271 lines
33 KiB
XML
271 lines
33 KiB
XML
<document id="ABFEB5D426D9D86BC3117B6C62D3A7D9" ID="10.11646/zootaxa.4138.3.9" ID-DOI="10.11646/zootaxa.4138.3.7" ID-GBIF-Dataset="80cf4fb0-6c8e-4acf-ae03-b9504ff9bab6" ID-ISSN="1175-5326" ID-Zenodo-Dep="256358" ID-ZooBank="C7391621-50DB-4070-9BCF-3D00B49F291C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468826893632" checkinUser="plazi" docAuthor="Krysko, Kenneth L., Granatosky, Michael C., Nuñez, Leroy P. & Smith, Daniel J." docDate="2016" docId="B96C87BE23055C26FF6BF9FCB4B40595" docLanguage="en" docName="zootaxa.4138.3.9.pdf" docOrigin="Zootaxa 4138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drymarchon couperi Holbrook 1842" docType="treatment" docVersion="8" lastPageNumber="562" masterDocId="4555FFC623095C2BFFFCFF8FB74A0271" masterDocTitle="A cryptic new species of Indigo Snake (genus Drymarchon) from the Florida Platform of the United States" masterLastPageNumber="569" masterPageNumber="549" pageNumber="561" updateTime="1698365206289" updateUser="plazi">
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<mods:title id="BD485E889DA8FF0AE0DD87A7C635D919">A cryptic new species of Indigo Snake (genus Drymarchon) from the Florida Platform of the United States</mods:title>
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<mods:namePart id="F9FA1DFE70E3042C9C6C1D696EF9DB19">Krysko, Kenneth L.</mods:namePart>
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<mods:namePart id="3205F7A9976D3BDE52CE03C14807CFA6">Granatosky, Michael C.</mods:namePart>
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<mods:namePart id="16CB6025380FB9DB3BD9A1C59CD58190">Nuñez, Leroy P.</mods:namePart>
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<mods:namePart id="15C63ACD41ABEC8E2FF80463F32EF519">Smith, Daniel J.</mods:namePart>
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<mods:title id="EB8C0A2F77C9EFD430C563FE165DE4CB">Zootaxa</mods:title>
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<mods:date id="07C5AC86923D420875AB6906FB97E6A9">2016</mods:date>
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<treatment id="B96C87BE23055C26FF6BF9FCB4B40595" ID-DOI="http://doi.org/10.5281/zenodo.5632172" ID-GBIF-Taxon="123196026" ID-Zenodo-Dep="5632172" LSID="urn:lsid:plazi:treatment:B96C87BE23055C26FF6BF9FCB4B40595" httpUri="http://treatment.plazi.org/id/B96C87BE23055C26FF6BF9FCB4B40595" lastPageId="13" lastPageNumber="562" pageId="12" pageNumber="561">
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<subSubSection id="79DF652323055C27FF6BF9FCB6CD04DE" pageId="12" pageNumber="561" type="nomenclature">
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<paragraph id="317A36A823055C27FF6BF9FCB53A04FF" blockId="12.[151,624,1651,1747]" box="[151,624,1651,1678]" pageId="12" pageNumber="561">
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<heading id="6A3281C423055C27FF6BF9FCB53A04FF" bold="true" box="[151,624,1651,1678]" fontSize="11" level="1" pageId="12" pageNumber="561" reason="1">
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<taxonomicName id="F6C54D2B23055C27FF6BF9FCB53A04FF" authority="Holbrook, 1842" authorityName="Holbrook" authorityYear="1842" box="[151,624,1651,1678]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="species" species="couperi">
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<emphasis id="03B1EABA23055C27FF6BF9FCB53A04FF" bold="true" box="[151,624,1651,1678]" pageId="12" pageNumber="561">
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<emphasis id="03B1EABA23055C27FF6BF9FCB6DE04FC" bold="true" box="[151,404,1651,1677]" italics="true" pageId="12" pageNumber="561">Drymarchon couperi</emphasis>
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||
(
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||
<bibRefCitation id="55544B5923055C27FE5FF9FBB52204FF" author="Holbrook" box="[419,616,1652,1678]" pageId="12" pageNumber="561" refString="Holbrook, J. E. (1842) North American Herpetology; or, a Description of the Reptiles Inhabiting the United States. [Second edition.] Vol. 3. Dobson, Philadelphia, USA, ii, 128 pp., 30 pls." type="book" year="1842">Holbrook, 1842</bibRefCitation>
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||
)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="317A36A823055C27FF6BF918B6CD04DE" blockId="12.[151,624,1651,1747]" box="[151,391,1686,1711]" pageId="12" pageNumber="561">Eastern Indigo Snake</paragraph>
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||
</subSubSection>
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||
