369 lines
49 KiB
XML
369 lines
49 KiB
XML
<document id="7C5CF41E6ADDF8523459C8BC6AC348BA" ID-CLB-Dataset="46036" ID-DOI="10.5281/zenodo.201683" ID-GBIF-Dataset="5e3fc953-9cac-4d28-b33f-0c366534f471" ID-ISSN="1175-5326" ID-Zenodo-Dep="201683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460407527141" checkinUser="plazi" docAuthor="Dow, Rory A." docDate="2011" docId="BA4387BAFF969B0DFF53CFF4C66CFD04" docLanguage="en" docName="zt03093p046.pdf" docOrigin="Zootaxa 3093" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mortonagrion" docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="467AFFC2FF9F9B07FFC4CD7BC333FFA3" masterDocTitle="Mortonagrion indraneil spec. nov. from Borneo, and a redescription of M. arthuri Fraser (Odonata: Zygoptera: Coenagrionidae)" masterLastPageNumber="46" masterPageNumber="35" pageNumber="44" updateTime="1698299976102" updateUser="ExternalLinkService">
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<mods:title id="EF66D3F7242DB1120F4DE24A3D91D670">Mortonagrion indraneil spec. nov. from Borneo, and a redescription of M. arthuri Fraser (Odonata: Zygoptera: Coenagrionidae)</mods:title>
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<mods:namePart id="567AA187A9B2BD7E968A63F0EEE18783">Dow, Rory A.</mods:namePart>
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<mods:title id="B1547DC23DCC801D4870C411DB6AA123">Zootaxa</mods:title>
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<mods:part id="1DEF852517803D84D33162FF658E55B0">
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<mods:date id="FF26F6A2265E48BB8E634D2F6F7442B9">2011</mods:date>
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<mods:number id="29F2CD885D4B5E6C806A26A8AF444654">3093</mods:number>
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<mods:identifier id="5A77BC66FDB4251F1A5EA92F64470113" type="DOI">10.5281/zenodo.201683</mods:identifier>
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<mods:identifier id="5F24E76EED51006DA328629A4C0C47DA" type="Zenodo-Dep">201683</mods:identifier>
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<treatment id="BA4387BAFF969B0DFF53CFF4C66CFD04" ID-DOI="http://doi.org/10.5281/zenodo.6185155" ID-GBIF-Taxon="119422290" ID-Zenodo-Dep="6185155" LSID="urn:lsid:plazi:treatment:BA4387BAFF969B0DFF53CFF4C66CFD04" httpUri="http://treatment.plazi.org/id/BA4387BAFF969B0DFF53CFF4C66CFD04" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44">
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<subSubSection id="7AF06527FF969B0EFF53CFF4C08EFD0A" box="[151,957,655,681]" pageId="9" pageNumber="44" type="nomenclature">
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<paragraph id="325536ACFF969B0EFF53CFF4C08EFD0A" blockId="9.[151,957,655,681]" box="[151,957,655,681]" pageId="9" pageNumber="44">
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<heading id="691D81C0FF969B0EFF53CFF4C08EFD0A" bold="true" box="[151,957,655,681]" fontSize="11" level="1" pageId="9" pageNumber="44" reason="1">
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<emphasis id="009EEABEFF969B0EFF53CFF4C08EFD0A" bold="true" box="[151,957,655,681]" pageId="9" pageNumber="44">
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Relationship of
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<taxonomicName id="F5EA4D2FFF969B0EFE98CFF4C134FD0A" ID-CoL="5VC2" box="[348,519,655,681]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFE98CFF4C134FD0A" bold="true" box="[348,519,655,681]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="F5EA4D2FFF969B0EFD89CFF4C1D5FD0A" box="[589,742,655,681]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFD89CFF4C1D5FD0A" bold="true" box="[589,742,655,681]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F5EA4D2FFF969B0EFCE0CFF4C08EFD0A" box="[804,957,655,681]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFCE0CFF4C08EFD0A" bold="true" box="[804,957,655,681]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7AF06527FF969B0DFF53CFACC66CFD04" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44" type="reference_group">
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<paragraph id="325536ACFF969B0EFF53CFACC091FBD9" blockId="9.[151,1437,726,2010]" pageId="9" pageNumber="44">
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<taxonomicName id="F5EA4D2FFF969B0EFF53CFACC205FD4D" ID-CoL="5VC2" box="[151,310,727,750]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFF53CFACC205FD4D" box="[151,310,727,750]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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has had a confused history since it was erected. A number of species have at various times been placed in
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<taxonomicName id="F5EA4D2FFF969B0EFEC0CF87C290FCB0" ID-CoL="5VC2" box="[260,419,764,787]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFEC0CF87C290FCB0" box="[260,419,764,787]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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, and at other times placed in the allied genera
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<taxonomicName id="F5EA4D2FFF969B0EFC76CF80C773FCB0" ID-CoL="QXZ" box="[946,1088,763,787]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFC76CF80C773FCB0" box="[946,1088,763,787]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F5EA4D2FFF969B0EFBB3CF80C630FCB0" ID-CoL="32N3" box="[1143,1283,763,787]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFBB3CF80C630FCB0" box="[1143,1283,763,787]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
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</taxonomicName>
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. The issue of the relationships among these genera has never been satisfactorily resolved, for instance see
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<bibRefCitation id="567B4B5DFF969B0EFB0CCE65C65AFC95" author="Pinhey" box="[1224,1385,798,822]" pageId="9" pageNumber="44" refString="Pinhey, E. (1974) A revision of the African Agriocnemis Selys and Mortonagrion Fraser (Odonata Coenagrionidae). Occasional papers of the National Museums and Monuments of Rhodesia Series B Natural Sciences, 5 (4), 171 - 278." type="journal article" year="1974">Pinhey (1974)</bibRefCitation>
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and
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<bibRefCitation id="567B4B5DFF969B0EFF53CE39C245FCF9" author="Hamalainen" box="[151,374,834,858]" pageId="9" pageNumber="44" refString="Hamalainen, M. (1989) Synonymic notes on some species of the genus Mortonagrion (Zygoptera: Coenagrionidae). Indian Odonatology, 2, 1 - 4." type="journal article" year="1989">Hämäläinen (1989)</bibRefCitation>
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. Until now the sole character that has been considered to reliably distinguish
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<taxonomicName id="F5EA4D2FFF969B0EFB39CE3FC6AFFCF8" ID-CoL="5VC2" box="[1277,1436,836,859]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFB39CE3FC6AFFCF8" box="[1277,1436,836,859]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="F5EA4D2FFF969B0EFF11CE1DC250FCDD" ID-CoL="QXZ" box="[213,355,870,894]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFF11CE1DC250FCDD" box="[213,355,870,894]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F5EA4D2FFF969B0EFE5FCE1DC11BFCDD" ID-CoL="32N3" box="[411,552,870,894]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFE5FCE1DC11BFCDD" box="[411,552,870,894]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
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</taxonomicName>
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is a detail of wing venation: “
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<emphasis id="009EEABEFF969B0EFC45CE1DC0AEFCDD" box="[897,925,870,894]" italics="true" pageId="9" pageNumber="44">ab</emphasis>
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continued on as
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<emphasis id="009EEABEFF969B0EFB9ACE1DC023FC00" italics="true" pageId="9" pageNumber="44">1a in the same straight line, no angulated medio-anal link intervening at the junction</emphasis>
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” (
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<bibRefCitation id="567B4B5DFF969B0EFCEECEF1C0DDFC01" author="Fraser" box="[810,1006,906,930]" pageId="9" pageNumber="44" refString="Fraser, F. C. (1933) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor & Francis, London, xii + 423 pp." type="book" year="1933">Fraser 1933: 409</bibRefCitation>
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). The species currently placed in
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<taxonomicName id="F5EA4D2FFF969B0EFAA7CEF7C236FC65" ID-CoL="5VC2" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFAA7CEF7C236FC65" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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, excluding
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<taxonomicName id="F5EA4D2FFF969B0EFE4DCECBC127FC65" ID-CoL="44FB8" box="[393,532,942,967]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="aborense">
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<emphasis id="009EEABEFF969B0EFE4DCECBC127FC65" box="[393,532,942,967]" italics="true" pageId="9" pageNumber="44">M. aborense</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F5EA4D2FFF969B0EFD88CECBC1FAFC65" ID-CoL="44FB9" box="[588,713,942,967]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="alcyone">
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<emphasis id="009EEABEFF969B0EFD88CECBC1FAFC65" box="[588,713,942,967]" italics="true" pageId="9" pageNumber="44">M. alcyone</emphasis>
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</taxonomicName>
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and the African
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<taxonomicName id="F5EA4D2FFF969B0EFC43CECBC7B6FC65" ID-CoL="44FBP" authority="Fraser, 1954" authorityName="Fraser" authorityYear="1954" box="[903,1157,942,967]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="stygia">
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<emphasis id="009EEABEFF969B0EFC43CECBC0DCFC65" box="[903,1007,943,967]" italics="true" pageId="9" pageNumber="44">M. stygia</emphasis>
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Fraser, 1954
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</taxonomicName>
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, share a number of other characteristics, for instance: complicated postocular markings in the males, mesostigmal plates strongly raised up along one or more edges in females (e.g.
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<figureCitation id="AAD12A29FF969B0EFDA2CE8DC1E4FBAD" box="[614,727,1014,1038]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@2.[151,1436,193,1024]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Prothorax and mesostigmal region of female Mortonagrion (scale bar 100 μm unless otherwise noted): (7) M. alcyone mesostigmal plates in dorsal view; (8) M. amoena posterior pronotal lobe and mesostigmal plates, lateral; (9) M. arthuri posterior pronotal lobe and mesostigmal plates (scale bar 200 μm), lateral; (10) M. indraneil paratype (SAR 06 _ COE 353) posterior pronotal lobe and mesostigmal plates, lateral." httpUri="https://zenodo.org/record/201685/files/figure.png" pageId="9" pageNumber="44">Figs 8–10</figureCitation>
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) and triangular paraprocts (e.g.
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<figureCitation id="AAD12A29FF969B0EFBFACE8DC7BCFBAD" box="[1086,1167,1014,1038]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 32</figureCitation>
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) in females (rather than the rather formless paraprocts typical in the
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<taxonomicName id="F5EA4D2FFF969B0EFD52C961C07EFB91" ID-CoL="624DX" box="[662,845,1050,1074]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
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). These species certainly appear to be more closely related to each other than to any other species, but none of the characters mentioned is unique to this group of species, all being found in some species typically placed in
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<taxonomicName id="F5EA4D2FFF969B0EFCC9C918C0A9FBD8" ID-CoL="QXZ" box="[781,922,1123,1147]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFCC9C918C0A9FBD8" box="[781,922,1123,1147]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="325536ACFF969B0EFF01C9F3C2EAFA1D" blockId="9.[151,1437,726,2010]" pageId="9" pageNumber="44">
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<taxonomicName id="F5EA4D2FFF969B0EFF01C9F3C263FB3D" ID-CoL="44FB8" box="[197,336,1158,1183]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="aborense">
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<emphasis id="009EEABEFF969B0EFF01C9F3C263FB3D" box="[197,336,1158,1183]" italics="true" pageId="9" pageNumber="44">M. aborense</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F5EA4D2FFF969B0EFE41C9F3C132FB3D" ID-CoL="44FB9" box="[389,513,1158,1183]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="alcyone">
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<emphasis id="009EEABEFF969B0EFE41C9F3C132FB3D" box="[389,513,1158,1183]" italics="true" pageId="9" pageNumber="44">M. alcyone</emphasis>
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</taxonomicName>
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both share some characters with
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<taxonomicName id="F5EA4D2FFF969B0EFCB7C9FDC631FB3D" ID-CoL="GGWJ" authority="Selys, 1877" authorityName="Selys" authorityYear="1877" box="[883,1282,1158,1182]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="rubescens">
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<emphasis id="009EEABEFF969B0EFCB7C9FDC745FB3D" box="[883,1142,1158,1182]" italics="true" pageId="9" pageNumber="44">Argiocnemis rubescens</emphasis>
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Selys, 1877
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</taxonomicName>
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, the genotype of
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<taxonomicName id="F5EA4D2FFF969B0EFF71C9D0C272FB60" ID-CoL="32N3" box="[181,321,1195,1219]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFF71C9D0C272FB60" box="[181,321,1195,1219]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
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</taxonomicName>
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, that are not shared with the other
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<taxonomicName id="F5EA4D2FFF969B0EFD00C9D7C050FB60" ID-CoL="5VC2" box="[708,867,1196,1219]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="009EEABEFF969B0EFD00C9D7C050FB60" box="[708,867,1196,1219]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
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</taxonomicName>
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species: all have simple postocular markings, relatively simple broad triangular mesostigmal plates and formless paraprocts in the female, and very similar male anal appendages; they also share a typical scheme of colour changes from immaturity to maturity. See
|
||
<bibRefCitation id="567B4B5DFF969B0EFAD0C989C3D2FA8D" author="Hamalainen" pageId="9" pageNumber="44" refString="Hamalainen, M. (1989) Synonymic notes on some species of the genus Mortonagrion (Zygoptera: Coenagrionidae). Indian Odonatology, 2, 1 - 4." type="journal article" year="1989">Hämäläinen (1989)</bibRefCitation>
|
||
for a summary of the complicated history of
|
||
<taxonomicName id="F5EA4D2FFF969B0EFD28C86DC042FA8D" ID-CoL="44FB8" box="[748,881,1302,1326]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF969B0EFD28C86DC042FA8D" box="[748,881,1302,1326]" italics="true" pageId="9" pageNumber="44">A. aborense</emphasis>
|
||
</taxonomicName>
|
||
; however it differs significantly in genital ligula structure from all other species examined or for which illustrations were available from all three genera (see below).
|
||
<taxonomicName id="F5EA4D2FFF969B0EFF30C825C258FAD5" ID-CoL="44FB9" box="[244,363,1374,1398]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF969B0EFF30C825C258FAD5" box="[244,363,1374,1398]" italics="true" pageId="9" pageNumber="44">A. alcyone</emphasis>
|
||
</taxonomicName>
|
||
appears to have become associated with
|
||
<emphasis id="009EEABEFF969B0EFCFBC81BC0EDFAD4" box="[831,990,1376,1399]" italics="true" pageId="9" pageNumber="44">Mortanagrion</emphasis>
|
||
almost by accident, as it does not even consistently share the venational character, for instance the
|
||
<typeStatus id="ED51880EFF969B0EFCFCC8F9C0ADFA39" box="[824,926,1410,1434]" pageId="9" pageNumber="44" type="lectotype">lectotype</typeStatus>
|
||
male in the BMNH has an angulated medioanal link joining ab with 1A.
|
||
</paragraph>
|
||
<paragraph id="325536ACFF969B0EFF01C8B7C051F989" blockId="9.[151,1437,726,2010]" pageId="9" pageNumber="44">
|
||
<taxonomicName id="F5EA4D2FFF969B0EFF01C8B7C21EFA40" ID-CoL="44FBP" box="[197,301,1484,1507]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="stygia">
|
||
<emphasis id="009EEABEFF969B0EFF01C8B7C21EFA40" box="[197,301,1484,1507]" italics="true" pageId="9" pageNumber="44">M. stygia</emphasis>
|
||
</taxonomicName>
|
||
has mature colouration more typical of many
|
||
<taxonomicName id="F5EA4D2FFF969B0EFCF0C8B0C0F2FA40" ID-CoL="QXZ" box="[820,961,1483,1507]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFCF0C8B0C0F2FA40" box="[820,961,1483,1507]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
|
||
</taxonomicName>
|
||
species, but the female variably has triangular paraprocts. The genital ligula is of
|
||
<taxonomicName id="F5EA4D2FFF969B0EFD99C895C1D9F9A5" ID-CoL="QXZ" box="[605,746,1518,1542]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFD99C895C1D9F9A5" box="[605,746,1518,1542]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="F5EA4D2FFF969B0EFD37C88BC0A1F9A4" ID-CoL="5VC2" box="[755,914,1520,1543]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFD37C88BC0A1F9A4" box="[755,914,1520,1543]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
form (see below). It is difficult to come to any firm conclusion on the placement of this species at the moment.
|
||
</paragraph>
|
||
<paragraph id="325536ACFF969B0DFF01CB4DC6AFFE0F" blockId="9.[151,1437,726,2010]" lastBlockId="10.[151,1436,151,680]" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44">
|
||
Probably the most promising character to distinguish
|
||
<taxonomicName id="F5EA4D2FFF969B0EFCEDCB4DC086F9ED" ID-CoL="32N3" box="[809,949,1590,1614]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFCEDCB4DC086F9ED" box="[809,949,1590,1614]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="F5EA4D2FFF969B0EFC39CB4DC7B9F9ED" ID-CoL="QXZ" box="[1021,1162,1590,1614]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFC39CB4DC7B9F9ED" box="[1021,1162,1590,1614]" italics="true" pageId="9" pageNumber="44">Agriocnemis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5EA4D2FFF969B0EFB01CB43C657F9EC" ID-CoL="5VC2" box="[1221,1380,1592,1615]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFB01CB43C657F9EC" box="[1221,1380,1592,1615]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
, and possibly the latter two genera from each other, is the structure of the genital ligula. The genital ligula of
|
||
<taxonomicName id="F5EA4D2FFF969B0EFAE8CB20C28AF935" ID-CoL="B7KL" authority="Brauer, 1868" authorityName="Brauer" authorityYear="1868" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="femina">
|
||
<emphasis id="009EEABEFF969B0EFAE8CB20C223F935" italics="true" pageId="9" pageNumber="44">Agriocnemis femina</emphasis>
|
||
(Brauer, 1868)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFE0CCB05C128F935" box="[456,539,1662,1686]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 25</figureCitation>
|
||
),
|
||
<taxonomicName id="F5EA4D2FFF969B0EFDEACB05C0BDF935" ID-CoL="5DKM3" box="[558,910,1662,1686]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="rubescens" subSpecies="rubeola">
|
||
<emphasis id="009EEABEFF969B0EFDEACB05C0BDF935" box="[558,910,1662,1686]" italics="true" pageId="9" pageNumber="44">Argiocnemis rubescens rubeola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFC59CB05C0C3F935" box="[925,1008,1662,1686]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 26</figureCitation>
|
||
),
|
||
<taxonomicName id="F5EA4D2FFF969B0EFBC0CB05C7A3F935" ID-CoL="32N3" box="[1028,1168,1662,1686]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFBC0CB05C7A3F935" box="[1028,1168,1662,1686]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<figureCitation id="AAD12A29FF969B0EFB01CB05C624F935" box="[1221,1303,1662,1686]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 27</figureCitation>
|
||
; a problematic form distinct from, but possibly with a name buried in the synonymy of,
|
||
<taxonomicName id="F5EA4D2FFF969B0EFC28CBD8C748F918" ID-CoL="GGWJ" box="[1004,1147,1699,1723]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="rubescens">
|
||
<emphasis id="009EEABEFF969B0EFC28CBD8C748F918" box="[1004,1147,1699,1723]" italics="true" pageId="9" pageNumber="44">A. rubescens</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="F5EA4D2FFF969B0EFB55CBDFC6A8F918" ID-CoL="44FB8" box="[1169,1435,1699,1723]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF969B0EFB55CBDFC6A8F918" box="[1169,1435,1699,1723]" italics="true" pageId="9" pageNumber="44">Mortonagrion aborense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFF5BCBBDC3C0F97D" box="[159,243,1734,1758]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 28</figureCitation>
|
||
),
|
||
<taxonomicName id="F5EA4D2FFF969B0EFECDCBB3C2B5F97D" ID-CoL="44FB9" box="[265,390,1734,1759]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF969B0EFECDCBB3C2B5F97D" box="[265,390,1734,1759]" italics="true" pageId="9" pageNumber="44">M. alcyone</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFE52CBBDC2D8F97D" box="[406,491,1734,1758]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 29</figureCitation>
|
||
),
|
||
<taxonomicName id="F5EA4D2FFF969B0EFDC4CBB3C145F97D" ID-CoL="44FBC" box="[512,630,1734,1759]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="arthuri">
|
||
<emphasis id="009EEABEFF969B0EFDC4CBB3C145F97D" box="[512,630,1734,1759]" italics="true" pageId="9" pageNumber="44">M. arthuri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFD42CBBDC1EAF97D" box="[646,729,1734,1758]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 30</figureCitation>
|
||
) and
|
||
<taxonomicName id="F5EA4D2FFF969B0EFCDDCBB3C094F97D" ID-CoL="44FBK" box="[793,935,1734,1759]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="indraneil">
|
||
<emphasis id="009EEABEFF969B0EFCDDCBB3C094F97D" box="[793,935,1734,1759]" italics="true" pageId="9" pageNumber="44">M. indraneil</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF969B0EFC73CBBDC73AF97D" box="[951,1033,1734,1758]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="9" pageNumber="44">Fig. 31</figureCitation>
|
||
) were examined by Scanning Electron Microscopy (SEM). With the exception of
|
||
<taxonomicName id="F5EA4D2FFF969B0EFD0ECB97C06AF8A0" ID-CoL="44FB8" box="[714,857,1771,1795]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF969B0EFD0ECB97C06AF8A0" box="[714,857,1771,1795]" italics="true" pageId="9" pageNumber="44">M. aborense</emphasis>
|
||
</taxonomicName>
|
||
, the genital ligula of all the species examined is broadly similar; the terminal segment has simple-shaped broad apical lateral lobes, at rest directed back on either side of the shaft. In
|
||
<taxonomicName id="F5EA4D2FFF969B0EFEBDCA4FC124F8E8" ID-CoL="5VC2" box="[377,535,1844,1867]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFEBDCA4FC124F8E8" box="[377,535,1844,1867]" italics="true" pageId="9" pageNumber="44">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5EA4D2FFF969B0EFD8BCA48C1EFF8E8" ID-CoL="32N3" box="[591,732,1843,1867]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFD8BCA48C1EFF8E8" box="[591,732,1843,1867]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
there are rows of short setae or spines dorsolaterally on either side of the shaft; in
|
||
<taxonomicName id="F5EA4D2FFF969B0EFEB7CA2DC160F8CD" ID-CoL="B7KL" box="[371,595,1878,1902]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="femina">
|
||
<emphasis id="009EEABEFF969B0EFEB7CA2DC160F8CD" box="[371,595,1878,1902]" italics="true" pageId="9" pageNumber="44">Agriocnemis femina</emphasis>
|
||
</taxonomicName>
|
||
the spines are absent from the shaft, but there is a small clump of spines on each side of the segment that lies between the apical lobes of the terminal segment. In
|
||
<taxonomicName id="F5EA4D2FFF969B0EFB4ACA00C6A6F830" ID-CoL="GGWJ" box="[1166,1429,1915,1939]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="rubescens">
|
||
<emphasis id="009EEABEFF969B0EFB4ACA00C6A6F830" box="[1166,1429,1915,1939]" italics="true" pageId="9" pageNumber="44">Argiocnemis rubescens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5EA4D2FFF969B0EFF53CAE5C210F815" ID-CoL="32N3" box="[151,291,1950,1974]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF969B0EFF53CAE5C210F815" box="[151,291,1950,1974]" italics="true" pageId="9" pageNumber="44">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName id="F5EA4D2FFF969B0EFE46CADBC2CCF815" ID-CoL="44FB9" box="[386,511,1950,1975]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF969B0EFE46CADBC2CCF815" box="[386,511,1950,1975]" italics="true" pageId="9" pageNumber="44">M. alcyone</emphasis>
|
||
</taxonomicName>
|
||
the terminal part of the lateral lobes has a fringe of narrow teeth along its margin, but these teeth are absent in the other species examined.
|
||
<bibRefCitation id="567B4B5DFF969B0EFCD8CAB9C08EF879" author="Pinhey" box="[796,957,1986,2010]" pageId="9" pageNumber="44" refString="Pinhey, E. (1974) A revision of the African Agriocnemis Selys and Mortonagrion Fraser (Odonata Coenagrionidae). Occasional papers of the National Museums and Monuments of Rhodesia Series B Natural Sciences, 5 (4), 171 - 278." type="journal article" year="1974">Pinhey (1974)</bibRefCitation>
|
||
illustrated the genital ligula of all African species known to him, including
|
||
<taxonomicName id="F5EA4D2FFF959B0DFDC2CDE2C15DFF13" ID-CoL="44FBP" box="[518,622,153,176]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="stygia">
|
||
<emphasis id="009EEABEFF959B0DFDC2CDE2C15DFF13" box="[518,622,153,176]" italics="true" pageId="10" pageNumber="45">M. stygia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="F5EA4D2FFF959B0DFD6ECDE3C768FF0C" ID-CoL="B7LD" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[682,1115,151,176]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="009EEABEFF959B0DFD6ECDE3C090FF13" box="[682,931,152,176]" italics="true" pageId="10" pageNumber="45">Agriocnemis pygmaea</emphasis>
|
||
(Rambur, 1842)
|
||
</taxonomicName>
|
||
which is also widely distributed in Asia; the genital ligula of all of these species is of the general
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC70CDC7C772FF77" ID-CoL="QXZ" box="[948,1089,188,212]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFC70CDC7C772FF77" box="[948,1089,188,212]" italics="true" pageId="10" pageNumber="45">Agriocnemis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="F5EA4D2FFF959B0DFB8DCDC6C7DBFF77" ID-CoL="5VC2" box="[1097,1256,189,212]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFB8DCDC6C7DBFF77" box="[1097,1256,189,212]" italics="true" pageId="10" pageNumber="45">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
form. However with the equipment available to him Pinhey may not have been able to see certain features of the genital ligula, such as spines or setae, but it seems likely that a fringe of teeth such as occurs in
|
||
<taxonomicName id="F5EA4D2FFF959B0DFBF0CC7EC782FEBF" ID-CoL="44FB9" box="[1076,1201,260,284]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF959B0DFBF0CC7EC782FEBF" box="[1076,1201,260,284]" italics="true" pageId="10" pageNumber="45">M. alcyone</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5EA4D2FFF959B0DFB04CC7FC3FAFEE3" ID-CoL="GGWJ" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="rubescens">
|
||
<emphasis id="009EEABEFF959B0DFB04CC7FC3FAFEE3" italics="true" pageId="10" pageNumber="45">Argiocnemis rubescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5EA4D2FFF959B0DFEC5CC53C2BEFEE3" ID-CoL="32N3" box="[257,397,296,320]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFEC5CC53C2BEFEE3" box="[257,397,296,320]" italics="true" pageId="10" pageNumber="45">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
sp. would have been visible to him; presumably no such fringe occurs in any of the species examined by Pinhey. As noted by
|
||
<bibRefCitation id="567B4B5DFF959B0DFDD0CC37C19DFEC7" author="Pinhey" box="[532,686,332,356]" pageId="10" pageNumber="45" refString="Pinhey, E. (1974) A revision of the African Agriocnemis Selys and Mortonagrion Fraser (Odonata Coenagrionidae). Occasional papers of the National Museums and Monuments of Rhodesia Series B Natural Sciences, 5 (4), 171 - 278." type="journal article" year="1974">Pinhey (1974</bibRefCitation>
|
||
: 271 as
|
||
<taxonomicName id="F5EA4D2FFF959B0DFCC3CC36C0D2FEC7" ID-CoL="44FBN" authority="Ris, 1930" authorityName="Ris" authorityYear="1930" box="[775,993,332,356]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="simile">
|
||
<emphasis id="009EEABEFF959B0DFCC3CC36C05CFEC7" box="[775,879,332,356]" italics="true" pageId="10" pageNumber="45">M. simile</emphasis>
|
||
Ris, 1930
|
||
</taxonomicName>
|
||
) and illustrated by
|
||
<bibRefCitation id="567B4B5DFF959B0DFB7DCC37C652FEC7" author="Asahina" box="[1209,1377,332,356]" pageId="10" pageNumber="45" refString="Asahina, S. (1982) A list of the Odonata from Thailand. Part I. Agrionidae. Kontyu, Toyko, 50 (3), 454 - 466." type="journal article" year="1982">Asahina (1982</bibRefCitation>
|
||
: 457,
|
||
<figureCitation id="AAD12A29FF959B0DFF53CC14C235FE24" box="[151,262,367,391]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@2.[151,1436,193,1024]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Prothorax and mesostigmal region of female Mortonagrion (scale bar 100 μm unless otherwise noted): (7) M. alcyone mesostigmal plates in dorsal view; (8) M. amoena posterior pronotal lobe and mesostigmal plates, lateral; (9) M. arthuri posterior pronotal lobe and mesostigmal plates (scale bar 200 μm), lateral; (10) M. indraneil paratype (SAR 06 _ COE 353) posterior pronotal lobe and mesostigmal plates, lateral." httpUri="https://zenodo.org/record/201685/files/figure.png" pageId="10" pageNumber="45">Figs 9–10</figureCitation>
|
||
as
|
||
<taxonomicName id="F5EA4D2FFF959B0DFEE8CC0AC157FE24" ID-CoL="44FBD" authority="Fraser, 1922" authorityName="Fraser" authorityYear="1922" box="[300,612,367,392]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="binocellata">
|
||
<emphasis id="009EEABEFF959B0DFEE8CC0AC2E3FE2B" box="[300,464,368,392]" italics="true" pageId="10" pageNumber="45">M. binocellata</emphasis>
|
||
Fraser, 1922
|
||
</taxonomicName>
|
||
) the genital ligula of
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC91CC0AC0D3FE2B" ID-CoL="44FB8" box="[853,992,368,392]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF959B0DFC91CC0AC0D3FE2B" box="[853,992,368,392]" italics="true" pageId="10" pageNumber="45">M. aborense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="AAD12A29FF959B0DFC34CC14C771FE24" box="[1008,1090,367,391]" captionStart="FIGURES 25 – 32" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[281,1315,195,1850]" captionTargetId="figure@6.[281,1315,195,1850]" captionTargetPageId="6" captionText="FIGURES 25 – 32. Genital ligula of Agriocnemis, Argiocnemis and Mortonagrion species (scale bar 100 μm): (25) Agriocnemis femina; (26) Argiocnemis rubescens rubeola; (27) Argiocnemis species; (28) Mortonagrion aborense; (29) Mortonagrion alcyone; (30) Mortonagrion arthuri; (31) Mortonagrion indraneil paratype (SAR 05 _ COE 152). Mortonagrion female anal appendages (32) Mortonagrion indraneil female paratype (SAR 06 _ COE 353), scale bar 200 μm." httpUri="https://zenodo.org/record/201689/files/figure.png" pageId="10" pageNumber="45">Fig. 28</figureCitation>
|
||
) is of a different form from all the other species considered, with, as Pinhey puts it “flagella united into a single short, broad scoop over the stem.”
|
||
</paragraph>
|
||
<paragraph id="325536ACFF959B0DFF01CCC3C66CFD04" blockId="10.[151,1436,151,680]" pageId="10" pageNumber="45">
|
||
I consider the placement of
|
||
<taxonomicName id="F5EA4D2FFF959B0DFDC9CCC3C140FE73" ID-CoL="44FB8" box="[525,627,440,464]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF959B0DFDC9CCC3C140FE73" box="[525,627,440,464]" italics="true" pageId="10" pageNumber="45">aborense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5EA4D2FFF959B0DFD6BCCC3C035FE73" ID-CoL="44FB9" box="[687,774,440,464]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF959B0DFD6BCCC3C035FE73" box="[687,774,440,464]" italics="true" pageId="10" pageNumber="45">alcyone</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="F5EA4D2FFF959B0DFCF5CCC2C0E3FE73" ID-CoL="5VC2" box="[817,976,441,464]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFCF5CCC2C0E3FE73" box="[817,976,441,464]" italics="true" pageId="10" pageNumber="45">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
to be extremely doubtful, with
|
||
<taxonomicName id="F5EA4D2FFF959B0DFA81CCC3C6AFFE73" ID-CoL="44FB9" box="[1349,1436,440,464]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="alcyone">
|
||
<emphasis id="009EEABEFF959B0DFA81CCC3C6AFFE73" box="[1349,1436,440,464]" italics="true" pageId="10" pageNumber="45">alcyone</emphasis>
|
||
</taxonomicName>
|
||
belonging with the
|
||
<taxonomicName id="F5EA4D2FFF959B0DFEABCCA7C2C8FE57" ID-CoL="32N3" box="[367,507,476,500]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFEABCCA7C2C8FE57" box="[367,507,476,500]" italics="true" pageId="10" pageNumber="45">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
species which it closely resembles and
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC73CCA7C72FFE57" ID-CoL="44FB8" box="[951,1052,476,500]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="aborense">
|
||
<emphasis id="009EEABEFF959B0DFC73CCA7C72FFE57" box="[951,1052,476,500]" italics="true" pageId="10" pageNumber="45">aborense</emphasis>
|
||
</taxonomicName>
|
||
possibly deserving its own genus. However in order to prevent further confusion it appears best to leave the status of these species unchanged at this time.
|
||
<taxonomicName id="F5EA4D2FFF959B0DFF13CF5EC245FD9F" ID-CoL="5VC2" box="[215,374,549,572]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFF13CF5EC245FD9F" box="[215,374,549,572]" italics="true" pageId="10" pageNumber="45">Mortonagrion</emphasis>
|
||
</taxonomicName>
|
||
may itself prove to be merely a specialised group within
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC3BCF5FC7BFFD9F" ID-CoL="QXZ" box="[1023,1164,548,572]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFC3BCF5FC7BFFD9F" box="[1023,1164,548,572]" italics="true" pageId="10" pageNumber="45">Agriocnemis</emphasis>
|
||
</taxonomicName>
|
||
, but this should be considered as part of a full review of the relationships between
|
||
<emphasis id="009EEABEFF959B0DFC87CF32C74DFDC3" box="[835,1150,584,608]" italics="true" pageId="10" pageNumber="45">
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC87CF32C0D7FDC3" ID-CoL="5VC2" box="[835,996,585,608]" class="Insecta" family="Coenagrionidae" genus="Mortonagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">Mortonagrion</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5EA4D2FFF959B0DFC35CF33C74DFDC3" ID-CoL="QXZ" box="[1009,1150,584,608]" class="Insecta" family="Coenagrionidae" genus="Agriocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">Agriocnemis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="F5EA4D2FFF959B0DFB73CF33C670FDC3" ID-CoL="32N3" box="[1207,1347,584,608]" class="Insecta" family="Coenagrionidae" genus="Argiocnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="009EEABEFF959B0DFB73CF33C670FDC3" box="[1207,1347,584,608]" italics="true" pageId="10" pageNumber="45">Argiocnemis</emphasis>
|
||
</taxonomicName>
|
||
; such a review is a major undertaking that must include the African as well as Asian species, SEM studies of the genital ligula and molecular studies. A final resolution of this issue will not be based on characters of wing venation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |