treatments-xml/data/B9/4D/5B/B94D5B1AFFF4F66C587423C359CCACB6.xml

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<mods:title id="A641167C8552FD8045BBB3045E300C80">Six new species of Danais Vent. (Rubiaceae, Danaideae) from Madagascar</mods:title>
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<mods:namePart id="AF65A827EAC3A5E0BFA129668082714B">Taylor, Charlotte M.</mods:namePart>
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<mods:namePart id="FC5E406F472ED92745BB843B13384331">Rogers, Zachary S.</mods:namePart>
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<mods:title id="CCF7478466B76DB75CE31856F9AC28B0">Candollea</mods:title>
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<paragraph id="315BEA0CFFF4F663587423C35929ABD7" blockId="7.[98,749,925,949]" box="[98,606,925,949]" pageId="7" pageNumber="174">
<heading id="6A135D60FFF4F663587423C35929ABD7" box="[98,606,925,949]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="174" reason="3">
<emphasis id="0390361EFFF4F663587423C35B01ABD7" bold="true" box="[98,118,925,949]" pageId="7" pageNumber="174">3.</emphasis>
<taxonomicName id="F6E4918FFFF4F663589E23C35AADABD7" ID-CoL="343PZ" authority="C. M. Taylor" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[136,474,925,949]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="species" species="laciniata" status="sp. nov.">
<emphasis id="0390361EFFF4F663589E23C35A33ABD7" bold="true" box="[136,324,925,949]" italics="true" pageId="7" pageNumber="174">Danais laciniata</emphasis>
C. M. Taylor
</taxonomicName>
, 
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<heading id="6A135D60FFF4F6635A7023C3599AABD7" box="[614,749,925,949]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="174" reason="3">
(
<figureCitation id="A9DFF689FFF4F6635A7823C359D9ABD7" box="[622,686,925,949]" captionStart="Fig" captionStartId="5.[98,123,1867,1887]" captionTargetBox="[97,1475,275,1818]" captionTargetId="figure-8@5.[163,1409,277,1816]" captionTargetPageId="5" captionText="Fig. 3. – Danais disticha C. M. Taylor. A. Flowering branch; B. Portion of inflorescence with three flowers at anthesis. – Danais laciniata C. M. Taylor. C. Fruiting branch; D. Stem node with two reflexed stipules, and petioles and bases of two leaves; E. Stipule, abaxial face. [A-B: Rakoto 29, MO; C: Antilahimena 2627, MO; D-E: Schatz &amp; Antilahimena 4007, MO] [Drawing: C. M. Taylor]" figureDoi="http://doi.org/10.5281/zenodo.6045564" httpUri="https://zenodo.org/record/6045564/files/figure.png" pageId="7" pageNumber="174">Fig. 3</figureCitation>
C-E).
</heading>
</paragraph>
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<subSubSection id="79FEB987FFF4F663589E239B5A4FACE2" pageId="7" pageNumber="174" type="materials_examined">
<paragraph id="315BEA0CFFF4F663589E239B5A4FACE2" blockId="7.[135,748,965,1152]" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F663589E239B5BADABBF" bold="true" box="[136,218,965,989]" pageId="7" pageNumber="174">Typus:</emphasis>
<materialsCitation id="818CE051FFF4F66358F7239B59C5AC02" ID-GBIF-Occurrence="3463315311" collectingDate="2002-11-20" collectionCode="MO" collectorName="Schatz &amp; Antilahimena" country="Madagascar" elevation="150" location="Ankirindro Massif, slopes above the village Ambodivato" municipality="Toamasina" pageId="7" pageNumber="174" specimenCode="MO-5989728" specimenCount="1" stateProvince="Analanjirofo" typeStatus="holotype">
<emphasis id="0390361EFFF4F66358F7239B592EABBF" bold="true" box="[225,601,965,989]" pageId="7" pageNumber="174">
<collectingCountry id="49F3AA9CFFF4F66358F7239B5A09ABBE" box="[225,382,965,989]" pageId="7" pageNumber="174">MADAGASCAR</collectingCountry>
. Prov. 
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:
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<collectingRegion id="F32024EEFFF4F6635A77239B599BABBF" box="[609,748,965,989]" pageId="7" pageNumber="174">Analanjirofo</collectingRegion>
Region, 
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, ca. 
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NW of Ambinanitelo along the Vohimaro River
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, 
<geoCoordinate id="54D08CCBFFF4F66358C724795A95AC5D" box="[209,482,1063,1087]" pageId="7" pageNumber="174">15°19’13”S 49°33’29”E</geoCoordinate>
, 
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<elevation id="BAC90D3FFFF4F66359FB24795941AC5D" box="[493,566,1063,1087]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="174" unit="m" value="150.0">150 m</elevation>
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, 
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, fr., 
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<collectorName id="9C118FDAFFF4F663589124165BBAAC02" box="[135,205,1096,1120]" pageId="7" pageNumber="174">Schatz</collectorName>
&amp; 
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4007
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(
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: 
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!
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; 
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: 
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, 
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, 
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, 
<collectionCode id="57F572C9FFF4F66358E524365A5AACE2" box="[243,301,1128,1152]" pageId="7" pageNumber="174">TAN</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F663589E24CF5B9EAD68" pageId="7" pageNumber="174" type="diagnosis">
<paragraph id="315BEA0CFFF4F663589E24CF5B9EAD68" blockId="7.[135,748,1169,1290]" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F663589E24CF5B9EAD68" italics="true" pageId="7" pageNumber="174">Haec species acongenerispubescentiahispida, foliisternatis, stipulis reflexis laciniatis, inflorescentia terminali, pedicellis longis atque lobis calycinis longis linearibus distinguitur.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F663589125455E1FAA90" pageId="7" pageNumber="174" type="description">
<paragraph id="315BEA0CFFF4F663589125455E1FAA90" blockId="7.[97,750,1307,1951]" lastBlockId="7.[824,1475,276,754]" pageId="7" pageNumber="174">
Woody 
<emphasis id="0390361EFFF4F66358CA25455A69AD51" box="[220,286,1307,1331]" italics="true" pageId="7" pageNumber="174">lianas</emphasis>
or climbing 
<emphasis id="0390361EFFF4F66359BF25455A86AD51" box="[425,497,1307,1331]" italics="true" pageId="7" pageNumber="174">shrubs</emphasis>
, climbing to 
<quantity id="F61C47E9FFF4F6635A94254559DCAD51" box="[642,683,1307,1331]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="174" unit="m" value="7.0">7 m</quantity>
high; stems moderately to densely hispid or pilose, subterete. 
<emphasis id="0390361EFFF4F6635AB52565599BAD31" box="[675,748,1339,1363]" italics="true" pageId="7" pageNumber="174">Leaves</emphasis>
opposite or ternate with arrangement often variable on a single stem; blades obovate to oblanceolate or elliptic, 2.5-8.5 × 
<quantity id="F61C47E9FFF4F663587425C35BC6ADD7" box="[98,177,1437,1461]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="174" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 cm</quantity>
, at base acute to cuneate, at apex acute to acuminate with tips 
<quantity id="F61C47E9FFF4F66358D425E05A53ADB4" box="[194,292,1470,1494]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="7" pageNumber="174" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3-10 mm</quantity>
, drying papery, adaxially moderately hispid to strigose throughout, abaxially moderately to densely hispid to strigose with pubescence denser on principal veins, with margins plane; secondary veins 7 to 11 pairs, not looping to interconnect with each other, adaxially costa and secondary veins prominulous and remaining venation plane, abaxially costa prominent, secondary veins prominulous, intersecondary veins inconspicuous or hardly developed, and tertiary venation loosely reticulated and plane to prominulous. 
<emphasis id="0390361EFFF4F66359F1269D594DAEB9" box="[487,570,1731,1755]" italics="true" pageId="7" pageNumber="174">Petioles</emphasis>
<quantity id="F61C47E9FFF4F6635A56269D59E4AEB9" box="[576,659,1731,1755]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="7" pageNumber="174" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 mm</quantity>
, densely hispid to pilose. 
<emphasis id="0390361EFFF4F663591B26BD5A28AE99" box="[269,351,1763,1787]" italics="true" pageId="7" pageNumber="174">Stipules</emphasis>
interpetiolar, generally persistent with leaves and becoming reflexed, deeply laciniate, with unlobed basal portion truncate to triangular, 
<quantity id="F61C47E9FFF4F66359C1277B5937AF5F" box="[471,576,1829,1853]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="7" pageNumber="174" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">1-2.5 mm</quantity>
, with laciniae or lobes 3 to 6, narrowly triangular, 
<quantity id="F61C47E9FFF4F66359D7271B5960AF3F" box="[449,535,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="7" pageNumber="174" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2-4 mm</quantity>
, acute to acuminate, entire to ciliate. 
<emphasis id="0390361EFFF4F663591D27385AD7AF1C" box="[267,416,1894,1918]" italics="true" pageId="7" pageNumber="174">Inflorescences</emphasis>
and 
<emphasis id="0390361EFFF4F66359C727385969AF1C" box="[465,542,1894,1918]" italics="true" pageId="7" pageNumber="174">flowers</emphasis>
not seen. 
<emphasis id="0390361EFFF4F6635A9127385BDFAFFD" italics="true" pageId="7" pageNumber="174">Infructescences</emphasis>
terminal and sometimes also axillary at distalmost 1 to 4 nodes, cymose, rounded in outline, 3-5 × 
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(including peduncles and fruits), several-flowered (2 to 5), with fruits umbellate or usually with primary axis developed and fruits produced at 2 nodes, with axes hispid to pilose; bracts narrowly elliptic to narrowly triangular or stipuliform, 
<quantity id="F61C47E9FFF4F6635D0B21C85EF5A9CC" box="[1309,1410,406,430]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="7" pageNumber="174" unit="mm" value="9.0" valueMax="13.0" valueMin="5.0">5-13 mm</quantity>
. 
<emphasis id="0390361EFFF4F6635D9A21C85818A9AC" italics="true" pageId="7" pageNumber="174">Capsules</emphasis>
in umbelliform groups of 2 to 5 and all pedicellate on pedicels 
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, subglobose to oblate and laterally somewhat flattened, 8-12 × 
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, stiffly chartaceous to thinly woody, moderately to densely pilose to hispid, dehiscent from apex for 1/4-1/2 of body, beak portion flat to broadly conical, with persistent 
<emphasis id="0390361EFFF4F6635BCE22065F32AA12" box="[984,1093,600,624]" italics="true" pageId="7" pageNumber="174">calyx limb</emphasis>
divided nearly to base, hispid, lobes 5, narrowly triangular, 
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, subequal or unequal by up to 1/3 on an individual fruit, acute to acuminate. 
<emphasis id="0390361EFFF4F6635D0A22C75E2FAAD3" box="[1308,1368,665,689]" italics="true" pageId="7" pageNumber="174">Seeds</emphasis>
elliptic to suborbicular, 
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, flattened, with circumferential wing subentire to weakly dentate and with 1 deep incision.
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F6635B48235C5888ABDE" pageId="7" pageNumber="174" type="etymology">
<paragraph id="315BEA0CFFF4F6635B48235C5888ABDE" blockId="7.[823,1474,770,956]" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F6635B48235C58B7AB78" box="[862,960,770,794]" italics="true" pageId="7" pageNumber="174">Names. –</emphasis>
The scientific name created here for this species refers to the laciniate stipule form, which is unusual in the genus. The vernacular name “Vahidambo” was noted on 
<emphasis id="0390361EFFF4F6635D9B231D58A2AB19" italics="true" pageId="7" pageNumber="174">Antilahimena 2627</emphasis>
, and is probably in the northern Betsimisaraka dialect. This name translated into English means “pig liana” or “wild pig vine”.
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F6635B4B23925E08AC27" pageId="7" pageNumber="174" type="biology_ecology">
<paragraph id="315BEA0CFFF4F6635B4B23925E08AC27" blockId="7.[822,1473,972,1093]" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F6635B4B23925F1BAB86" box="[861,1132,972,996]" italics="true" pageId="7" pageNumber="174">Habitat and distribution</emphasis>
. 
<emphasis id="0390361EFFF4F6635C6D23925FFEAB86" box="[1147,1161,972,996]" italics="true" pageId="7" pageNumber="174">–</emphasis>
This species is documented from humid forests, at least sometimes in highly disturbed sites, at 
<quantity id="F61C47E9FFF4F6635B47245358B0AC47" box="[849,967,1037,1061]" metricMagnitude="2" metricUnit="m" metricValue="2.2399999999999998" metricValueMax="2.98" metricValueMin="1.5" pageId="7" pageNumber="174" unit="m" value="224.0" valueMax="298.0" valueMin="150.0">150-298 m</quantity>
in eastern Madagascar, in Toamasina Province (Analanjirofo Region, 
<figureCitation id="A9DFF689FFF4F6635C3E24735F10AC27" box="[1064,1127,1069,1093]" captionStart="Fig" captionStartId="10.[113,137,1227,1247]" captionTargetBox="[114,763,276,1209]" captionTargetId="figure-591@10.[114,763,276,1209]" captionTargetPageId="10" captionText="Fig. 5. – Distribution of Danais laciniata C. M. Taylor (asteriks) and Danais masoalana C. M. Taylor (circles), mapped over the outlines of the five simplified bioclimatic zones of Madagascar of SCHTZ (2000, following CORNET, 1974)." figureDoi="http://doi.org/10.5281/zenodo.6045577" httpUri="https://zenodo.org/record/6045577/files/figure.png" pageId="7" pageNumber="174">Fig. 5</figureCitation>
, humid bioclimatic zone).
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F6635B4A240B5FAAAD90" pageId="7" pageNumber="174" type="conservation">
<paragraph id="315BEA0CFFF4F6635B4A240B5FAAAD90" blockId="7.[822,1475,1109,1522]" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F6635B4A240B5F44AC0F" box="[860,1075,1109,1133]" italics="true" pageId="7" pageNumber="174">Conservation status</emphasis>
. 
<emphasis id="0390361EFFF4F6635C57240B5F39AC0F" box="[1089,1102,1109,1133]" italics="true" pageId="7" pageNumber="174">–</emphasis>
An EOO cannot be calculated for 
<taxonomicName id="F6E4918FFFF4F6635B2E242B589EACEF" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[824,1001,1141,1165]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFF4F6635B2E242B589EACEF" box="[824,1001,1141,1165]" italics="true" pageId="7" pageNumber="174">Danais laciniata</emphasis>
</taxonomicName>
because the species is known only from two localities. Its AOO is very restricted, certainly &lt;
<quantity id="F61C47E9FFF4F6635D3A24C85E1AACCC" box="[1324,1389,1174,1198]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="174" unit="km" value="10.0">10 km</quantity>
<superScript id="C6914744FFF4F6635D7B24CB5E02ACC1" attach="left" box="[1389,1397,1173,1187]" fontSize="6" pageId="7" pageNumber="174">2</superScript>
, which meets the criterion for Critically Endangered. The two localities where this species is known are ca. 
<quantity id="F61C47E9FFF4F6635CFE24895E69AC8D" box="[1256,1310,1239,1263]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="174" unit="km" value="4.0">4 km</quantity>
apart, and neither falls within Madagascar’s current system of protected areas. Vegetation at one of the localities was highly fragmented and disturbed in 2002 according to the label information, and the other site is most likely similarly degraded. The habitat quality is certainly expected to worsen in the future, and 
<taxonomicName id="F6E4918FFFF4F6635B2125C75890ADD3" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[823,999,1433,1457]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFF4F6635B2125C75890ADD3" box="[823,999,1433,1457]" italics="true" pageId="7" pageNumber="174">Danais laciniata</emphasis>
</taxonomicName>
is assigned a preliminary conservation status of Critically Endangered due to its limited distribution and threatened habitat [CR B2ab(i,ii,iii,iv)].
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFF4F66C5B4A265C5AE4AC17" lastPageId="8" lastPageNumber="175" pageId="7" pageNumber="174" type="discussion">
<paragraph id="315BEA0CFFF4F66C5B4A265C59B4A90F" blockId="7.[822,1474,1538,1951]" lastBlockId="8.[113,764,276,1236]" lastPageId="8" lastPageNumber="175" pageId="7" pageNumber="174">
<emphasis id="0390361EFFF4F6635B4A265C58F4AE58" italics="true" pageId="7" pageNumber="174">Notes on morphology, taxonomic identity and similar species</emphasis>
. 
<emphasis id="0390361EFFF4F6635B86267C5F3FAE58" box="[912,1096,1570,1594]" italics="true" pageId="7" pageNumber="174">–Danaislaciniata</emphasis>
can be recognized by its hispid or pilose pubescence, its leaves that are ternately arranged on at least some nodes and with the blades generally widest above the middle, its laciniate reflexed stipules, its rather lax umbelliform infructescences, its well developed persistent calyx lobes, and its relatively large fruits and seeds. Although the inflorescences and flowers of this species have not been seen, the fruit characters along with the climbing habit support its classification in 
<taxonomicName id="F6E4918FFFF4F6635BF5277B5F58AF5F" box="[995,1071,1829,1853]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="genus">
<emphasis id="0390361EFFF4F6635BF5277B5F58AF5F" box="[995,1071,1829,1853]" italics="true" pageId="7" pageNumber="174">Danais</emphasis>
</taxonomicName>
and the combination of the distinctive characters listed above distinguish 
<taxonomicName id="F6E4918FFFF4F6635CBD27185E5CAF3C" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[1195,1323,1862,1886]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFF4F6635CBD27185E5CAF3C" box="[1195,1323,1862,1886]" italics="true" pageId="7" pageNumber="174">D. laciniata</emphasis>
</taxonomicName>
from all other 
<taxonomicName id="F6E4918FFFF4F6635B2E273858ABAF1C" box="[824,988,1894,1918]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="0390361EFFF4F6635B2E273858F3AF1C" box="[824,900,1894,1918]" italics="true" pageId="7" pageNumber="174">Danais</emphasis>
species.
</taxonomicName>
The relatively long, perhaps flexuous pedicels also appear to be distinctive for this species, but in 
<taxonomicName id="F6E4918FFFF4F6635D5827D95EEDAFFD" box="[1358,1434,1927,1951]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="174" phylum="Tracheophyta" rank="genus">
<emphasis id="0390361EFFF4F6635D5827D95EEDAFFD" box="[1358,1434,1927,1951]" italics="true" pageId="7" pageNumber="174">Danais</emphasis>
</taxonomicName>
the fruiting pedicels are frequently much longer than flowering pedicels (e.g., 
<taxonomicName id="F6E4918FFFFBF66C5910216B5AD8A92F" authority="Baker" authorityName="Baker" box="[262,431,309,333]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="vestita">
<emphasis id="0390361EFFFBF66C5910216B5A1FA92F" box="[262,360,309,333]" italics="true" pageId="8" pageNumber="175">D. vestita</emphasis>
Baker
</taxonomicName>
), so details of the inflorescences cannot necessarily be inferred from the infructescences.
</paragraph>
<paragraph id="315BEA0CFFFBF66C588121275AE4AC17" blockId="8.[113,764,276,1236]" pageId="8" pageNumber="175">
These plants may be the same as the fruiting specimen 
<emphasis id="0390361EFFFBF66C5AC621275A5FA9D3" italics="true" pageId="8" pageNumber="175">J. S. Miller &amp; al. 3326</emphasis>
that was studied by 
<bibRefCitation id="557597FDFFFBF66C59ED21C75BE1A9B3" author="PUFF, C. &amp; R. BUCHNER" firstAuthor="PUFF" journalOrPublisher="Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia" page="44" pageId="8" pageNumber="175" pagination="11 - 64" part="16" refId="ref10969" refString="PUFF, C. &amp; R. BUCHNER (1994). Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores. Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia 16: 11 - 64." title="Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores" type="journal article" year="1994">PUFF &amp; BUCHNER (1994: 44)</bibRefCitation>
, but neither that collection nor duplicates have been located. 
<bibRefCitation id="557597FDFFFBF66C586721875A02A993" author="PUFF, C. &amp; R. BUCHNER" box="[113,373,473,497]" firstAuthor="PUFF" journalOrPublisher="Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia" pageId="8" pageNumber="175" pagination="11 - 64" part="16" refId="ref10969" refString="PUFF, C. &amp; R. BUCHNER (1994). Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores. Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia 16: 11 - 64." title="Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores" type="journal article" year="1994">PUFF &amp; BUCHNER (1994)</bibRefCitation>
included that specimen provisionally in their circumscription of 
<taxonomicName id="F6E4918FFFFBF66C599321A75929AA73" authority="Baker" authorityName="Baker" box="[389,606,505,529]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="vestita">
<emphasis id="0390361EFFFBF66C599321A7596EAA73" box="[389,537,505,529]" italics="true" pageId="8" pageNumber="175">Danais vestita</emphasis>
Baker
</taxonomicName>
but considered it anomalous due to its ternate leaves, sparser pubescence that dried whitish rather than rusty brown, fewer-flowered inflorescences, and more northern collection locality. Additional specimens of both 
<taxonomicName id="F6E4918FFFFBF66C58E922245A14AAF0" box="[255,355,634,658]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="vestita">
<emphasis id="0390361EFFFBF66C58E922245A14AAF0" box="[255,355,634,658]" italics="true" pageId="8" pageNumber="175">D. vestita</emphasis>
</taxonomicName>
and this new species are now available and clarify that the distinctive features cited by 
<bibRefCitation id="557597FDFFFBF66C5A7422C45B93AAB0" author="PUFF, C. &amp; R. BUCHNER" journalOrPublisher="Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia" pageId="8" pageNumber="175" pagination="11 - 64" part="16" refId="ref10969" refString="PUFF, C. &amp; R. BUCHNER (1994). Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores. Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia 16: 11 - 64." title="Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores" type="journal article" year="1994">PUFF &amp; BUCHNER (1994)</bibRefCitation>
for each of these are consistent. 
<taxonomicName id="F6E4918FFFFBF66C5A2F22E459B8AAB0" box="[569,719,698,722]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="vestita">
<emphasis id="0390361EFFFBF66C5A2F22E459B8AAB0" box="[569,719,698,722]" italics="true" pageId="8" pageNumber="175">Danais vestita</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="F6E4918FFFFBF66C58C122855A24AA91" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[215,339,731,755]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFFBF66C58C122855A24AA91" box="[215,339,731,755]" italics="true" pageId="8" pageNumber="175">D. laciniata</emphasis>
</taxonomicName>
in its pubescence, which is densely pilosulous or hirtellous with shorter trichomes; its leaf blades that are widest below the middle and truncate to cordulate at the base; its stipules that are erect or sometimes weakly spreading; and its inflorescences that are mainly axillary with the bracts linear and generally 
<quantity id="F61C47E9FFFBF66C58F623225A4EABF6" box="[224,313,892,916]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="8" pageNumber="175" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">1-5 mm</quantity>
long.Specimens of 
<taxonomicName id="F6E4918FFFFBF66C5A0B232259D5ABF6" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[541,674,892,916]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFFBF66C5A0B232259D5ABF6" box="[541,674,892,916]" italics="true" pageId="8" pageNumber="175">D. laciniata</emphasis>
</taxonomicName>
will be identified by 
<bibRefCitation id="557597FDFFFBF66C58EA23C25973ABD6" author="PUFF, C. &amp; R. BUCHNER" box="[252,516,924,948]" firstAuthor="PUFF" journalOrPublisher="Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia" pageId="8" pageNumber="175" pagination="11 - 64" part="16" refId="ref10969" refString="PUFF, C. &amp; R. BUCHNER (1994). Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores. Bull. Mus. Natl. Hist. Nat., sect. B, Adansonia 16: 11 - 64." title="Revision of Danais Vent. (Rubiaceae) in Madagascar and the Comores" type="journal article" year="1994">PUFF &amp; BUCHNER (1994)</bibRefCitation>
’s key as 
<taxonomicName id="F6E4918FFFFBF66C5A7723C25BA5ABB6" authority="Homolle" authorityName="Homolle" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="andribensis">
<emphasis id="0390361EFFFBF66C5A7723C2598CABD6" box="[609,763,924,948]" italics="true" pageId="8" pageNumber="175">D. andribensis</emphasis>
Homolle
</taxonomicName>
, of central and southern Madagascar; however 
<taxonomicName id="F6E4918FFFFBF66C586723825A71AB96" box="[113,262,988,1012]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="andribensis">
<emphasis id="0390361EFFFBF66C586723825A71AB96" box="[113,262,988,1012]" italics="true" pageId="8" pageNumber="175">D. andribensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="F6E4918FFFFBF66C599F23825976AB96" authorityName="C. M. Taylor. C. Fruiting" authorityYear="2013" box="[393,513,988,1012]" class="Magnoliopsida" family="Rubiaceae" genus="Danais" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="175" phylum="Tracheophyta" rank="species" species="laciniata">
<emphasis id="0390361EFFFBF66C599F23825976AB96" box="[393,513,988,1012]" italics="true" pageId="8" pageNumber="175">D. laciniata</emphasis>
</taxonomicName>
in its pubescence, which is pilosulous to puberulous or entirely lacking, its opposite leaves with the blades widest below the middle and rounded to cordulate at the base, its consistently bilobed stipules, and its smaller capsules (
<quantity id="F61C47E9FFFBF66C58C124035A5EAC17" box="[215,297,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="8" pageNumber="175" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4-8 mm</quantity>
in diam.).
</paragraph>
</subSubSection>
<subSubSection id="79FEB987FFFBF66C588124DA59CCACB6" pageId="8" pageNumber="175" type="materials_examined">
<paragraph id="315BEA0CFFFBF66C588124DA59CCACB6" blockId="8.[113,764,276,1236]" pageId="8" pageNumber="175">
<emphasis id="0390361EFFFBF66C588124DA5A7DACFB" box="[151,266,1156,1177]" italics="true" pageId="8" pageNumber="175">Paratypi. –</emphasis>
<materialsCitation id="818CE051FFFBF66C590524DA59C0ACB6" ID-GBIF-Occurrence="3463315308" collectingDate="2004-08-29" collectionCode="MO, P, TAN" collectorName="Antilahimena" country="Madagascar" elevation="298" location="Maroantsetra, Ambinanitelo, Marovovonana" municipality="Toamasina" pageId="8" pageNumber="175" specimenCount="1" stateProvince="Analanjirofo">
<emphasis id="0390361EFFFBF66C590524DA5919ACFB" bold="true" box="[275,622,1156,1177]" pageId="8" pageNumber="175">
<collectingCountry id="49F3AA9CFFFBF66C590524DA5AD6ACFA" box="[275,417,1156,1177]" pageId="8" pageNumber="175">MADAGASCAR</collectingCountry>
. Prov. 
<collectingMunicipality id="D13F7076FFFBF66C59E724DA591DACFB" box="[497,618,1156,1177]" pageId="8" pageNumber="175">Toamasina</collectingMunicipality>
:
</emphasis>
<collectingRegion id="F32024EEFFFBF66C5A6E24DA598FACFB" box="[632,760,1156,1177]" pageId="8" pageNumber="175">Analanjirofo</collectingRegion>
, 
<location id="343BBCD7FFFBF66C586724FF597BACD4" LSID="urn:lsid:plazi:treatment:B94D5B1AFFF4F66C587423C359CCACB6:343BBCD7FFFBF66C586724FF597BACD4" box="[113,524,1185,1206]" country="Madagascar" municipality="Toamasina" name="Maroantsetra, Ambinanitelo, Marovovonana" pageId="8" pageNumber="175" stateProvince="Analanjirofo">Maroantsetra, Ambinanitelo, Marovovonana</location>
, 
<geoCoordinate id="54D08CCBFFFBF66C5A0224FF598FACD4" box="[532,760,1185,1206]" pageId="8" pageNumber="175">15°19’34”S 49°31’15’E</geoCoordinate>
, 
<quantity id="F61C47E9FFFBF66C586424E15BC6ACB6" box="[114,177,1215,1236]" metricMagnitude="2" metricUnit="m" metricValue="2.98" pageId="8" pageNumber="175" unit="m" value="298.0">
<elevation id="BAC90D3FFFFBF66C586424E15BC6ACB6" box="[114,177,1215,1236]" metricMagnitude="2" metricUnit="m" metricValue="2.98" pageId="8" pageNumber="175" unit="m" value="298.0">298 m</elevation>
</quantity>
, 
<date id="455ACCCCFFFBF66C58AC24E15A4DACB6" box="[186,314,1215,1236]" pageId="8" pageNumber="175" value="2004-08-29">
<collectingDate id="551E3524FFFBF66C58AC24E15A4DACB6" box="[186,314,1215,1236]" pageId="8" pageNumber="175" value="2004-08-29">29.VIII.2004</collectingDate>
</date>
, fr., 
<emphasis id="0390361EFFFBF66C597524E1596CACB6" box="[355,539,1215,1236]" italics="true" pageId="8" pageNumber="175">
<collectorName id="9C118FDAFFFBF66C597524E15A93ACB6" box="[355,484,1215,1236]" pageId="8" pageNumber="175">Antilahimena</collectorName>
2627
</emphasis>
(
<collectionCode id="57F572C9FFFBF66C5A3E24E15920ACB6" box="[552,599,1215,1236]" pageId="8" pageNumber="175">MO</collectionCode>
!, 
<collectionCode id="57F572C9FFFBF66C5A7324E15904ACB6" box="[613,627,1215,1236]" pageId="8" pageNumber="175">P</collectionCode>
, 
<collectionCode id="57F572C9FFFBF66C5A6A24E159C7ACB6" box="[636,688,1215,1236]" pageId="8" pageNumber="175">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>