293 lines
43 KiB
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293 lines
43 KiB
XML
<document id="2D51CC7207455720B16B48C319D33DDC" ID-CLB-Dataset="32179" ID-DOI="10.5281/zenodo.886217" ID-GBIF-Dataset="e87ff7fb-7b80-4e0c-bdf1-384a2ec7bd25" ID-Zenodo-Dep="886217" ID-ZooBank="urn:lsid:zoobank.org:pub:70097B8F-3EF2-4740-97D4-AD2357234011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1504701458027" checkinUser="plazi" docAuthor="Maurice Kottelat" docDate="2017" docId="B85F8784FFAA100DFBB1B179FB6E0778" docLanguage="en" docName="2017 Malihkaia.pdf" docOrigin="Raffles Bulletin Of Zoology 6" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Malihkaia Maurice Kottelat 2017, new genus" docType="treatment" docUuid="4F3A40F7-1C83-4FBC-A91E-292BF09CD49B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="84" masterDocId="4466FFFCFFAB1009FFB7B546FFB50F78" masterDocTitle="A new genus and three new species of nemacheilid loaches from northern Irrawaddy drainage, Myanmar (Teleostei: Cypriniformes)" masterLastPageNumber="99" masterPageNumber="80" pageNumber="81" updateTime="1698468881364" updateUser="ExternalLinkService">
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<mods:title id="C815DBECF157D256CDFAE7F51C4D245C">A new genus and three new species of nemacheilid loaches from northern Irrawaddy drainage, Myanmar (Teleostei: Cypriniformes)</mods:title>
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<mods:name id="52C95B50BBAAB4175091412EC3A4322A" type="personal">
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<mods:namePart id="A401B32C724A91CAEB8D52E572576BFA">Maurice Kottelat</mods:namePart>
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<mods:title id="45A15CE7B4212D4000F00EB76AB4205F">Raffles Bulletin Of Zoology</mods:title>
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<mods:part id="3EDBD207DF9DFC04BFF3104D64101354">
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<mods:date id="2102231B46AC86DB53880F95124E0D23">2017</mods:date>
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<mods:number id="426149867207F796B8658B5E4DC6BD54">2017-05-12</mods:number>
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<treatment id="B85F8784FFAA100DFBB1B179FB6E0778" ID-DOI="http://doi.org/10.5281/zenodo.5632151" ID-GBIF-Taxon="133186468" ID-Zenodo-Dep="5632151" LSID="urn:lsid:zoobank.org:act:4F3A40F7-1C83-4FBC-A91E-292BF09CD49B" httpUri="http://treatment.plazi.org/id/B85F8784FFAA100DFBB1B179FB6E0778" lastPageId="4" lastPageNumber="84" pageId="1" pageNumber="81">
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<subSubSection id="78EC6519FFAA1008FBB1B179FB4A0B20" box="[1030,1279,1087,1112]" pageId="1" pageNumber="81" type="nomenclature">
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<paragraph id="30493692FFAA1008FBB1B179FB4A0B20" blockId="1.[1030,1279,1087,1112]" box="[1030,1279,1087,1112]" pageId="1" pageNumber="81">
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<heading id="6B0181FEFFAA1008FBB1B179FB4A0B20" bold="true" box="[1030,1279,1087,1112]" centered="true" fontSize="10" level="3" pageId="1" pageNumber="81" reason="2">
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<emphasis id="0282EA80FFAA1008FBB1B179FB4A0B20" bold="true" box="[1030,1279,1087,1112]" pageId="1" pageNumber="81">
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<taxonomicName id="F7F64D11FFAA1008FBB1B179FBCD0B2F" authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[1030,1144,1087,1111]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="81" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="0282EA80FFAA1008FBB1B179FBCD0B2F" bold="true" box="[1030,1144,1087,1111]" italics="true" pageId="1" pageNumber="81">Malihkaia</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="19B157FBFFAA1008FB3FB106FB4A0B20" box="[1160,1279,1088,1112]" pageId="1" pageNumber="81" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="78EC6519FFAA1008FCF4B1C6FA0A0BE0" box="[835,1471,1151,1176]" pageId="1" pageNumber="81" type="materials_examined">
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<paragraph id="30493692FFAA1008FCF4B1C6FA0A0BE0" blockId="1.[835,1471,1151,1176]" box="[835,1471,1151,1176]" pageId="1" pageNumber="81">
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<emphasis id="0282EA80FFAA1008FCF4B1C6FC6E0BEF" bold="true" box="[835,987,1151,1176]" pageId="1" pageNumber="81">Type species.</emphasis>
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<taxonomicName id="F7F64D11FFAA1008FC53B139FB120BEF" authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[996,1191,1151,1175]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="81" phylum="Chordata" rank="species" species="aligera" status="sp. nov.">
