treatments-xml/data/B9/24/25/B92425C49A4E53A5900B9BBE4F397B67.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1029.63940" ID-GBIF-Dataset="27e8615f-ba3f-4d59-99a1-ecb8493d10f2" ID-PMC="PMC8050014" ID-Pensoft-Pub="1313-2970-1029-155" ID-Pensoft-UUID="50846BA2C2A85039BEFA6C1E2E3526FB" ID-PubMed="33935554" ID-ZooBank="9AE12DC8CDA44BA481C2B655C649AEC5" ModsDocID="1313-2970-1029-155" checkinTime="1617901766197" checkinUser="pensoft" docAuthor="Hava, Jiri, Abdel-Dayem, Mahmoud S. &amp; Aldhafer, Hathal M." docDate="2021" docId="B92425C49A4E53A5900B9BBE4F397B67" docLanguage="en" docName="ZooKeys 1029: 155-174" docOrigin="ZooKeys 1029" docPubDate="2021-04-08" docSource="http://dx.doi.org/10.3897/zookeys.1029.63940" docTitle="Thorictus riyadhensis Hava &amp; Abdel-Dayem 2021, sp. nov." docType="treatment" docUuid="6018C111-2A2F-43D8-A506-E885A4B9120A" docUuidSource="ZooBank" docVersion="4" id="50846BA2C2A85039BEFA6C1E2E3526FB" lastPageNumber="155" masterDocId="50846BA2C2A85039BEFA6C1E2E3526FB" masterDocTitle="The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia" masterLastPageNumber="174" masterPageNumber="155" pageNumber="155" updateTime="1668152078424" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hava, Jiri</mods:namePart>
<mods:affiliation>Forestry and Game Management Research Institute, Strnady 136, CZ- 252 02 Praha 5 - Zbraslav, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Abdel-Dayem, Mahmoud S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6276-1740</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
<mods:nameIdentifier type="email">mseleem@ksu.edu.sa</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aldhafer, Hathal M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4911-2332</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1029</mods:number>
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<mods:extent unit="page">
<mods:start>155</mods:start>
<mods:end>174</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1029.63940</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1029.63940</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1029-155</mods:identifier>
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<treatment ID-GBIF-Taxon="180881644" LSID="urn:lsid:zoobank.org:act:6018C111-2A2F-43D8-A506-E885A4B9120A" httpUri="http://treatment.plazi.org/id/B92425C49A4E53A5900B9BBE4F397B67" lastPageNumber="155" pageId="0" pageNumber="155">
<subSubSection pageId="0" pageNumber="155" type="nomenclature">
<paragraph pageId="0" pageNumber="155">
<taxonomicName LSID="http://zoobank.org/6018C111-2A2F-43D8-A506-E885A4B9120A" authority="Háva &amp; Abdel-Dayem" authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis" status="sp. nov.">
Thorictus riyadhensis 
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="155">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus dorsal aspect of Thorictus riyadhensis sp. nov. Hava &amp; Abdel-Dayem a specimen from Wadi Hanifah b specimen from Rhodet Khorim." figureDoi="10.3897/zookeys.1029.63940.figure3" httpUri="https://binary.pensoft.net/fig/529721" pageId="0" pageNumber="155">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Thorictus riyadhensis Hava &amp; Abdel-Dayem, sp. nov. a head lateral eye b ventral setation on pronotum and metepisternum; Thorictus arabicus Hava, 2010 c head lateral eye d ventral setation on pronotum and metepisternum." figureDoi="10.3897/zookeys.1029.63940.figure4" httpUri="https://binary.pensoft.net/fig/529722" pageId="0" pageNumber="155">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="materials_examined">
<paragraph pageId="0" pageNumber="155">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="155">
<materialsCitation ID-GBIF-Occurrence="3069288307" collectingDate="2015-04-29" collectingMethod="pitfall trap" collectorName="Abdel-Dayem M" country="Saudi Arabia" elevation="695" latitude="24.86682" location="Wadi Hanifah" longLatPrecision="1" longitude="46.45959" specimenCount="1" stateProvince="Riyadh Province" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="155">
<typeStatus>Holotype</typeStatus>
.
