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<document id="2DA3752DFBC00DDDF823E03A9778D886" ID-CLB-Dataset="263641" ID-DOI="10.1093/zoolinnean/zly087" ID-GBIF-Dataset="648e1853-2069-4326-8728-bf45d07dc2f4" ID-ISSN="0024-4082" ID-Zenodo-Dep="8225921" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1691512074212" checkinUser="felipe" docAuthor="Ruschel, Tatiana Petersen &amp; Campos, Luiz Alexandre" docDate="2019" docId="B937B004A13EFE104B72FC72FA5E7B14" docLanguage="en" docName="ZoolJLinnSoc.185.4.1150-1187.pdf" docOrigin="Zoological Journal of the Linnean Society 185 (4)" docSource="https://academic.oup.com/zoolinnean/article/185/4/1150/5299641" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Hemidictya frondosa Burmeister 1835" docType="treatment" docVersion="4" lastPageNumber="1182" masterDocId="450EC87CA121FE30481FFF9AFFD87F0F" masterDocTitle="Phylogeny and biogeography of the leaf-winged cicadas (Hemiptera: Auchenorrhyncha: Cicadidae)" masterLastPageNumber="1187" masterPageNumber="1150" pageNumber="1181" updateTime="1691597201196" updateUser="ExternalLinkService">
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<mods:title id="F65A309741E8EC4DC74CA22AD14AC3EA">Phylogeny and biogeography of the leaf-winged cicadas (Hemiptera: Auchenorrhyncha: Cicadidae)</mods:title>
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<mods:namePart id="4B52E911C206FF6B6C07B42118DACA51">Ruschel, Tatiana Petersen</mods:namePart>
<mods:affiliation id="D5C0DA97E402DCC11F0236B87D340BD9">Programa de Pós – Graduação em Biologia Animal, Departamento de Zoologia, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves 9500, Prédio 43435, Sala 201, 91501 – 970, Porto Alegre, RS, Brazil</mods:affiliation>
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<mods:namePart id="6A34000A197BB4479E9E8951246343DE">Campos, Luiz Alexandre</mods:namePart>
<mods:affiliation id="1BAD3147E1906487E2AC3A9E6EF637D6">Programa de Pós – Graduação em Biologia Animal, Departamento de Zoologia, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves 9500, Prédio 43435, Sala 201, 91501 – 970, Porto Alegre, RS, Brazil</mods:affiliation>
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<mods:number id="AD083B619855E59CFACA2988225A1891">185</mods:number>
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<treatment id="B937B004A13EFE104B72FC72FA5E7B14" ID-DOI="http://doi.org/10.5281/zenodo.8225942" ID-Zenodo-Dep="8225942" LSID="urn:lsid:plazi:treatment:B937B004A13EFE104B72FC72FA5E7B14" httpUri="http://treatment.plazi.org/id/B937B004A13EFE104B72FC72FA5E7B14" lastPageId="32" lastPageNumber="1182" pageId="31" pageNumber="1181">
<subSubSection id="79845299A13EFE2F4B72FC72FA947B0F" box="[877,1356,999,1024]" pageId="31" pageNumber="1181" type="nomenclature">
<paragraph id="31210112A13EFE2F4B72FC72FA947B0F" blockId="31.[877,1356,999,1024]" box="[877,1356,999,1024]" pageId="31" pageNumber="1181">
<heading id="6A69B67EA13EFE2F4B72FC72FA947B0F" box="[877,1356,999,1024]" centered="true" fontSize="9" level="2" pageId="31" pageNumber="1181" reason="2">
<taxonomicName id="F69E7A91A13EFE2F4B72FC72FA947B0F" ID-CoL="3KNJY" ID-ENA="2606459" authority="Burmeister, 1835" authorityName="Burmeister" authorityYear="1835" box="[877,1356,999,1024]" class="Insecta" family="Cicadidae" genus="Hemidictya" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1181" phylum="Arthropoda" rank="species" species="frondosa">
<emphasis id="03EADD00A13EFE2F4B72FC72FBA27CF0" box="[877,1146,1000,1023]" italics="true" pageId="31" pageNumber="1181">
<smallCapsWord id="37C797CEA13EFE2F4B72FC72FC247CF0" baselines="1017,1018" box="[877,1020,1000,1023]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Hemidictya" pageId="31" pageNumber="1181">HEMIDICTYA</smallCapsWord>
<smallCapsWord id="37C797CEA13EFE2F4C1DFC71FBA27CF0" baselines="1018" box="[1026,1146,1003,1023]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="frondosa" pageId="31" pageNumber="1181">FRONDOSA</smallCapsWord>
</emphasis>
<bibRefCitation id="550F7CE3A13EFE2F4C9DFC7DFA947B0F" author="Burmeister" box="[1154,1356,999,1024]" firstAuthor="Burmeister" pageId="31" pageNumber="1181" pagination="170 - 183" refId="ref22548" refString="Burmeister HCC. 1835. Bierte (8.) Familie. Gingzirpen Stridulantia Handbuch der Entomologie. In: Burmeister HCC, ed. Handbuch der entomologie. Berlin: Jena, 170 - 183." type="book chapter" year="1835">Burmeister, 1835</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="79845299A13EFE2F4C0CFBB3FB7E7B4D" box="[1043,1190,1065,1090]" pageId="31" pageNumber="1181" type="description">
<paragraph id="31210112A13EFE2F4C0CFBB3FB7E7B4D" blockId="31.[1043,1190,1065,1090]" box="[1043,1190,1065,1090]" pageId="31" pageNumber="1181">
(
<figureCitation id="A9A51D97A13EFE2F4C04FBB3FB467B4D" box="[1051,1182,1065,1090]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStart-2="Figure 15" captionStartId-0="20.[164,242,1309,1331]" captionStartId-1="21.[147,225,1412,1434]" captionStartId-2="22.[163,241,1848,1870]" captionTargetBox-0="[323,1281,202,1269]" captionTargetBox-1="[192,1377,196,1371]" captionTargetBox-2="[227,1378,940,1807]" captionTargetId-0="figure-283@20.[321,1284,195,1270]" captionTargetId-1="figure-222@21.[191,1378,195,1372]" captionTargetId-2="figure-411@22.[225,1380,938,1809]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="Figure 13. Lectotype (A) and paralectotype (B) male of Hemidictya frondosa (ZMHB) in dorsal and lateral view, respectively. Scale: 1 mm." captionText-1="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." captionText-2="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8225956" figureDoi-1="http://doi.org/10.5281/zenodo.8225962" figureDoi-2="http://doi.org/10.5281/zenodo.8225964" httpUri-0="https://zenodo.org/record/8225956/files/figure.png" httpUri-1="https://zenodo.org/record/8225962/files/figure.png" httpUri-2="https://zenodo.org/record/8225964/files/figure.png" pageId="31" pageNumber="1181">
<smallCapsWord id="37C797CEA13EFE2F4C04FBB3FB897B4F" baselines="1084,1083" box="[1051,1105,1065,1090]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="31" pageNumber="1181">FIGS</smallCapsWord>
13–15
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="79845299A13EFE2F4B36FBCBFACE7B69" box="[809,1302,1105,1127]" pageId="31" pageNumber="1181" type="reference_group">
<paragraph id="31210112A13EFE2F4B36FBCBFACE7B69" blockId="31.[809,1424,1105,1189]" box="[809,1302,1105,1127]" pageId="31" pageNumber="1181">
<treatmentCitationGroup id="118E263CA13EFE2F4B36FBCBFACE7B69" box="[809,1302,1105,1127]" pageId="31" pageNumber="1181">
<treatmentCitation id="B03F2703A13EFE2F4B36FBCBFACE7B69" author="Burmeister HCC" box="[809,1302,1105,1127]" page="178" pageId="31" pageNumber="1181" year="1835">
<taxonomicName id="F69E7A91A13EFE2F4B36FBCBFACE7B69" ID-CoL="3KNJY" authority="Burmeister, 1835: 178" authorityName="Burmeister" authorityPageNumber="178" authorityYear="1835" box="[809,1302,1105,1127]" class="Insecta" family="Cicadidae" genus="Hemidictya" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1181" phylum="Arthropoda" rank="species" species="frondosa">
<emphasis id="03EADD00A13EFE2F4B36FBCBFBCE7B69" box="[809,1046,1105,1126]" italics="true" pageId="31" pageNumber="1181">Hemidictya frondosa</emphasis>
<bibRefCitation id="550F7CE3A13EFE2F4C02FBCBFACE7B69" author="Burmeister" box="[1053,1302,1105,1127]" firstAuthor="Burmeister" page="178" pageId="31" pageNumber="1181" pagination="170 - 183" refId="ref22548" refString="Burmeister HCC. 1835. Bierte (8.) Familie. Gingzirpen Stridulantia Handbuch der Entomologie. In: Burmeister HCC, ed. Handbuch der entomologie. Berlin: Jena, 170 - 183." type="book chapter" year="1835">Burmeister, 1835: 178</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="79845299A13EFE2F4B36FBEBFB337A0E" pageId="31" pageNumber="1181" type="materials_examined">
<paragraph id="31210112A13EFE2F4B36FBEBFAF97BAB" blockId="31.[809,1424,1105,1189]" pageId="31" pageNumber="1181">
<materialsCitation id="81F60B4FA13EFE2F4B36FBEBFAF97BAB" collectionCode="ZMHB" country="Brazil" location="Rio de Janeiro" pageId="31" pageNumber="1181" specimenCode="5671" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
<emphasis id="03EADD00A13EFE2F4B36FBEBFB547B8A" box="[809,1164,1136,1158]" italics="true" pageId="31" pageNumber="1181">
<typeStatus id="EE25BFB0A13EFE2F4B36FBEBFC4A7B89" box="[809,914,1137,1158]" pageId="31" pageNumber="1181">Lectotype</typeStatus>
male here designated:
</emphasis>
<collectingRegion id="F35ACFF0A13EFE2F4C87FBEAFAE37B89" box="[1176,1339,1136,1158]" country="Brazil" name="Rio de Janeiro" pageId="31" pageNumber="1181">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="49894182A13EFE2F4D5AFBEAFA557B89" box="[1349,1421,1136,1158]" name="Brazil" pageId="31" pageNumber="1181">Brazil</collectingCountry>
, male (
<collectionCode id="578F99D7A13EFE2F4B70FB15FC187BAA" box="[879,960,1167,1189]" pageId="31" pageNumber="1181">ZMHB</collectionCode>
). Type Cat. no. 5671 (
<figureCitation id="A9A51D97A13EFE2F4CACFB14FACA7BAB" box="[1203,1298,1166,1188]" captionStart="Figure 13" captionStartId="20.[164,242,1309,1331]" captionTargetBox="[323,1281,202,1269]" captionTargetId="figure-283@20.[321,1284,195,1270]" captionTargetPageId="20" captionText="Figure 13. Lectotype (A) and paralectotype (B) male of Hemidictya frondosa (ZMHB) in dorsal and lateral view, respectively. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225956" httpUri="https://zenodo.org/record/8225956/files/figure.png" pageId="31" pageNumber="1181">Fig. 13A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="31210112A13EFE2F4B36FB57FB337A0E" blockId="31.[809,1424,1229,1282]" pageId="31" pageNumber="1181">
<materialsCitation id="81F60B4FA13EFE2F4B36FB57FB047A0E" collectionCode="ZMHB" country="Brazil" location="Rio de Janeiro" pageId="31" pageNumber="1181" specimenCode="5671" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="paralectotype">
<emphasis id="03EADD00A13EFE2F4B36FB57FB397BED" box="[809,1249,1229,1250]" italics="true" pageId="31" pageNumber="1181">
<typeStatus id="EE25BFB0A13EFE2F4B36FB57FC147BED" box="[809,972,1229,1250]" pageId="31" pageNumber="1181">Paralectotype</typeStatus>
male here designated:
</emphasis>
same data as lectotype. Type Cat. no. 5671 (
<figureCitation id="A9A51D97A13EFE2F4C62FB76FB047A0E" box="[1149,1244,1260,1282]" captionStart="Figure 13" captionStartId="20.[164,242,1309,1331]" captionTargetBox="[323,1281,202,1269]" captionTargetId="figure-283@20.[321,1284,195,1270]" captionTargetPageId="20" captionText="Figure 13. Lectotype (A) and paralectotype (B) male of Hemidictya frondosa (ZMHB) in dorsal and lateral view, respectively. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225956" httpUri="https://zenodo.org/record/8225956/files/figure.png" pageId="31" pageNumber="1181">Fig. 13B</figureCitation>
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="79845299A13EFE2F4B36FAB1FA897AB4" pageId="31" pageNumber="1181" type="diagnosis">
<paragraph id="31210112A13EFE2F4B36FAB1FA897AB4" blockId="31.[809,1425,1322,1467]" pageId="31" pageNumber="1181">
<emphasis id="03EADD00A13EFE2F4B36FAB1FC727A4F" box="[809,938,1323,1344]" italics="true" pageId="31" pageNumber="1181">Diagnosis:</emphasis>
Apical area reticulated separate from proximal area by a median line; ulnar cell (u3) subdivided; clavus (clv) pigmented; basal cell subrectangular; apex of claspers posteriorly directed; aedeagus curved with two apical parallel spines.
</paragraph>
</subSubSection>
<subSubSection id="79845299A13EFE104B36FA7EFB487EE0" lastPageId="32" lastPageNumber="1182" pageId="31" pageNumber="1181" type="description">
<paragraph id="31210112A13EFE2F4B36FA7EFC7D7917" blockId="31.[809,1425,1508,1560]" pageId="31" pageNumber="1181">
<emphasis id="03EADD00A13EFE2F4B36FA7EFC747AF6" box="[809,940,1508,1529]" italics="true" pageId="31" pageNumber="1181">Coloration:</emphasis>
Green in live specimens, yellow in dried specimens.
</paragraph>
<paragraph id="31210112A13EFE104B36F9D8FCDA7E5A" blockId="31.[809,1425,1601,1899]" lastBlockId="32.[163,779,197,1630]" lastPageId="32" lastPageNumber="1182" pageId="31" pageNumber="1181">
<emphasis id="03EADD00A13EFE2F4B36F9D8FC607958" box="[809,952,1602,1623]" italics="true" pageId="31" pageNumber="1181">Description:</emphasis>
Male. Head (
<figureCitation id="A9A51D97A13EFE2F4C40F9DBFB1C7959" box="[1119,1220,1601,1623]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="31" pageNumber="1181">Fig. 14A</figureCitation>
) narrow, smaller than mesonotum. Lateral margin of eyes reaching the lateral margin of pronotum, not salient relative to the lateral margins of head. Posterior margin of eyes not reaching the anterior margin of pronotum. Supra-antennal plates not prominent. Region of ocelli flat, without tubercles, the lateral ocelli not higher than the median. Lateral ocelli widely spaced, the distance between them about the same distance between each lateral ocellus and eye. Postclypeus long, anterior outline V-shaped, rectangular in ventral view and slightly salient in lateral view. Flagellum with three segments, all almost the same length, distal article black at apex in
<specimenCount id="2798CA9BA101FE104943FEB8FE2F7E38" box="[348,503,289,311]" pageId="32" pageNumber="1182" type="generic">one specimen</specimenCount>
. Anteclypeus and carina bear tuft of setae. Labium short, reaching mesocoxae.
</paragraph>
<paragraph id="31210112A101FE1048A4FEC5FF2E7CF7" blockId="32.[163,779,197,1630]" pageId="32" pageNumber="1182">
Pronotum (
<figureCitation id="A9A51D97A101FE104924FEC4FE407E7B" box="[315,408,350,372]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14A</figureCitation>
) with a narrow and deep median grove; lateral lobes larger than paramedian lobes. Paranota developed. Pronotal collar wide, lateral angles truncate, reaching the articulation of forewings. Mesonotum (
<figureCitation id="A9A51D97A101FE10494EFE43FE197EE1" box="[337,449,473,495]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14A</figureCitation>
) with a long triangular scutellum reaching tergite 2 and bearing a central crest. Parapsidal suture poorly marked, lateral and submedian sigilla brown. Operculum (
<figureCitation id="A9A51D97A101FE104A78FDAFFD117D45" box="[615,713,565,587]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14B</figureCitation>
) thin and short, not covering the tympanal cavity and not reaching sternite II. Meracanthus (
<figureCitation id="A9A51D97A101FE104A30FDE8FD557D87" box="[559,653,626,648]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14B</figureCitation>
) exceeding the size of operculum, apex directed posteriorly. Gutter across the margin of operculum. Profemora with three small spines, the primary leaning forward against the ventral margin of femur. Tarsi with three articles. Forewings (
<figureCitation id="A9A51D97A101FE104936FC96FE547C2E" box="[297,396,780,802]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14D</figureCitation>
) semi-opaque and wide, covering the whole abdomen and pleurae in lateral view, apex acute. Veins C and R+Sc apart from each other. Basal cell subretangular, longer than wide. Ulnar cell (u3) subdivided and angled to medial cell (mc). Apical area reticulated, separated from the proximal area by a median line. Clavus (clv) pigmented. Posterior margin absent.
</paragraph>
<paragraph id="31210112A101FE1048A4FB9BFEC17B9E" blockId="32.[163,779,197,1630]" pageId="32" pageNumber="1182">
Abdomen fusiform, apex triangular. Sternite I not completely covered by metacoxae, touching sternite II, whose median portion of anterior margin is nearly straight. Sternite VII subrectangular, obtuse at apex (
<figureCitation id="A9A51D97A101FE1048B4FBE6FED27B9E" box="[171,266,1148,1170]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14C</figureCitation>
).
</paragraph>
<paragraph id="31210112A101FE1048A4FB00FD407A45" blockId="32.[163,779,197,1630]" pageId="32" pageNumber="1182">
Pygofer (
<figureCitation id="A9A51D97A101FE10493AFB00FE5F7BA0" box="[293,391,1178,1200]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14E</figureCitation>
) with distal shoulders developed and rounded. Claspers posteriorly developed in U-shape (
<figureCitation id="A9A51D97A101FE104907FB42FEA67BE2" box="[280,382,1240,1262]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14G</figureCitation>
), process close not enfolding the aedeagus, apex obtuse. Upper lobe undeveloped. Aedeagus (
<figureCitation id="A9A51D97A101FE104939FA8FFE717A25" box="[294,425,1301,1323]" captionStart="Figure 14" captionStartId="21.[147,225,1412,1434]" captionTargetBox="[192,1377,196,1371]" captionTargetId="figure-222@21.[191,1378,195,1372]" captionTargetPageId="21" captionText="Figure 14. Hemidictya frondosa (male). A, head, pronotum and mesonotum in dorsal view; B, operculum and meracanthus in ventral view; C, sternite VII in ventral view; D, forewing in dorsal view; E, pygofer in lateral view; F, aedeagus in frontal view; G, claspers in ventral view. Scale: A, D = 2 mm, B, C, E = 1 mm, F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8225962" httpUri="https://zenodo.org/record/8225962/files/figure.png" pageId="32" pageNumber="1182">Fig. 14E, F</figureCitation>
) C-shaped, angle of proximal curvature obtuse, bearing two apical spines.
</paragraph>
<paragraph id="31210112A101FE1048A4FAC8FE477952" blockId="32.[163,779,197,1630]" pageId="32" pageNumber="1182">
Female (
<figureCitation id="A9A51D97A101FE104939FAC8FE767A67" box="[294,430,1362,1384]" captionStart="Figure 15" captionStartId="22.[163,241,1848,1870]" captionTargetBox="[227,1378,940,1807]" captionTargetId="figure-411@22.[225,1380,938,1809]" captionTargetPageId="22" captionText="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225964" httpUri="https://zenodo.org/record/8225964/files/figure.png" pageId="32" pageNumber="1182">Fig. 15A–D</figureCitation>
). Somatic characteristics as described for male, except the operculum that almost covers the tympanal cavity, meracanthus reaching sternite II (
<figureCitation id="A9A51D97A101FE10493DFA34FE587ACB" box="[290,384,1454,1476]" captionStart="Figure 15" captionStartId="22.[163,241,1848,1870]" captionTargetBox="[227,1378,940,1807]" captionTargetId="figure-411@22.[225,1380,938,1809]" captionTargetPageId="22" captionText="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225964" httpUri="https://zenodo.org/record/8225964/files/figure.png" pageId="32" pageNumber="1182">Fig. 15A</figureCitation>
), the latter not projected anteriorly at midline; posterior margin of sternite VII concave (
<figureCitation id="A9A51D97A101FE1048B4FA76FED5790E" box="[171,269,1515,1538]" captionStart="Figure 15" captionStartId="22.[163,241,1848,1870]" captionTargetBox="[227,1378,940,1807]" captionTargetId="figure-411@22.[225,1380,938,1809]" captionTargetPageId="22" captionText="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225964" httpUri="https://zenodo.org/record/8225964/files/figure.png" pageId="32" pageNumber="1182">Fig. 15B</figureCitation>
). Tergite 9 with the mesial margin concave becoming convex in the middle (
<figureCitation id="A9A51D97A101FE104A00F990FD5A792F" box="[543,642,1546,1568]" captionStart="Figure 15" captionStartId="22.[163,241,1848,1870]" captionTargetBox="[227,1378,940,1807]" captionTargetId="figure-411@22.[225,1380,938,1809]" captionTargetPageId="22" captionText="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225964" httpUri="https://zenodo.org/record/8225964/files/figure.png" pageId="32" pageNumber="1182">Fig. 15B</figureCitation>
). The tip of ovipositor sheath with same length of dorsal beak of tergite 9 (
<figureCitation id="A9A51D97A101FE10490DF9D2FE487952" box="[274,400,1607,1630]" captionStart="Figure 15" captionStartId="22.[163,241,1848,1870]" captionTargetBox="[227,1378,940,1807]" captionTargetId="figure-411@22.[225,1380,938,1809]" captionTargetPageId="22" captionText="Figure 15. Hemidictya frondosa (female) (DZUP). A, operculum in ventral view; B, terminalia in ventral view, lateral view (C) and postero-lateral view (D). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8225964" httpUri="https://zenodo.org/record/8225964/files/figure.png" pageId="32" pageNumber="1182">Fig. 15C, D</figureCitation>
).
</paragraph>
<paragraph id="31210112A101FE1048BCF91DFB487EE0" blockId="32.[163,779,1670,1876]" lastBlockId="32.[827,1443,197,495]" pageId="32" pageNumber="1182">
<emphasis id="03EADD00A101FE1048BCF91DFE7F7993" box="[163,423,1671,1692]" italics="true" pageId="32" pageNumber="1182">Measurements (mm):</emphasis>
male,
<emphasis id="03EADD00A101FE104A1CF91DFDCF7993" box="[515,535,1671,1692]" italics="true" pageId="32" pageNumber="1182">N</emphasis>
= 3, mean (range). Length of body: 12.79 (12.48–13.10); width of head including eyes: 3.66 (3.55–3.90); length of head: 1.60 (1.42–1.70); width of pronotum including pronotal collar: 5.20 (4.97–5.67); length of pronotum including pronotal collar: 1.69 (1.63–1.76); width of mesonotum: 4.29 (3.97–4.66); length of mesonotum: 4.34 (4.18–4.53); width of forewing: 9.41 (9.20–9.67); length of forewing: 18.72 (17.94–20.28). Female,
<emphasis id="03EADD00A101FE104D09FF7EFAF27FF6" box="[1302,1322,228,249]" italics="true" pageId="32" pageNumber="1182">N</emphasis>
= 2, mean (range). Length of body: 16.53 (16.38–16.69); width of head including eyes: 4.67 (4.30–5.04); length of head: 2.02 (1.90–2.14); width of pronotum including pronotal collar: 6.85 (6.50–7.20); length of pronotum including pronotal collar: 2.15 (2.00–2.30); width of mesonotum: 5.50 (5.20–5.80); length of mesonotum: 5.75 (5.50– 6.00); width of forewing: 12.24 (11.70–12.79); length of forewing: 25.42 (23.40–27.45).
</paragraph>
</subSubSection>
<subSubSection id="79845299A101FE104B24FD8DFB7D7C4D" pageId="32" pageNumber="1182" type="materials_examined">
<paragraph id="31210112A101FE104B24FD8DFB7D7C4D" blockId="32.[827,1444,535,834]" pageId="32" pageNumber="1182">
<emphasis id="03EADD00A101FE104B24FD8DFBFA7D23" box="[827,1058,535,556]" italics="true" pageId="32" pageNumber="1182">Material examined:</emphasis>
<materialsCitation id="81F60B4FA101FE104C31FD82FC217D85" collectingDate="1983-12-03" collectingDateMax="1983-12-05" collectingDateMin="1983-12-03" collectingMethod="Malaise" collectionCode="DZUP" collectorName="Exc. Dep. Zool. Polonoroeste" location="Chapada dos Guimaraes" municipality="Chapada dos Guimaraes" pageId="32" pageNumber="1182" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso">
<collectingRegion id="F35ACFF0A101FE104C31FD82FB1F7D22" box="[1070,1223,536,557]" country="Brazil" name="Mato Grosso" pageId="32" pageNumber="1182">Mato Grosso</collectingRegion>
:
<collectingMunicipality id="D1459B68A101FE104CCCFD8DFA7A7D22" box="[1235,1442,535,557]" pageId="32" pageNumber="1182">Chap. Guimarães</collectingMunicipality>
– MT \
<date id="452027D2A101FE104BBFFDACFB8F7D43" box="[928,1111,566,588]" pageId="32" pageNumber="1182" value="1983-12-03" valueMax="1983-12-05" valueMin="1983-12-03">
<collectingDate id="5564DE3AA101FE104BBFFDACFB8F7D43" box="[928,1111,566,588]" pageId="32" pageNumber="1182" value="1983-12-03" valueMax="1983-12-05" valueMin="1983-12-03">03-05.XII.1983</collectingDate>
</date>
\
<collectorName id="9C6B64C4A101FE104C63FDADFAED7D43" box="[1148,1333,566,588]" pageId="32" pageNumber="1182">Exc. Dep. Zool.</collectorName>
UFPR \ (
<collectorName id="9C6B64C4A101FE104B5DFDCFFC027D64" box="[834,986,597,619]" pageId="32" pageNumber="1182">Polonoroeste</collectorName>
) \
<collectingMethod id="E8DF7905A101FE104C1DFDCFFB877D64" box="[1026,1119,597,619]" pageId="32" pageNumber="1182">Malaise</collectingMethod>
\\ Dpto. Zoll.\UF-Paraná,
<specimenCount id="2798CA9BA101FE104B24FDEEFC547D86" box="[827,908,627,649]" pageId="32" pageNumber="1182" type="male">1 male</specimenCount>
(
<collectionCode id="578F99D7A101FE104B80FDEEFC367D85" box="[927,1006,628,650]" collectionName="Brazil, Parana, Curitiba, Universidade Federal do Parana, Museu de Entomologia Pe. Jesus Santiago Moure" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="32" pageNumber="1182" type="University or college">DZUP</collectionCode>
);
</materialsCitation>
<materialsCitation id="81F60B4FA101FE104C1BFDEEFB767DA7" collectingDate="1963-11" collectionCode="DZUP" collectorName="M. Alvarenga" country="Brazil" location="Chapada dos Guimaraes" municipality="Chapada dos Guimaraes" pageId="32" pageNumber="1182" specimenCount="2" specimenCount-male="2" stateProvince="Mato Grosso">
<collectingMunicipality id="D1459B68A101FE104C1BFDEEFB5F7D86" box="[1028,1159,628,649]" pageId="32" pageNumber="1182">CHAPADA</collectingMunicipality>
–
<collectingRegion id="F35ACFF0A101FE104CB6FDEEFB0A7D86" box="[1193,1234,628,649]" pageId="32" pageNumber="1182">MT</collectingRegion>
\
<collectingCountry id="49894182A101FE104CEAFDE9FAE77D86" box="[1269,1343,627,649]" name="Brazil" pageId="32" pageNumber="1182">Brasil</collectingCountry>
<date id="452027D2A101FE104D56FDE9FA527D86" box="[1353,1418,627,649]" pageId="32" pageNumber="1182" value="1963-11">
<collectingDate id="5564DE3AA101FE104D56FDE9FA527D86" box="[1353,1418,627,649]" pageId="32" pageNumber="1182" value="1963-11">XI.63</collectingDate>
</date>
\
<collectorName id="9C6B64C4A101FE104B24FD09FC047DA7" box="[827,988,658,680]" pageId="32" pageNumber="1182">M. Alvarenga</collectorName>
,
<specimenCount id="2798CA9BA101FE104BF6FD08FB9D7DA7" box="[1001,1093,658,680]" pageId="32" pageNumber="1182" type="male">2 males</specimenCount>
(
<collectionCode id="578F99D7A101FE104C4AFD08FB7A7DA7" box="[1109,1186,658,680]" collectionName="Brazil, Parana, Curitiba, Universidade Federal do Parana, Museu de Entomologia Pe. Jesus Santiago Moure" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="32" pageNumber="1182" type="University or college">DZUP</collectionCode>
);
</materialsCitation>
<materialsCitation id="81F60B4FA101FE104CA8FD08FB817C0B" collectingDate="1980-11-14" collectingDateMax="1980-11-16" collectingDateMin="1980-11-14" collectionCode="UFMG" collectorName="M. A. Vulcano" country="Brazil" location="Parque Estadual do Rio Doce" municipality="Marlieria" pageId="32" pageNumber="1182" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais">
<collectingRegion id="F35ACFF0A101FE104CA8FD08FA827DA7" box="[1207,1370,658,680]" country="Brazil" name="Minas Gerais" pageId="32" pageNumber="1182">Minas Gerais</collectingRegion>
:
<collectingCountry id="49894182A101FE104D79FD08FA467DA8" box="[1382,1438,658,679]" pageId="32" pageNumber="1182">BRA</collectingCountry>
, MG,
<collectingMunicipality id="D1459B68A101FE104B6BFD2BFC3C7DC8" box="[884,996,689,711]" pageId="32" pageNumber="1182">Marliéria</collectingMunicipality>
\ Timóteo,
<location id="344157C9A101FE104C6EFD2BFCAD7DEA" LSID="urn:lsid:plazi:treatment:B937B004A13EFE104B72FC72FA5E7B14:344157C9A101FE104C6EFD2BFCAD7DEA" country="BRA" municipality="Marlieria" name="Parque Estadual \ do Rio Doce" pageId="32" pageNumber="1182" stateProvince="Minas Gerais">Parque Estadual \ do Rio Doce</location>
,
<date id="452027D2A101FE104B9EFD4AFBDF7DEA" box="[897,1031,720,741]" pageId="32" pageNumber="1182" value="1980-11-14">
<collectingDate id="5564DE3AA101FE104B9EFD4AFBDF7DEA" box="[897,1031,720,741]" pageId="32" pageNumber="1182" value="1980-11-14">14.XI.1980</collectingDate>
</date>
-\
<date id="452027D2A101FE104C3EFD55FB7A7DEA" box="[1057,1186,719,741]" pageId="32" pageNumber="1182" value="1980-11-16">
<collectingDate id="5564DE3AA101FE104C3EFD55FB7A7DEA" box="[1057,1186,719,741]" pageId="32" pageNumber="1182" value="1980-11-16">16.XI.1980</collectingDate>
</date>
,
<collectorName id="9C6B64C4A101FE104CB0FD4AFA897DEA" box="[1199,1361,719,741]" pageId="32" pageNumber="1182">M. A. Vulcano</collectorName>
\
<emphasis id="03EADD00A101FE104D76FD4AFA447DEB" box="[1385,1436,719,741]" italics="true" pageId="32" pageNumber="1182">et al</emphasis>
. leg.,
<specimenCount id="2798CA9BA101FE104B66FD74FC3D7C0B" box="[889,997,750,772]" pageId="32" pageNumber="1182" type="female">1 female</specimenCount>
(
<collectionCode id="578F99D7A101FE104BE7FD74FB897C0B" box="[1016,1105,750,772]" collectionName="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="32" pageNumber="1182">UFMG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="81F60B4FA101FE104C76FD74FB7D7C4D" collectingDate="1965-12" collectionCode="DZUP" collectorName="A. Maller" country="Brazil" location="Linhares" municipality="Linhares" pageId="32" pageNumber="1182" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<collectingRegion id="F35ACFF0A101FE104C76FD74FAFC7C0C" box="[1129,1316,750,772]" country="Brazil" name="Espirito Santo" pageId="32" pageNumber="1182">Espírito Santo</collectingRegion>
:
<collectingMunicipality id="D1459B68A101FE104D2CFD74FA797C0B" box="[1331,1441,750,772]" pageId="32" pageNumber="1182">Linhares</collectingMunicipality>
\ ES-
<collectingCountry id="49894182A101FE104B99FC97FC167C2C" box="[902,974,781,803]" name="Brazil" pageId="32" pageNumber="1182">Brasil</collectingCountry>
\ XII-196 5\
<collectorName id="9C6B64C4A101FE104C67FC97FB307C2C" box="[1144,1256,781,803]" pageId="32" pageNumber="1182">A. Maller</collectorName>
\\ Dpto Zoo l\ UF-PARANÁ,
<specimenCount id="2798CA9BA101FE104BC0FCB6FBE77C4E" box="[991,1087,811,833]" pageId="32" pageNumber="1182" type="female">1 female</specimenCount>
(
<collectionCode id="578F99D7A101FE104C51FCB6FB427C4D" box="[1102,1178,812,834]" collectionName="Brazil, Parana, Curitiba, Universidade Federal do Parana, Museu de Entomologia Pe. Jesus Santiago Moure" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="32" pageNumber="1182" type="University or college">DZUP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="79845299A101FE104B24FCF0FB747C90" pageId="32" pageNumber="1182" type="distribution">
<paragraph id="31210112A101FE104B24FCF0FB747C90" blockId="32.[827,1442,874,927]" pageId="32" pageNumber="1182">
<emphasis id="03EADD00A101FE104B24FCF0FC0C7C70" box="[827,980,874,895]" italics="true" pageId="32" pageNumber="1182">Distribution:</emphasis>
<collectingCountry id="49894182A101FE104BFEFCF0FBF27C8F" box="[993,1066,874,896]" name="Brazil" pageId="32" pageNumber="1182">Brazil</collectingCountry>
(
<collectingRegion id="F35ACFF0A101FE104C25FCF0FB0D7C70" box="[1082,1237,874,895]" country="Brazil" name="Mato Grosso" pageId="32" pageNumber="1182">Mato Grosso</collectingRegion>
*,
<collectingRegion id="F35ACFF0A101FE104CF1FCF0FA4A7C8F" box="[1262,1426,874,896]" country="Brazil" name="Minas Gerais" pageId="32" pageNumber="1182">Minas Gerais</collectingRegion>
*,
<collectingRegion id="F35ACFF0A101FE104B24FC13FC3A7C91" box="[827,994,905,927]" country="Brazil" name="Espirito Santo" pageId="32" pageNumber="1182">Espírito Santo</collectingRegion>
*,
<collectingRegion id="F35ACFF0A101FE104BE6FC13FB787C90" box="[1017,1184,905,927]" country="Brazil" name="Rio de Janeiro" pageId="32" pageNumber="1182">Rio de Janeiro</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="79845299A101FE104B24FC5DFA5E7B14" pageId="32" pageNumber="1182" type="discussion">
<paragraph id="31210112A101FE104B24FC5DFA5E7B14" blockId="32.[827,1443,967,1051]" pageId="32" pageNumber="1182">
<emphasis id="03EADD00A101FE104B24FC5DFB7E7CD3" box="[827,1190,967,988]" italics="true" pageId="32" pageNumber="1182">
Remarks:
<taxonomicName id="F69E7A91A101FE104BA6FC5DFB7E7CD3" authorityName="Burmeister" authorityYear="1835" box="[953,1190,967,988]" class="Insecta" family="Cicadidae" genus="Hemidictya" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1182" phylum="Arthropoda" rank="species" species="frondosa">Hemidictya frondosa</taxonomicName>
</emphasis>
is included in clade C as sister group to
<taxonomicName id="F69E7A91A101FE104C1DFC7DFB627CF4" baseAuthorityName="Brancsik" baseAuthorityYear="1893" box="[1026,1210,998,1020]" class="Insecta" family="Cicadidae" genus="Hovana" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1182" phylum="Arthropoda" rank="species" species="distanti">
<emphasis id="03EADD00A101FE104C1DFC7DFB627CF4" box="[1026,1210,998,1020]" italics="true" pageId="32" pageNumber="1182">Hovana distanti</emphasis>
</taxonomicName>
. The species present one homoplastic synapomorphy: uncus absent (48 1).
</paragraph>
</subSubSection>
</treatment>
</document>
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