<subSubSection id="79DF652323055C27FF6BF935B66404A2" box="[151,302,1722,1747]" pageId="12" pageNumber="561" type="description">
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<paragraph id="317A36A823055C27FF6BF935B66404A2" blockId="12.[151,624,1651,1747]" box="[151,302,1722,1747]" pageId="12" pageNumber="561">
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||
(
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||
<figureCitation id="A9FE2A2D23055C27FF62F935B66C04A2" box="[158,294,1722,1747]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="13.[151,250,813,835]" captionStartId-1="14.[151,250,1975,1997]" captionTargetBox-0="[174,1406,210,791]" captionTargetBox-1="[400,1191,195,1923]" captionTargetId-0="figure@13.[150,1436,195,791]" captionTargetId-1="figure@14.[379,1192,195,1923]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 8. Holotype (ANSP 3937) of Eastern Indigo Snake, Drymarchon couperi, presumably from Wayne County, Georgia. A) dorsal and B) ventral views." captionText-1="FIGURE 9. Holotype (ANSP 3937) of Eastern Indigo Snake, Drymarchon couperi, presumably from Wayne County, Georgia. A) lateral, B) dorsal, and C) ventral views of head region." httpUri-0="https://zenodo.org/record/256366/files/figure.png" httpUri-1="https://zenodo.org/record/256367/files/figure.png" pageId="12" pageNumber="561">Figures 8–9</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="79DF652323055C26FF6BF88DB3200198" lastPageId="13" lastPageNumber="562" pageId="12" pageNumber="561" type="reference_group">
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<paragraph id="317A36A823055C27FF6BF88DB6AE0569" blockId="12.[151,668,1794,2031]" box="[151,484,1794,1816]" pageId="12" pageNumber="561">
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<taxonomicName id="F6C54D2B23055C27FF6BF88DB6AE0569" authority="Holbrook, 1842" authorityName="Holbrook" authorityYear="1842" box="[151,484,1794,1816]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="species" species="couperi">
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<emphasis id="03B1EABA23055C27FF6BF88DB6770569" box="[151,317,1794,1816]" italics="true" pageId="12" pageNumber="561">Coluber couperi</emphasis>
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<bibRefCitation id="55544B5923055C27FEB8F88DB6AE0569" author="Holbrook" box="[324,484,1794,1816]" pageId="12" pageNumber="561" refString="Holbrook, J. E. (1842) North American Herpetology; or, a Description of the Reptiles Inhabiting the United States. [Second edition.] Vol. 3. Dobson, Philadelphia, USA, ii, 128 pp., 30 pls." type="book" year="1842">Holbrook, 1842</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
<paragraph id="317A36A823055C27FF6BF8ADB5520546" blockId="12.[151,668,1794,2031]" box="[151,536,1825,1847]" pageId="12" pageNumber="561">
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||
<emphasis id="03B1EABA23055C27FF6BF8ADB7A10546" box="[151,235,1826,1847]" italics="true" pageId="12" pageNumber="561">
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||
<collectingRegion id="F301F84A23055C27FF6BF8ADB7A10546" box="[151,235,1826,1847]" country="United States of America" name="Georgia" pageId="12" pageNumber="561">
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||
<collectingCountry id="49D2763823055C27FF6BF8ADB7A10546" box="[151,235,1826,1847]" name="Georgia" pageId="12" pageNumber="561">Georgia</collectingCountry>
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</collectingRegion>
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</emphasis>
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||
<taxonomicName id="F6C54D2B23055C27FF0DF8ADB5520546" authority="Baird & Girard 1853" authorityName="Baird & Girard" authorityYear="1853" box="[241,536,1825,1847]" class="Reptilia" family="Colubridae" genus="Coluber" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="species" species="couperi">
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<emphasis id="03B1EABA23055C27FF0DF8ADB6740546" box="[241,318,1826,1847]" italics="true" pageId="12" pageNumber="561">couperi</emphasis>
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||
<bibRefCitation id="55544B5923055C27FEB8F8AEB5520546" author="Baird" box="[324,536,1825,1847]" pageId="12" pageNumber="561" refString="Baird, S. F. & Girard, C. (1853) Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1. - Serpents. Smithsonian Institution, Washington, D. C., USA, xvi + 172 pp." type="book" year="1853">Baird & Girard 1853</bibRefCitation>
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||
</taxonomicName>
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||
</paragraph>
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||
<paragraph id="317A36A823055C27FF6BF8B0B6A40524" blockId="12.[151,668,1794,2031]" box="[151,494,1855,1878]" pageId="12" pageNumber="561">
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||
<taxonomicName id="F6C54D2B23055C27FF6BF8B0B6700527" box="[151,314,1855,1878]" class="Reptilia" family="Colubridae" genus="Spilotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="species" species="couperi">
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<emphasis id="03B1EABA23055C27FF6BF8B0B6700527" box="[151,314,1855,1878]" italics="true" pageId="12" pageNumber="561">Spilotes couperi</emphasis>
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</taxonomicName>
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(part)
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<bibRefCitation id="55544B5923055C27FE83F8B0B6A40524" author="Cope" box="[383,494,1855,1877]" pageId="12" pageNumber="561" refString="Cope, E. D. (1860) Catalogue of the Colubridae in the Museum of the Academy of Natural Sciences of Philadelphia. Part 3. Proceedings of the Academy of Natural Sciences of Philadelphia, 12, 553 - 566." type="journal article" year="1860">Cope 1860</bibRefCitation>
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||
</paragraph>
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||
<paragraph id="317A36A823055C26FF6BF8D1B3200198" blockId="12.[151,668,1794,2031]" lastBlockId="13.[151,1437,904,2021]" lastPageId="13" lastPageNumber="562" pageId="12" pageNumber="561">
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||
<taxonomicName id="F6C54D2B23055C27FF6BF8D1B6CD0505" box="[151,391,1886,1908]" class="Reptilia" family="Colubridae" genus="Spilotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="subSpecies" species="corais" subSpecies="couperii">
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<emphasis id="03B1EABA23055C27FF6BF8D1B6CD0505" box="[151,391,1886,1908]" italics="true" pageId="12" pageNumber="561">Spilotes corais couperii</emphasis>
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</taxonomicName>
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(part)
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||
<bibRefCitation id="55544B5923055C27FE30F8D1B52C0505" author="Lonnberg" box="[460,614,1886,1908]" pageId="12" pageNumber="561" refString="Lonnberg, E. (1894) Notes on reptiles and batrachians collected in Florida in 1892 and 1893. Proceedings of the United States National Museum, 17, 317 - 339. http: // dx. doi. org / 10.5479 / si. 00963801.17 - 1003.317" type="journal article" year="1894">Lönnberg 1894</bibRefCitation>
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||
<taxonomicName id="F6C54D2B23055C27FF6BF8F2B6F505E2" box="[151,447,1917,1939]" class="Insecta" family="Cerambycidae" genus="Compsosoma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="561" phylum="Arthropoda" rank="subSpecies" species="corais" subSpecies="couperii">
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<emphasis id="03B1EABA23055C27FF6BF8F2B6F505E2" box="[151,447,1917,1939]" italics="true" pageId="12" pageNumber="561">Compsosoma corais couperii</emphasis>
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||
</taxonomicName>
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||
(part)
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||
<bibRefCitation id="55544B5923055C27FDF8F8F3B53905E3" author="Cope" box="[516,627,1916,1938]" pageId="12" pageNumber="561" refString="Cope, E. D. (1900) The crocodilians, lizards, and snakes of North America. Report of the U. S. National Museum 1898, 151 - 1294 pp." type="journal article" year="1900">Cope 1900</bibRefCitation>
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||
<taxonomicName id="F6C54D2B23055C27FF6BF814B6CA05C0" box="[151,384,1947,1969]" class="Reptilia" family="Colubridae" genus="Spilotes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="subSpecies" species="corais" subSpecies="couperi">
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||
<emphasis id="03B1EABA23055C27FF6BF814B6CA05C0" box="[151,384,1947,1969]" italics="true" pageId="12" pageNumber="561">Spilotes corais couperi</emphasis>
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||
</taxonomicName>
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||
(part)
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||