</emphasis>
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
• 
<specimenCount type="generic">1 ex</specimenCount>
; 
<collectingRegion country="Saudi Arabia" name="Ar Riyad">Riyadh Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:B92425C49A4E53A5900B9BBE4F397B67:D8FEB6CEE537F04DB430DE5A0B0506CB" country="Saudi Arabia" latitude="24.86682" longLatPrecision="1" longitude="46.45959" name="Wadi Hanifah" stateProvince="Riyadh Province">Wadi Hanifah</location>
, WHS04 [location near Waseel]; 
<geoCoordinate degrees="24.86682" direction="north" orientation="latitude" precision="1" value="24.86682">24.86682°N</geoCoordinate>
, 
<geoCoordinate degrees="46.45959" direction="east" orientation="longitude" precision="1" value="46.45959">46.45959°E</geoCoordinate>
; alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.94942" unit="m" value="694.942">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.94942" unit="m" value="694.942">694.942 m</elevation>
</quantity>
; 
<collectingDate value="2015-04-29">29 Apr. 2015</collectingDate>
; 
<collectorName>Abdel-Dayem M</collectorName>
et al. leg.; 
<collectingMethod>pitfall trap</collectingMethod>
; KSMA
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3069288303" collectingDate="2011-12-10" collectingDateMax="2016-01-31" collectingDateMin="2011-12-10" collectingMethod="pitfall trap" collectorName="Abdel-Dayem M" country="Saudi Arabia" elevation="670" latitude="25.432383" location="Rawdat Al Sabalh" longLatPrecision="1" longitude="47.23105" municipality="Az Zulfi" specimenCount="6" stateProvince="Riyadh Province" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="155">
<typeStatus>Paratypes</typeStatus>
.
</emphasis>
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
• 
<specimenCount type="generic">1 ex</specimenCount>
; 
<collectingRegion country="Saudi Arabia" name="Ar Riyad">Riyadh Province</collectingRegion>
, 
<collectingMunicipality>Az Zulfi</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:B92425C49A4E53A5900B9BBE4F397B67:0372CF854272B63AFE81508F6A0A1524" country="Saudi Arabia" latitude="25.432383" longLatPrecision="1" longitude="47.23105" municipality="Az Zulfi" name="Rawdat Al Sabalh" stateProvince="Riyadh Province">Rawdat Al Sabalh</location>
; 
<geoCoordinate degrees="26" direction="north" minutes="22.040" orientation="latitude" precision="1" value="26.367332">26°22.040'N</geoCoordinate>
, 
<geoCoordinate degrees="44" direction="east" minutes="59.137" orientation="longitude" precision="1" value="44.98562">44°59.137'E</geoCoordinate>
; alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</elevation>
</quantity>
; 
<collectingDate value="2016-01-31">31 Jan. 2016</collectingDate>
; 
<collectorName>Abdel-Dayem M</collectorName>
et al. leg.; 
<collectingMethod>pitfall trap</collectingMethod>
; KSMA • 
<specimenCount type="generic">1 ex</specimenCount>
; 
<collectingRegion country="Saudi Arabia" name="Ar Riyad">Riyadh Province</collectingRegion>
, Rhodet Khorim; 
<geoCoordinate degrees="25" direction="north" minutes="25.943" orientation="latitude" precision="1" value="25.432383">25°25.943'N</geoCoordinate>
, 
<geoCoordinate degrees="47" direction="east" minutes="13.863" orientation="longitude" precision="1" value="47.23105">47°13.863'E</geoCoordinate>
; alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.72" unit="m" value="572.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.72" unit="m" value="572.0">572 m</elevation>
</quantity>
; 
<collectingDate value="2012-04-28">28 Apr. 2012</collectingDate>
; 
<collectingMethod>pitfall trap</collectingMethod>
, 
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Calotropis procera</emphasis>
</taxonomicName>
; KSMA • 
<specimenCount type="generic">1 ex</specimenCount>
; same collection data as preceding; 
<collectingDate value="2012-06-09">9 Jun. 2012</collectingDate>
; 
<collectingMethod>pitfall trap</collectingMethod>
, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia gerrardii" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="gerrardii">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia gerrardii</emphasis>
</taxonomicName>
; KSMA • 
<specimenCount type="generic">1 ex</specimenCount>
; same collection data as preceding; 
<collectingDate value="2012-05-26">26 May 2012</collectingDate>
; 
<collectingMethod>pitfall trap</collectingMethod>
, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia gerrardii" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="gerrardii">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia gerrardii</emphasis>
</taxonomicName>
; JHAC • 
<specimenCount type="generic">1 ex</specimenCount>
; same collection data as preceding; 
<collectingDate value="2012-05-26">26 May 2012</collectingDate>
; PT, 
<taxonomicName authorityName="Roem &amp; Schult" authorityYear="1819" class="Dicotyledoneae" family="Solanaceae" genus="Lycium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lycium shawii" order="Tubiflorae" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="shawii">
<emphasis italics="true" pageId="0" pageNumber="155">Lycium shawii</emphasis>
</taxonomicName>
; KSMA • 
<specimenCount type="generic">1 ex</specimenCount>
; same collection data as preceding; 
<collectingDate value="2011-12-10">10 Dec. 2011</collectingDate>
; 
<collectingMethod>pitfall trap</collectingMethod>
; KSMA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="description">
<paragraph pageId="0" pageNumber="155">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="155">Body small, brown covered by long, yellow setae on dorsal surfaces and short setae on ventral surfaces. Measurements (mm): TL 2.9, PW 1.9, EW 1.9. Head finely punctate with long yellow setae. Labial palpi entirely brown. Antennae brown, with 11 antennomeres; antennal club compact, with three antennomeres. Pronotum as finely punctate as head and covered by long yellow setation. Lateral margin of pronotum not dentate. Pronotum in posterior part near scutellum without bumps. Ventral posterior pronotal angles with long yellow setation. Scutellum not visible from above. Elytra very finely punctate covered by long yellow setation. Each elytron in anterior part near humeri with one large bump. Epipleuron finely punctate, anterior angles with long yellow setation. Prosternum finely punctate. Mesosternum finely punctate, mesosternal bulge. Metasternum finely punctate. Visible abdominal ventrites very finely punctate and covered by long yellow setae. First abdominal ventrite without anterior, longitudinal striation. Legs brown, covered by yellow setae.</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure3" httpUri="https://binary.pensoft.net/fig/529721" pageId="0" pageNumber="155" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 3.</emphasis>
Habitus dorsal aspect of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
sp. nov. 
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem 
<emphasis bold="true" pageId="0" pageNumber="155">a</emphasis>
specimen from Wadi Hanifah 
<emphasis bold="true" pageId="0" pageNumber="155">b</emphasis>
specimen from Rhodet Khorim.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="variability">
<paragraph pageId="0" pageNumber="155">Variability.</paragraph>
<paragraph pageId="0" pageNumber="155">Body measurements (mm): TL 2.5-2.9, PW 1.5-1.9, EW 1.5-1.9.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="differential diagnosis">
<paragraph pageId="0" pageNumber="155">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="155">
The new species belong to the 
<taxonomicName authorityName="Germar" authorityYear="1834" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus castaneus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus castaneus</emphasis>
</taxonomicName>
species group, from Saudi Arabia. There is currently only one known species, 
<taxonomicName authorityName="Hava" authorityYear="2010" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus arabicus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus arabicus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
, 2010 but the new species differs from it in the abovementioned characteristics.
</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure4" httpUri="https://binary.pensoft.net/fig/529722" pageId="0" pageNumber="155" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 4.</emphasis>
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="155">a</emphasis>
head lateral eye 
<emphasis bold="true" pageId="0" pageNumber="155">b</emphasis>
ventral setation on pronotum and metepisternum; 
<taxonomicName authorityName="Hava" authorityYear="2010" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus arabicus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus arabicus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
, 2010 
<emphasis bold="true" pageId="0" pageNumber="155">c</emphasis>
head lateral eye 
<emphasis bold="true" pageId="0" pageNumber="155">d</emphasis>
ventral setation on pronotum and metepisternum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="etymology">
<paragraph pageId="0" pageNumber="155">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="155">Named according to type locality: Riyadh Province.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="remarks">
<paragraph pageId="0" pageNumber="155">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="155">
The species was erroneously published by 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.653.10252" author="Abdel-Dayem, MS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="155" pagination="1 - 78" refId="B1" refString="Abdel-Dayem, MS, Fad, HH, El-Torkey, AM, Elgharbawy, AA, Aldryhim, YN, Kondratieff, BC, Al, Ansi AN, Aldhafer, HM, 2017. The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia. ZooKeys 653: 1 - 78, DOI: https://doi.org/10.3897/zookeys.653.10252" title="The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia." url="https://doi.org/10.3897/zookeys.653.10252" volume="653" year="2017">Abdel-Dayem et al. (2017)</bibRefCitation>
as 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus foreli" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="foreli">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus foreli</emphasis>
</taxonomicName>
Wasmann, 1894. 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus foreli" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="foreli">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus foreli</emphasis>
</taxonomicName>
is a North African species, occurring in Algeria, Morocco, and Tunisia (
<bibRefCitation DOI="https://doi.org/10.1163/9789004286610" author="Hava, J" journalOrPublisher="Annals of the Ditsong National Museum of Natural History" pageId="0" pageNumber="155" refId="B9" refString="Hava, J, 2015a. World Catalogue of Insects (Vol. 13). Dermestidae (Coleoptera). Leiden/Boston: Brill, 419 pp. https://doi.org/10.1163/9789004286610" title="World Catalogue of Insects (Vol. 13). Dermestidae (Coleoptera). Leiden / Boston: Brill, 419 pp." url="https://doi.org/10.1163/9789004286610" year="2015 a">
<normalizedToken originalValue="Háva">Hava</normalizedToken>
2015a
</bibRefCitation>
), so it is excluded from the fauna of Saudi Arabia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="ecological notes">
<paragraph pageId="0" pageNumber="155">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="155">
The holotype was found in Wadi Hanifah at an area with loam soil covered by 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Carthamus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carthamus oxyacantha" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="oxyacantha">
<emphasis italics="true" pageId="0" pageNumber="155">Carthamus oxyacantha</emphasis>
</taxonomicName>
M. Bieb. (
<taxonomicName class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) and 
<taxonomicName class="Insecta" family="Brassicaceae" genus="Zilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zilla spinosa" order="Brassicales" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis italics="true" pageId="0" pageNumber="155">Zilla spinosa</emphasis>
</taxonomicName>
(L.) Prantl (
<taxonomicName class="Insecta" family="Brassicaceae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Brassicales" pageId="0" pageNumber="155" phylum="Arthropoda" rank="family">Brassicaceae</taxonomicName>
) (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Thorictus riyadhensis Hava &amp; Abdel-Dayem, sp. nov. holotype in Wadi Hanifah, Ad Diriyah, Riyadh Province, at an elevation of 695 m. Carthamus oxyacantha M. Bieb. (Asteraceae) in the bottom right corner, and Zilla spinosa (L.) Prantl (Brassicaceae) in the foreground." figureDoi="10.3897/zookeys.1029.63940.figure5" httpUri="https://binary.pensoft.net/fig/529723" pageId="0" pageNumber="155">5</figureCitation>
). The ant species 
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1804" class="Insecta" family="Staphylinidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Camponotus thoracicus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="thoracicus">
<emphasis italics="true" pageId="0" pageNumber="155">Camponotus thoracicus</emphasis>
</taxonomicName>
(Fabricius 1804), 
<taxonomicName authorityName="Collingwood &amp; Agosti" authorityYear="1996" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis holgerseni" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="holgerseni">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis holgerseni</emphasis>
</taxonomicName>
(Collingwood &amp; Agosti, 1996), 
<taxonomicName baseAuthorityName="Andre" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis lividus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="lividus">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis lividus</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1881), 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Lepisiota" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lepisiota simplex" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="155">Lepisiota simplex</emphasis>
</taxonomicName>
(Forel, 1892), and 
<taxonomicName class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium abeillei" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="abeillei">
<emphasis italics="true" pageId="0" pageNumber="155">Monomorium abeillei</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1881) were collected with the holotype in the same pitfall trap. The paratype was found in loam area covered with 
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Calotropis procera</emphasis>
</taxonomicName>
(Aiton) W.T. Aiton (
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Apocynaceae</taxonomicName>
) and 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Pulicaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pulicaria undulata" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis italics="true" pageId="0" pageNumber="155">Pulicaria undulata</emphasis>
</taxonomicName>
(L.) C. A. Mey. (
<taxonomicName class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) at Rawdat Al Sabalh (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitat of Thorictus riyadhensis Hava &amp; Abdel-Dayem, sp. nov. paratype in Rawdat Al Sabalh, Az Zulfi, Riyadh Province, at 670 m elevation. Calotropis procera (Aiton) W. T. Aiton (Apocynaceae) in background, and Pulicaria undulata (L.) C. A. Mey. (Asteraceae) in the foreground and among the shrubs." figureDoi="10.3897/zookeys.1029.63940.figure6" httpUri="https://binary.pensoft.net/fig/529724" pageId="0" pageNumber="155">6</figureCitation>
). While at Rhodet Khorim, the paratypes were collected from a sandy area using pitfall traps under canopies of 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia gerrardii" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="gerrardii">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia gerrardii</emphasis>
</taxonomicName>
Benth. (
<taxonomicName class="Magnoliopsida" family="Fabaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), 
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Calotropis procera</emphasis>
</taxonomicName>
(Aiton) W.T. Aiton, and 
<taxonomicName authorityName="Roem &amp; Schult" authorityYear="1819" class="Dicotyledoneae" family="Solanaceae" genus="Lycium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lycium shawii" order="Tubiflorae" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="shawii">
<emphasis italics="true" pageId="0" pageNumber="155">Lycium shawii</emphasis>
</taxonomicName>
Roem. &amp; Schult. (
<taxonomicName class="Dicotyledoneae" family="Solanaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Solanaceae</taxonomicName>
) (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat of Thorictus riyadhensis Hava &amp; Abdel-Dayem, sp. nov. paratype in Rhodet Khorim, Ramah, Riyadh Province, at an elevation of 572 m. Trees of Acacia gerrardii Benth. (Fabaceae) in the background, shrubs of Calotropis procera (Aiton) W. T. Aiton (Apocynaceae) and Ziziphus nummularia (Burm. f) Wight &amp; Arn. (Rhamnaceae), shrub of Lycium shawii Roem. &amp; Schult. (Solanaceae) in the left corner of the bottom, and Rhazya stricta Decne. (Apocynaceae) in the foreground and middle." figureDoi="10.3897/zookeys.1029.63940.figure7" httpUri="https://binary.pensoft.net/fig/529725" pageId="0" pageNumber="155">7</figureCitation>
). Ant species of 
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Insecta" family="Staphylinidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Camponotus sericeus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sericeus">
<emphasis italics="true" pageId="0" pageNumber="155">Camponotus sericeus</emphasis>
</taxonomicName>
(Fabricius, 1798), 
<taxonomicName authorityName="Forel" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Camponotus xerxes" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="xerxes">
<emphasis italics="true" pageId="0" pageNumber="155">Camponotus xerxes</emphasis>
</taxonomicName>
Forel, 1904, 
<taxonomicName authorityName="Sharaf &amp; Aldawood" authorityYear="2015" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis fisheri" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis fisheri</emphasis>
</taxonomicName>
Sharaf &amp; Aldawood, 2015, 
<taxonomicName baseAuthorityName="Andre" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis lividus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="lividus">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis lividus</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1881), 
<taxonomicName baseAuthorityName="Andre" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis niger" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="niger">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis niger</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1881), 
<taxonomicName baseAuthorityName="Andre" baseAuthorityYear="1881" class="Insecta" family="Formicidae" genus="Cataglyphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataglyphis viaticoides" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="viaticoides">
<emphasis italics="true" pageId="0" pageNumber="155">Cataglyphis viaticoides</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1881), 
<taxonomicName authorityName="Collingwood &amp; Agosti" authorityYear="1996" class="Magnoliopsida" family="Asteraceae" genus="Lepisiota" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lepisiota dammama" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="dammama">
<emphasis italics="true" pageId="0" pageNumber="155">Lepisiota dammama</emphasis>
</taxonomicName>
Collingwood &amp; Agosti, 1996, 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Lepisiota" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lepisiota simplex" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="simplex">
<emphasis italics="true" pageId="0" pageNumber="155">Lepisiota simplex</emphasis>
</taxonomicName>
(Forel, 1892), 
<taxonomicName authorityName="Forel" authorityYear="1911" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium chefketi" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="chefketi">
<emphasis italics="true" pageId="0" pageNumber="155">Tetramorium chefketi</emphasis>
</taxonomicName>
Forel, 1911, 
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium sericeiventre" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sericeiventre">
<emphasis italics="true" pageId="0" pageNumber="155">Tetramorium sericeiventre</emphasis>
</taxonomicName>
Emery, 1877, and 
<taxonomicName authorityName="Forel" authorityYear="1902" class="Insecta" family="Formicidae" genus="Trichomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trichomyrmex mayri" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="mayri">
<emphasis italics="true" pageId="0" pageNumber="155">Trichomyrmex mayri</emphasis>
</taxonomicName>
(Forel, 1902) were caught with the paratypes from the same pitfall traps. 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
was collected during December and January, and from April to June.
</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure5" httpUri="https://binary.pensoft.net/fig/529723" pageId="0" pageNumber="155" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 5.</emphasis>
Habitat of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. holotype in Wadi Hanifah, Ad Diriyah, Riyadh Province, at an elevation of 695 m. 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Carthamus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carthamus oxyacantha" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="oxyacantha">
<emphasis italics="true" pageId="0" pageNumber="155">Carthamus oxyacantha</emphasis>
</taxonomicName>
M. Bieb. (
<taxonomicName class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) in the bottom right corner, and 
<taxonomicName class="Insecta" family="Brassicaceae" genus="Zilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zilla spinosa" order="Brassicales" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis italics="true" pageId="0" pageNumber="155">Zilla spinosa</emphasis>
</taxonomicName>
(L.) Prantl (
<taxonomicName class="Insecta" family="Brassicaceae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Brassicales" pageId="0" pageNumber="155" phylum="Arthropoda" rank="family">Brassicaceae</taxonomicName>
) in the foreground.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1029.63940.figure6" httpUri="https://binary.pensoft.net/fig/529724" pageId="0" pageNumber="155" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 6.</emphasis>
Habitat of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. paratype in Rawdat Al Sabalh, Az Zulfi, Riyadh Province, at 670 m elevation. 
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Calotropis procera</emphasis>
</taxonomicName>
(Aiton) W.T. Aiton (
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Apocynaceae</taxonomicName>
) in background, and 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Pulicaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pulicaria undulata" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="undulata">
<emphasis italics="true" pageId="0" pageNumber="155">Pulicaria undulata</emphasis>
</taxonomicName>
(L.) C. A. Mey. (
<taxonomicName class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) in the foreground and among the shrubs.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1029.63940.figure7" httpUri="https://binary.pensoft.net/fig/529725" pageId="0" pageNumber="155" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 7.</emphasis>
Habitat of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus riyadhensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="riyadhensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus riyadhensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. paratype in Rhodet Khorim, Ramah, Riyadh Province, at an elevation of 572 m. Trees of 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia gerrardii" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="gerrardii">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia gerrardii</emphasis>
</taxonomicName>
Benth. (
<taxonomicName class="Magnoliopsida" family="Fabaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
) in the background, shrubs of 
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Calotropis procera</emphasis>
</taxonomicName>
(Aiton) W. T. Aiton (
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Apocynaceae</taxonomicName>
) and 
<taxonomicName class="Dicotyledoneae" family="Rhamnaceae" genus="Ziziphus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ziziphus nummularia" order="Rhamnales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="nummularia">
<emphasis italics="true" pageId="0" pageNumber="155">Ziziphus nummularia</emphasis>
</taxonomicName>
(Burm. f) Wight &amp; Arn. (
<taxonomicName class="Dicotyledoneae" family="Rhamnaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rhamnales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Rhamnaceae</taxonomicName>
), shrub of 
<taxonomicName authorityName="Roem &amp; Schult" authorityYear="1819" class="Dicotyledoneae" family="Solanaceae" genus="Lycium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lycium shawii" order="Tubiflorae" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="shawii">
<emphasis italics="true" pageId="0" pageNumber="155">Lycium shawii</emphasis>
</taxonomicName>
Roem. &amp; Schult. (
<taxonomicName class="Dicotyledoneae" family="Solanaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Solanaceae</taxonomicName>
) in the left corner of the bottom, and 
<taxonomicName authorityName="Decne" authorityYear="1835" class="Dicotyledoneae" family="Apocynaceae" genus="Rhazya" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rhazya stricta" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="stricta">
<emphasis italics="true" pageId="0" pageNumber="155">Rhazya stricta</emphasis>
</taxonomicName>
Decne. (
<taxonomicName authorityName="Decaisne" authorityYear="1835" class="Dicotyledoneae" family="Apocynaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Apocynaceae</taxonomicName>
) in the foreground and middle.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="distribution">
<paragraph pageId="0" pageNumber="155">Geographical distribution.</paragraph>
<paragraph pageId="0" pageNumber="155">
This new 
<taxonomicName authorityName="Agassiz" authorityYear="1846" lsidName="" pageId="0" pageNumber="155" rank="tribe" tribe="Thorictini">Thorictini</taxonomicName>
species is known from Najd Plateau, Central Saudi Arabia, Riyadh Province (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of Saudi Arabian provinces and distribution of Thorictinae species." figureDoi="10.3897/zookeys.1029.63940.figure1" httpUri="https://binary.pensoft.net/fig/529719" pageId="0" pageNumber="155">1</figureCitation>
), at Wadi Hanifah (Ad Diriyah), Rawdat Al Sabalh (Az Zulfi), and Rhodet Khorim (Ramah).
</paragraph>
</subSubSection>
</treatment>
</document>