,1725,1751]" captionStart="Fig" captionStartId="10.[174,208,1925,1945]" captionTargetBox="[273,1353,681,1898]" captionTargetId="figure-201@10.[273,1353,681,1898]" captionTargetPageId="10" captionText="Fig. 7. Habitus of Psammoecus harmandi Grouvelle in dorsal (A, C and D) and in ventral (B). (C) lighter colored specimen (D) darker colored specimen. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064351" httpUri="https://zenodo.org/record/8064351/files/figure.png" pageId="9" pageNumber="10">Fig. 7C</figureCitation>
): maculae sometimes connected to each other (
<figureCitation id="AADB89D31246FFFCFB58F91FFAC4AF96" box="[1278,1371,1757,1782]" captionStart="Fig" captionStartId="10.[174,208,1925,1945]" captionTargetBox="[273,1353,681,1898]" captionTargetId="figure-201@10.[273,1353,681,1898]" captionTargetPageId="10" captionText="Fig. 7. Habitus of Psammoecus harmandi Grouvelle in dorsal (A, C and D) and in ventral (B). (C) lighter colored specimen (D) darker colored specimen. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064351" httpUri="https://zenodo.org/record/8064351/files/figure.png" pageId="9" pageNumber="10">Fig. 7A</figureCitation>
) or forming a wide v-shaped covering posterior half of elytra (
<figureCitation id="AADB89D31246FFFCFCDBF8DFFC45AE56" box="[893,986,1821,1846]" captionStart="Fig" captionStartId="10.[174,208,1925,1945]" captionTargetBox="[273,1353,681,1898]" captionTargetId="figure-201@10.[273,1353,681,1898]" captionTargetPageId="10" captionText="Fig. 7. Habitus of Psammoecus harmandi Grouvelle in dorsal (A, C and D) and in ventral (B). (C) lighter colored specimen (D) darker colored specimen. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064351" httpUri="https://zenodo.org/record/8064351/files/figure.png" pageId="9" pageNumber="10">Fig. 7D</figureCitation>
). Antennae yellowish-brown basally, 9th and 10th antennomere black, 11th yellowish-brown, sometimes darkened.
</paragraph>
</subSubSection>
<subSubSection id="7AFAC6DD1246FFFCFCCFF8BFFBB9AED6" pageId="9" pageNumber="10" type="distribution">
<paragraph id="325F95561246FFFCFCCFF8BFFBB9AED6" blockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFCCFF8BFFC68AEF6" box="[873,1015,1917,1942]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
:
<collectingCountry id="4AF7D5C61246FFFCFBACF8BFFBF9AEF6" box="[1034,1126,1917,1942]" name="Taiwan" pageId="9" pageNumber="10">Taiwan</collectingCountry>
;
<collectingCountry id="4AF7D5C61246FFFCFBD2F8BFFB5BAEF6" box="[1140,1220,1917,1942]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
;
<collectingCountry id="4AF7D5C61246FFFCFB75F8BFFAA1AEF6" box="[1235,1342,1917,1942]" name="Thailand" pageId="9" pageNumber="10">Thailand</collectingCountry>
(new record);
<collectingCountry id="4AF7D5C61246FFFCFC36F85FFC45AED6" box="[912,986,1949,1974]" name="Nepal" pageId="9" pageNumber="10">Nepal</collectingCountry>
;
<collectingCountry id="4AF7D5C61246FFFCFC42F85FFBBDAED6" box="[996,1058,1949,1974]" name="India" pageId="9" pageNumber="10">India</collectingCountry>
.
</paragraph>
</subSubSection>
<footnote id="51FB89581246FFFCFF2EF81FFE07AE8F" box="[136,408,2013,2031]" pageId="9" pageNumber="10">
<paragraph id="325F95561246FFFCFF2EF81FFE07AE8F" blockId="9.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="9" pageNumber="10">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<subSubSection id="7AFAC6DD1245FFFEFF4CFF3FFE6FA836" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="325F95561245FFFFFF4CFF3FFE6BA856" blockId="10.[174,775,253,630]" pageId="10" pageNumber="11">
<emphasis id="009449441245FFFFFF4CFF3FFEC8A876" box="[234,343,253,278]" italics="true" pageId="10" pageNumber="11">Remarks</emphasis>
: This species occurs sympatrically with
<taxonomicName id="F5E0EED51245FFFFFF43FEDFFE0FA856" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[229,400,285,310]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="009449441245FFFFFF43FEDFFF68A856" box="[229,247,285,310]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFEA3FEDFFE0FA856" box="[261,400,285,310]" italics="true" pageId="10" pageNumber="11">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F1245FFFFFE3EFEDFFE6BA856" box="[408,500,285,310]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="325F95561245FFFFFF4CFEFFFBBFA856" blockId="10.[174,775,253,630]" lastBlockId="10.[850,1452,253,630]" pageId="10" pageNumber="11">
<taxonomicName id="F5E0EED51245FFFFFF4CFEFFFD9DA836" authorityName="Yoshida and Hirowatari" authorityYear="2014" box="[234,514,317,342]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="009449441245FFFFFF4CFEFFFD9DA836" box="[234,514,317,342]" italics="true" pageId="10" pageNumber="11">Psammoecus boreas</emphasis>
</taxonomicName>
, now synonymized with
<taxonomicName id="F5E0EED51245FFFFFF49FE9FFE0FA816" authorityName=", Grouvelle" authorityYear="1912" box="[239,400,349,374]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFF49FE9FFE9EA816" box="[239,257,349,374]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFEB0FE9FFE0FA816" box="[278,400,349,374]" italics="true" pageId="10" pageNumber="11">harmandi</emphasis>
</taxonomicName>
, was described from
<collectingCountry id="4AF7D5C61245FFFFFD1EFE9FFC9AA816" box="[696,773,349,374]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
based on diagnostic differences from
<taxonomicName id="F5E0EED51245FFFFFDD1FEBFFC98A8F6" authorityName=", Grouvelle" authorityYear="1912" box="[631,775,381,406]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFDD1FEBFFC98A8F6" box="[631,775,381,406]" italics="true" pageId="10" pageNumber="11">P. harmandi</emphasis>
</taxonomicName>
according to a redescription of
<taxonomicName id="F5E0EED51245FFFFFD87FE5FFD2DA8D6" authorityName=", Grouvelle" authorityYear="1912" box="[545,690,413,438]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFD87FE5FFDACA8D6" box="[545,563,413,438]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFDE5FE5FFD2DA8D6" box="[579,690,413,438]" italics="true" pageId="10" pageNumber="11">harmandi</emphasis>
</taxonomicName>
by
<bibRefCitation id="5671E8A71245FFFFFD46FE5FFE9AA8B6" author="Pal TK" pageId="10" pageNumber="11" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal (1985)</bibRefCitation>
.
<bibRefCitation id="5671E8A71245FFFFFEB2FE7FFD02A8B6" author="Yoshida T &amp; Hirowatari T." box="[276,669,445,470]" pageId="10" pageNumber="11" pagination="15 - 45" refId="ref13985" refString="Yoshida T, Hirowatari T. 2014. A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae). ZooKeys 403: 15 - 45. doi: 10.3897 / zookeys. 403.7145." type="journal article" year="2014">Yoshida and Hirowatari (2014)</bibRefCitation>
pointed out that
<taxonomicName id="F5E0EED51245FFFFFEBFFE1FFE2FA896" authorityName=", Grouvelle" authorityYear="1912" box="[281,432,477,502]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFEBFFE1FFEB4A896" box="[281,299,477,502]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFE9BFE1FFE2FA896" box="[317,432,477,502]" italics="true" pageId="10" pageNumber="11">harmandi</emphasis>
</taxonomicName>
differed from
<taxonomicName id="F5E0EED51245FFFFFDC0FE1FFD40A896" authorityName="Yoshida and Hirowatari" authorityYear="2014" box="[614,735,477,502]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="009449441245FFFFFDC0FE1FFDE7A896" box="[614,632,477,502]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFD2CFE1FFD40A896" box="[650,735,477,502]" italics="true" pageId="10" pageNumber="11">boreas</emphasis>
</taxonomicName>
by its shorter antennae and slightly transverse 10th antennomere. Although we could not examine the
<typeStatus id="ED5B2BF41245FFFFFF79FDFFFE8CAB36" box="[223,275,573,598]" pageId="10" pageNumber="11">type</typeStatus>
of
<taxonomicName id="F5E0EED51245FFFFFEE7FDFFFE45AB36" authorityName=", Grouvelle" authorityYear="1912" box="[321,474,573,598]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFEE7FDFFFECCAB36" box="[321,339,573,598]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFEC0FDFFFE45AB36" box="[358,474,573,598]" italics="true" pageId="10" pageNumber="11">harmandi</emphasis>
</taxonomicName>
, we had an opportunity to examine a male specimen of
<taxonomicName id="F5E0EED51245FFFFFD93FD9FFD58AB16" authorityName=", Grouvelle" authorityYear="1912" box="[565,711,605,630]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="009449441245FFFFFD93FD9FFDD8AB16" box="[565,583,605,630]" italics="true" pageId="10" pageNumber="11">P</emphasis>
.
<emphasis id="009449441245FFFFFDFEFD9FFD58AB16" box="[600,711,605,630]" italics="true" pageId="10" pageNumber="11">harmandi</emphasis>
</taxonomicName>
from northern
<collectingCountry id="4AF7D5C61245FFFFFC66FF3FFC65A876" box="[960,1018,253,278]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
and found these two species to be indistinguishable.
</paragraph>
<paragraph id="325F95561245FFFFFC29FEFFFC21A896" blockId="10.[850,1452,253,630]" pageId="10" pageNumber="11">
This species is widely distributed; specimens from
<collectingCountry id="4AF7D5C61245FFFFFC33FE9FFC7AA816" box="[917,997,349,374]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
,
<collectingCountry id="4AF7D5C61245FFFFFC55FE9FFBD1A816" box="[1011,1102,349,374]" name="Taiwan" pageId="10" pageNumber="11">Taiwan</collectingCountry>
,
<collectingCountry id="4AF7D5C61245FFFFFBFAFE9FFB53A816" box="[1116,1228,349,374]" name="Thailand" pageId="10" pageNumber="11">Thailand</collectingCountry>
,
<collectingCountry id="4AF7D5C61245FFFFFB7CFE9FFAB8A816" box="[1242,1319,349,374]" name="Nepal" pageId="10" pageNumber="11">Nepal</collectingCountry>
, and
<collectingCountry id="4AF7D5C61245FFFFFAC9FE9FFA34A816" box="[1391,1451,349,374]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
have been studied. Except for
<collectingRegion id="F0245BB41245FFFFFB74FEBFFADBA8F6" box="[1234,1348,381,406]" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
(
<collectingCountry id="4AF7D5C61245FFFFFAF3FEBFFA3BA8F6" box="[1365,1444,381,406]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
) where the northern limit of its distribution might be reached, they are exclusively distributed in high altitudes.
</paragraph>
<paragraph id="325F95561245FFFEFC29FE3FFE6FA836" blockId="10.[850,1452,253,630]" lastBlockId="11.[136,736,253,342]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Compared to the coloration described by
<bibRefCitation id="5671E8A71245FFFFFCF4FDDFFC78AB56" author="Pal TK" box="[850,999,541,566]" pageId="10" pageNumber="11" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal (1985)</bibRefCitation>
, the Taiwanese specimens show considerable intraspecific variation. We have not seen specimens outside of
<collectingCountry id="4AF7D5C61245FFFFFB0BFD9FFA9CAB16" box="[1197,1283,605,630]" name="Taiwan" pageId="10" pageNumber="11">Taiwan</collectingCountry>
with strongly enlarged black maculae that cover the posterior half of the elytra (
<figureCitation id="AADB89D31244FFFEFEDDFEDFFE7AA856" box="[379,485,285,310]" captionStart="Fig" captionStartId="10.[174,208,1925,1945]" captionTargetBox="[273,1353,681,1898]" captionTargetId="figure-201@10.[273,1353,681,1898]" captionTargetPageId="10" captionText="Fig. 7. Habitus of Psammoecus harmandi Grouvelle in dorsal (A, C and D) and in ventral (B). (C) lighter colored specimen (D) darker colored specimen. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064351" httpUri="https://zenodo.org/record/8064351/files/figure.png" pageId="11" pageNumber="12">Fig. 7D</figureCitation>
). Examples for the variability are given in figure 7.
</paragraph>
</subSubSection>
</treatment>
</document>
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