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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.820.31527" ID-GBIF-Dataset="19e4c0b1-d8aa-4ff8-a8e0-d639580dc280" ID-PMC="PMC6365488" ID-Pensoft-Pub="1313-2970-820-139" ID-PubMed="30740017" ID-ZBK="83F75B4D6E544E2B94B86525010CF725" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-820-139" ModsDocOrigin="ZooKeys 820" ModsDocTitle="Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)" checkinTime="1548887832952" checkinUser="pensoft" docAuthor="Yoshitomi, Hiroyuki &amp; Sogoh, Koichi" docDate="2019" docId="B5B65A52EF22DC3B15FD4681299518AF" docLanguage="en" docName="ZooKeys 820: 139-149" docOrigin="ZooKeys 820" docSource="http://dx.doi.org/10.3897/zookeys.820.31527" docTitle="Eumorphus mirabilis Yoshitomi &amp; Sogoh, 2019, sp. n." docType="treatment" docUuid="82BE8F98-3293-4D4D-966C-5DA213E52B60" docUuidSource="ZooBank" docVersion="5" lastPageNumber="147" masterDocId="ED1CFF82FF84FF97FF8CEE45FFCAFF91" masterDocTitle="Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)" masterLastPageNumber="149" masterPageNumber="139" pageNumber="144" updateTime="1668166862078" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sogoh, Koichi</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>820</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>149</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.820.31527</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.820.31527</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-820-139</mods:identifier>
<mods:identifier type="ZBK">83F75B4D6E544E2B94B86525010CF725</mods:identifier>
<mods:identifier type="ZooBank">83F75B4D6E544E2B94B86525010CF725</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154126362" LSID="urn:lsid:zoobank.org:act:82BE8F98-3293-4D4D-966C-5DA213E52B60" httpUri="http://treatment.plazi.org/id/B5B65A52EF22DC3B15FD4681299518AF" lastPageId="8" lastPageNumber="147" pageId="5" pageNumber="144">
<subSubSection pageId="5" pageNumber="144" type="nomenclature">
<paragraph pageId="5" pageNumber="144">
<taxonomicName LSID="http://zoobank.org/82BE8F98-3293-4D4D-966C-5DA213E52B60" class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus mirabilis" order="Coleoptera" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="mirabilis">Eumorphus mirabilis</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="144">sp. n.</taxonomicNameLabel>
Figs 2A, B; 3D, E; 4D, E; 5D, E; 6
<normalizedToken originalValue="IL">I-L</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="materials_examined">
<paragraph pageId="5" pageNumber="144">Material examined.</paragraph>
<paragraph pageId="5" pageNumber="144">Holotype, male (EUMJ), &quot;(C. SULAWESI) Puncak Dingin, alt. ca. 1700 m, 14.XI.1985, S. Nagai leg.&quot; Paratypes, female (EUMJ), same locality and collector, but 19.X.1985; 1 male and 1 female (NSMT), &quot;Puncak Palopo, C. Celebes, IV.1989&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="diagnosis">
<paragraph pageId="5" pageNumber="144">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="144">
This species can be easily separated from all other
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="5" pageNumber="144" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
species in having cordiform elytra possessing one pair of spots and the pronotum laterally serrated.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="147" pageId="5" pageNumber="144" type="description">
<paragraph pageId="5" pageNumber="144">Description.</paragraph>
<paragraph pageId="5" pageNumber="144">Male.</paragraph>
<paragraph pageId="5" pageNumber="144">Coloration of body black, with blueish luster; elytral spots reddish orange.</paragraph>
<paragraph pageId="5" pageNumber="144">Antennae long, reaching ca. basal 1/4 of elytra; antennal club wide.</paragraph>
<paragraph lastPageId="6" lastPageNumber="145" pageId="5" pageNumber="144">
Pronotum (Fig. 4D) glossy, widely depressed in lateral parts; lateral margins irregularly serrate; lateral sulci relatively long, reaching ca. basal 1/3 of PLM; anterior angles nearly right-angled; posterior angles sharply pointed, projecting postero-ventrally; PLM/PLS 0.82-0.87 (0.84); PWP/PWA 1.84-1.99 (1.91); PWP/PLM 1.86-1.93 (1.90); PWP/PLS 1.53-1.68 (1.60). Elytra (Fig. 2A) cordiform, widest at anterior 1/4; lateral flattened margins wide, undulate vertically; elytral spots one pair, present in anterior 1/4, relatively large and oval, each distant from suture by half of its diameter; elytra along suture (Fig. 3D) distinctly elevated in basal 1/3; EL/EW 0.86-1.07 (0.97); EL/PLM 4.02-4.41 (4.21); EW/PWP 2.21-2.42 (2.31); TL/EW 1.07-1.31 (1.19). Fore tibiae (Fig. 5D) slender, slightly curved laterally in apical parts; apical carinae weakly projecting dorsally, covered with yellowish brown pubescence in dorsal and
<pageBreakToken pageId="6" pageNumber="145" start="start">inner</pageBreakToken>
parts; fore tibial tooth short, situated in apical 1/4. Metatibial apices shortly prolonged in form of bifid projection; with outer angles shorter than inner ones.
</paragraph>
<caption pageId="6" pageNumber="145">
<paragraph pageId="6" pageNumber="145">
Figure 2. Habitus of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus mirabilis" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="mirabilis">Eumorphus mirabilis</taxonomicName>
sp. n. A Holotype, male B paratype, female. Scale bar: 5.0 mm.
</paragraph>
</caption>
<caption pageId="6" pageNumber="145">
<paragraph pageId="6" pageNumber="145">
Figure 3. Elytra of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
spp. in dorso-lateral view. A
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, male B
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, female C
<taxonomicName lsidName="E. wegneri" pageId="6" pageNumber="145" rank="species" species="wegneri">E. wegneri</taxonomicName>
, male D
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., male E
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., female. Scale bar: 5.0 mm.
</paragraph>
</caption>
<caption pageId="6" pageNumber="145">
<paragraph pageId="6" pageNumber="145">
Figure 4. Head and pronotum of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
spp. A
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, male B
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, female C
<taxonomicName lsidName="E. wegneri" pageId="6" pageNumber="145" rank="species" species="wegneri">E. wegneri</taxonomicName>
, male D
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., male E
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., female. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption pageId="6" pageNumber="145">
<paragraph pageId="6" pageNumber="145">
Figure 5. Fore tibia of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
spp. in dorsal view. A
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, male B
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
, female C
<taxonomicName lsidName="E. wegneri" pageId="6" pageNumber="145" rank="species" species="wegneri">E. wegneri</taxonomicName>
, male D
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., male E
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n., female. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="145">
Aedeagus (Fig. 6
<normalizedToken originalValue="IL">I-L</normalizedToken>
) 5.4 mm long, relatively slender; median lobe gently curved ventrally; apical branch short, trapezoidal in apical part, with small projection in dorsal part of apical branch; subapical branch slender and long, evenly curved antero-laterally; tegmen movable, with relatively long tegminal strut; endophallus consisting of bifid long right membranous lobe and long left membranous lobe.
</paragraph>
<caption pageId="6" pageNumber="145">
<paragraph pageId="6" pageNumber="145">
Figure 6. Aedeagus of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
spp.
<normalizedToken originalValue="AD">A-D</normalizedToken>
<taxonomicName lsidName="E. costatus" pageId="6" pageNumber="145" rank="species" species="costatus">E. costatus</taxonomicName>
<normalizedToken originalValue="EH">E-H</normalizedToken>
<taxonomicName lsidName="E. wegneri" pageId="6" pageNumber="145" rank="species" species="wegneri">E. wegneri</taxonomicName>
<normalizedToken originalValue="IL">I-L</normalizedToken>
<taxonomicName lsidName="E. mirabilis" pageId="6" pageNumber="145" rank="species" species="mirabilis">E. mirabilis</taxonomicName>
sp. n. A, E, I ventral B, F, J lateral C, G, K dorsal D, H, I apical. Abbreviations: ab: apical branch ed: endophallus sb: subapical branch ts: tegminal strut. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146">
<pageBreakToken pageId="7" pageNumber="146" start="start">Female</pageBreakToken>
. Similar to male, but sexual dimorphism distinct in the following characteristics: posterior angles of pronotum (Fig. 4E) ca. 50°; fore tibiae (Fig. 5E) slender, slightly curved laterally in apical parts, covered with yellowish brown pubescence in apical half; metatibial apices simple; PLM/PLS 0.77-0.78 (0.78); PWP/PWA 1.60-1.68 (1.64); PWP/PLM 1.78-1.91 (1.85); PWP/PLS 1.37-1.50 (1.44); EL/EW
<pageBreakToken pageId="8" pageNumber="147" start="start">0.99</pageBreakToken>
-1.07 (1.03); EL/PLM 4.45-4.46 (4.45); EW/PWP 2.33-2.35 (2.34); TL/EW 1.21-1.32 (1.26).
</paragraph>
<paragraph pageId="8" pageNumber="147">Measurements. Male (n = 2). TL 16.55-18.65 (17.60) mm; PW 6.37-6.43 (6.40) mm; PW 3.20-3.50 (3.35) mm; PL 3.30-3.45 (3.38) mm; PL 3.80-4.20 (4.00) mm; EL 13.25-15.20 (14.23) mm; EW 14.20-15.40 (14.80) mm. Female (n = 2). TL 16.23-16.38 (16.31) mm; PW 5.35-5.70 (5.53) mm; PW 3.35-3.40 (3.38) mm; PL 2.98-3.00 (2.99) mm; PL 3.80-3.90 (3.85) mm; EL 13.25-13.38 (13.32) mm; EW 12.45-13.40 (12.93) mm.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="147" type="distribution">
<paragraph pageId="8" pageNumber="147">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="147">Indonesia (Sulawesi: Central).</paragraph>
</subSubSection>
</treatment>
</document>