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<emphasis id="0282EA80FFAA1008FC53B139FB120BEF" box="[996,1191,1151,1175]" italics="true" pageId="1" pageNumber="81">Malihkaia aligera</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="19B157FBFFAA1008FB01B1C6FA8D0BEF" box="[1206,1336,1151,1175]" pageId="1" pageNumber="81" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="A8CD2A17FFAA1008FAFEB139FA070BE0" box="[1353,1458,1151,1176]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStartId-0="2.[113,148,1530,1552]" captionStartId-1="3.[135,170,1977,1999]" captionTargetBox-0="[157,1430,218,1481]" captionTargetBox-1="[166,1447,297,1943]" captionTargetId-0="figure@2.[114,1452,189,1512]" captionTargetId-1="figure@3.[135,1473,278,1960]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Malihkaia aligera, MHNG 2766.051, holotype, male, 65.7 mm SL; Myanmar: Irrawaddy drainage; Mali Hka River near Putao." captionText-1="Fig. 2. Malihkaia aligera, CMK 25508, paratypes; a, male, 73.0 mm SL; b, female, 77.2 mm SL; Myanmar: Irrawaddy drainage: Mali Hka River near Putao." httpUri-0="https://zenodo.org/record/886223/files/figure.png" httpUri-1="https://zenodo.org/record/886225/files/figure.png" pageId="1" pageNumber="81">Figs. 1–2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="78EC6519FFAA100DFCF4B1F9FE830A17" lastPageId="4" lastPageNumber="84" pageId="1" pageNumber="81" type="diagnosis">
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<paragraph id="30493692FFAA1008FCF4B1F9FA760AEF" blockId="1.[835,1475,1215,1431]" pageId="1" pageNumber="81">
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<emphasis id="0282EA80FFAA1008FCF4B1F9FC0D0BAF" bold="true" box="[835,952,1215,1239]" pageId="1" pageNumber="81">Diagnosis.</emphasis>
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<taxonomicName id="F7F64D11FFAA1008FC08B1F9FB980BAF" authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[959,1069,1215,1239]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="81" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="0282EA80FFAA1008FC08B1F9FB980BAF" box="[959,1069,1215,1239]" italics="true" pageId="1" pageNumber="81">Malihkaia</emphasis>
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</taxonomicName>
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is distinguished from all other genera of nemacheilids by the unique morphology of the lips and the unique pectoral fn sexual dimorphism. Lips thick, with numerous, closely set, deep furrows (
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<figureCitation id="A8CD2A17FFAA1008FB66B059FAA70A40" box="[1233,1298,1311,1336]" captionStart="Fig. 3" captionStartId="4.[113,150,653,675]" captionTargetBox="[113,752,211,636]" captionTargetId="figure@4.[113,752,189,636]" captionTargetPageId="4" captionText="Fig. 3. Malihkaia aligera, CMK 25508, 73.0 mm SL; ventral view of head." httpUri="https://zenodo.org/record/886227/files/figure.png" pageId="1" pageNumber="81">Fig. 3</figureCitation>
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). Upper lip with a small median notch, with transverse furrows on its whole length, edge crenulated. Lower lip with a median interruption; with transverse furrows on its whole length, edge crenulated.
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</paragraph>
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<paragraph id="30493692FFAA1008FCF4B086FBE30800" blockId="1.[835,1475,1471,1912]" pageId="1" pageNumber="81">
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In the male, the pectoral fn is strongly falcate (
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<figureCitation id="A8CD2A17FFAA1008FAF5B0F9FA310AA0" box="[1346,1412,1471,1496]" captionStart="Fig. 2" captionStartId="3.[135,170,1977,1999]" captionTargetBox="[166,1447,297,1943]" captionTargetId="figure@3.[135,1473,278,1960]" captionTargetPageId="3" captionText="Fig. 2. Malihkaia aligera, CMK 25508, paratypes; a, male, 73.0 mm SL; b, female, 77.2 mm SL; Myanmar: Irrawaddy drainage: Mali Hka River near Putao." httpUri="https://zenodo.org/record/886225/files/figure.png" pageId="1" pageNumber="81">Fig. 2</figureCitation>
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). The unbranched ray is fexible and shaped as in other species of
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<taxonomicName id="F7F64D11FFAA1008FCD3B0B9FBBF096F" authorityName="Swainson" authorityYear="1839" box="[868,1034,1535,1559]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="81" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
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. The frst branched ray is rigid, arched and curled upwards (
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<figureCitation id="A8CD2A17FFAA1008FB9FB359FBDE0940" box="[1064,1131,1567,1592]" captionStart="Fig. 4" captionStartId="4.[813,848,865,887]" captionTargetBox="[813,1435,189,848]" captionTargetId="figure@4.[813,1452,189,848]" captionTargetPageId="4" captionText="Fig. 4. Malihkaia aligera, CMK 25508, 73.0 mm SL; left pectoral fn of male; dorsal (above) and ventral (below) view. Extremity of fourth branched ray damaged." httpUri="https://zenodo.org/record/886229/files/figure.png" pageId="1" pageNumber="81">Fig. 4</figureCitation>
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). It is about 5 times wider than the other branched rays. It is fattened dorso-ventrally and very elongate, reaching beyond pelvic-fn base. It is frst branched at about midlength. The posterior main branch is thicker than the anterior main branch; it is branched again twice, the third branching about at distal one-sixth of the ray. The anterior branch is branched again only once, in a position intermediate to that of branching points 2 and 3 of posterior main branch. There is no membrane between all the branches and sub-branches, but there is a membrane around the tip of the ray.
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</paragraph>
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<paragraph id="30493692FFAA100BFCF4B2D9FE1B09A7" blockId="1.[835,1474,1951,2104]" lastBlockId="2.[113,753,1607,1759]" lastPageId="2" lastPageNumber="82" pageId="1" pageNumber="81">The second and third branched rays are very slender, branched only at their distal third, the anterior branch is thicker than the posterior one and it is branched again once; the posterior branch has no sub-branches. The membrane between the branches is very narrow. The spaces and relative ‘depth’ of the branching increase regularly in the following rays. The fourth and following branched rays have two branches that are each branched once. There is some variation in the branching apparently related to size and small injuries; the above is based on intact fns.</paragraph>
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<caption id="6489661AFFA9100BFFC6B0BCFA2A0968" box="[113,1439,1530,1552]" httpUri="https://zenodo.org/record/886223/files/figure.png" pageId="2" pageNumber="82" targetBox="[157,1430,218,1481]" targetPageId="2">
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<paragraph id="30493692FFA9100BFFC6B0BCFA2A0968" blockId="2.[113,1439,1530,1552]" box="[113,1439,1530,1552]" pageId="2" pageNumber="82">
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Fig. 1.
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<taxonomicName id="F7F64D11FFA9100BFF0EB0BCFEDD0977" authorityName="Maurice Kottelat" authorityYear="2017" box="[185,360,1530,1551]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="82" phylum="Chordata" rank="species" species="aligera">
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<emphasis id="0282EA80FFA9100BFF0EB0BCFEDD0977" box="[185,360,1530,1551]" italics="true" pageId="2" pageNumber="82">Malihkaia aligera</emphasis>
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</taxonomicName>
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, MHNG 2766.051, holotype, male, 65.7 mm SL; Myanmar: Irrawaddy drainage; Mali Hka River near Putao.
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</paragraph>
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</caption>
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<paragraph id="30493692FFA9100BFFC6B241FEA80887" blockId="2.[113,753,1799,2047]" pageId="2" pageNumber="82">
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The lower surface of the unbranched ray and the unbranched part of frst branched ray and the membrane between the three frst rays are covered by an unculiferous pad (
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<bibRefCitation id="54674B63FFA9100BFD21B20EFF4008F8" author="Conway KW & Lujan NK & Lundberg JG & Mayden RL & Siegel DS" journalOrPublisher="Journal of Morphology" pageId="2" pageNumber="82" pagination="1127 - 1149" part="273" refString="Conway KW, Lujan NK, Lundberg JG, Mayden RL & Siegel DS (2012) Microanatomy of the paired-fn pads of ostariophysan fshes (Teleostei: Ostariophysi). Journal of Morphology, 273: 1127 - 1149." title="Microanatomy of the paired-fn pads of ostariophysan fshes (Teleostei: Ostariophysi)" type="journal article" year="2012">Conway et al., 2012</bibRefCitation>
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), thickest under the frst branched ray. There are also pads under the membranes between the other rays, but less developed. There also are granulations (unculi?) on the dorsal surface of the rays and thickened tissue on the membranes.
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</paragraph>
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<paragraph id="30493692FFA9100BFC9AB30EFA880767" blockId="2.[813,1453,1607,2079]" pageId="2" pageNumber="82">In females, the pectoral fn is rounded; there are unculiferous pads under the rays, thickest under the unbranched and frst branched ones, decreasing in extant and thickness posteriorly. The unbranched ray is thicker than the following rays and rigid; dorsally, the membrane between the frst two rays is thickened. The frst branched ray is thickened but fexible, round in cross-section, thicker than the following rays; the anterior branch is unbranched. The posterior branch is branched once, the anterior sub-branch is unbranched and the posterior one branched again. In the second branched ray, the anterior branch is unbranched and the posterior branch is branched again. The remaining rays are all branched twice as in other nemacheilids. There is some little variation in the branching, related with small injuries and regrowth; the above is based on one intact fn on each female.</paragraph>
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<caption id="6489661AFFA8100AFF30B2FFFED60894" httpUri="https://zenodo.org/record/886225/files/figure.png" pageId="3" pageNumber="83" targetBox="[166,1447,297,1943]" targetPageId="3">
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<paragraph id="30493692FFA8100AFF30B2FFFED60894" blockId="3.[135,1473,1977,2028]" pageId="3" pageNumber="83">
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Fig. 2.
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<taxonomicName id="F7F64D11FFA8100AFF67B2FFFE3508B6" authorityName="Maurice Kottelat" authorityYear="2017" box="[208,384,1977,1998]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="83" phylum="Chordata" rank="species" species="aligera">
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<emphasis id="0282EA80FFA8100AFF67B2FFFE3508B6" box="[208,384,1977,1998]" italics="true" pageId="3" pageNumber="83">Malihkaia aligera</emphasis>
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</taxonomicName>
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, CMK 25508, paratypes; a, male, 73.0 mm SL; b, female, 77.2 mm SL; Myanmar: Irrawaddy drainage: Mali Hka River near Putao.
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</paragraph>
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</caption>
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<caption id="6489661AFFAF100DFFC6B7CBFF4D0DB8" httpUri="https://zenodo.org/record/886227/files/figure.png" pageId="4" pageNumber="84" targetBox="[113,752,211,636]" targetPageId="4">
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<paragraph id="30493692FFAF100DFFC6B7CBFF4D0DB8" blockId="4.[113,751,653,704]" pageId="4" pageNumber="84">
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Fig. 3.
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<taxonomicName id="F7F64D11FFAF100DFF08B7CBFECD0DDA" authorityName="Maurice Kottelat" authorityYear="2017" box="[191,376,653,674]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="species" species="aligera">
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<emphasis id="0282EA80FFAF100DFF08B7CBFECD0DDA" box="[191,376,653,674]" italics="true" pageId="4" pageNumber="84">Malihkaia aligera</emphasis>
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</taxonomicName>
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, CMK 25508, 73.0 mm SL; ventral view of head.
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</paragraph>
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</caption>
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<paragraph id="30493692FFAF100DFFC6B7BEFD610CF7" blockId="4.[113,752,759,912]" pageId="4" pageNumber="84">In both sexes, in both pelvic and pectoral fns, the anterior edge of the unbranched ray has a thick tissue cover. In both sexes, the pelvic fn has thickened tissue on membranes between the unbranched ray and branched rays 1–3, dorsally and ventrally, also covering part of these rays ventrally.</paragraph>
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<caption id="6489661AFFAF100DFC9AB627FBE60CC9" httpUri="https://zenodo.org/record/886229/files/figure.png" pageId="4" pageNumber="84" targetBox="[813,1435,189,848]" targetPageId="4">
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<paragraph id="30493692FFAF100DFC9AB627FBE60CC9" blockId="4.[813,1451,865,945]" pageId="4" pageNumber="84">
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Fig. 4.
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<taxonomicName id="F7F64D11FFAF100DFCC5B627FB950C0E" authorityName="Maurice Kottelat" authorityYear="2017" box="[882,1056,865,886]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="species" species="aligera">
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<emphasis id="0282EA80FFAF100DFCC5B627FB950C0E" box="[882,1056,865,886]" italics="true" pageId="4" pageNumber="84">Malihkaia aligera</emphasis>
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</taxonomicName>
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, CMK 25508, 73.0 mm SL; left pectoral fn of male; dorsal (above) and ventral (below) view. Extremity of fourth branched ray damaged.
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</paragraph>
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</caption>
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<paragraph id="30493692FFAF100DFFC6B6F1FE830A17" blockId="4.[113,753,951,1392]" pageId="4" pageNumber="84">Additional characters useful to distinguish the genus are: processus dentiformis present; suborbital fap present in males; 8½ branched dorsal-fn rays; 9+8 branched caudal-fn rays; axillary pelvic lobe present; anus situated about 1.5–2 eye diameter in front of anal fn; body entirely scaled; lateral line complete; air bladder without posterior chamber; body with 9–12 bars extending from dorsal midline to level of pectoral fn, bars of quite regular width and shape in front of dorsal fin, less regular posteriorly, much wider than interspaces; and black mark at caudal-fn base made of a more or less squarish blotch in middle of base, a more or less triangular blotch over dorsal procurrent rays, fainter pigments over base of uppermost and lowermost 4–6 principal rays of caudal fn.</paragraph>
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</subSubSection>
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<subSubSection id="78EC6519FFAF100DFFC6B0D1FD8F0AB7" pageId="4" pageNumber="84" type="etymology">
|
||
<paragraph id="30493692FFAF100DFFC6B0D1FD8F0AB7" blockId="4.[113,752,1431,1487]" pageId="4" pageNumber="84">
|
||
<emphasis id="0282EA80FFAF100DFFC6B0D1FF420AD7" bold="true" box="[113,247,1431,1455]" pageId="4" pageNumber="84">Etymology.</emphasis>
|
||
Named for
|
||
<collectingCountry id="48E17602FFAF100DFE33B0D1FE0D0AD7" box="[388,440,1431,1455]" name="Serbia" pageId="4" pageNumber="84">Mali</collectingCountry>
|
||
Hka River, where the
|
||
<typeStatus id="EF4D8830FFAF100DFD76B0DEFD450AC8" box="[705,752,1432,1456]" pageId="4" pageNumber="84">type</typeStatus>
|
||
species was discovered. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78EC6519FFAF100DFFC6B0B1FED708C8" pageId="4" pageNumber="84" type="discussion">
|
||
<paragraph id="30493692FFAF100DFFC6B0B1FED708C8" blockId="4.[113,753,1527,1968]" pageId="4" pageNumber="84">
|
||
<emphasis id="0282EA80FFAF100DFFC6B0B1FF680977" bold="true" box="[113,221,1527,1551]" pageId="4" pageNumber="84">Remarks.</emphasis>
|
||
The deeply furrowed lower lip of
|
||
<taxonomicName id="F7F64D11FFAF100DFD81B0B1FD450977" authorityName="Maurice Kottelat" authorityYear="2017" box="[566,752,1527,1551]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="species" species="aligera">
|
||
<emphasis id="0282EA80FFAF100DFD81B0B1FD450977" box="[566,752,1527,1551]" italics="true" pageId="4" pageNumber="84">Malihkaia aligera</emphasis>
|
||
</taxonomicName>
|
||
is unique in South and Southeast Asian
|
||
<taxonomicName id="F7F64D11FFAF100DFDF3B351FD5B0957" authorityName="Swainson" authorityYear="1839" box="[580,750,1559,1583]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
.
|
||
<taxonomicName id="F7F64D11FFAF100DFFC6B371FEA40937" authorityName="Peters" authorityYear="1861" box="[113,273,1591,1615]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFFC6B371FEA40937" box="[113,273,1591,1615]" italics="true" pageId="4" pageNumber="84">Acanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<taxonomicName id="F7F64D11FFAF100DFE25B371FDE20937" authorityName="Grant" authorityYear="2007" box="[402,599,1591,1615]" class="Actinopterygii" family="Nemacheilidae" genus="Paracanthocobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFE25B371FDE20937" box="[402,599,1591,1615]" italics="true" pageId="4" pageNumber="84">Paracanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="54674B63FFAF100DFD23B371FF0C0917" author="Kottelat M" journalOrPublisher="Raffes Bulletin of Zoology, Supplement" pageId="4" pageNumber="84" pagination="1 - 199" part="26" refString="Kottelat M (2012 b) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidea). Raffes Bulletin of Zoology, Supplement 26: 1 - 199." title="Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidea)" type="journal article" year="2012" yearSuffix="b">Kottelat, 2012b</bibRefCitation>
|
||
: 74, 2013: 198) has papillated lips (
|
||
<bibRefCitation id="54674B63FFAF100DFDFBB311FD590917" author="Kottelat M" bookContentInfo="262 pp." box="[588,748,1623,1647]" journalOrPublisher="Pfeil, Munchen" pageId="4" pageNumber="84" refString="Kottelat M (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Vietnam. Pfeil, Munchen, 262 pp." title="Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Vietnam" type="book" year="1990">Kottelat, 1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="54674B63FFAF100DFFC6B331FF0209F7" author="Kottelat M" box="[113,183,1655,1679]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="84" pagination="45 - 52" part="3327" refString="Kottelat M (2012 a) Acanthocobitis pictilis, a new species of loach from Myanmar and Thailand (Teleostei: Nemacheilidae). Zootaxa, 3327: 45 - 52." title="Acanthocobitis pictilis, a new species of loach from Myanmar and Thailand (Teleostei: Nemacheilidae)" type="journal article" year="2012" yearSuffix="a">2012a</bibRefCitation>
|
||
). On the upper lip, the papillae are organised in several rows; on the lower lip there is a broad median interruption and, on each side, a widened, strongly papillated pad, covered by papillae. Species of
|
||
<taxonomicName id="F7F64D11FFAF100DFE35B391FD9D0997" authorityName="Peters" authorityYear="1861" box="[386,552,1751,1775]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFE35B391FD9D0997" box="[386,552,1751,1775]" italics="true" pageId="4" pageNumber="84">Acanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
also have a very different colour pattern, made of obliquely organised bars, usually of irregular shape and organisation. All species of
|
||
<taxonomicName id="F7F64D11FFAF100DFFC6B271FEA40837" authorityName="Peters" authorityYear="1861" box="[113,273,1847,1871]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFFC6B271FEA40837" box="[113,273,1847,1871]" italics="true" pageId="4" pageNumber="84">Acanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
have a small black spot (sometime ocellated) at the upper extremity of the base of the caudal fn. Finally, species of
|
||
<taxonomicName id="F7F64D11FFAF100DFF5AB231FE2608F7" authorityName="Peters" authorityYear="1861" box="[237,403,1911,1935]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFF5AB231FE2608F7" box="[237,403,1911,1935]" italics="true" pageId="4" pageNumber="84">Acanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
have a longer dorsal fn, with 9–18½ branched rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78EC6519FFAF100DFC9AB6AFFB6E0778" pageId="4" pageNumber="84" type="reference_group">
|
||
<paragraph id="30493692FFAF100DFC9AB6AFFB5A0A78" blockId="4.[813,1452,1000,1281]" pageId="4" pageNumber="84">
|
||
In
|
||
<taxonomicName id="F7F64D11FFAF100DFCFAB6AEFC440B78" ID-CoL="D7R" authorityName="Swainson" authorityYear="1839" box="[845,1009,1000,1024]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
, the deeply furrowed lips are otherwise known only in some species of
|
||
<taxonomicName id="F7F64D11FFAF100DFB3DB14EFAAE0B58" authorityName="Prokofiev" authorityYear="2010" box="[1162,1307,1032,1056]" class="Actinopterygii" family="Nemacheilidae" genus="Labiatophysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFB3DB14EFAAE0B58" box="[1162,1307,1032,1056]" italics="true" pageId="4" pageNumber="84">Labiatophysa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F7F64D11FFAF100DFA9DB14EFA1E0B58" ID-CoL="82D8" authorityName="Rendahl" authorityYear="1933" box="[1322,1451,1032,1056]" class="Actinopterygii" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFA9DB14EFA1E0B58" box="[1322,1451,1032,1056]" italics="true" pageId="4" pageNumber="84">Triplophysa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7F64D11FFAF100DFCEAB16EFC5E0B38" authorityName="Prokofiev" authorityYear="2010" box="[861,1003,1064,1088]" class="Actinopterygii" family="Nemacheilidae" genus="Tarimichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFCEAB16EFC5E0B38" box="[861,1003,1064,1088]" italics="true" pageId="4" pageNumber="84">Tarimichthys</emphasis>
|
||
</taxonomicName>
|
||
, all endemic to the Tibetan plateau, the Tarim and adjacent endorheic basins in
|
||
<collectingCountry id="48E17602FFAF100DFB57B10EFA970B18" box="[1248,1314,1096,1120]" name="China" pageId="4" pageNumber="84">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="48E17602FFAF100DFA98B10EFA1E0B18" box="[1327,1451,1096,1120]" name="Kazakhstan" pageId="4" pageNumber="84">Kazakhstan</collectingCountry>
|
||
and
|
||
<collectingCountry id="48E17602FFAF100DFCE8B12FFC560BF9" box="[863,995,1129,1153]" name="Kyrgyzstan" pageId="4" pageNumber="84">Kyrgyzstan</collectingCountry>
|
||
, and adjacent waters. These genera are distinguished, among others, by the absence of scales (most species), males with densely set unculi on the side of the head below the eye and on the dorsal surface of anterior pectoral-fn rays (
|
||
<bibRefCitation id="54674B63FFAF100DFC5AB1AEFB280A78" author="Prokofiev AM" box="[1005,1181,1256,1280]" journalOrPublisher="Senckenbergiana Biologica" pageId="4" pageNumber="84" pagination="181 - 208" part="83" refString="Prokofiev AM (2004) Revision of the species-complex of Triplophysa labiata with description of a new species, T. kaznakowi sp. n. (Osteichthyes, Balitoridae, Nemacheilinae). Senckenbergiana Biologica, 83 (2): 181 - 208." title="Revision of the species-complex of Triplophysa labiata with description of a new species, T. kaznakowi sp. n. (Osteichthyes, Balitoridae, Nemacheilinae)" type="journal article" year="2004">Prokofev, 2004</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="54674B63FFAF100DFB1DB1AEFB510A78" author="Prokofiev AM" box="[1194,1252,1256,1280]" journalOrPublisher="Journal of Ichthyology" pageId="4" pageNumber="84" pagination="827 - 913" part="50" refString="Prokofiev AM (2010) Morphological classification of loaches (Nemacheilinae). Journal of Ichthyology, 50 (10): 827 - 913." title="Morphological classification of loaches (Nemacheilinae)" type="journal article" year="2010">2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="30493692FFAF100DFC9AB06EFB6E0778" blockId="4.[813,1452,1320,2048]" pageId="4" pageNumber="84">
|
||
Cobitiforms have developed a great variety of sexual dimorphism, but most of it associated with the pectoral fns and involving thickening and modifcation of rays (see, e.g.,
|
||
<bibRefCitation id="54674B63FFAF100DFC9AB0C0FBA10AD8" author="Slechtova V & Bohlen J & Perdices A" box="[813,1044,1414,1440]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="4" pageNumber="84" pagination="812 - 831" part="47" refString="Slechtova V, Bohlen J & Perdices A (2008) Molecular phylogeny of the freshwater fish family Cobitidae (Cypriniformes: Teleostei): delimitation of genera, mitochondrial introgression and evolution of sexual dimorphism. Molecular Phylogenetics and Evolution, 47: 812 - 831." title="Molecular phylogeny of the freshwater fish family Cobitidae (Cypriniformes: Teleostei): delimitation of genera, mitochondrial introgression and evolution of sexual dimorphism" type="journal article" year="2008">Šlechtová et al., 2008</bibRefCitation>
|
||
for sexual dimorphism in
|
||
<taxonomicName id="F7F64D11FFAF100DFA86B0CEFACB0AD8" ID-CoL="3RJL" box="[1329,1406,1416,1440]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFA86B0CEFACB0AD8" box="[1329,1406,1416,1440]" italics="true" pageId="4" pageNumber="84">Cobitis</emphasis>
|
||
</taxonomicName>
|
||
and related genera;
|
||
<bibRefCitation id="54674B63FFAF100DFC7EB0EEFB130AB8" author="Kottelat M & Lim KKP" box="[969,1190,1448,1472]" journalOrPublisher="Raffes Bulletin of Zoology" pageId="4" pageNumber="84" pagination="201 - 220" part="40" refString="Kottelat M & Lim KKP (1992) A synopsis of the Malayan species of Lepidocephalichthys, with descriptions of two new species (Teleostei: Cobitidae). Raffes Bulletin of Zoology, 40 (2): 201 - 220." title="A synopsis of the Malayan species of Lepidocephalichthys, with descriptions of two new species (Teleostei: Cobitidae)" type="journal article" year="1992">Kottelat & Lim, 1992</bibRefCitation>
|
||
for
|
||
<taxonomicName id="F7F64D11FFAF100DFB79B0EEFA110AB8" ID-CoL="62WWB" authorityName="Bleeker" authorityYear="1863" box="[1230,1444,1448,1472]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFB79B0EEFA110AB8" box="[1230,1444,1448,1472]" italics="true" pageId="4" pageNumber="84">Lepidocephalichthys</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="54674B63FFAF100DFC9AB08EFB910A98" author="Kottelat M & Tan HH" box="[813,1060,1480,1504]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="84" pagination="63 - 72" part="1967" refString="Kottelat M & Tan HH (2008) Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae). Zootaxa, 1967: 63 - 72." title="Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae)" type="journal article" year="2008">Kottelat & Tan, 2008</bibRefCitation>
|
||
for
|
||
<taxonomicName id="F7F64D11FFAF100DFBEDB08EFB580A98" ID-CoL="592C" authorityName="Nalbant" authorityYear="1994" box="[1114,1261,1480,1504]" class="Actinopterygii" family="Cobitidae" genus="Kottelatlimia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFBEDB08EFB580A98" box="[1114,1261,1480,1504]" italics="true" pageId="4" pageNumber="84">Kottelatlimia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7F64D11FFAF100DFA9DB08EFA110AA7" ID-CoL="MG8" box="[1322,1444,1480,1503]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFA9DB08EFA110AA7" box="[1322,1444,1480,1503]" italics="true" pageId="4" pageNumber="84">Acantopsis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="54674B63FFAF100DFC9AB0AEFC7E0978" author="Kottelat M" bookContentInfo="262 pp." box="[813,971,1512,1536]" journalOrPublisher="Pfeil, Munchen" pageId="4" pageNumber="84" refString="Kottelat M (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Vietnam. Pfeil, Munchen, 262 pp." title="Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Vietnam" type="book" year="1990">Kottelat, 1990</bibRefCitation>
|
||
for Indochinese
|
||
<taxonomicName id="F7F64D11FFAF100DFB3CB0AEFA840978" ID-CoL="D7R" authorityName="Swainson" authorityYear="1839" box="[1163,1329,1512,1536]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
;
|
||
<bibRefCitation id="54674B63FFAF100DFA88B0AEFC6C0958" author="Bohlen J & Slechtova V" journalOrPublisher="Ichthyological Exploration of Freshwaters 22" pageId="4" pageNumber="84" pagination="1 - 10" part="1" refString="Bohlen J & Slechtova V (2011) A new genus and two new species of loaches (Teleostei: Nemacheilidae) from Myanmar. Ichthyological Exploration of Freshwaters 22 (1): 1 - 10." title="A new genus and two new species of loaches (Teleostei: Nemacheilidae) from Myanmar" type="journal article" year="2011">Bohlen & Šlechtová, 2011</bibRefCitation>
|
||
for
|
||
<taxonomicName id="F7F64D11FFAF100DFBB0B34EFB7D0958" ID-CoL="74SL" box="[1031,1224,1544,1568]" class="Actinopterygii" family="Nemacheilidae" genus="Pteronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFBB0B34EFB7D0958" box="[1031,1224,1544,1568]" italics="true" pageId="4" pageNumber="84">Pteronemacheilus</emphasis>
|
||
</taxonomicName>
|
||
); sexual dimorphism may also involve, for example, thickening of parts of the body (e.g., in
|
||
<taxonomicName id="F7F64D11FFAF100DFC7BB30EFA360918" ID-CoL="7BCX" authority=", Kottelat & Freyhof, 2007" authorityName="Kottelat & Freyhof" authorityYear="2007" box="[972,1411,1608,1632]" class="Actinopterygii" family="Cobitidae" genus="Sabanejewia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFC7BB30EFBE00918" box="[972,1109,1608,1632]" italics="true" pageId="4" pageNumber="84">Sabanejewia</emphasis>
|
||
,
|
||
<bibRefCitation id="54674B63FFAF100DFBD1B30EFA360918" author="Kottelat M & Freyhof J" bookContentInfo="646 pp." box="[1126,1411,1608,1632]" journalOrPublisher="Cornol & Freyhof, Berlin, xiv" pageId="4" pageNumber="84" refString="Kottelat M & Freyhof J (2007) Handbook of European freshwater fshes. Kottelat, Cornol & Freyhof, Berlin, xiv + 646 pp." title="Handbook of European freshwater fshes. Kottelat" type="book" year="2007">Kottelat & Freyhof, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
) or vertical torsion of the caudal peduncle (in
|
||
<taxonomicName id="F7F64D11FFAF100DFB53B32EFAD909F8" ID-CoL="5RMV" authorityName="Bohlen & Harant" authorityYear="2011" box="[1252,1388,1640,1664]" class="Actinopterygii" family="Cobitidae" genus="Microcobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFB53B32EFAD909F8" box="[1252,1388,1640,1664]" italics="true" pageId="4" pageNumber="84">Microcobitis</emphasis>
|
||
</taxonomicName>
|
||
; pers. obs., unpubl.). The modifed pectoral fn of male
|
||
<taxonomicName id="F7F64D11FFAF100DFA8CB3CFFA1E09D8" authorityName="Maurice Kottelat" authorityYear="2017" box="[1339,1451,1672,1696]" class="Actinopterygii" family="Psilorhynchidae" genus="Malihkaia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="species" species="aligera">
|
||
<emphasis id="0282EA80FFAF100DFA8CB3CFFA1E09D8" box="[1339,1451,1672,1696]" italics="true" pageId="4" pageNumber="84">M. aligera</emphasis>
|
||
</taxonomicName>
|
||
seems unique in
|
||
<taxonomicName id="F7F64D11FFAF100DFC50B3EEFB3209B8" ID-CoL="D7R" authorityName="Swainson" authorityYear="1839" box="[999,1159,1704,1728]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
but is very reminiscent of what is observed in some