<bibRefCitation id="55544B5923055C27FE39F813B50905C0" author="Brown" box="[453,579,1947,1969]" pageId="12" pageNumber="561" refString="Brown, A. E. (1901) A review of the genera and species of American snakes, north of Mexico. Proceedings of the Academy of Natural Science of Philadelphia, 53, 10 - 119." type="journal article" year="1901">Brown 1901</bibRefCitation>
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||
<taxonomicName id="F6C54D2B23055C27FF6BF835B6FA05A1" box="[151,432,1978,2000]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="subSpecies" species="corais" subSpecies="couperi">
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||
<emphasis id="03B1EABA23055C27FF6BF835B6FA05A1" box="[151,432,1978,2000]" italics="true" pageId="12" pageNumber="561">Drymarchon corais couperi</emphasis>
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||
</taxonomicName>
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||
(part)
|
||
<bibRefCitation id="55544B5923055C27FE09F835B53105A1" author="Amaral" box="[501,635,1978,2000]" pageId="12" pageNumber="561" refString="Amaral, A. do (1929) Estudos sobre ofidios neotropicos. XXI. Revisao do genero Drymarchon Fitzinger. Memorias do Instituto Butantan, 4, 323 - 330." type="journal article" year="1929">Amaral 1929</bibRefCitation>
|
||
<taxonomicName id="F6C54D2B23055C27FF6BF856B620059E" box="[151,362,2009,2031]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="561" phylum="Chordata" rank="species" species="couperi">
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||
<emphasis id="03B1EABA23055C27FF6BF856B620059E" box="[151,362,2009,2031]" italics="true" pageId="12" pageNumber="561">Drymarchon couperi</emphasis>
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||
</taxonomicName>
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||
(part)
|
||
<bibRefCitation id="55544B5923055C27FE4CF856B5D6059E" author="Conant" box="[432,668,2009,2031]" pageId="12" pageNumber="561" refString="Conant, R. & Collins, J. T. (1991) A Field Guide to Reptiles and Amphibians: Eastern and Central North America. Third edition. Houghton Mifflin Company, Boston, Massachusetts, USA, 592 pp." type="book" year="1991">Conant & Collins 1991</bibRefCitation>
|
||
<emphasis id="03B1EABA23045C26FF6BFC06B64101D0" bold="true" box="[151,267,905,929]" pageId="13" pageNumber="562">
|
||
<typeStatus id="EE7E880A23045C26FF6BFC06B64C01D0" box="[151,262,905,929]" pageId="13" pageNumber="562" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
Academy of Natural Sciences of Drexel University (ANSP 3937); although there is no known collector, locality, nor date assigned to this specimen, it is presumably collected by J.H. Couper (
|
||
<bibRefCitation id="55544B5923045C26FB8DFC23B26D01B4" author="McCranie" box="[1137,1319,940,965]" pageId="13" pageNumber="562" refString="McCranie, J. R. (1980) Drymarchon, D. corais. Catalogue of American Amphibians and Reptiles, 267, 1 - 4." type="journal article" year="1980">McCranie 1980</bibRefCitation>
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||
), from the Altamaha River (
|
||
<bibRefCitation id="55544B5923045C26FEA4FC5FB50C0198" author="Baird" box="[344,582,976,1001]" pageId="13" pageNumber="562" refString="Baird, S. F. & Girard, C. (1853) Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1. - Serpents. Smithsonian Institution, Washington, D. C., USA, xvi + 172 pp." type="book" year="1853">Baird & Girard 1853</bibRefCitation>
|
||
), Wayne County,
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||
<collectingRegion id="F301F84A23045C26FCF1FC5FB4200198" box="[781,874,976,1001]" country="United States of America" name="Georgia" pageId="13" pageNumber="562">
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||
<collectingCountry id="49D2763823045C26FCF1FC5FB4200198" box="[781,874,976,1001]" name="Georgia" pageId="13" pageNumber="562">Georgia</collectingCountry>
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||
</collectingRegion>
|
||
,
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||
<collectingCountry id="49D2763823045C26FC89FC5EB4E70198" box="[885,941,977,1001]" name="United States of America" pageId="13" pageNumber="562">USA</collectingCountry>
|
||
(
|
||
<bibRefCitation id="55544B5923045C26FC47FC5FB3140198" author="Schmidt" box="[955,1118,976,1001]" pageId="13" pageNumber="562" refString="Schmidt, K. P. (1953) A Check List of North American Amphibians and Reptiles. American Society of Ichthyologists and Herpetologists. Chicago, Illinois, USA, 280 pp." type="book" year="1953">Schmidt 1953</bibRefCitation>
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||
).
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||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="65BA662023045C26FF6BFCA2B69F0110" httpUri="https://zenodo.org/record/256366/files/figure.png" pageId="13" pageNumber="562" targetBox="[174,1406,210,791]" targetPageId="13">
|
||
<paragraph id="317A36A823045C26FF6BFCA2B69F0110" blockId="13.[151,1437,812,866]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF6BFCA2B6580133" bold="true" box="[151,274,813,835]" pageId="13" pageNumber="562">FIGURE 8.</emphasis>
|
||
Holotype (ANSP 3937) of Eastern Indigo Snake,
|
||
<taxonomicName id="F6C54D2B23045C26FCF2FCA3B4A80132" box="[782,994,812,835]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="562" phylum="Chordata" rank="species" species="couperi">
|
||
<emphasis id="03B1EABA23045C26FCF2FCA3B4A80132" box="[782,994,812,835]" italics="true" pageId="13" pageNumber="562">Drymarchon couperi</emphasis>
|
||
</taxonomicName>
|
||
, presumably from Wayne County, Georgia. A) dorsal and B) ventral views.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="79DF652323045C26FF3BFC7AB3D60627" pageId="13" pageNumber="562" type="etymology">
|
||
<paragraph id="317A36A823045C26FF3BFC7AB3D60627" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF3BFC7AB604067C" bold="true" box="[199,334,1013,1037]" pageId="13" pageNumber="562">Etymology.</emphasis>
|
||
<taxonomicName id="F6C54D2B23045C26FEA4FC7AB6AC067C" box="[344,486,1013,1037]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="562" phylum="Chordata" rank="genus">
|
||
<emphasis id="03B1EABA23045C26FEA4FC7AB6AC067C" box="[344,486,1013,1037]" italics="true" pageId="13" pageNumber="562">Drymarchon</emphasis>
|
||
</taxonomicName>
|
||
is derived from the Greek
|
||
<taxonomicName id="F6C54D2B23045C26FCD5FC79B4C8067C" box="[809,898,1014,1037]" class="Aves" family="Petroicidae" genus="Drymos" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="562" phylum="Chordata" rank="genus">
|
||
<emphasis id="03B1EABA23045C26FCD5FC79B4C8067C" box="[809,898,1014,1037]" italics="true" pageId="13" pageNumber="562">Drymos</emphasis>
|
||
</taxonomicName>
|
||
(meaning Oak Coppice or Forest) and
|
||
<emphasis id="03B1EABA23045C26FAADFC7AB2D6067C" box="[1361,1436,1013,1037]" italics="true" pageId="13" pageNumber="562">archos</emphasis>
|
||
(meaning Commander or Chief), referring to it being the chief of the oak woodland. The species epithet
|
||
<taxonomicName id="F6C54D2B23045C26FAD0FB96B2CB0641" box="[1324,1409,1049,1072]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="562" phylum="Chordata" rank="species" species="couperi">
|
||
<emphasis id="03B1EABA23045C26FAD0FB96B2CB0641" box="[1324,1409,1049,1072]" italics="true" pageId="13" pageNumber="562">couperi</emphasis>
|
||
</taxonomicName>
|
||
is derived from the surname of J.H. Couper, who supposedly collected the
|
||
<typeStatus id="EE7E880A23045C26FC3DFBB2B3690627" box="[961,1059,1085,1110]" pageId="13" pageNumber="562" type="holotype">holotype</typeStatus>
|
||
specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="79DF652323045C26FF3BFBEFB35B06EC" pageId="13" pageNumber="562" type="distribution">
|
||
<paragraph id="317A36A823045C26FF3BFBEFB35B06EC" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF3BFBEFB62A0608" bold="true" box="[199,352,1120,1145]" pageId="13" pageNumber="562">Distribution.</emphasis>
|
||
Occurs from southeastern
|
||
<collectingRegion id="F301F84A23045C26FD53FBEFB4460608" box="[687,780,1120,1145]" country="United States of America" name="Georgia" pageId="13" pageNumber="562">
|
||
<collectingCountry id="49D2763823045C26FD53FBEFB4460608" box="[687,780,1120,1145]" name="Georgia" pageId="13" pageNumber="562">Georgia</collectingCountry>
|
||
</collectingRegion>
|
||
, southward into
|
||
<collectingRegion id="F301F84A23045C26FC22FBEFB3650608" box="[990,1071,1120,1145]" country="United States of America" name="Florida" pageId="13" pageNumber="562">Florida</collectingRegion>
|
||
from Suwannee and Alachua counties, southeastward to Marion, Osceola, and Indian River counties (
|
||
<figureCitation id="A9FE2A2D23045C26FC43FB0AB34906EC" box="[959,1027,1157,1182]" captionStart="FIGURE 7" captionStartId="12.[151,250,1343,1365]" captionTargetBox="[151,1423,193,1321]" captionTargetId="figure@12.[151,1435,193,1322]" captionTargetPageId="12" captionText="FIGURE 7. Distribution of Indigo Snake (Drymarchon) samples on the subaerially exposed Florida Platform in Florida and Georgia. Atlantic Lineage (blue triangles; with dots = genetic samples, without dots = this study using morphology based on cluster analysis); Gulf Lineage (red circles; with dots = genetic samples, without dots = this study using morphology based on cluster analysis); and stars represent holotype specimens for D. couperi (blue) and D. kolpobasileus (red). Genetic data taken from Krysko et al. (2016)." httpUri="https://zenodo.org/record/256365/files/figure.png" pageId="13" pageNumber="562">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="79DF652323045C26FF3BFB27B621070A" pageId="13" pageNumber="562" type="diagnosis">
|
||
<paragraph id="317A36A823045C26FF3BFB27B621070A" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF3BFB27B60B06B0" bold="true" box="[199,321,1192,1217]" pageId="13" pageNumber="562">Diagnosis.</emphasis>
|
||
<taxonomicName id="F6C54D2B23045C26FEBBFB26B57B06B1" box="[327,561,1193,1217]" class="Reptilia" family="Colubridae" genus="Drymarchon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="562" phylum="Chordata" rank="species" species="couperi">
|
||
<emphasis id="03B1EABA23045C26FEBBFB26B57B06B1" box="[327,561,1193,1217]" italics="true" pageId="13" pageNumber="562">Drymarchon couperi</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by a suite of molecular and morphological features, including relatively longer and deeper head dimensions, longer and shallower 7th infralabials, and longer temporal scales. Overall, the presence of a longer and shallow 7th infralabial scale provides the best univariate predictor for this species (
|
||
<tableCitation id="7C47031323045C26FF0BFA95B6050742" box="[247,335,1306,1331]" captionStart="TABLE 3" captionStartId="10.[151,239,756,779]" captionTargetBox="[159,1385,839,975]" captionTargetPageId="10" captionText="TABLE 3. Summary statistics for morphological variables (mean ± standard deviation) arranged by species. All values are presented as geometric shape variables." httpUri="http://table.plazi.org/id/65BA662023035C21FF6BFD7BB52E015A" pageId="13" pageNumber="562" tableUuid="65BA662023035C21FF6BFD7BB52E015A">Table 3</tableCitation>
|
||
;
|
||
<figureCitation id="A9FE2A2D23045C26FEA1FA95B6E80742" box="[349,418,1306,1331]" captionStart="FIGURE 5" captionStartId="10.[151,250,599,621]" captionTargetBox="[160,1429,220,541]" captionTargetId="figure@10.[150,1437,213,550]" captionTargetPageId="10" captionText="FIGURE 5. Differences in infralabial and temporal scale morphology between the A) Eastern Indigo Snake, Drymarchon couperi (Holbrook, 1842), and B) Gulf Coast Indigo Snake, D. kolpobasileus, found in this study. Illustrations originally taken and modified from Cope (1900). Note that D. kolpobasileus has relatively deeper and shorter 7 th infralabial scales, shorter temporal scales, and overall shorter and shallower head dimensions (not depicted)." httpUri="https://zenodo.org/record/256363/files/figure.png" pageId="13" pageNumber="562">Fig. 5</figureCitation>
|
||
). Based on both DNA (
|
||
<bibRefCitation id="55544B5923045C26FD4CFA95B4C20742" author="Krysko" box="[688,904,1306,1331]" pageId="13" pageNumber="562" refString="Krysko, K. L., Nunez, L. P., Lippi, C. A., Smith, D. J. & Granatosky, M. C. (2016) Pliocene-Pleistocene lineage diversifications in the Eastern Indigo Snake (Drymarchon couperi) in the southeastern United States. Molecular Phylogenetics and Evolution, 98, 111 - 122. http: // dx. doi. org / 10.1016 / j. ympev. 2015.12.022" type="journal article" year="2016">
|
||
Krysko
|
||
<emphasis id="03B1EABA23045C26FCF1FA94B40C0743" box="[781,838,1306,1330]" italics="true" pageId="13" pageNumber="562">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
) and morphology (specimens examined in this study) this species includes populations from southeastern
|
||
<collectingRegion id="F301F84A23045C26FCC4FAB1B4D80726" box="[824,914,1342,1367]" country="United States of America" name="Georgia" pageId="13" pageNumber="562">
|
||
<collectingCountry id="49D2763823045C26FCC4FAB1B4D80726" box="[824,914,1342,1367]" name="Georgia" pageId="13" pageNumber="562">Georgia</collectingCountry>
|
||
</collectingRegion>
|
||
southward along the Atlantic coast to central peninsular
|
||
<collectingRegion id="F301F84A23045C26FEEFFAEDB62D070A" box="[275,359,1378,1403]" country="United States of America" name="Florida" pageId="13" pageNumber="562">Florida</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="79DF652323045C26FF3BFA09B4B40595" pageId="13" pageNumber="562" type="description">
|
||
<paragraph id="317A36A823045C26FF3BFA09B50104B4" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF3BFA09B6AC07EE" bold="true" box="[199,486,1414,1439]" pageId="13" pageNumber="562">
|
||
Description of
|
||
<typeStatus id="EE7E880A23045C26FE84FA08B6AB07EE" box="[376,481,1415,1439]" pageId="13" pageNumber="562" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male; Total Length
|
||
<quantity id="F63D9B4D23045C26FCE9FA09B4C807EF" box="[789,898,1414,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.6309999999999998" pageId="13" pageNumber="562" unit="cm" value="163.1">163.1 cm</quantity>
|
||
; Snout-vent Length
|
||
<quantity id="F63D9B4D23045C26FB94FA08B39F07EF" box="[1128,1237,1415,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.331" pageId="13" pageNumber="562" unit="cm" value="133.1">133.1 cm</quantity>
|
||
; Tail Length 30.0 cm; Head Length
|
||
<quantity id="F63D9B4D23045C26FEA3FA25B68C07B3" box="[351,454,1450,1475]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" pageId="13" pageNumber="562" unit="mm" value="51.3">51.3 mm</quantity>
|
||
; Head Width
|
||
<quantity id="F63D9B4D23045C26FDA3FA25B58C07B3" box="[607,710,1450,1475]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="13" pageNumber="562" unit="mm" value="25.2">25.2 mm</quantity>
|
||
. The relative head length (geometric shape variable; see
|
||
<tableCitation id="7C47031323045C26FAC3FA25B2D607B2" box="[1343,1436,1450,1475]" captionStart="TABLE 2" captionStartId="8.[119,195,491,510]" captionText="TABLE 2. Summary statistics for morphological variables (mean ± stanđarđ đeviation) arrangeđ by species anđ county of origin. All values are presenteđ as geometric shape variables. Note that holotype specimens are inđicateđ in bolđ." pageId="13" pageNumber="562">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="7C47031323045C26FF37FA40B7930796" box="[203,217,1487,1511]" captionStart="TABLE 3" captionStartId="10.[151,239,756,779]" captionTargetBox="[159,1385,839,975]" captionTargetPageId="10" captionText="TABLE 3. Summary statistics for morphological variables (mean ± standard deviation) arranged by species. All values are presented as geometric shape variables." httpUri="http://table.plazi.org/id/65BA662023035C21FF6BFD7BB52E015A" pageId="13" pageNumber="562" tableUuid="65BA662023035C21FF6BFD7BB52E015A">3</tableCitation>
|
||
) is short (3.80) and relative head height is narrow (1.98). Supralabials 8/8 (left/right); Infralabials 9/9; Oculars 1+2/1+2; Temporals 2+2/2+2 (divided). The 7th infralabial is relatively long (0.65) and narrow (0.26) (geometric shape variable; see
|
||
<tableCitation id="7C47031323045C26FE02F993B52A0445" box="[510,608,1564,1589]" captionStart="TABLE 2" captionStartId="8.[119,195,491,510]" captionText="TABLE 2. Summary statistics for morphological variables (mean ± stanđarđ đeviation) arrangeđ by species anđ county of origin. All values are presenteđ as geometric shape variables. Note that holotype specimens are inđicateđ in bolđ." pageId="13" pageNumber="562">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="7C47031323045C26FD61F993B5E10445" box="[669,683,1564,1588]" captionStart="TABLE 3" captionStartId="10.[151,239,756,779]" captionTargetBox="[159,1385,839,975]" captionTargetPageId="10" captionText="TABLE 3. Summary statistics for morphological variables (mean ± standard deviation) arranged by species. All values are presented as geometric shape variables." httpUri="http://table.plazi.org/id/65BA662023035C21FF6BFD7BB52E015A" pageId="13" pageNumber="562" tableUuid="65BA662023035C21FF6BFD7BB52E015A">3</tableCitation>
|
||
) and the temporal scale is relatively short (0.78). The 5th and 7th supralabials are in contact with each other. Two pairs of chin shields, both of which are in contact with each other; posterior pair slightly narrower than anterior pair. The rostral visible from above, broader than high. Dorsal scales smooth and in rows at Mid-Body 17, Anterior 17 and Posterior 15; Ventral Scales 184; cloaca undivided; Subcaudal Scale Total 59 (all divided).
|
||
</paragraph>
|
||
<paragraph id="317A36A823045C26FF3BF95FB2570524" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">The specimen has faded from its likely original solid black dorsum. There is light pigment (likely reddish in real life) on the rostral, loreals, labials and chin shields that extend posteriorly onto the first three to four ventrals. The first 40 ventral scales are mottled or bicolored with different degrees of black (posteriorly) and light colored (distally) pigmentation with the remaining ventrals predominantly black. Subcaudals are entirely black.</paragraph>
|
||
<paragraph id="317A36A823045C26FF3BF8D0B4B40595" blockId="13.[151,1437,904,2021]" pageId="13" pageNumber="562">
|
||
<emphasis id="03B1EABA23045C26FF3BF8D0B6910509" bold="true" box="[199,475,1887,1912]" pageId="13" pageNumber="562">Intraspecific variation.</emphasis>
|
||
Ventrals range from 182–197 (mean = 188,
|
||
<emphasis id="03B1EABA23045C26FC1EF8EEB4BA0509" box="[994,1008,1889,1912]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 30); subcaudals range from 60–67 (mean = 63,
|
||
<emphasis id="03B1EABA23045C26FEDBF80AB67F05ED" box="[295,309,1925,1948]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 19); supralabials are arranged 8/8 (
|
||
<emphasis id="03B1EABA23045C26FD23F80AB5A705ED" box="[735,749,1925,1948]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 28) and 7/7 (
|
||
<emphasis id="03B1EABA23045C26FC63F80AB4E705ED" box="[927,941,1925,1948]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 2); infralabials are arranged 9/9 (
|
||
<emphasis id="03B1EABA23045C26FABDF80AB20505ED" box="[1345,1359,1925,1948]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 30); temporals are arranged 2 + 2 (
|
||
<emphasis id="03B1EABA23045C26FE1AF826B6BE05B1" box="[486,500,1961,1984]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 30), some individuals with smaller divided scales; oculars are arranged 1 + 2 (
|
||
<emphasis id="03B1EABA23045C26FA8BF826B2CF05B1" box="[1399,1413,1961,1984]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 30); and DSR are 15-17-15 (
|
||
<emphasis id="03B1EABA23045C26FE24F842B6AC0595" box="[472,486,1997,2020]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 17), 17-17-15 (
|
||
<emphasis id="03B1EABA23045C26FD52F842B5F60595" box="[686,700,1997,2020]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 12), and 17-19-15 (
|
||
<emphasis id="03B1EABA23045C26FC48F842B4880595" box="[948,962,1997,2020]" italics="true" pageId="13" pageNumber="562">n</emphasis>
|
||
= 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |