treatments-xml/data/BA/22/14/BA221469C04EA15B9FDDFB3B2F59B3CE.xml

<document id="9C41B136FE994883316994B2F4809FC2" ID-CLB-Dataset="31603" ID-DOI="10.11646/zootaxa.4344.2.7" ID-GBIF-Dataset="458df93d-4b4d-46c0-a9ee-d004cfde7863" ID-ISSN="1175-5326" ID-Zenodo-Dep="1043104" ID-ZooBank="BACD8FC5-66FB-4514-B4AA-629D5A8F84AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510041172352" checkinUser="plazi" docAuthor="Zhang, Hao-Miao, Guan, Zhao-Ying &amp; Wang, Wen-Zhi" docDate="2017" docId="BA221469C04EA15B9FDDFB3B2F59B3CE" docLanguage="en" docName="zootaxa.4344.2.7.pdf" docOrigin="Zootaxa 4344 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gomphidictinus tongi Zhang &amp; Guan &amp; Wang 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="328" masterDocId="461B6C11C04FA15C9F4AFFA52A7FB621" masterDocTitle="Updated information on genus Gomphidictinus (Odonata: Gomphidae) in China with description of Gomphidictinus tongi sp. nov." masterLastPageNumber="332" masterPageNumber="321" pageNumber="322" updateTime="1698480864195" updateUser="ExternalLinkService">
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<mods:title id="40C676A6E18C9EA7A3D671D7CD5BCBBB">Updated information on genus Gomphidictinus (Odonata: Gomphidae) in China with description of Gomphidictinus tongi sp. nov.</mods:title>
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<mods:namePart id="C6FAE28D96EA82DA920677E4C83A4704">Zhang, Hao-Miao</mods:namePart>
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<mods:namePart id="C57D4656436EA13D20C56B1A0EA09CA1">Guan, Zhao-Ying</mods:namePart>
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<mods:namePart id="AC3D01BAB694D3BD1FFD40CFF01D8EAF">Wang, Wen-Zhi</mods:namePart>
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<taxonomicName id="F58BDEFCC04EA15D9FDDFB3B2BE2B299" ID-CoL="9X2PW" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[151,413,1182,1208]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04EA15D9FDDFB3B2BE2B299" bold="true" box="[151,413,1182,1208]" italics="true" pageId="1" pageNumber="322">Gomphidictinus tongi</emphasis>
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<materialsCitation id="82E3AF22C04EA15D9FDDFB4029FCB367" ID-GBIF-Occurrence="1699405280" collectingDate="2014-05-28" collectorName="Hao-miao Zhang" country="China" county="Lingshui County" elevation="930" latitude="18.7231" location="Collection of Kunming Institute" longLatPrecision="1" longitude="109.86955" municipality="Mt. Diaoluoshan" pageId="1" pageNumber="322" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
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<typeStatus id="ED301BDDC04EA15D9FDDFB402B79B2DF" box="[151,262,1253,1278]" pageId="1" pageNumber="322" type="holotype">Holotype</typeStatus>
.
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(deposited in the 
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Collection of 
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of Zoology, 
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, 
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, 
<collectingCountry id="4A9CE5EFC04EA15D9E93FAAC285FB303" box="[473,544,1289,1314]" name="China" pageId="1" pageNumber="322">China</collectingCountry>
), 
<collectingCountry id="4A9CE5EFC04EA15D9D7FFAAC2804B303" box="[565,635,1289,1314]" name="China" pageId="1" pageNumber="322">China</collectingCountry>
, 
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, 
<collectingRegion id="F04F6B9DC04EA15D9C19FAAC2E6CB303" box="[851,1043,1289,1314]" country="China" name="Hainan" pageId="1" pageNumber="322">Hainan Province</collectingRegion>
, 
<collectingMunicipality id="D2503F05C04EA15D9B55FAAF2EA2B303" box="[1055,1245,1289,1314]" pageId="1" pageNumber="322">Mt. Diaoluoshan</collectingMunicipality>
(
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, 
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; 
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a.s.l.
</elevation>
), 
<date id="463583BFC04EA15D9EBBFA8B28F6B367" box="[497,649,1326,1350]" pageId="1" pageNumber="322" value="2014-05-28">
<collectingDate id="56717A57C04EA15D9EBBFA8B28F6B367" box="[497,649,1326,1350]" pageId="1" pageNumber="322" value="2014-05-28">28 May 2014</collectingDate>
</date>
, 
<collectorName id="9F7EC0A9C04EA15D9DDEFA88292DB367" box="[660,850,1325,1350]" pageId="1" pageNumber="322">Hao-miao Zhang</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="3234A57FC04EA15D9F8DFAF429CCB3AF" blockId="1.[151,1437,1253,2034]" pageId="1" pageNumber="322">
<materialsCitation id="82E3AF22C04EA15D9F8DFAF428B7B34B" ID-GBIF-Occurrence="1699405278" box="[199,712,1361,1386]" collectingDate="2014-05-28" collectorName="Hao-miao Zhang" country="China" county="Lingshui County" elevation="930" latitude="18.7231" location="Collection of Kunming Institute" longLatPrecision="1" longitude="109.86955" municipality="Mt. Diaoluoshan" pageId="1" pageNumber="322" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan Province" typeStatus="paratype">
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<typeStatus id="ED301BDDC04EA15D9F8DFAF42B3EB34B" box="[199,321,1361,1386]" pageId="1" pageNumber="322" type="paratype">Paratypes</typeStatus>
.
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<specimenCount id="248D6EF6C04EA15D9E04FAF72BF6B34B" box="[334,393,1362,1386]" pageId="1" pageNumber="322" type="male">2♂♂</specimenCount>
, 
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, same data as holotype
</materialsCitation>
; 
<materialsCitation id="82E3AF22C04EA15D9D9CFAF729CCB3AF" ID-GBIF-Occurrence="1699405279" collectingDate="2017-05-30" collectorName="Haomiao Zhang &amp; All" country="China" location="Chongzuo" pageId="1" pageNumber="322" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="248D6EF6C04EA15D9D9CFAF72885B34B" box="[726,762,1362,1386]" pageId="1" pageNumber="322" type="male">1♂</specimenCount>
, 
<collectingCountry id="4A9CE5EFC04EA15D9C43FAF42930B34B" box="[777,847,1361,1386]" name="China" pageId="1" pageNumber="322">China</collectingCountry>
, 
<collectingRegion id="F04F6B9DC04EA15D9C16FAF429C3B34B" box="[860,956,1361,1386]" country="China" name="Guangxi" pageId="1" pageNumber="322">Guangxi</collectingRegion>
Province, 
<location id="3754F3A4C04EA15D9B72FAF42ED1B34B" LSID="urn:lsid:plazi:treatment:BA221469C04EA15B9FDDFB3B2F59B3CE:3754F3A4C04EA15D9B72FAF42ED1B34B" box="[1080,1198,1361,1386]" country="China" name="Chongzuo" pageId="1" pageNumber="322" stateProvince="Guangxi">Chongzuo</location>
, 
<date id="463583BFC04EA15D9BF1FAF72F27B34B" box="[1211,1368,1362,1386]" pageId="1" pageNumber="322" value="2017-05-30">
<collectingDate id="56717A57C04EA15D9BF1FAF72F27B34B" box="[1211,1368,1362,1386]" pageId="1" pageNumber="322" value="2017-05-30">30 May 2017</collectingDate>
</date>
, 
<collectorName id="9F7EC0A9C04EA15D9A2EFAF72B61B3AF" pageId="1" pageNumber="322">Haomiao Zhang</collectorName>
leg. 
<collectorName id="9F7EC0A9C04EA15D9E1EFAD02B07B3AF" box="[340,376,1397,1422]" pageId="1" pageNumber="322">All</collectorName>
<typeStatus id="ED301BDDC04EA15D9E35FAD32B95B3AF" box="[383,490,1398,1422]" pageId="1" pageNumber="322" type="paratype">paratypes</typeStatus>
deposited at the same place as 
<typeStatus id="ED301BDDC04EA15D9C03FAD029D0B3AF" box="[841,943,1397,1422]" pageId="1" pageNumber="322" type="holotype">holotype</typeStatus>
.
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</paragraph>
</subSubSection>
<subSubSection id="7A91F6F4C04EA15D9F8DFA3C2E98B03F" pageId="1" pageNumber="322" type="etymology">
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<emphasis id="00FF796DC04EA15D9F8DFA3C2B31B393" bold="true" box="[199,334,1433,1458]" pageId="1" pageNumber="322">Etymology.</emphasis>
The species name, a noun in the genitive case, is dedicated to Professor Xiao-li Tong, from South 
<collectingCountry id="4A9CE5EFC04EA15D9FDDFA182AA5B3F7" box="[151,218,1469,1494]" name="China" pageId="1" pageNumber="322">China</collectingCountry>
Agricultural University, the Ph.D. supervisor of the first and second authors. He passed on professional knowledge of entomology to Hao-miao Zhang and gave great help for his study on 
<taxonomicName id="F58BDEFCC04EA15D9B74FA442EDEB3DB" authorityName="Fabricius" authorityYear="1793" box="[1086,1185,1505,1530]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="322" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
.
</paragraph>
<paragraph id="3234A57FC04EA15D9F8DF9A32E98B03F" blockId="1.[151,1437,1253,2034]" box="[199,1255,1541,1566]" pageId="1" pageNumber="322">
A big and robust dragonfly. Body black with yellow maculation and hyaline wings (
<figureCitation id="AAB0B9FAC04EA15D9B3EF9A02EA5B03F" box="[1140,1242,1541,1566]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1994,2017]" captionTargetBox="[317,1285,228,1967]" captionTargetId="figure@3.[245,1352,193,1978]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Habitus of Gomphidictinus tongi sp. nov. in lateral view: 5) holotype male; 6) paratype female." httpUri="https://zenodo.org/record/1043110/files/figure.png" pageId="1" pageNumber="322">Figs 5–6</figureCitation>
).
</paragraph>
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<subSubSection id="7A91F6F4C04EA1599F8DF98C2F41B0ED" lastPageId="5" lastPageNumber="326" pageId="1" pageNumber="322" type="description">
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<emphasis id="00FF796DC04EA15D9F8DF98C2867B063" bold="true" box="[199,536,1577,1602]" pageId="1" pageNumber="322">
Description of the 
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.
</emphasis>
<emphasis id="00FF796DC04EA15D9D6BF98C2821B063" box="[545,606,1577,1602]" italics="true" pageId="1" pageNumber="322">Head</emphasis>
. Eyes dark green when alive (
<figureCitation id="AAB0B9FAC04EA15D9C8AF98C2E79B063" box="[960,1030,1577,1602]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="1" pageNumber="322">Fig. 7</figureCitation>
). Face black with yellow markings (
<figureCitation id="AAB0B9FAC04EA15D9FD5F9E82A99B047" box="[159,230,1613,1638]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="1" pageNumber="322">Fig. 7</figureCitation>
). Labium almost black with indistinct yellow spots at the posterior of each lobe; mandible bases yellow; labrum black; anteclypeus entirely yellow; postclypeus largely black with paired yellow spots at the lateral sides; antefrons black below, with the upper half yellow, top of frons with anterior 2/3 yellow; vertex above lateral ocelli triangularly projected, black with paired triangularly projections behind lateral ocelli; occiput black with a very short median tubercle, occipital margin almost straight.
</paragraph>
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<emphasis id="00FF796DC04EA15D9F8DF8A42B60B13B" box="[199,287,1793,1818]" italics="true" pageId="1" pageNumber="322">Thorax.</emphasis>
Generally black with yellow markings (
<figureCitation id="AAB0B9FAC04EA15D9DA0F8A4294FB13B" box="[746,816,1793,1818]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1994,2017]" captionTargetBox="[317,1285,228,1967]" captionTargetId="figure@3.[245,1352,193,1978]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Habitus of Gomphidictinus tongi sp. nov. in lateral view: 5) holotype male; 6) paratype female." httpUri="https://zenodo.org/record/1043110/files/figure.png" pageId="1" pageNumber="322">Fig. 5</figureCitation>
). Prothorax black. Synthorax with mesothoracic collar stripes connecting at midpoint, dorsal stripes not connecting with collar stripes; antehumeral spot small, comma shaped; sides of synthorax mostly black with yellow markings: mesepimeron with a broad yellow stripe, metepisternum with a triangular shaped spot near the upper margin, metepimeron largely yellow, mesokatepisternum with a lower spot, metakatepisternum largely yellow. Poststernum with paired large spots. Legs black with yellow markings. Coxae with yellow spots in all legs; fore leg femur with yellow stripes in the inner side, mid and hind legs entirely black.
</paragraph>
<caption id="66F4F5F7C04DA15E9FDDF9AB2B14B062" httpUri="https://zenodo.org/record/1043108/files/figure.png" pageId="2" pageNumber="323" targetBox="[260,1330,215,1536]" targetPageId="2">
<paragraph id="3234A57FC04DA15E9FDDF9AB2B14B062" blockId="2.[151,1436,1550,1603]" pageId="2" pageNumber="323">
<emphasis id="00FF796DC04DA15E9FDDF9AB2B40B005" bold="true" box="[151,319,1550,1573]" pageId="2" pageNumber="323">FIGURES 3–4.</emphasis>
Comparison of wings between genus 
<taxonomicName id="F58BDEFCC04DA15E9D96F9AA2901B004" authorityName="Fraser" authorityYear="1942" box="[732,894,1551,1573]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="00FF796DC04DA15E9D96F9AA2901B004" box="[732,894,1551,1573]" italics="true" pageId="2" pageNumber="323">Gomphidictinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F58BDEFCC04DA15E9CF2F9AA2E55B004" authorityName="Selys" authorityYear="1854" box="[952,1066,1551,1573]" class="Insecta" family="Gomphidae" genus="Gomphidia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="00FF796DC04DA15E9CF2F9AA2E55B004" box="[952,1066,1551,1573]" italics="true" pageId="2" pageNumber="323">Gomphidia</emphasis>
</taxonomicName>
: 3) 
<taxonomicName id="F58BDEFCC04DA15E9B16F9AA2F0AB004" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[1116,1397,1551,1573]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC04DA15E9B16F9AA2F0AB004" box="[1116,1397,1551,1573]" italics="true" pageId="2" pageNumber="323">Gomphidictinus perakensis</emphasis>
</taxonomicName>
; 4) 
<taxonomicName id="F58BDEFCC04DA15E9FDDF9882B1AB062" authorityName="Martin" authorityYear="1904" box="[151,357,1581,1603]" class="Insecta" family="Gomphidae" genus="Gomphidia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="00FF796DC04DA15E9FDDF9882B1AB062" box="[151,357,1581,1603]" italics="true" pageId="2" pageNumber="323">Gomphidia kruegeri</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3234A57FC04DA15E9F8DF9D42B90B11C" blockId="2.[151,1437,1648,2033]" pageId="2" pageNumber="323">
<emphasis id="00FF796DC04DA15E9F8DF9D42B6CB0A9" box="[199,275,1649,1672]" italics="true" pageId="2" pageNumber="323">Wings.</emphasis>
Hyaline, slightly tinted with amber brown, venation black. Forewings: 22 antenodals above Sc and 18 antenodals below Sc; 18 (left) and 19 (right) postnodals above R1. Hindwing: 15 (left) and 16 (right) above Sc, 15 antenodals below Sc; 18 (left) and 20 (right) postnodals above R1. Triangle 4-celled in forewings and 3-celled in hindwings, supertriangle 3- or 5-celled in forewings, 3-celled in hindwings, subtriangle 3-celled in forewings and 1-celled in hindwings. Anal triangle 5-celled. Anal loop 5-celled. Pterostigma long, 
<quantity id="F573089AC04DA15E9B21F8A42EDBB139" box="[1131,1188,1793,1816]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="323" unit="mm" value="7.0">7mm</quantity>
in length, color dark brown, covering 9 cells below.
</paragraph>
<paragraph id="3234A57FC04DA15E9F8DF8ED285DB1D0" blockId="2.[151,1437,1648,2033]" pageId="2" pageNumber="323">
<emphasis id="00FF796DC04DA15E9F8DF8ED2B47B140" box="[199,312,1864,1889]" italics="true" pageId="2" pageNumber="323">Abdomen.</emphasis>
Black with pale marking as follows (
<figureCitation id="AAB0B9FAC04DA15E9D91F8ED2960B140" box="[731,799,1864,1889]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1994,2017]" captionTargetBox="[317,1285,228,1967]" captionTargetId="figure@3.[245,1352,193,1978]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Habitus of Gomphidictinus tongi sp. nov. in lateral view: 5) holotype male; 6) paratype female." httpUri="https://zenodo.org/record/1043110/files/figure.png" pageId="2" pageNumber="323">Fig. 5</figureCitation>
): S1 with a large lateral spot close the lower margin, and very fine stripe along the hind margin dorsally; S2 with a large anterior central arrow-shaped spot dorsally, laterally with two spots, anterior one covering auricle; S3 with a large basal spot, triangular shaped; bases of S4–6 with paired spots, connecting at mid-carina; basal half of S7 with a large spot; S8–10 and anal appendages almost black, S10 with paired tiny spots dorsally.
</paragraph>
<caption id="66F4F5F7C04CA15F9FDDF86F2F79B1C0" box="[151,1286,1994,2017]" httpUri="https://zenodo.org/record/1043110/files/figure.png" pageId="3" pageNumber="324" targetBox="[317,1285,228,1967]" targetPageId="3">
<paragraph id="3234A57FC04CA15F9FDDF86F2F79B1C0" blockId="3.[151,1286,1994,2017]" box="[151,1286,1994,2017]" pageId="3" pageNumber="324">
<emphasis id="00FF796DC04CA15F9FDDF86F2B45B1C1" bold="true" box="[151,314,1994,2017]" pageId="3" pageNumber="324">FIGURES 5–6.</emphasis>
Habitus of 
<taxonomicName id="F58BDEFCC04CA15F9EFBF86E28F2B1C0" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[433,653,1995,2017]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04CA15F9EFBF86E28F2B1C0" box="[433,653,1995,2017]" italics="true" pageId="3" pageNumber="324">Gomphidictinus tongi</emphasis>
</taxonomicName>
<emphasis id="00FF796DC04CA15F9DD9F86E289BB1C1" bold="true" box="[659,740,1995,2016]" pageId="3" pageNumber="324">
<taxonomicNameLabel id="1BCCC416C04CA15F9DD9F86E289BB1C1" box="[659,740,1995,2016]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in lateral view: 5) holotype male; 6) paratype female.
</paragraph>
</caption>
<caption id="66F4F5F7C04BA1589FDDF9D42808B0E5" httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325" targetBox="[237,1363,217,1612]" targetPageId="4">
<paragraph id="3234A57FC04BA1589FDDF9D42808B0E5" blockId="4.[151,1436,1649,1732]" pageId="4" pageNumber="325">
<emphasis id="00FF796DC04BA1589FDDF9D42B39B0A7" bold="true" box="[151,326,1649,1672]" pageId="4" pageNumber="325">FIGURES 7–14.</emphasis>
<taxonomicName id="F58BDEFCC04BA1589E06F9D42858B0A6" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[332,551,1649,1671]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04BA1589E06F9D42858B0A6" box="[332,551,1649,1671]" italics="true" pageId="4" pageNumber="325">Gomphidictinus tongi</emphasis>
</taxonomicName>
<emphasis id="00FF796DC04BA1589D67F9D72802B0A6" bold="true" box="[557,637,1650,1671]" pageId="4" pageNumber="325">
<taxonomicNameLabel id="1BCCC416C04BA1589D67F9D72802B0A6" box="[557,637,1650,1671]" pageId="4" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (7–12) and paratype female (13–14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view.
</paragraph>
</caption>
<paragraph id="3234A57FC04BA1589F8DF9572BF9B1C3" blockId="4.[151,1436,1777,2018]" pageId="4" pageNumber="325">
<emphasis id="00FF796DC04BA1589F8DF95728D3B12B" box="[199,684,1777,1802]" italics="true" pageId="4" pageNumber="325">Accessory genitalia and anal appendages.</emphasis>
Vesicle with a huge ventral projection. Median segment of penis organ with a sharp basal spine, distal segment with twisted flagellae (
<figureCitation id="AAB0B9FAC04BA1589CAAF8B02E55B10F" box="[992,1066,1813,1838]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325">Fig. 8</figureCitation>
). Anterior hamuli stout, gently expanded apically, hooked backwards in apex (
<figureCitation id="AAB0B9FAC04BA1589DEEF89C2897B173" box="[676,744,1849,1874]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325">Fig. 9</figureCitation>
); posterior hamuli broad and strongly curved posteriorly (
<figureCitation id="AAB0B9FAC04BA1589A25F89C2AC9B157" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325">Fig. 10</figureCitation>
). Anal appendages black. Cerci about 1.65 times as long as S10, conical and tapering distally in lateral view, with a ventral spine basally (
<figureCitation id="AAB0B9FAC04BA1589E93F8242854B1BB" box="[473,555,1921,1946]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325">Fig. 11</figureCitation>
); in dorsal view, depressed at apical 2/5, with a row of 7–9 denticles (
<figureCitation id="AAB0B9FAC04BA1589A71F8242FF0B1BB" box="[1339,1423,1921,1946]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="4" pageNumber="325">Fig. 12</figureCitation>
). Epiproct very short, thus forming a big hollow from the ventral spine of cerci, the dorsal surface with two very small spines apically.
</paragraph>
<caption id="66F4F5F7C04AA1599FDDFBDD292CB2AF" box="[151,851,1144,1166]" httpUri="https://zenodo.org/record/1043114/files/figure.png" pageId="5" pageNumber="326" targetBox="[253,1343,203,1113]" targetPageId="5">
<paragraph id="3234A57FC04AA1599FDDFBDD292CB2AF" blockId="5.[151,851,1144,1166]" box="[151,851,1144,1166]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599FDDFBDD2B60B2AC" bold="true" box="[151,287,1144,1166]" pageId="5" pageNumber="326">FIGURE 15.</emphasis>
Habitus of 
<taxonomicName id="F58BDEFCC04AA1599EDCFBDD28D5B2AF" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[406,682,1144,1166]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC04AA1599EDCFBDD28D5B2AF" box="[406,682,1144,1166]" italics="true" pageId="5" pageNumber="326">Gomphidictinus perakensis</emphasis>
</taxonomicName>
, from Malaysia.
</paragraph>
</caption>
<paragraph id="3234A57FC04AA1599F8DFB1E2BFFB2D6" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DFB1E2BEAB2F5" bold="true" box="[199,405,1211,1236]" pageId="5" pageNumber="326">
<typeStatus id="ED301BDDC04AA1599F8DFB1E2B4BB2F5" box="[199,308,1211,1236]" pageId="5" pageNumber="326" type="paratype">Paratype</typeStatus>
female:
</emphasis>
<emphasis id="00FF796DC04AA1599ED4FB1E2BA5B2F5" box="[414,474,1211,1236]" italics="true" pageId="5" pageNumber="326">Head</emphasis>
. Maculation very similar to 
<typeStatus id="ED301BDDC04AA1599C68FB1E29FBB2F5" box="[802,900,1211,1236]" pageId="5" pageNumber="326" type="holotype">holotype</typeStatus>
male. Occiput with a pair of lobed projections posteriorly (
<figureCitation id="AAB0B9FAC04AA1599E55FB7A2B0CB2D6" box="[287,371,1247,1272]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="5" pageNumber="326">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="3234A57FC04AA1599F8DFAA6285DB361" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DFAA62B5FB33D" box="[199,288,1283,1308]" italics="true" pageId="5" pageNumber="326">Thorax.</emphasis>
Almost identical to 
<typeStatus id="ED301BDDC04AA1599D40FAA62813B33D" box="[522,620,1283,1308]" pageId="5" pageNumber="326" type="holotype">holotype</typeStatus>
male, but yellow spots on head and coxae slightly larger than 
<typeStatus id="ED301BDDC04AA1599A70FAA62FE3B33D" box="[1338,1436,1283,1308]" pageId="5" pageNumber="326" type="holotype">holotype</typeStatus>
male, wings brown color darkened.
</paragraph>
<paragraph id="3234A57FC04AA1599F8DFAEE2B16B036" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DFAEE2B47B345" box="[199,312,1355,1380]" italics="true" pageId="5" pageNumber="326">Abdomen.</emphasis>
Black with pale marking as follows (
<figureCitation id="AAB0B9FAC04AA1599DAAFAEE2959B345" box="[736,806,1355,1380]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1994,2017]" captionTargetBox="[317,1285,228,1967]" captionTargetId="figure@3.[245,1352,193,1978]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Habitus of Gomphidictinus tongi sp. nov. in lateral view: 5) holotype male; 6) paratype female." httpUri="https://zenodo.org/record/1043110/files/figure.png" pageId="5" pageNumber="326">Fig. 6</figureCitation>
): S1 with a large lateral spot posteriorly and a very fine stripe along the hind margin dorsally; S2 with a triangular central spot dorsally, lateral side with a big rectangular spot; S3 with anterior semicircle spot dorsally and anterior lateral small stripes; S4–6 with anterior spot dorsally; S7 with a large basal spot; S8–10 and cerci almost black except for the small paired spots on S10 dorsally. Cerci long, about 1.5 times as long as S10. Vulvar lamina very long, reach almost the end of S9, deeply depressed medially (
<figureCitation id="AAB0B9FAC04AA1599E42FA5A2B23B036" box="[264,348,1535,1560]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,264,1649,1672]" captionTargetBox="[237,1363,217,1612]" captionTargetId="figure@4.[217,1369,193,1628]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Gomphidictinus tongi sp. nov., holotype male (7 – 12) and paratype female (13 – 14): 7) head in frontal view; 8) penis in lateral view; 9) anterior hamuli; 10) posterior hamuli; 11) cerci in lateral view; 12) cerci in dorsal view; 13) head in frontal view; 14) vulvar lamina in ventral view." httpUri="https://zenodo.org/record/1043112/files/figure.png" pageId="5" pageNumber="326">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="3234A57FC04AA1599F8DF9862E41B041" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DF9862860B01D" bold="true" box="[199,543,1571,1596]" pageId="5" pageNumber="326">
Variation in 
<typeStatus id="ED301BDDC04AA1599E2AF9862BB4B01D" box="[352,459,1571,1596]" pageId="5" pageNumber="326" type="paratype">paratype</typeStatus>
males.
</emphasis>
Two 
<typeStatus id="ED301BDDC04AA1599D2BF98128BEB01D" box="[609,705,1572,1596]" pageId="5" pageNumber="326" type="paratype">paratype</typeStatus>
males have collar stripes interrupted at midpoint. Paired yellow spots larger. Fore leg femur with less yellow markings. Anal triangle 5- or 6-celled.
</paragraph>
<paragraph id="3234A57FC04AA1599F8DF9CE2F41B0ED" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DF9CE2BB6B0A5" bold="true" box="[199,457,1643,1668]" pageId="5" pageNumber="326">Measurements (mm).</emphasis>
<typeStatus id="ED301BDDC04AA1599E9FF9CE2842B0A5" box="[469,573,1643,1668]" pageId="5" pageNumber="326" type="holotype">Holotype</typeStatus>
male: total length 85.0, abdomen (including anal appendages) 63.0, hind wing 51.5; 
<typeStatus id="ED301BDDC04AA1599E56F9352B01B089" box="[284,382,1680,1704]" pageId="5" pageNumber="326" type="paratype">Paratype</typeStatus>
males: total length 78.0–79.0, abdomen (including anal appendages) 58.0–60.0, hind wing 48.0–49.0; 
<typeStatus id="ED301BDDC04AA1599E5FF9112B08B0ED" box="[277,375,1716,1740]" pageId="5" pageNumber="326" type="paratype">Paratype</typeStatus>
female: total length 81.0, abdomen (including anal appendages) 61.0, hind wing 54.0;
</paragraph>
</subSubSection>
<subSubSection id="7A91F6F4C04AA1599F8DF97228CCB0D1" box="[199,691,1751,1776]" pageId="5" pageNumber="326" type="distribution">
<paragraph id="3234A57FC04AA1599F8DF97228CCB0D1" blockId="5.[151,1437,1211,2028]" box="[199,691,1751,1776]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DF9722B1FB0D1" bold="true" box="[199,352,1751,1776]" pageId="5" pageNumber="326">Distribution.</emphasis>
<collectingCountry id="4A9CE5EFC04AA1599E2DF9722BD5B0D1" box="[359,426,1751,1776]" name="China" pageId="5" pageNumber="326">China</collectingCountry>
(
<collectingRegion id="F04F6B9DC04AA1599EF3F9722873B0D1" box="[441,524,1751,1776]" country="China" name="Hainan" pageId="5" pageNumber="326">Hainan</collectingRegion>
and 
<collectingRegion id="F04F6B9DC04AA1599D08F97228D8B0D1" box="[578,679,1751,1776]" country="China" name="Guangxi" pageId="5" pageNumber="326">Guangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="7A91F6F4C04AA15B9F8DF95E2F59B3CE" lastPageId="7" lastPageNumber="328" pageId="5" pageNumber="326" type="diagnosis">
<paragraph id="3234A57FC04AA1599F8DF95E298BB1E6" blockId="5.[151,1437,1211,2028]" pageId="5" pageNumber="326">
<emphasis id="00FF796DC04AA1599F8DF95E2B3EB135" bold="true" box="[199,321,1787,1812]" pageId="5" pageNumber="326">Diagnosis.</emphasis>
Male of 
<taxonomicName id="F58BDEFCC04AA1599EEFF9592B81B132" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[421,510,1788,1811]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04AA1599EEFF9592B81B132" box="[421,510,1788,1811]" italics="true" pageId="5" pageNumber="326">G. tongi</emphasis>
</taxonomicName>
<emphasis id="00FF796DC04AA1599D4EF95E2822B135" bold="true" box="[516,605,1787,1812]" pageId="5" pageNumber="326">
<taxonomicNameLabel id="1BCCC416C04AA1599D4EF95E2822B135" box="[516,605,1787,1812]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="F58BDEFCC04AA1599DA7F9592914B135" authorityName="Karube" authorityYear="2016" box="[749,875,1787,1812]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="00FF796DC04AA1599DA7F9592914B135" box="[749,875,1787,1812]" italics="true" pageId="5" pageNumber="326">G. kompieri</emphasis>
</taxonomicName>
, but can be distinguished from it by the following characters: 1) the epiproct shorter, with a bigger hollow between cerci and epiproct in lateral view; 2) the spine on the median segment of the penis is situated more basally; 3) the subtriangle of the hindwings is not reticulated. Male of 
<taxonomicName id="F58BDEFCC04AA1599FBDF8CD2B30B15E" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[247,335,1896,1919]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04AA1599FBDF8CD2B30B15E" box="[247,335,1896,1919]" italics="true" pageId="5" pageNumber="326">G. tongi</emphasis>
</taxonomicName>
<emphasis id="00FF796DC04AA1599E12F8C22BCDB1A1" bold="true" box="[344,434,1895,1920]" pageId="5" pageNumber="326">
<taxonomicNameLabel id="1BCCC416C04AA1599E12F8C22BCDB1A1" box="[344,434,1895,1920]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is quite different from 
<taxonomicName id="F58BDEFCC04AA1599DF0F8CD292DB1A1" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[698,850,1895,1920]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC04AA1599DF0F8CD292DB1A1" box="[698,850,1895,1920]" italics="true" pageId="5" pageNumber="326">G. perakensis</emphasis>
</taxonomicName>
by the absence of the whale-tail shaped tubercle on ventral surface of S1 and the shape of the male anal appendages. Female of 
<taxonomicName id="F58BDEFCC04AA1599CADF8292E40B182" authority="Zhang &amp; Guan &amp; Wang, 2017" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[999,1087,1932,1955]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="00FF796DC04AA1599CADF8292E40B182" box="[999,1087,1932,1955]" italics="true" pageId="5" pageNumber="326">G. tongi</emphasis>
</taxonomicName>
<emphasis id="00FF796DC04AA1599B0FF82E2EE0B185" bold="true" box="[1093,1183,1931,1956]" pageId="5" pageNumber="326">
<taxonomicNameLabel id="1BCCC416C04AA1599B0FF82E2EE0B185" box="[1093,1183,1931,1956]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from the other congeners by longer vulvar lamina which is reaching the end of S9.
</paragraph>
<paragraph id="3234A57FC04AA1599F8DF8712994B1CD" blockId="5.[151,1437,1211,2028]" box="[199,1003,2003,2028]" pageId="5" pageNumber="326">
More detailed characters between the three species are listed in 
<tableCitation id="7F0990C4C04AA1599CDBF876299BB1CD" box="[913,996,2003,2028]" captionStart="TABLE 1" captionStartId="6.[151,239,152,175]" captionText="TABLE 1. Comparison of male characters among the known Gomphidictinus species." pageId="5" pageNumber="326">Table 1</tableCitation>
.
</paragraph>
<caption id="66F4F5F7C049A15A9FDDFF3D2E51B68E" box="[151,1070,151,176]" pageId="6" pageNumber="327">
<paragraph id="3234A57FC049A15A9FDDFF3D2E51B68E" blockId="6.[151,1070,151,176]" box="[151,1070,151,176]" pageId="6" pageNumber="327">
<emphasis id="00FF796DC049A15A9FDDFF3D2B75B68E" bold="true" box="[151,266,152,175]" pageId="6" pageNumber="327">TABLE 1.</emphasis>
Comparison of male characters among the known 
<taxonomicName id="F58BDEFCC049A15A9C63FF3229ACB68E" authorityName="Fraser" authorityYear="1942" box="[809,979,151,175]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="00FF796DC049A15A9C63FF3229ACB68E" box="[809,979,151,175]" italics="true" pageId="6" pageNumber="327">Gomphidictinus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="3234A57FC049A15A9FD5FF6F2EC1B4D9" pageId="6" pageNumber="327">
<table id="408B57DFC0495EA39FDDFF6F2F4DB4D9" box="[151,1330,202,760]" gridcols="4" gridrows="11" pageId="6" pageNumber="327">
<tr id="8CBBA73DC0495EA39FDDFF6F2F4DB6C1" box="[151,1330,202,224]" gridrow="0" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFF6F2BEFB6C1" box="[151,400,202,224]" gridcol="0" gridrow="0" pageId="6" pageNumber="327">
<emphasis id="00FF796DC049A15A9FD5FF6F2B78B6C1" box="[159,263,202,224]" italics="true" pageId="6" pageNumber="327">Character</emphasis>
</th>
<th id="CF6ACE41C0495EA39EFCFF6F297FB6C1" box="[438,768,202,224]" gridcol="1" gridrow="0" pageId="6" pageNumber="327">
<taxonomicName id="F58BDEFCC049A15A9EFCFF6E2879B6C1" authorityName="Zhang &amp; Guan &amp; Wang" authorityYear="2017" box="[438,518,203,224]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="tongi">
<emphasis id="00FF796DC049A15A9EFCFF6E2879B6C1" box="[438,518,203,224]" italics="true" pageId="6" pageNumber="327">G. tongi</emphasis>
</taxonomicName>
</th>
<th id="CF6ACE41C0495EA39C52FF6F2E49B6C1" box="[792,1078,202,224]" gridcol="2" gridrow="0" pageId="6" pageNumber="327">
<taxonomicName id="F58BDEFCC049A15A9C52FF6E29F1B6C1" authorityName="Karube" authorityYear="2016" box="[792,910,202,224]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="00FF796DC049A15A9C52FF6E29F1B6C1" box="[792,910,202,224]" italics="true" pageId="6" pageNumber="327">G. kompieri</emphasis>
</taxonomicName>
</th>
<th id="CF6ACE41C0495EA39B2FFF6F2F4DB6C1" box="[1125,1330,202,224]" gridcol="3" gridrow="0" pageId="6" pageNumber="327">
<taxonomicName id="F58BDEFCC049A15A9B2FFF6E2E92B6C1" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[1125,1261,202,224]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC049A15A9B2FFF6E2E92B6C1" box="[1125,1261,202,224]" italics="true" pageId="6" pageNumber="327">G. perakensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="8CBBA73DC0495EA39FDDFF512F4DB72B" box="[151,1330,244,266]" gridrow="1" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFF512BEFB72B" box="[151,400,244,266]" gridcol="0" gridrow="1" pageId="6" pageNumber="327">Length (Holotype, mm)</th>
<td id="CF6ACE41C0495EA39EFCFF51297FB72B" box="[438,768,244,266]" gridcol="1" gridrow="1" pageId="6" pageNumber="327">Abd. 63; Hw. 51.5</td>
<td id="CF6ACE41C0495EA39C52FF512E49B72B" box="[792,1078,244,266]" gridcol="2" gridrow="1" pageId="6" pageNumber="327">Abd. 64.1; Hw. 54</td>
<td id="CF6ACE41C0495EA39B2FFF512F4DB72B" box="[1125,1330,244,266]" gridcol="3" gridrow="1" pageId="6" pageNumber="327">Abd. 51; Hw. 48</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFEB82F4DB712" box="[151,1330,285,307]" gridrow="2" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFEB82BEFB712" box="[151,400,285,307]" gridcol="0" gridrow="2" pageId="6" pageNumber="327">Antefrons</th>
<td id="CF6ACE41C0495EA39EFCFEB8297FB712" box="[438,768,285,307]" gridcol="1" gridrow="2" pageId="6" pageNumber="327">Upper margin with yellow stripe</td>
<td id="CF6ACE41C0495EA39C52FEB82E49B712" box="[792,1078,285,307]" gridcol="2" gridrow="2" pageId="6" pageNumber="327">Entirely black</td>
<td id="CF6ACE41C0495EA39B2FFEB82F4DB712" box="[1125,1330,285,307]" gridcol="3" gridrow="2" pageId="6" pageNumber="327">Entirely black</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFEE32F4DB77D" box="[151,1330,326,348]" gridrow="3" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFEE32BEFB77D" box="[151,400,326,348]" gridcol="0" gridrow="3" pageId="6" pageNumber="327">Top of frons</th>
<td id="CF6ACE41C0495EA39EFCFEE3297FB77D" box="[438,768,326,348]" gridcol="1" gridrow="3" pageId="6" pageNumber="327">Complete yellow band</td>
<td id="CF6ACE41C0495EA39C52FEE32E49B77D" box="[792,1078,326,348]" gridcol="2" gridrow="3" pageId="6" pageNumber="327">Complete yellow band</td>
<td id="CF6ACE41C0495EA39B2FFEE32F4DB77D" box="[1125,1330,326,348]" gridcol="3" gridrow="3" pageId="6" pageNumber="327">Paired yellow spots</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFED52F4DB785" box="[151,1330,368,420]" gridrow="4" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFED52BEFB785" box="[151,400,368,420]" gridcol="0" gridrow="4" pageId="6" pageNumber="327">Triangle</th>
<td id="CF6ACE41C0495EA39EFCFED5297FB785" box="[438,768,368,420]" gridcol="1" gridrow="4" pageId="6" pageNumber="327">4-celled in forewings and 3- celled in hindwings</td>
<td id="CF6ACE41C0495EA39C52FED52E49B785" box="[792,1078,368,420]" gridcol="2" gridrow="4" pageId="6" pageNumber="327">4-celled in all wings</td>
<td id="CF6ACE41C0495EA39B2FFED52F4DB785" box="[1125,1330,368,420]" gridcol="3" gridrow="4" pageId="6" pageNumber="327">4-celled in all wings</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFE1D2F4DB7CD" box="[151,1330,440,492]" gridrow="5" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFE1D2BEFB7CD" box="[151,400,440,492]" gridcol="0" gridrow="5" pageId="6" pageNumber="327">Supertriangle</th>
<td id="CF6ACE41C0495EA39EFCFE1D297FB7CD" box="[438,768,440,492]" gridcol="1" gridrow="5" pageId="6" pageNumber="327">3-celled (except a 5-celled forewing in holotype)</td>
<td id="CF6ACE41C0495EA39C52FE1D2E49B7CD" box="[792,1078,440,492]" gridcol="2" gridrow="5" pageId="6" pageNumber="327">4-celled</td>
<td id="CF6ACE41C0495EA39B2FFE1D2F4DB7CD" box="[1125,1330,440,492]" gridcol="3" gridrow="5" pageId="6" pageNumber="327">4-celled</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFDA52F4DB415" box="[151,1330,512,564]" gridrow="6" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFDA52BEFB415" box="[151,400,512,564]" gridcol="0" gridrow="6" pageId="6" pageNumber="327">Subtriangle</th>
<td id="CF6ACE41C0495EA39EFCFDA5297FB415" box="[438,768,512,564]" gridcol="1" gridrow="6" pageId="6" pageNumber="327">3-celled in forewings and 1- celled in hindwings</td>
<td id="CF6ACE41C0495EA39C52FDA52E49B415" box="[792,1078,512,564]" gridcol="2" gridrow="6" pageId="6" pageNumber="327">3-celled in forewings and 2- celled in hindwings</td>
<td id="CF6ACE41C0495EA39B2FFDA52F4DB415" box="[1125,1330,512,564]" gridcol="3" gridrow="6" pageId="6" pageNumber="327">3-celled</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFDED2F4DB47F" box="[151,1330,584,606]" gridrow="7" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFDED2BEFB47F" box="[151,400,584,606]" gridcol="0" gridrow="7" pageId="6" pageNumber="327">Anal triangle</th>
<td id="CF6ACE41C0495EA39EFCFDED297FB47F" box="[438,768,584,606]" gridcol="1" gridrow="7" pageId="6" pageNumber="327">5- or 6-celled</td>
<td id="CF6ACE41C0495EA39C52FDED2E49B47F" box="[792,1078,584,606]" gridcol="2" gridrow="7" pageId="6" pageNumber="327">7 to 10-celled</td>
<td id="CF6ACE41C0495EA39B2FFDED2F4DB47F" box="[1125,1330,584,606]" gridcol="3" gridrow="7" pageId="6" pageNumber="327">5- or 6-celled</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFDD42F4DB4A6" box="[151,1330,625,647]" gridrow="8" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFDD42BEFB4A6" box="[151,400,625,647]" gridcol="0" gridrow="8" pageId="6" pageNumber="327">Ventral surface of S1</th>
<td id="CF6ACE41C0495EA39EFCFDD4297FB4A6" box="[438,768,625,647]" gridcol="1" gridrow="8" pageId="6" pageNumber="327">Without tubercle</td>
<td id="CF6ACE41C0495EA39C52FDD42E49B4A6" box="[792,1078,625,647]" gridcol="2" gridrow="8" pageId="6" pageNumber="327">Without tubercle</td>
<td id="CF6ACE41C0495EA39B2FFDD42F4DB4A6" box="[1125,1330,625,647]" gridcol="3" gridrow="8" pageId="6" pageNumber="327">With tubercle</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFD3F2F4DB4EE" box="[151,1330,666,719]" gridrow="9" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFD3F2BEFB4EE" box="[151,400,666,719]" gridcol="0" gridrow="9" pageId="6" pageNumber="327">Posterior hamulus</th>
<td id="CF6ACE41C0495EA39EFCFD3F297FB4EE" box="[438,768,666,719]" gridcol="1" gridrow="9" pageId="6" pageNumber="327">Strongly curved posteriorly</td>
<td id="CF6ACE41C0495EA39C52FD3F2E49B4EE" box="[792,1078,666,719]" gridcol="2" gridrow="9" pageId="6" pageNumber="327">Strongly curved posteriorly</td>
<td id="CF6ACE41C0495EA39B2FFD3F2F4DB4EE" box="[1125,1330,666,719]" gridcol="3" gridrow="9" pageId="6" pageNumber="327">Not strongly curved posteriorly</td>
</tr>
<tr id="8CBBA73DC0495EA39FDDFD472F4DB4D9" box="[151,1330,738,760]" gridrow="10" pageId="6" pageNumber="327">
<th id="CF6ACE41C0495EA39FDDFD472BEFB4D9" box="[151,400,738,760]" gridcol="0" gridrow="10" pageId="6" pageNumber="327">Spine on penis</th>
<td id="CF6ACE41C0495EA39EFCFD47297FB4D9" box="[438,768,738,760]" gridcol="1" gridrow="10" pageId="6" pageNumber="327">More basally, erected</td>
<td id="CF6ACE41C0495EA39C52FD472E49B4D9" box="[792,1078,738,760]" gridcol="2" gridrow="10" pageId="6" pageNumber="327">Basally, erected</td>
<td id="CF6ACE41C0495EA39B2FFD472F4DB4D9" box="[1125,1330,738,760]" gridcol="3" gridrow="10" pageId="6" pageNumber="327">Gradient</td>
</tr>
</table>
</paragraph>
<caption id="66F4F5F7C049A15A9FDDF8102E25B1CB" httpUri="https://zenodo.org/record/1043116/files/figure.png" pageId="6" pageNumber="327" targetBox="[235,1360,842,1936]" targetPageId="6">
<paragraph id="3234A57FC049A15A9FDDF8102E25B1CB" blockId="6.[151,1436,1973,2026]" pageId="6" pageNumber="327">
<emphasis id="00FF796DC049A15A9FDDF8102B28B1EA" bold="true" box="[151,343,1973,1996]" pageId="6" pageNumber="327">FIGURES 16–20.</emphasis>
<taxonomicName id="F58BDEFCC049A15A9E2AF8102807B1EA" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[352,632,1973,1995]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC049A15A9E2AF8102807B1EA" box="[352,632,1973,1995]" italics="true" pageId="6" pageNumber="327">Gomphidictinus perakensis</emphasis>
</taxonomicName>
, from Malaysia: 16) basal abdominal segments in lateral view; 17) same in dorsal ventral view; 18) cerci in lateral view; 19) cerci in dorsal view; 20) penis in lateral view.
</paragraph>
</caption>
<caption id="66F4F5F7C048A15B9FDDFB18293CB2F2" box="[151,835,1213,1235]" httpUri="https://zenodo.org/record/1043118/files/figure.png" pageId="7" pageNumber="328" targetBox="[251,1332,205,1174]" targetPageId="7">
<paragraph id="3234A57FC048A15B9FDDFB18293CB2F2" blockId="7.[151,835,1213,1235]" box="[151,835,1213,1235]" pageId="7" pageNumber="328">
<emphasis id="00FF796DC048A15B9FDDFB182B60B2F3" bold="true" box="[151,287,1213,1235]" pageId="7" pageNumber="328">FIGURE 21.</emphasis>
Habitus of 
<taxonomicName id="F58BDEFCC048A15B9EDCFB1828D5B2F2" baseAuthorityName="Laidlaw" baseAuthorityYear="1902" box="[406,682,1213,1235]" class="Insecta" family="Gomphidae" genus="Gomphidictinus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="perakensis">
<emphasis id="00FF796DC048A15B9EDCFB1828D5B2F2" box="[406,682,1213,1235]" italics="true" pageId="7" pageNumber="328">Gomphidictinus perakensis</emphasis>
</taxonomicName>
, from Yunnan.
</paragraph>
</caption>
<paragraph id="3234A57FC048A15B9F8DFB5A2F59B3CE" blockId="7.[151,1437,1279,1520]" pageId="7" pageNumber="328">
<emphasis id="00FF796DC048A15B9F8DFB5A2BE8B339" bold="true" box="[199,407,1279,1304]" pageId="7" pageNumber="328">Notes on biology.</emphasis>
Only a few adults were collected from Mt. Diaoluoshan Mountain in 
<collectingRegion id="F04F6B9DC048A15B9B8BFB5A2F6DB339" box="[1217,1298,1279,1304]" country="China" name="Hainan" pageId="7" pageNumber="328">Hainan</collectingRegion>
Island. The population is quite small and all the males were hidden in the very dense forested streams. Males held the territory quite near water, usually perch on the short trees or top of sticks, which is also seen in 
<taxonomicName id="F58BDEFCC048A15B9BC3FAE22F78B341" authorityName="Selys" authorityYear="1854" box="[1161,1287,1351,1376]" class="Insecta" family="Gomphidae" genus="Gomphidia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="00FF796DC048A15B9BC3FAE22F78B341" box="[1161,1287,1351,1376]" italics="true" pageId="7" pageNumber="328">Gomphidia</emphasis>
</taxonomicName>
males. They always perch but seldom fly, and only appear in the sunny days when the sunlight can pass through the dense tropical rain forest. Species co-occurring include the endemic 
<taxonomicName id="F58BDEFCC048A15B9C1DFA2A2F11B389" authority="Wilson &amp; Reels, 2001" authorityName="Wilson &amp; Reels" authorityYear="2001" box="[855,1390,1423,1448]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="00FF796DC048A15B9C1DFA2A2E18B389" box="[855,1127,1423,1448]" italics="true" pageId="7" pageNumber="328">Chlorogomphus gracilis</emphasis>
Wilson &amp; Reels, 2001
</taxonomicName>
, the famous 
<taxonomicName id="F58BDEFCC048A15B9FB8FA162800B3ED" authority="Kirby, 1900" authorityName="Kirby" authorityYear="1900" box="[242,639,1459,1484]" class="Insecta" family="Pseudolestidae" genus="Pseudolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="00FF796DC048A15B9FB8FA162B8FB3ED" box="[242,496,1459,1484]" italics="true" pageId="7" pageNumber="328">Pseudolestes mirabilis</emphasis>
Kirby, 1900
</taxonomicName>
and various gomphids including 
<taxonomicName id="F58BDEFCC048A15B9CB6FA162E9EB3ED" authorityName="Martin" authorityYear="1904" box="[1020,1249,1459,1484]" class="Insecta" family="Gomphidae" genus="Gomphidia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="00FF796DC048A15B9CB6FA162E9EB3ED" box="[1020,1249,1459,1484]" italics="true" pageId="7" pageNumber="328">Gomphidia kruegeri</emphasis>
</taxonomicName>
. In 
<collectingRegion id="F04F6B9DC048A15B9A5BFA162F0EB3ED" box="[1297,1393,1459,1484]" country="China" name="Guangxi" pageId="7" pageNumber="328">Guangxi</collectingRegion>
the stream is less shady and the 
<typeStatus id="ED301BDDC048A15B9E9EFA7D284BB3D1" box="[468,564,1496,1520]" pageId="7" pageNumber="328" type="paratype">paratype</typeStatus>
male and two males of 
<taxonomicName id="F58BDEFCC048A15B9C0AFA7D29C2B3D1" authorityName="Martin" authorityYear="1904" box="[832,957,1495,1520]" class="Insecta" family="Gomphidae" genus="Gomphidia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="00FF796DC048A15B9C0AFA7D29C2B3D1" box="[832,957,1495,1520]" italics="true" pageId="7" pageNumber="328">G. kruegeri</emphasis>
</taxonomicName>
were collected in the same tree.
</paragraph>
</subSubSection>
</treatment>
</document>