150 lines
20 KiB
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150 lines
20 KiB
XML
<document id="00BDB36728A7E5ACF6DB6B4152F0CFEB" ID-DOI="10.5281/zenodo.3738587" ID-GBIF-Dataset="9c4d25fb-5ff7-4ebd-87f9-9d53b2d53038" ID-ISSN="0253-116X" ID-Zenodo-Dep="3738587" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585850417291" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2019" docId="B861878DFFB69B1861FECDBCFCB8F096" docLanguage="en" docName="Linzer.biol.Beitr.51.2.773-788.pdf.imf" docOrigin="Linzer biologische Beiträge 51 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Ischnosoma cuspidatum Assing 2019, nov.sp." docType="treatment" docVersion="6" lastPageNumber="777" masterDocId="4458FFF5FFB59B1C6169C96BFFD3F322" masterDocTitle="New species and additional records of Ischnosoma from Georgia (Coleoptera, Staphylinidae, Tachyporinae)" masterLastPageNumber="788" masterPageNumber="773" pageNumber="776" updateTime="1698735196410" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="0E4D179CE5B715FA36187261623D3839">
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<mods:title id="DC6C6551D414A324EC552C10CB91A5B4">New species and additional records of Ischnosoma from Georgia (Coleoptera, Staphylinidae, Tachyporinae)</mods:title>
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<mods:name id="B51E3F31AFF2F8E71D2185D9040D3518" type="personal">
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<mods:namePart id="AC5A7EF0751F14BA06964B593028571B">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="7C5386C63B917E20631E589BF4AD90F5">text</mods:typeOfResource>
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<mods:title id="CDB2EAD08384152F697EA1D7318C7323">Linzer biologische Beiträge</mods:title>
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<mods:part id="182D37A07B52D7F509371BEC88AA857B">
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<mods:date id="3BCFBAD3CCF8D7709DC80A75E198E540">2019</mods:date>
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<mods:number id="F99038F1CD964222C7EFB5DAA88ADC5A">2019-12-20</mods:number>
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<mods:number id="BAD85C82ACF5BD072AEB0C945515B413">51</mods:number>
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<mods:identifier id="98245F666A1D8314001DD14E5A3B0E12" type="DOI">10.5281/zenodo.3738587</mods:identifier>
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<mods:identifier id="0A5FE68E75ECB88A29CA3D07C5CF28E2" type="ISSN">0253-116X</mods:identifier>
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<treatment id="B861878DFFB69B1861FECDBCFCB8F096" ID-DOI="http://doi.org/10.5281/zenodo.3811211" ID-GBIF-Taxon="163422848" ID-Zenodo-Dep="3811211" LSID="urn:lsid:plazi:treatment:B861878DFFB69B1861FECDBCFCB8F096" httpUri="http://treatment.plazi.org/id/B861878DFFB69B1861FECDBCFCB8F096" lastPageId="4" lastPageNumber="777" pageId="3" pageNumber="776">
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<subSubSection id="78D26510FFB69B1F61FECDBCFEC6F667" pageId="3" pageNumber="776" type="nomenclature">
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<paragraph id="3077369BFFB69B1F61FECDBCFD0AF7CF" blockId="3.[151,729,1239,1261]" box="[151,729,1239,1261]" pageId="3" pageNumber="776">
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<heading id="6B3F81F7FFB69B1F61FECDBCFD0AF7CF" box="[151,729,1239,1261]" fontSize="8" level="2" pageId="3" pageNumber="776" reason="2">
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<emphasis id="02BCEA89FFB69B1F61FECDBCFD0AF7CF" bold="true" box="[151,729,1239,1261]" pageId="3" pageNumber="776">
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<taxonomicName id="F7C84D18FFB69B1F61FECDBCFE46F7CF" ID-CoL="8Y4X6" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[151,405,1239,1261]" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="cuspidatum" status="sp. nov.">
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<emphasis id="02BCEA89FFB69B1F61FECDBCFE46F7CF" bold="true" box="[151,405,1239,1261]" italics="true" pageId="3" pageNumber="776">Ischnosoma cuspidatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="198F57F2FFB69B1F60F2CDB3FE3BF7CF" box="[411,488,1240,1261]" pageId="3" pageNumber="776" rank="species">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="A8F32A1EFFB69B1F609CCDBCFDEBF7CE" box="[501,568,1239,1261]" captionStart="Figs 1-8" captionStartId="5.[151,191,1499,1521]" captionTargetBox="[157,1060,198,1474]" captionTargetId="figure@5.[155,1063,198,1484]" captionTargetPageId="5" captionText="Figs 1-8: Ischnosoma acre (1), I. cuspidatum (2), I. convergens (3-4), I. parallelum (5-6), and I. myops (7-8): (1-3, 5, 7) habitus; (4, 6, 8) head and pronotum in lateral view. Scale bars: 1-3, 5, 7: 1.0 mm; 4, 6, 8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3738593" httpUri="https://zenodo.org/record/3738593/files/figure.png" pageId="3" pageNumber="776">Figs 2</figureCitation>
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, 47-51,
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<figureCitation id="A8F32A1EFFB69B1F63E2CDB3FD02F7CF" box="[651,721,1239,1261]" captionStart="Map 2" captionStartId="4.[151,196,1515,1536]" captionTargetBox="[153,1047,957,1503]" captionTargetId="figure@4.[153,1047,957,1503]" captionTargetPageId="4" captionText="Map 2: Confirmed distribution of Ischnosoma myops (black circles) and type locality of I. cuspidatum (white star)." figureDoi="http://doi.org/10.5281/zenodo.3738591" httpUri="https://zenodo.org/record/3738591/files/figure.png" pageId="3" pageNumber="776">Map 2</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3077369BFFB69B1F61FECC69FEC6F667" blockId="3.[151,1058,1282,1349]" pageId="3" pageNumber="776">
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<materialsCitation id="80A03CC6FFB69B1F61FECC69FEC6F667" ID-GBIF-Occurrence="2595615714" collectingDate="2019-05-09" collectionCode="T, V" elevation="830" latitude="42.20528" location="Kakheti" longLatPrecision="19" longitude="42.20528" pageId="3" pageNumber="776" specimenCount="2" specimenCount-male="2" stateProvince="Kakheti" typeStatus="holotype">
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<collectionCode id="56D9AE5EFFB69B1F61FECC69FF76F635" box="[151,165,1282,1303]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="776">T</collectionCode>
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y p e m a t e r i a l:
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<typeStatus id="EF738839FFB69B1F60E4CC69FE32F63A" box="[397,481,1282,1304]" pageId="3" pageNumber="776" type="holotype">Holotype</typeStatus>
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<specimenCount id="26CEFD12FFB69B1F6084CC69FDD3F635" box="[493,512,1282,1303]" pageId="3" pageNumber="776" type="male">♂</specimenCount>
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: "
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<geoCoordinate id="55FC505CFFB69B1F637ECC69FDAAF635" box="[535,633,1282,1303]" degrees="42" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="776" precision="15" seconds="19" value="42.20528">N42°12'19</geoCoordinate>
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<geoCoordinate id="55FC505CFFB69B1F63EDCC69FD36F63A" box="[644,741,1282,1304]" degrees="45" direction="east" minutes="27" orientation="longitude" pageId="3" pageNumber="776" precision="15" seconds="45" value="45.4625">E45°27'45</geoCoordinate>
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, GG
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<collectingRegion id="F20CF879FFB69B1F6276CC69FCB4F63A" box="[799,871,1282,1304]" country="Georgia" name="Kakheti" pageId="3" pageNumber="776">Kakheti</collectingRegion>
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Lechuri N,
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<quantity id="F7309B7EFFB69B1F628DCC69FBF1F635" box="[996,1058,1282,1303]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="776" unit="m" value="830.0">
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<elevation id="BBE5D1A8FFB69B1F628DCC69FBF1F635" box="[996,1058,1282,1303]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="776" unit="m" value="830.0">830 m</elevation>
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</quantity>
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<date id="4476105BFFB69B1F61FECC73FF3AF60C" box="[151,233,1304,1326]" pageId="3" pageNumber="776" value="2019-05-09">
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<collectingDate id="5432E9B3FFB69B1F61FECC73FF3AF60C" box="[151,233,1304,1326]" pageId="3" pageNumber="776" value="2019-05-09">9.5.2019</collectingDate>
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</date>
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, Brachat & Meybohm (2) /
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<typeStatus id="EF738839FFB69B1F609ECC72FD85F60D" box="[503,598,1305,1327]" pageId="3" pageNumber="776" type="holotype">Holotypus</typeStatus>
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<specimenCount id="26CEFD12FFB69B1F6337CC72FDBCF60C" box="[606,623,1305,1326]" pageId="3" pageNumber="776" type="male">♂</specimenCount>
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<taxonomicName id="F7C84D18FFB69B1F6311CC72FC85F60C" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[632,854,1305,1326]" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="776" phylum="Arthropoda" rank="species" species="cuspidatum" status="sp. nov.">
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<emphasis id="02BCEA89FFB69B1F6311CC72FC85F60C" box="[632,854,1305,1326]" italics="true" pageId="3" pageNumber="776">Ischnosoma cuspidatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="198F57F2FFB69B1F6237CC72FC41F60C" box="[862,914,1305,1326]" pageId="3" pageNumber="776" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<collectionCode id="56D9AE5EFFB69B1F62ADCC72FC04F60C" box="[964,983,1305,1326]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="776" type="Museum">V</collectionCode>
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. Assing 2019" (cAss).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="78D26510FFB69B1861FECC3FFCB8F096" lastPageId="4" lastPageNumber="777" pageId="3" pageNumber="776" type="description">
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<paragraph id="3077369BFFB69B1F61FECC3FFCE1F6A5" blockId="3.[151,1058,1363,1415]" pageId="3" pageNumber="776">E t y m o l o g y: The specific epithet (Latin, adjective: acute) alludes to the sharply acute apex of the ventral process of the aedeagus in ventral view.</paragraph>
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<paragraph id="3077369BFFB69B1F61FECCFCFD48F6C5" blockId="3.[151,1058,1430,1511]" pageId="3" pageNumber="776">
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D e s c r i p t i o n: Body length
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<quantity id="F7309B7EFFB69B1F6361CCFDFD88F68E" box="[520,603,1430,1452]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="776" unit="mm" value="4.3">4.3 mm</quantity>
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; length of forebody
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<quantity id="F7309B7EFFB69B1F625FCCFDFC59F68E" box="[822,906,1430,1452]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="776" unit="mm" value="1.9">1.9 mm</quantity>
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. Habitus as in
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<figureCitation id="A8F32A1EFFB69B1F61FECCD8FF06F6EA" box="[151,213,1459,1481]" captionStart="Figs 1-8" captionStartId="5.[151,191,1499,1521]" captionTargetBox="[157,1060,198,1474]" captionTargetId="figure@5.[155,1063,198,1484]" captionTargetPageId="5" captionText="Figs 1-8: Ischnosoma acre (1), I. cuspidatum (2), I. convergens (3-4), I. parallelum (5-6), and I. myops (7-8): (1-3, 5, 7) habitus; (4, 6, 8) head and pronotum in lateral view. Scale bars: 1-3, 5, 7: 1.0 mm; 4, 6, 8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3738593" httpUri="https://zenodo.org/record/3738593/files/figure.png" pageId="3" pageNumber="776">Fig. 2</figureCitation>
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. Coloration: body reddish, with the antero-median portions of tergite IV-VI slightly darker; legs dark-yellow; antennae pale-reddish.
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</paragraph>
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<paragraph id="3077369BFFB69B1F61FECC9CFD91F50B" blockId="3.[151,1058,1526,1577]" pageId="3" pageNumber="776">
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Eyes (
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<figureCitation id="A8F32A1EFFB69B1F61B1CC9DFEF0F52E" box="[216,291,1526,1548]" captionStart="Figs 47-51" captionStartId="11.[151,191,1169,1191]" captionTargetBox="[156,1059,207,1148]" captionTargetId="figure@11.[153,1059,203,1157]" captionTargetPageId="11" captionText="Figs 47-51: Ischnosoma cuspidatum: (47) head and pronotum in lateral view; (48) male sternite VII; (49) male sternite VIII; (50-51) aedeagus in lateral and in ventral view. Scale bars: 47: 0.5 mm; 48-51: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3738607" httpUri="https://zenodo.org/record/3738607/files/figure.png" pageId="3" pageNumber="776">Fig. 47</figureCitation>
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) slightly longer than postocular region in lateral view, composed of some 40-50 ommatidia. Antenna
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<quantity id="F7309B7EFFB69B1F60C7CF78FDDBF50B" box="[430,520,1555,1577]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="776" unit="mm" value="1.45">1.45 mm</quantity>
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long.
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</paragraph>
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<paragraph id="3077369BFFB19B1861FEC9BFFD42F206" blockId="4.[151,1058,211,292]" pageId="4" pageNumber="777">Pronotum 1.15 times as broad as long and 1.83 times as broad as head; disc with moderately dense micropunctation, mostly without microsculpture (nearly obsolete traces of microsculpture visible only near anterior margin).</paragraph>
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<paragraph id="3077369BFFB19B1861FEC858FE31F244" blockId="4.[151,1059,307,359]" pageId="4" pageNumber="777">Elytra 0.82 times as long as pronotum, without microsculpture; sutural and lateral series each composed of six punctures.</paragraph>
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<paragraph id="3077369BFFB19B1861FEC81DFC2DF28B" blockId="4.[151,1058,374,425]" pageId="4" pageNumber="777">Abdomen with rather coarse and dense punctation, punctures somewhat sparser in postero-median portions of tergites; posterior margin of tergite VII without palisade fringe.</paragraph>
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<paragraph id="3077369BFFB19B1861FEC8D2FCDEF166" blockId="4.[151,1058,441,580]" pageId="4" pageNumber="777">
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♂: sternite VII (
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<figureCitation id="A8F32A1EFFB19B186053C8D2FE57F2ED" box="[314,388,441,463]" captionStart="Figs 47-51" captionStartId="11.[151,191,1169,1191]" captionTargetBox="[156,1059,207,1148]" captionTargetId="figure@11.[153,1059,203,1157]" captionTargetPageId="11" captionText="Figs 47-51: Ischnosoma cuspidatum: (47) head and pronotum in lateral view; (48) male sternite VII; (49) male sternite VIII; (50-51) aedeagus in lateral and in ventral view. Scale bars: 47: 0.5 mm; 48-51: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3738607" httpUri="https://zenodo.org/record/3738607/files/figure.png" pageId="4" pageNumber="777">Fig. 48</figureCitation>
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) weakly transverse, with a distinct cluster of dense setae in postero-median portion; sternite VIII (
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<figureCitation id="A8F32A1EFFB19B18609FC8BDFD97F2CE" box="[502,580,470,492]" captionStart="Figs 47-51" captionStartId="11.[151,191,1169,1191]" captionTargetBox="[156,1059,207,1148]" captionTargetId="figure@11.[153,1059,203,1157]" captionTargetPageId="11" captionText="Figs 47-51: Ischnosoma cuspidatum: (47) head and pronotum in lateral view; (48) male sternite VII; (49) male sternite VIII; (50-51) aedeagus in lateral and in ventral view. Scale bars: 47: 0.5 mm; 48-51: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3738607" httpUri="https://zenodo.org/record/3738607/files/figure.png" pageId="4" pageNumber="777">Fig. 49</figureCitation>
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) distinctly oblong, in postero-median portion with a cluster of moderately dense and rather short setae, posterior margin strongly concave; aedeagus (
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<figureCitation id="A8F32A1EFFB19B18600CCB7AFE05F104" box="[357,470,528,551]" captionStart="Figs 47-51" captionStartId="11.[151,191,1169,1191]" captionTargetBox="[156,1059,207,1148]" captionTargetId="figure@11.[153,1059,203,1157]" captionTargetPageId="11" captionText="Figs 47-51: Ischnosoma cuspidatum: (47) head and pronotum in lateral view; (48) male sternite VII; (49) male sternite VIII; (50-51) aedeagus in lateral and in ventral view. Scale bars: 47: 0.5 mm; 48-51: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3738607" httpUri="https://zenodo.org/record/3738607/files/figure.png" pageId="4" pageNumber="777">Figs 50-51</figureCitation>
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)
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<quantity id="F7309B7EFFB19B18608CCB7AFD92F104" box="[485,577,529,551]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="777" unit="mm" value="0.68">0.68 mm</quantity>
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long; ventral process apically extremely acute in ventral view; parameres with numerous setae in apical half.
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</paragraph>
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<paragraph id="3077369BFFB19B1861FECB38FCABF015" blockId="4.[151,1058,595,823]" pageId="4" pageNumber="777">
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C o m p a r a t i v e n o t e s:
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<taxonomicName id="F7C84D18FFB19B18609ACB38FD3BF14A" authorityName="Assing" authorityYear="2019" box="[499,744,595,616]" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="777" phylum="Arthropoda" rank="species" species="cuspidatum">
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<emphasis id="02BCEA89FFB19B18609ACB38FD3BF14A" box="[499,744,595,616]" italics="true" pageId="4" pageNumber="777">Ischnosoma cuspidatum</emphasis>
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</taxonomicName>
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is distinguished from
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<emphasis id="02BCEA89FFB19B1862A3CB3FFBC8F14B" box="[970,1051,596,617]" italics="true" pageId="4" pageNumber="777">I. major</emphasis>
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, its geographically closest micropterous congener (see map
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<quantity id="F7309B7EFFB19B18627ECB1AFC90F1A5" box="[791,835,625,647]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="777" unit="in" value="2.0">2 in</quantity>
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ASSING & SCHÜLKE 2017) by slightly larger body size, the (near) absence of microsculpture on the pronotum and the elytra, a male sternite VII with a truncate posterior margin and with a less extensive cluster of setae in the postero-median portion, a more slender male sternite VIII with a more deeply concave posterior margin and a distinct cluster of dense setae in the postero-median portion, and by a more acute and differently shaped apex of the ventral process of the aedeagus. For illustrations of
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<emphasis id="02BCEA89FFB19B18633ECA4AFD7AF014" box="[599,681,801,822]" italics="true" pageId="4" pageNumber="777">I. major</emphasis>
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see
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<bibRefCitation id="54594B6AFFB19B1863BCCA4AFCA7F015" author="KOCIAN M" box="[725,884,801,823]" pageId="4" pageNumber="777" pagination="241 - 299" refId="ref7181" refString="KOCIAN M. (1997): A revision of Western Palearctic species of the genus Ischnosoma STEPHENS (Coleoptera, Staphylinidae: Tachyporinae). - Acta Universitatis Carolinae Biologica 40 (1996): 241 - 299." type="journal article" year="1997">KOCIAN (1997)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="3077369BFFB19B1861FECA2CFCB8F096" blockId="4.[151,1058,838,948]" pageId="4" pageNumber="777">
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D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated to the north of Lechuri in the southern slopes of the Greater Caucasus,
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<collectingRegion id="F20CF879FFB19B1862E9CA08FBCDF05B" box="[896,1054,867,889]" country="Georgia" name="Kakheti" pageId="4" pageNumber="777">Kakheti region</collectingRegion>
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, Northeast
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<collectingCountry id="48DF760BFFB19B18606ACAEAFE86F0B5" box="[259,341,897,919]" name="Georgia" pageId="4" pageNumber="777">Georgia</collectingCountry>
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(
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<figureCitation id="A8F32A1EFFB19B18600DCAEAFE7FF0B5" box="[356,428,897,919]" captionStart="Map 2" captionStartId="4.[151,196,1515,1536]" captionTargetBox="[153,1047,957,1503]" captionTargetId="figure@4.[153,1047,957,1503]" captionTargetPageId="4" captionText="Map 2: Confirmed distribution of Ischnosoma myops (black circles) and type locality of I. cuspidatum (white star)." figureDoi="http://doi.org/10.5281/zenodo.3738591" httpUri="https://zenodo.org/record/3738591/files/figure.png" pageId="4" pageNumber="777">Map 2</figureCitation>
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). The
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<typeStatus id="EF738839FFB19B18609BCAEAFD98F0B5" box="[498,587,897,919]" pageId="4" pageNumber="777" type="holotype">holotype</typeStatus>
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was sifted from leaf litter in a stream valley with deciduous forest at an altitude of
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<quantity id="F7309B7EFFB19B186375CAF5FD8FF096" box="[540,604,926,948]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="4" pageNumber="777" unit="m" value="830.0">830 m</quantity>
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(MEYBOHM pers. comm.).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |