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<document id="7F7DF78D48FE5544C8B5CECEE710EF4E" ID-CLB-Dataset="55752" ID-DOI="10.11646/zootaxa.5115.3.7" ID-GBIF-Dataset="b6e931ce-d152-4431-83dd-1e1f107edc8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="6358479" ID-ZooBank="F9C7819F-1752-4F28-B5A6-15530B058DA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647346788143" checkinUser="plazi" docAuthor="Golovatch, Sergei I., Gallo, Jéssica S. &amp; Bichuette, Maria E." docDate="2022" docId="BA1D87E1335CFFC1FF0EFF69FCBAFF4E" docLanguage="en" docName="zootaxa.5115.3.7.pdf" docOrigin="Zootaxa 5115 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moojenodesmus schubarti Golovatch &amp; Gallo &amp; Bichuette 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="433" masterDocId="4624FF993351FFCEFF99FFFEFF9AFF9B" masterDocTitle="Two new species of the millipede family Trichopolydesmidae from Bahia state northeastern Brazil, including a remarkable threatened troglobiont (Diplopoda, Polydesmida)" masterLastPageNumber="437" masterPageNumber="419" pageNumber="432" updateTime="1699318044195" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BFDCF76C8B70624BAA18485ED4D64C98" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0E2B5014B35302767772930A74D88C80">
<mods:title id="5573E4649E4BE486D1B11278317AD91E">Two new species of the millipede family Trichopolydesmidae from Bahia state northeastern Brazil, including a remarkable threatened troglobiont (Diplopoda, Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="3EBE24FE9FB7BDAE2B79524AC1D583F1" type="personal">
<mods:role id="F24585A990923862B7F2100998ACBC0A">
<mods:roleTerm id="8780B6B6DBF94EAB80C86F6AD95953FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF316E6B302B3065AC66C875E5C514C0">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="40996F6CE413B9143FE6343B447BF183">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name id="B95B429A51B18434F2451A109264F06B" type="personal">
<mods:role id="EDE2CBE45DF0E23DF8C7849A6FD5C298">
<mods:roleTerm id="FE2D7B0226C04BAF0EACF312DCD0D5C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E15E5A1A587D72E45C94B08B44CD209E">Gallo, Jéssica S.</mods:namePart>
<mods:nameIdentifier id="607FA5BAFC03C0874E3678258E5C0A88" type="ORCID">0000-0001-9350-4686</mods:nameIdentifier>
<mods:affiliation id="A74BEABDA6F3D9DCD7535272DF609668">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brazil; &amp; jessicasgallo @ gmail. com; https: // orcid. org / 0000 - 0001 - 9350 - 4686</mods:affiliation>
<mods:nameIdentifier id="D7E7DFE9D806CB4422F625846ADA67A7" type="email">jessicasgallo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="ECEA527BF17394AD2A79665C896A4FA3" type="personal">
<mods:role id="9B2385D8B3CE20C23BDAB7B29204D120">
<mods:roleTerm id="D2DD582153DA232C095B075D647C6E74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6B356614633F51D7672E8BD5BF2A699">Bichuette, Maria E.</mods:namePart>
<mods:nameIdentifier id="DC2AAC90A3DC4B02F88980006B313C42" type="ORCID">0000-0002-9515-4832</mods:nameIdentifier>
<mods:affiliation id="308E86AB9AFD89180C164F1D7993C1A7">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brazil; &amp; lina. cave @ gmail. com; http: // orcid. org / 0000 - 0002 - 9515 - 4832</mods:affiliation>
<mods:nameIdentifier id="EAEA0988B76B902DF1039AD3B3046AB8" type="email">lina.cave@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="09FF833FEEC816F09B73EB226225D6C9">text</mods:typeOfResource>
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<mods:titleInfo id="B0A25B6D88ACE3ADEF7F2FFA56B94E19">
<mods:title id="8FF5E0DB322DE7A89EBDACD5B920992C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2F8D14098AA579DCA4CC954784B4E8AF">
<mods:date id="FAB933AE77817F4CD3CB52E0F4AF15CD">2022</mods:date>
<mods:detail id="B1975E78F1C152C50E41FE4A1053C7BC" type="pubDate">
<mods:number id="FCB3DD53DB3D72934373AA66C338AFCF">2022-03-15</mods:number>
</mods:detail>
<mods:detail id="25BF71123F177313FA3783A71040AA54" type="volume">
<mods:number id="64B34E9A336867CA4B98C67EA472267D">5115</mods:number>
</mods:detail>
<mods:detail id="16C4FDDD8279BFED38FC6CB4C31876EF" type="issue">
<mods:number id="A30CC14BFA1A86835BA5B70480E493B5">3</mods:number>
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<mods:classification id="7221AD23D17A8D0E138CDB236AE22D0B">journal article</mods:classification>
<mods:identifier id="4D1E16BDE28FE88E39EEBFF118006B9C" type="CLB-Dataset">55752</mods:identifier>
<mods:identifier id="21925D04B325F09C9C907953B641D1C6" type="DOI">10.11646/zootaxa.5115.3.7</mods:identifier>
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<treatment id="BA1D87E1335CFFC1FF0EFF69FCBAFF4E" ID-DOI="http://doi.org/10.5281/zenodo.6511721" ID-GBIF-Taxon="194952012" ID-Zenodo-Dep="6511721" LSID="urn:lsid:plazi:treatment:BA1D87E1335CFFC1FF0EFF69FCBAFF4E" httpUri="http://treatment.plazi.org/id/BA1D87E1335CFFC1FF0EFF69FCBAFF4E" lastPageId="15" lastPageNumber="433" pageId="13" pageNumber="432">
<subSubSection id="7AAE657C335CFFC3FF0EFF69FCACFF2A" box="[151,822,151,178]" pageId="13" pageNumber="432" type="nomenclature">
<paragraph id="320B36F7335CFFC3FF0EFF69FCACFF2A" blockId="13.[151,822,151,213]" box="[151,822,151,178]" pageId="13" pageNumber="432">
<heading id="6943819B335CFFC3FF0EFF69FCACFF2A" bold="true" box="[151,822,151,178]" fontSize="11" level="1" pageId="13" pageNumber="432" reason="1">
<emphasis id="00C0EAE5335CFFC3FF0EFF69FCACFF2A" bold="true" box="[151,822,151,178]" italics="true" pageId="13" pageNumber="432">
<taxonomicName id="F5B44D74335CFFC3FF0EFF69FD51FF2A" authority="Golovatch &amp; Gallo" authorityName="Golovatch &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[151,715,151,178]" class="Diplopoda" family="Trichopolydesmidae" genus="Moojenodesmus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="432" phylum="Arthropoda" rank="species" species="schubarti" status="sp. nov.">Moojenodesmus schubarti Golovatch &amp; Gallo</taxonomicName>
,
<taxonomicNameLabel id="1BF3579E335CFFC3FD41FF69FCACFF2A" box="[728,822,151,177]" pageId="13" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7AAE657C335CFFC3FF0EFF45FEB3FF4E" box="[151,297,187,213]" pageId="13" pageNumber="432" type="description">
<paragraph id="320B36F7335CFFC3FF0EFF45FEB3FF4E" blockId="13.[151,822,151,213]" box="[151,297,187,213]" pageId="13" pageNumber="432">
<figureCitation id="AA8F2A72335CFFC3FF0EFF45FF73FF4E" box="[151,233,187,213]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Figs 11</figureCitation>
&amp;
<figureCitation id="AA8F2A72335CFFC3FE94FF45FEB3FF4E" box="[269,297,187,213]" captionStart="FIGURE 12" captionStartId="15.[151,250,1628,1652]" captionTargetBox="[164,1422,248,1591]" captionTargetId="figure-44@15.[151,1436,237,1605]" captionTargetPageId="15" captionText="FIGURE 12. Moojenodesmus schubarti sp. nov., male paratype. A. Leg 9, lateral view. B. Right gonopod, posterior view. C &amp; D. Left gonopod, mesal and lateral views, respectively. Scale bars: 0.2 mm (A) and 0.1 mm (B–D). Abbreviations: ex, exomere; sl, solenomere." figureDoi="http://doi.org/10.5281/zenodo.6358508" httpUri="https://zenodo.org/record/6358508/files/figure.png" pageId="13" pageNumber="432">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7AAE657C335CFFC3FF0EFEFDFA80FEFE" pageId="13" pageNumber="432" type="materials_examined">
<paragraph id="320B36F7335CFFC3FF0EFEFDFBFDFEDA" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
<emphasis id="00C0EAE5335CFFC3FF0EFEFDFEE0FE86" bold="true" box="[151,378,259,285]" pageId="13" pageNumber="432">Material examined</emphasis>
.
<materialsCitation id="82DC3CAA335CFFC3FE10FEFDFBFDFEDA" ID-GBIF-Occurrence="3758809308" collectingDate="2013-12-08" collectionCode="LES" collectorName="M. E. Bichuette &amp; J. E. Gallao" country="Brazil" latitude="-14.435389" location="Lapa do Baixao" longLatPrecision="1" longitude="-43.624332" municipality="Iuiu Municipality" pageId="13" pageNumber="432" specimenCode="LES 0027864" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="ED0F8855335CFFC3FE10FEFDFE6BFE86" box="[393,497,259,285]" pageId="13" pageNumber="432" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="62129E8C335CFFC3FDA6FEFDFD7BFE85" box="[575,737,259,286]" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" pageId="13" pageNumber="432" type="Museum">LES 0027864</specimenCode>
),
<collectingCountry id="4AA37667335CFFC3FD6DFEFDFCA1FE86" box="[756,827,259,285]" name="Brazil" pageId="13" pageNumber="432">Brazil</collectingCountry>
,
<collectingRegion id="F070F815335CFFC3FCD1FEFDFC56FE86" box="[840,972,259,285]" country="Brazil" name="Bahia" pageId="13" pageNumber="432">Bahia State</collectingRegion>
,
<collectingMunicipality id="D26FAC8D335CFFC3FC41FEFDFB05FE86" box="[984,1183,259,285]" pageId="13" pageNumber="432">Iuiu Municipality</collectingMunicipality>
, near
<location id="376B602C335CFFC3FB7BFEFAFA02FE86" LSID="urn:lsid:plazi:treatment:BA1D87E1335CFFC1FF0EFF69FCBAFF4E:376B602C335CFFC3FB7BFEFAFA02FE86" box="[1250,1432,259,286]" country="Brazil" latitude="-14.435389" longLatPrecision="1" longitude="-43.624332" municipality="Iuiu Municipality" name="Lapa do Baixao" pageId="13" pageNumber="432" stateProvince="Bahia">Lapa do Baixão</location>
, epigean,
<geoCoordinate id="57805030335CFFC3FF65FED9FE02FEDA" box="[252,408,295,321]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="13" pageNumber="432" precision="1" seconds="07.4" value="-14.435389">S14°26’07.4”</geoCoordinate>
,
<geoCoordinate id="57805030335CFFC3FE3AFED9FDD0FEDA" box="[419,586,295,321]" degrees="43" direction="west" minutes="37" orientation="longitude" pageId="13" pageNumber="432" precision="1" seconds="27.6" value="-43.624332">W43°37’27.6”</geoCoordinate>
,
<date id="460A1037335CFFC3FDCFFED9FD49FEDA" box="[598,723,295,321]" pageId="13" pageNumber="432" value="2013-12-08">
<collectingDate id="564EE9DF335CFFC3FDCFFED9FD49FEDA" box="[598,723,295,321]" pageId="13" pageNumber="432" value="2013-12-08">8.XII.2013</collectingDate>
</date>
,
<collectorName id="9F415321335CFFC3FD46FED6FC0AFEDA" box="[735,912,295,322]" pageId="13" pageNumber="432">M. E. Bichuette</collectorName>
&amp;
<collectorName id="9F415321335CFFC3FC2DFED6FBADFEDA" box="[948,1079,295,322]" pageId="13" pageNumber="432">J. E. Gallão</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="320B36F7335CFFC3FF5EFEB2FA80FEFE" blockId="13.[151,1437,259,1834]" box="[199,1306,331,358]" pageId="13" pageNumber="432">
<materialsCitation id="82DC3CAA335CFFC3FF5EFEB2FD98FEFE" ID-GBIF-Occurrence="3758809302" box="[199,514,331,358]" collectionCode="ZMUM" pageId="13" pageNumber="432" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="ED0F8855335CFFC3FF5EFEB2FEA2FEFD" box="[199,312,332,358]" pageId="13" pageNumber="432" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="24B2FD7E335CFFC3FEDAFEB5FE14FEFE" box="[323,398,331,357]" pageId="13" pageNumber="432" type="male">1 male</specimenCount>
(
<collectionCode id="54A5AE32335CFFC3FE04FEB5FE60FEFE" box="[413,506,331,357]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="13" pageNumber="432">ZMUM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="82DC3CAA335CFFC3FD94FEB5FCBCFEFD" ID-GBIF-Occurrence="3758809301" box="[525,806,331,358]" location="LES" pageId="13" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="East" typeStatus="paratype">
<specimenCount id="24B2FD7E335CFFC3FD94FEB5FDF4FEFE" box="[525,622,331,357]" pageId="13" pageNumber="432" type="female">1 female</specimenCount>
(
<collectingRegion id="F070F815335CFFC3FDE4FEB5FD2AFEFE" box="[637,688,331,357]" country="Cameroon" name="East" pageId="13" pageNumber="432">LES</collectingRegion>
0027865)
</materialsCitation>
,
<materialsCitation id="82DC3CAA335CFFC3FCA8FEB2FA8CFEFE" ID-GBIF-Occurrence="3758809306" box="[817,1302,331,358]" location="LES" pageId="13" pageNumber="432" specimenCount="1" stateProvince="East" typeStatus="holotype">
same locality, taken together with
<typeStatus id="ED0F8855335CFFC3FB29FEB5FA8CFEFE" box="[1200,1302,331,357]" pageId="13" pageNumber="432" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7AAE657C335CFFC3FF5EFE91FE9EFE6E" pageId="13" pageNumber="432" type="materials_examined">
<paragraph id="320B36F7335CFFC3FF5EFE91FE9EFE6E" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
<emphasis id="00C0EAE5335CFFC3FF5EFE91FE97FE12" bold="true" box="[199,269,367,393]" pageId="13" pageNumber="432">Name</emphasis>
. To honour Otto Schubart (1900–1962), one of the leading specialists in the systematics of
<taxonomicName id="F5B44D74335CFFC3FABFFE91FA06FE12" authorityName="de Blainville in Gervais" authorityYear="1844" box="[1318,1436,367,393]" class="Diplopoda" kingdom="Animalia" pageId="13" pageNumber="432" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
of the time globally, Nestor of diplopodology in
<collectingCountry id="4AA37667335CFFC3FD23FE6DFC9BFE36" box="[698,769,403,429]" name="Brazil" pageId="13" pageNumber="432">Brazil</collectingCountry>
, and the author of the genus. Being resident of
<collectingCountry id="4AA37667335CFFC3FA8EFE6DFAC1FE36" box="[1303,1371,403,429]" name="Brazil" pageId="13" pageNumber="432">Brazil</collectingCountry>
since 1933, he made particularly numerous and profound contributions to the millipede fauna of that country (
<bibRefCitation id="56254B06335CFFC3FABFFE49FF62FE6E" author="Lindner, E. N. &amp; Reip, H. S. &amp; Spelda, J. &amp; Decker, P. &amp; Voigtlander, K." pageId="13" pageNumber="432" pagination="1 - 40" refId="ref9391" refString="Lindner, E. N., Reip, H. S., Spelda, J., Decker, P. &amp; Voigtlander, K. (2012) Dr. Otto Schubart (17 February 1900 - 8 November 1962) - Zum 50. Todestag. Schubartiana, 5, 1 - 40. [https: // www. researchgate. net / publication / 266614489 _ Dr _ Otto _ Schubart _ 17 _ Februar _ 1900 _ - _ 8 _ November _ 1962 _ - _ Zum _ 50 _ Todestag / link / 54379 ceb 0 cf 2 d 5 fa 292 b 46 da / download]" type="journal article" year="2012">
Lindner
<emphasis id="00C0EAE5335CFFC3FA11FE46FF2EFE6E" italics="true" pageId="13" pageNumber="432">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7AAE657C335CFFC3FF5EFE01FE3AFD1D" pageId="13" pageNumber="432" type="diagnosis">
<paragraph id="320B36F7335CFFC3FF5EFE01FE3AFD1D" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
<emphasis id="00C0EAE5335CFFC3FF5EFE01FEA0FD82" bold="true" box="[199,314,511,537]" pageId="13" pageNumber="432">Diagnosis</emphasis>
. Differs from congeners by 20 body rings in both sexes, the presence of a strong median hump (
<emphasis id="00C0EAE5335CFFC3FA1AFE01FA09FD82" bold="true" box="[1411,1427,511,537]" pageId="13" pageNumber="432">h</emphasis>
) above the antennae on the male head, coupled with a particularly simple gonopodal telopodite divided into two parts only: a prominent, rounded and slightly higher lateral lobe (= exomere,
<emphasis id="00C0EAE5335CFFC3FC58FDB9FC41FDFA" bold="true" box="[961,987,583,609]" pageId="13" pageNumber="432">ex</emphasis>
) and a barely discernible mesal solenomere (
<emphasis id="00C0EAE5335CFFC3FF46FD95FF68FD1E" bold="true" box="[223,242,619,645]" pageId="13" pageNumber="432">sl</emphasis>
) (
<figureCitation id="AA8F2A72335CFFC3FE92FD95FE0EFD1D" box="[267,404,619,646]" captionStart="FIGURE 12" captionStartId="15.[151,250,1628,1652]" captionTargetBox="[164,1422,248,1591]" captionTargetId="figure-44@15.[151,1436,237,1605]" captionTargetPageId="15" captionText="FIGURE 12. Moojenodesmus schubarti sp. nov., male paratype. A. Leg 9, lateral view. B. Right gonopod, posterior view. C &amp; D. Left gonopod, mesal and lateral views, respectively. Scale bars: 0.2 mm (A) and 0.1 mm (B–D). Abbreviations: ex, exomere; sl, solenomere." figureDoi="http://doi.org/10.5281/zenodo.6358508" httpUri="https://zenodo.org/record/6358508/files/figure.png" pageId="13" pageNumber="432">Fig. 12B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7AAE657C335CFFC3FF5EFD71FE5BF8B1" pageId="13" pageNumber="432" type="description">
<paragraph id="320B36F7335CFFC3FF5EFD71FB50FD56" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
<emphasis id="00C0EAE5335CFFC3FF5EFD71FECBFD32" bold="true" box="[199,337,655,681]" pageId="13" pageNumber="432">Description</emphasis>
. Length ca
<quantity id="F54C9B12335CFFC3FE4DFD71FDB3FD31" box="[468,553,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="432" unit="mm" value="3.5">3.5 mm</quantity>
(male holo- and
<typeStatus id="ED0F8855335CFFC3FD7CFD6EFCDDFD31" box="[741,839,656,682]" pageId="13" pageNumber="432" type="paratype">paratype</typeStatus>
) or
<quantity id="F54C9B12335CFFC3FCEBFD71FC52FD31" box="[882,968,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="432" unit="mm" value="3.8">3.8 mm</quantity>
(female
<typeStatus id="ED0F8855335CFFC3FBB0FD6EFB14FD31" box="[1065,1166,656,682]" pageId="13" pageNumber="432" type="paratype">paratype</typeStatus>
), width of midbody pro- and metazonae 0.7 and
<quantity id="F54C9B12335CFFC3FE06FD4DFE6FFD55" box="[415,501,691,718]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="432" unit="mm" value="0.8">0.8 mm</quantity>
(male, female), respectively. Coloration entirely pallid (
<figureCitation id="AA8F2A72335CFFC3FBF3FD4DFB26FD56" box="[1130,1212,691,717]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="320B36F7335CFFC3FF5EFD29FD30FB8A" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
Body with 20 rings (male, female). Tegument mainly dull, at most slightly shining, texture very delicately alveolate and scaly. Head densely pilose throughout; epicranial suture faint; isthmus between antennae ca 1.2 times broader than antennal socket (
<figureCitation id="AA8F2A72335CFFC3FE75FCE1FDC9FCA2" box="[492,595,799,825]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11C</figureCitation>
). A strong, median, rounded, bare hump (
<emphasis id="00C0EAE5335CFFC3FBB4FCE1FBA7FCA2" bold="true" box="[1069,1085,799,825]" pageId="13" pageNumber="432">h</emphasis>
) above antennae on male head (
<figureCitation id="AA8F2A72335CFFC3FF39FCBDFE9EFCC6" box="[160,260,835,861]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11B</figureCitation>
), but female head regularly convex, not modified. Antennae very short, strongly clavate due to a considerably enlarged antennomere 6, slightly produced past ring 2 (male) or collum (female) dorsally; in length, antennomere 6&gt; 2 = 3 = 5&gt; 4&gt; 7&gt; 1 (
<figureCitation id="AA8F2A72335CFFC3FD83FC75FD3DFC3E" box="[538,679,907,933]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Figs 11A–C</figureCitation>
); antennomere 6 with a large, compact, roundish, distodorsal group of bacilliform sensilla; a smaller distodorsal group of similar, but shorter sensilla present also on antennomere 5; a few minute distodorsal sensilla discernible on antennomere 4 as well (
<figureCitation id="AA8F2A72335CFFC3FC35FC2DFBAFFC76" box="[940,1077,979,1005]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11A–C</figureCitation>
). Genae regularly rounded (
<figureCitation id="AA8F2A72335CFFC3FAF6FC2DFF73FB8A" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11A, C</figureCitation>
), gnathochilarium without peculiarities.
</paragraph>
<paragraph id="320B36F7335CFFC3FF5EFBE2FE0FFA7D" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
In width, collum = ring 3 &lt;2 = 4 &lt;5(6) = 14(15) &lt;head; body gradually tapering towards telson on segments 16–20 (
<figureCitation id="AA8F2A72335CFFC3FF77FBC1FEE3FBC2" box="[238,377,1087,1113]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Figs 11D–F</figureCitation>
). Paraterga moderately developed, starting with collum, mostly clearly declivous, set high at about upper third of metazonae and always lying below a regularly convex dorsum, with squarish to narrowly rounded shoulders frontally, faintly and regularly rounded sides, and narrowly rounded caudal corners (
<figureCitation id="AA8F2A72335CFFC3FA8DFB79FAFFFB3A" box="[1300,1381,1159,1185]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11</figureCitation>
); the latter never pointed and always lying within rear tergal margin. Lateral margin of postcollum paraterga with two or three small setigerous indentations and a seta at caudal corner. Pore formula normal, ozopores inconspicuous, round, located laterally in front of caudalmost incision (
<figureCitation id="AA8F2A72335CFFC3FC9CFB0DFC10FA95" box="[773,906,1267,1294]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11D, E</figureCitation>
). Collum and following metaterga with regular, mostly subclavate, medium-sized to rather long setae borne on minute knobs, these knobs being arranged in three transverse rows typical of most polydesmoids; polygonal bosses missing (
<figureCitation id="AA8F2A72335CFFC3FC72FAC5FBCEFACE" box="[1003,1108,1339,1365]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11D</figureCitation>
). Stricture between pro- and metazonae wide, shallow and smooth. Limbus very thin, very faintly microdenticulate, nearly smooth. Pleurosternal carinae absent (
<figureCitation id="AA8F2A72335CFFC3FED3FA7DFE49FA06" box="[330,467,1411,1437]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11B, C</figureCitation>
). Epiproct short, conical, directed caudoventrally, tip subtruncate; pre-apical papillae small (
<figureCitation id="AA8F2A72335CFFC3FF7FFA59FEF4FA5A" box="[230,366,1447,1473]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11D–F</figureCitation>
). Hypoproct trapeziform, setiferous papillae at caudal corners evident, rather well separated; sides slightly concave.
</paragraph>
<paragraph id="320B36F7335CFFC3FF5EFA11FAB5F9CA" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
Sterna broad, without modifications, poorly setose (
<figureCitation id="AA8F2A72335CFFC3FC82FA11FC39F991" box="[795,931,1519,1546]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11C, F</figureCitation>
). Epigynal ridge very low. Legs moderately long and slender (
<figureCitation id="AA8F2A72335CFFC3FEFFF9EDFE20F9B6" box="[358,442,1555,1581]" captionStart="FIGURE 11" captionStartId="14.[151,250,1422,1446]" captionTargetBox="[151,1436,181,1398]" captionTargetId="figure-22@14.[151,1436,181,1398]" captionTargetPageId="14" captionText="FIGURE 11. Moojenodesmus schubarti sp. nov., male paratype. A–C. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively. Abbreviation: h, hump. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="13" pageNumber="432">Fig. 11</figureCitation>
), ca 1.2–1.3 times as long as midbody height (male, female); tarsi especially long and slender, claw very short; sphaerotrichomes missing (
<figureCitation id="AA8F2A72335CFFC3FD7AF9C9FCD6F9CA" box="[739,844,1591,1617]" captionStart="FIGURE 12" captionStartId="15.[151,250,1628,1652]" captionTargetBox="[164,1422,248,1591]" captionTargetId="figure-44@15.[151,1436,237,1605]" captionTargetPageId="15" captionText="FIGURE 12. Moojenodesmus schubarti sp. nov., male paratype. A. Leg 9, lateral view. B. Right gonopod, posterior view. C &amp; D. Left gonopod, mesal and lateral views, respectively. Scale bars: 0.2 mm (A) and 0.1 mm (B–D). Abbreviations: ex, exomere; sl, solenomere." figureDoi="http://doi.org/10.5281/zenodo.6358508" httpUri="https://zenodo.org/record/6358508/files/figure.png" pageId="13" pageNumber="432">Fig. 12A</figureCitation>
). Gonapophyses on male coxae 2 vestigial.
</paragraph>
<paragraph id="320B36F7335CFFC3FF5EF9A5FE5BF8B1" blockId="13.[151,1437,259,1834]" pageId="13" pageNumber="432">
Gonopod aperture subcordiform, broader than male prozona 7. Gonopods (
<figureCitation id="AA8F2A72335CFFC3FB8DF9A5FB01F9EE" box="[1044,1179,1627,1653]" captionStart="FIGURE 12" captionStartId="15.[151,250,1628,1652]" captionTargetBox="[164,1422,248,1591]" captionTargetId="figure-44@15.[151,1436,237,1605]" captionTargetPageId="15" captionText="FIGURE 12. Moojenodesmus schubarti sp. nov., male paratype. A. Leg 9, lateral view. B. Right gonopod, posterior view. C &amp; D. Left gonopod, mesal and lateral views, respectively. Scale bars: 0.2 mm (A) and 0.1 mm (B–D). Abbreviations: ex, exomere; sl, solenomere." figureDoi="http://doi.org/10.5281/zenodo.6358508" httpUri="https://zenodo.org/record/6358508/files/figure.png" pageId="13" pageNumber="432">Fig. 12B–D</figureCitation>
) rather simple, coxites very large, subglobose, sparsely setose laterally, cannulae typical hollow tubes; telopodites sunken deep inside a prominent gonocoel, apical parts being barely exposed; each telopodite indistinctly biramous, strongly curved caudad; prefemoral (= densely setose) parts short and held subtransversely to main body axis, but acropodites lying parallel to each other; telopodite consisting of a very short solenomere (
<emphasis id="00C0EAE5335CFFC3FC68F915FB9EF89E" bold="true" box="[1009,1028,1771,1797]" pageId="13" pageNumber="432">sl</emphasis>
) lying mesal to a prominent, round, lobe-shaped exomere (
<emphasis id="00C0EAE5335CFFC3FE0FF8F1FE2AF8B2" bold="true" box="[406,432,1807,1833]" pageId="13" pageNumber="432">ex</emphasis>
).
</paragraph>
</subSubSection>
<caption id="66CB667F335FFFC0FF0EFA70FD93FA75" ID-DOI="http://doi.org/10.5281/zenodo.6358504" ID-Zenodo-Dep="6358504" httpUri="https://zenodo.org/record/6358504/files/figure.png" pageId="14" pageNumber="433" startId="14.[151,250,1422,1446]" targetBox="[151,1436,181,1398]" targetPageId="14">
<paragraph id="320B36F7335FFFC0FF0EFA70FD93FA75" blockId="14.[151,1437,1422,1518]" pageId="14" pageNumber="433">
<emphasis id="00C0EAE5335FFFC0FF0EFA70FE84FA3D" bold="true" box="[151,286,1422,1446]" pageId="14" pageNumber="433">FIGURE 11.</emphasis>
<taxonomicName id="F5B44D74335FFFC0FEBCFA70FDAAFA3D" authority="Golovatch &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[293,560,1422,1446]" class="Diplopoda" family="Trichopolydesmidae" genus="Moojenodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="schubarti" status="sp. nov.">
<emphasis id="00C0EAE5335FFFC0FEBCFA70FDAAFA3D" box="[293,560,1422,1446]" italics="true" pageId="14" pageNumber="433">Moojenodesmus schubarti</emphasis>
</taxonomicName>
<emphasis id="00C0EAE5335FFFC0FDAEFA70FD12FA3D" bold="true" box="[567,648,1422,1446]" pageId="14" pageNumber="433">
<taxonomicNameLabel id="1BF3579E335FFFC0FDAEFA70FD12FA3D" box="[567,648,1422,1446]" pageId="14" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype.
<emphasis id="00C0EAE5335FFFC0FCA8FA70FCF8FA3D" bold="true" box="[817,866,1422,1446]" pageId="14" pageNumber="433">A–C</emphasis>
. Anterior part of body, dorsal, lateral and ventral views, respectively. D–F. Posterior half of body, dorsal, lateral and ventral views, respectively.
<emphasis id="00C0EAE5335FFFC0FB84FA4CFB36FA51" bold="true" box="[1053,1196,1458,1482]" pageId="14" pageNumber="433">Abbreviation</emphasis>
:
<emphasis id="00C0EAE5335FFFC0FB22FA4CFB53FA51" bold="true" box="[1211,1225,1458,1482]" pageId="14" pageNumber="433">h</emphasis>
, hump. Photographs by K.V. Makarov, taken not to scale.
</paragraph>
</caption>
<subSubSection id="7AAE657C335FFFC1FF5EF9E2FCBAFF4E" lastPageId="15" lastPageNumber="434" pageId="14" pageNumber="433" type="discussion">
<paragraph id="320B36F7335FFFC1FF5EF9E2FCBAFF4E" blockId="14.[151,1437,1564,2023]" lastBlockId="15.[151,1436,151,213]" lastPageId="15" lastPageNumber="434" pageId="14" pageNumber="433">
<emphasis id="00C0EAE5335FFFC0FF5EF9E2FEAEF9AD" bold="true" box="[199,308,1564,1590]" pageId="14" pageNumber="433">Remarks</emphasis>
. This species is a typical, very small, epigean
<taxonomicName id="F5B44D74335FFFC0FCDFF9E2FC66F9AD" authorityName="Schubart" authorityYear="1945" box="[838,1020,1564,1590]" class="Diplopoda" family="Trichopolydesmidae" genus="Moojenodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="00C0EAE5335FFFC0FCDFF9E2FC66F9AD" box="[838,1020,1564,1590]" italics="true" pageId="14" pageNumber="433">Moojenodesmus</emphasis>
</taxonomicName>
, the first to be recorded from
<collectingRegion id="F070F815335FFFC0FACDF9E2FA03F9AD" box="[1364,1433,1564,1590]" country="Brazil" name="Bahia" pageId="14" pageNumber="433">Bahia</collectingRegion>
. Like some of its congeners, this new species is amongst the smallest
<taxonomicName id="F5B44D74335FFFC0FC2DF9BEFBDFF9C1" box="[948,1093,1600,1626]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
on Earth. Thus, the relatively widespread, eurytopic, apparently mass and at least partly parthenogeneric
<emphasis id="00C0EAE5335FFFC0FC68F99BFBEBF9E5" box="[1009,1137,1636,1662]" italics="true" pageId="14" pageNumber="433">M. pumilus</emphasis>
<bibRefCitation id="56254B06335FFFC0FBE0F99AFABFF9E5" author="Schubart, O." box="[1145,1317,1636,1662]" pageId="14" pageNumber="433" pagination="321 - 440" refId="ref9621" refString="Schubart, O. (1944) Os diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart, 1944</bibRefCitation>
is only ca 2.0–2.2 (male) to 2.1–3.2 (female) mm long and has 19 (male) or 20 (female) body rings (
<bibRefCitation id="56254B06335FFFC0FBEFF976FAB5F939" author="Golovatch, S. I." box="[1142,1327,1672,1698]" pageId="14" pageNumber="433" pagination="207 - 226" refId="ref7882" refString="Golovatch, S. I. (1992) Review of the Neotropical fauna of the millipede family Fuhrmannodesmidae, with the description of four new species from near Manaus, Central Amazonia, Brazil (Diplopoda, Polydesmida). Amazoniana, 12 (2), 207 - 226." type="journal article" year="1992">Golovatch 1992</bibRefCitation>
). Furthermore, based on the gonopodal conformation alone,
<emphasis id="00C0EAE5335FFFC0FD4BF953FCCAF95D" box="[722,848,1708,1734]" italics="true" pageId="14" pageNumber="433">M. pumilus</emphasis>
fails to differ from
<taxonomicName id="F5B44D74335FFFC0FBB0F953FAEAF95C" authority="Schubart, 1945" authorityName="Schubart" authorityYear="1945" box="[1065,1392,1708,1735]" class="Diplopoda" family="Trichopolydesmidae" genus="Moojenodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="00C0EAE5335FFFC0FBB0F953FB24F95D" box="[1065,1214,1709,1734]" italics="true" pageId="14" pageNumber="433">M. pygmaeus</emphasis>
<bibRefCitation id="56254B06335FFFC0FB5DF952FAEAF95C" author="Schubart, O." box="[1220,1392,1708,1735]" pageId="14" pageNumber="433" pagination="1 - 156" refId="ref9643" refString="Schubart, O. (1945) Os Proterospermophora do Distrito Federal (Myriapoda, Diplopoda). Arquivos do Museu National, 38, 1 - 156." type="journal article" year="1945">Schubart, 1945</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="ED0F8855335FFFC0FF0EF92FFF5DF970" box="[151,199,1745,1771]" pageId="14" pageNumber="433">type</typeStatus>
species that seems to be bisexual and has only 18 body rings in both sexes (
<bibRefCitation id="56254B06335FFFC0FBB2F92EFB4EF971" author="Schubart, O." box="[1067,1236,1744,1770]" pageId="14" pageNumber="433" pagination="321 - 440" refId="ref9621" refString="Schubart, O. (1944) Os diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart 1944</bibRefCitation>
,
<bibRefCitation id="56254B06335FFFC0FB79F92EFA81F970" author="Schubart, O." box="[1248,1307,1744,1771]" pageId="14" pageNumber="433" pagination="1 - 156" refId="ref9643" refString="Schubart, O. (1945) Os Proterospermophora do Distrito Federal (Myriapoda, Diplopoda). Arquivos do Museu National, 38, 1 - 156." type="journal article" year="1945">1945</bibRefCitation>
;
<bibRefCitation id="56254B06335FFFC0FAB1F92EFF49F895" author="Golovatch, S. I." pageId="14" pageNumber="433" pagination="207 - 226" refId="ref7882" refString="Golovatch, S. I. (1992) Review of the Neotropical fauna of the millipede family Fuhrmannodesmidae, with the description of four new species from near Manaus, Central Amazonia, Brazil (Diplopoda, Polydesmida). Amazoniana, 12 (2), 207 - 226." type="journal article" year="1992">Golovatch 1992</bibRefCitation>
). The situation thus strongly resembles that concerning another minute polydesmidan,
<emphasis id="00C0EAE5335FFFC0FB5FF90AFF71F8A9" italics="true" pageId="14" pageNumber="433">Ammodesmus congoensis</emphasis>
<bibRefCitation id="56254B06335FFFC0FF68F8E6FDE0F8A8" author="VandenSpiegel, D. &amp; Golovatch, S. I." box="[241,634,1816,1843]" pageId="14" pageNumber="433" pagination="1 - 7" refId="ref10213" refString="VandenSpiegel, D. &amp; Golovatch, S. I. (2015) A new millipede of the family Ammodesmidae found in central Africa (Diplopoda, Polydesmida). ZooKeys, 483, 1 - 7. https: // doi. org / 10.3897 / zookeys. 483.9150" type="journal article" year="2015">VandenSpiegel &amp; Golovatch, 2015</bibRefCitation>
, a tiny member of the Afrotropical family
<taxonomicName id="F5B44D74335FFFC0FBC8F8E6FA96F8A9" authorityName="Cook" authorityYear="1896" box="[1105,1292,1816,1842]" class="Diplopoda" family="Ammodesmidae" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="family">Ammodesmidae</taxonomicName>
, superfamily
<taxonomicName id="F5B44D74335FFFC0FF0EF8C2FECEF8CD" authorityName="Silvestri" authorityYear="1896" box="[151,340,1852,1878]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="superFamily" superFamily="Pyrgodesmoidea">Pyrgodesmoidea</taxonomicName>
, which is only
<quantity id="F54C9B12335FFFC0FE60F8C2FDE5F8CC" box="[505,639,1852,1879]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="14" pageNumber="433" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.5–2.1 mm</quantity>
long and has either 18 or 19 body rings in both sexes (
<bibRefCitation id="56254B06335FFFC0FB4FF8C2FECAF8E0" author="VandenSpiegel, D. &amp; Golovatch, S. I." pageId="14" pageNumber="433" pagination="1 - 7" refId="ref10213" refString="VandenSpiegel, D. &amp; Golovatch, S. I. (2015) A new millipede of the family Ammodesmidae found in central Africa (Diplopoda, Polydesmida). ZooKeys, 483, 1 - 7. https: // doi. org / 10.3897 / zookeys. 483.9150" type="journal article" year="2015">VandenSpiegel &amp; Golovatch 2015</bibRefCitation>
). This suggests that either
<taxonomicName id="F5B44D74335FFFC0FDE5F89EFC30F8E1" authorityName="Schubart" authorityYear="1945" box="[636,938,1888,1914]" class="Diplopoda" family="Trichopolydesmidae" genus="Moojenodesmus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="433" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="00C0EAE5335FFFC0FDE5F89EFC30F8E1" box="[636,938,1888,1914]" italics="true" pageId="14" pageNumber="433">Moojenodesmus pygmaeus</emphasis>
</taxonomicName>
and
<emphasis id="00C0EAE5335FFFC0FC78F89FFBFAF8E1" box="[993,1120,1888,1914]" italics="true" pageId="14" pageNumber="433">M. pumilus</emphasis>
are indeed different species distinguished morphologically only through a lesser number of body rings in the former species or, this being not less likely now, both actually represent a single species,
<emphasis id="00C0EAE5335FFFC0FC8BF857FC0BF859" box="[786,913,1960,1986]" italics="true" pageId="14" pageNumber="433">M. pumilus</emphasis>
by priority, that shows intraspecific variations in body segment counts like those already confirmed in
<emphasis id="00C0EAE5335FFFC0FC84F832FBA6F87D" box="[797,1084,1996,2022]" italics="true" pageId="14" pageNumber="433">Ammodesmus congoensis</emphasis>
. Further studies are necessary, especially molecular-based ones, to solve the riddle. Whether such variations are related to body miniaturization or parthenogenesis, or both, is likewise open to question yet.
</paragraph>
</subSubSection>
</treatment>
</document>
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