|
||
<taxonomicName id="F7F64D11FFAF100DFBE6B38EFB0E0998" ID-CoL="8D7" box="[1105,1211,1736,1760]" class="Actinopterygii" family="Cobitidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Cobitidae</taxonomicName>
|
||
, especially species of
|
||
<taxonomicName id="F7F64D11FFAF100DFC9AB3AEFC170987" ID-CoL="MG8" box="[813,930,1768,1791]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFC9AB3AEFC170987" box="[813,930,1768,1791]" italics="true" pageId="4" pageNumber="84">Acantopsis</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="F7F64D11FFAF100DFC7DB3AEFB8A0987" ID-CoL="MG8" box="[970,1087,1768,1791]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFC7DB3AEFB8A0987" box="[970,1087,1768,1791]" italics="true" pageId="4" pageNumber="84">Acantopsis</emphasis>
|
||
</taxonomicName>
|
||
, the branches of the frst branched ray are in contact and there is no membrane between them and the ray is covered by a dense layer of unculi (pers. obs.;
|
||
<bibRefCitation id="54674B63FFAF100DFC9AB20EFBBB0818" author="Kottelat M & Tan HH" box="[813,1038,1864,1888]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="84" pagination="63 - 72" part="1967" refString="Kottelat M & Tan HH (2008) Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae). Zootaxa, 1967: 63 - 72." title="Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae)" type="journal article" year="2008">Kottelat & Tan, 2008</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="F7F64D11FFAF100DFB8DB20EFB700818" ID-CoL="592C" authorityName="Nalbant" authorityYear="1994" box="[1082,1221,1864,1888]" class="Actinopterygii" family="Cobitidae" genus="Kottelatlimia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFB8DB20EFB700818" box="[1082,1221,1864,1888]" italics="true" pageId="4" pageNumber="84">Kottelatlimia</emphasis>
|
||
</taxonomicName>
|
||
, the frst branched ray is branched only at the tip and may appear as an unbranched ray (
|
||
<bibRefCitation id="54674B63FFAF100DFCE8B2CEFBE508D8" author="Kottelat M & Tan HH" box="[863,1104,1928,1952]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="84" pagination="63 - 72" part="1967" refString="Kottelat M & Tan HH (2008) Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae). Zootaxa, 1967: 63 - 72." title="Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae)" type="journal article" year="2008">Kottelat & Tan, 2008</bibRefCitation>
|
||
). A curled pectoral fn in males is observed in
|
||
<taxonomicName id="F7F64D11FFAF100DFC62B2EEFB8A08B8" ID-CoL="8D7" box="[981,1087,1960,1984]" class="Actinopterygii" family="Cobitidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Cobitidae</taxonomicName>
|
||
in
|
||
<taxonomicName id="F7F64D11FFAF100DFBDFB2EEFB6A08C7" ID-CoL="MG8" box="[1128,1247,1960,1983]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFBDFB2EEFB6A08C7" box="[1128,1247,1960,1983]" italics="true" pageId="4" pageNumber="84">Acantopsis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F7F64D11FFAF100DFAACB2EFFADC08B8" ID-CoL="6D7R" authorityName="Blyth" authorityYear="1860" box="[1307,1385,1961,1984]" class="Actinopterygii" family="Cobitidae" genus="Pangio" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="0282EA80FFAF100DFAACB2EFFADC08B8" box="[1307,1385,1961,1984]" italics="true" pageId="4" pageNumber="84">Pangio</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="54674B63FFAF100DFC9AB28EFBAB0898" author="Kottelat M & Tan HH" box="[813,1054,1992,2016]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="84" pagination="63 - 72" part="1967" refString="Kottelat M & Tan HH (2008) Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae). Zootaxa, 1967: 63 - 72." title="Kottelatlimia hipporhynchos, a new species of loach from southern Borneo (Teleostei: Cobitidae)" type="journal article" year="2008">Kottelat & Tan, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="54674B63FFAF100DFB9BB28EFAAA0898" author="Kottelat M & Lim KKP" box="[1068,1311,1992,2016]" journalOrPublisher="Raffes Bulletin of Zoology" pageId="4" pageNumber="84" pagination="203 - 249" part="41" refString="Kottelat M & Lim KKP (1993) A review of the eel-loaches of the genus Pangio (Teleostei: Cobitidae) from the Malay peninsula, with descriptions of six new species. Raffes Bulletin of Zoology, 41 (2): 203 - 249." title="A review of the eel-loaches of the genus Pangio (Teleostei: Cobitidae) from the Malay peninsula, with descriptions of six new species" type="journal article" year="1993">Kottelat & Lim, 1993</bibRefCitation>
|
||
) but has not been reported in
|
||
<taxonomicName id="F7F64D11FFAF100DFC50B2AEFB320778" ID-CoL="D7R" authorityName="Swainson" authorityYear="1839" box="[999,1159,2024,2048]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="84" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
before.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |