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<document id="2EAA45C79B57E601039C48A5661FBB1C" ID="10.11646/zootaxa.4140.1.1" ID-DOI="10.11646/zootaxa.4140.1.1" ID-GBIF-Dataset="01995ca4-7d04-4402-b21a-926957b1a391" ID-ISSN="1175-5326" ID-Zenodo-Dep="256507" ID-ZooBank="AD1F6B4E-3B0D-422C-B47C-0495BD385857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469426945103" checkinUser="plazi" docAuthor="Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. &amp; Blakemore, Robert J." docDate="2016" docId="B5727931FFAD791AFF3C221767248DEF" docLanguage="en" docName="zootaxa.4140.1.1.pdf" docOrigin="Zootaxa 4140 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Amynthas ghilarovi" docType="treatment" docVersion="22" lastPageNumber="30" masterDocId="494B0149FFB17907FFAB254162078C42" masterDocTitle="A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="29" updateTime="1698365898991" updateUser="plazi">
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<mods:title id="0A9B20C59E413DCA008C1183989F16E2">A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)</mods:title>
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<mods:namePart id="244E62E7DFB6D9FFFFD3BE3CDBE63E3D">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="CEF07971A4B6A9D8BE7900E3EC5E8E35">Nguyen, Anh D.</mods:namePart>
</mods:name>
<mods:name id="B34D64D4FA921E5FDDE3190F596E54F8" type="personal">
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<mods:roleTerm id="C58D6D1A6FA2054A5B5D9B0592DE79F4">Author</mods:roleTerm>
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<mods:namePart id="11BFF7F0BE099334D847A6156E358E9A">Tran, Binh T. T.</mods:namePart>
</mods:name>
<mods:name id="95FFC1B4E315844FBED0E460244A9E08" type="personal">
<mods:role id="21F0AD7CE25F2EBDEF7860139145CF23">
<mods:roleTerm id="CD092EB7D9AE18DAAE5FD4E07DAAE7AE">Author</mods:roleTerm>
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<mods:namePart id="2CC8FE0CD32783C980609678FF1A3B20">Blakemore, Robert J.</mods:namePart>
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<mods:title id="585F5A7BAF483A657E55F6D009E4DEBD">Zootaxa</mods:title>
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<mods:date id="CE7DAB74948609527F8D7578300CA60E">2016</mods:date>
<mods:detail id="8E14CFEE740978BD2F739DE38705CEB0" type="volume">
<mods:number id="C99A0EF3DB6DD931C044B912B73C480B">4140</mods:number>
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<mods:identifier id="280B1FB37AAC5ECD0E7332497B3E7EF5" type="DOI">10.11646/zootaxa.4140.1.1</mods:identifier>
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<mods:identifier id="4B1E294770705A8FDFA35C31CC3F5F56" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B5727931FFAD791AFF3C221767248DEF" ID-DOI="http://doi.org/10.5281/zenodo.5467142" ID-GBIF-Taxon="123207248" ID-Zenodo-Dep="5467142" LSID="urn:lsid:plazi:treatment:B5727931FFAD791AFF3C221767248DEF" httpUri="http://treatment.plazi.org/id/B5727931FFAD791AFF3C221767248DEF" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="75C19BACFFAD791BFF3C221760158B2D" box="[151,530,1878,1903]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="3D64C827FFAD791BFF3C221760158B2D" blockId="28.[151,530,1878,1903]" box="[151,530,1878,1903]" pageId="28" pageNumber="29">
<heading id="662C7F4BFFAD791BFF3C221760158B2D" bold="true" box="[151,530,1878,1903]" fontSize="10" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="FADBB3A4FFAD791BFF3C221760158B2D" authority="(Thai, 1982)" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[151,530,1878,1903]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="ghilarovi">
<emphasis id="0FAF1435FFAD791BFF3C221760158B2D" bold="true" box="[151,530,1878,1903]" pageId="28" pageNumber="29">
<emphasis id="0FAF1435FFAD791BFF3C2217637C8B2D" bold="true" box="[151,379,1878,1903]" italics="true" pageId="28" pageNumber="29">Amynthas ghilarovi</emphasis>
(
<bibRefCitation id="594AB5D6FFAD791BFE212217600D8B2D" author="Thai" box="[394,522,1878,1903]" pageId="28" pageNumber="29" refString="Thai, T. B. (1982) The fauna of earthworms f the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Jurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai, 1982</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFAD791BFF3C22DF61108BB0" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="3D64C827FFAD791BFF3C22DF60928B91" blockId="28.[151,1146,1950,2034]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="1DCBEF09FFAD791BFF3C22DF60928B91" pageId="28" pageNumber="29">
<taxonomicName id="FADBB3A4FFAD791BFF3C22DF600C8BF6" authority="Thai, 1982: 826" authorityName="Thai" authorityPageNumber="826" authorityYear="1982" box="[151,523,1950,1972]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="28" pageNumber="29" phylum="Annelida" rank="species" species="ghilarovi">
<emphasis id="0FAF1435FFAD791BFF3C22DF63658BF6" box="[151,354,1950,1972]" italics="true" pageId="28" pageNumber="29">Pheretima ghilarovi</emphasis>
<treatmentCitation id="BC7AEE36FFAD791BFEC322DF600C8BF6" author="Thai" box="[360,523,1950,1972]" page="826" pageId="28" pageNumber="29" year="1982">
<bibRefCitation id="594AB5D6FFAD791BFEC322DF63DE8BF6" author="Thai" box="[360,473,1950,1972]" pageId="28" pageNumber="29" refString="Thai, T. B. (1982) The fauna of earthworms f the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Jurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai, 1982</bibRefCitation>
: 826
</treatmentCitation>
</taxonomicName>
, fig. 2D; Do 1994: 77;
<treatmentCitation id="BC7AEE36FFAD791BFD5D22DF61C58BF6" author="Nakamura" box="[758,962,1950,1972]" page="63" pageId="28" pageNumber="29" year="1999">
<bibRefCitation id="594AB5D6FFAD791BFD5D22DF619A8BF6" author="Nakamura" box="[758,925,1950,1972]" pageId="28" pageNumber="29" refString="Nakamura, Y. (1999) Checklist of earthworms of Pheretima genus group (Megascolecidae: Oligochaeta) of the world. Edaphologia, 64, 1 - 78." type="journal article" year="1999">Nakamura 1999</bibRefCitation>
: 63
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFAD791BFC6622DF66718BF6" author="Thai" box="[973,1142,1950,1972]" page="309" pageId="28" pageNumber="29" year="2000">
<bibRefCitation id="594AB5D6FFAD791BFC6622DF66448BF6" author="Thai" box="[973,1091,1950,1972]" pageId="28" pageNumber="29" refString="Thai, T. B. (2000 a) Species diversity of earthworms in Vietnam. Proceedings of the national workshop on the basic issues in life science. Hanoi Science and Technics Publishing House, 307 - 311." type="proceedings paper" year="2000" yearSuffix="a">Thai 2000a</bibRefCitation>
: 309
</treatmentCitation>
.
<emphasis id="0FAF1435FFAD791BFF3C22FC63F48B91" box="[151,499,1981,2003]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="FADBB3A4FFAD791BFF3C22FC637D8B91" authority="(Amynthas)" baseAuthorityName="Amynthas" box="[151,378,1981,2003]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">Pheretima (Amynthas)</taxonomicName>
ghilarovi—
</emphasis>
<treatmentCitation id="BC7AEE36FFAD791BFE5822FC60968B91" author="Thai" box="[499,657,1981,2003]" page="125" pageId="28" pageNumber="29" year="1983">
<bibRefCitation id="594AB5D6FFAD791BFE5822FC60598B91" author="Thai" box="[499,606,1981,2003]" pageId="28" pageNumber="29" refString="Thai, T. B. (1983) Earthworms of Vietnam (Systematic, fauna, distribution and zoogeographic). PhD dissertation in Zooglogy. Lomonosov Moscow State University, Russia [in Russian]" type="book" year="1983">Thai 1983</bibRefCitation>
: 125
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3D64C827FFAD791BFF3C229D61108BB0" blockId="28.[151,1146,1950,2034]" box="[151,791,2012,2034]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="1DCBEF09FFAD791BFF3C229D61108BB0" box="[151,791,2012,2034]" pageId="28" pageNumber="29">
<emphasis id="0FAF1435FFAD791BFF3C229D63748BB0" box="[151,371,2012,2034]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="FADBB3A4FFAD791BFF3C229D62FC8BB0" box="[151,251,2012,2034]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="28" pageNumber="29" phylum="Annelida" rank="genus">Amynthas</taxonomicName>
ghilarovi—
</emphasis>
<treatmentCitation id="BC7AEE36FFAD791BFED8229D60488BB0" author="Blakemore" box="[371,591,2012,2034]" page="41" pageId="28" pageNumber="29" year="2007">
<bibRefCitation id="594AB5D6FFAD791BFED8229D602D8BB0" author="Blakemore" box="[371,554,2012,2034]" pageId="28" pageNumber="29" refString="Blakemore, R. J. (2007 a) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. www. annelida. net / earthworm / Pheretimoids. pdf (accessed 6 Jun 2016)" type="journal article" year="2007" yearSuffix="a">Blakemore 2007a</bibRefCitation>
: 41
</treatmentCitation>
;
<bibRefCitation id="594AB5D6FFAD791BFDF1229D61158BB0" author="Blakemore" box="[602,786,2012,2034]" pageId="28" pageNumber="29" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">Blakemore 2008b</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFAC791AFF3C25D661C58D5E" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="3D64C827FFAC791AFF3C25D660CD8CF2" blockId="29.[151,1436,151,429]" box="[151,714,151,176]" pageId="29" pageNumber="30">
<materialsCitation id="8DB3C27AFFAC791AFF3C25D660CD8CF2" ID-GBIF-Occurrence="1288043623" box="[151,714,151,176]" country="Vietnam" location="Cuc Phuong" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Ninh Binh" typeStatus="holotype">
<emphasis id="0FAF1435FFAC791AFF3C25D663328CF2" bold="true" box="[151,309,151,176]" pageId="29" pageNumber="30">
<typeStatus id="E2607685FFAC791AFF3C25D662D58CF2" box="[151,210,151,176]" pageId="29" pageNumber="30">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="45CC88B7FFAC791AFE9725D6639C8CF2" box="[316,411,151,176]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
(
<collectingRegion id="FF1F06C5FFAC791AFE0225D660228CF2" box="[425,549,151,176]" country="Vietnam" name="Ninh Binh" pageId="29" pageNumber="30">Ninh Binh</collectingRegion>
:
<location id="38049EFCFFAC791AFD8425D960B98CF2" LSID="urn:lsid:plazi:treatment:B5727931FFAD791AFF3C221767248DEF:38049EFCFFAC791AFD8425D960B98CF2" box="[559,702,151,176]" country="Vietnam" name="Cuc Phuong" pageId="29" pageNumber="30" stateProvince="Ninh Binh">Cuc Phuong</location>
).
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FFAC791AFF6C25FD60928C97" blockId="29.[151,1436,151,429]" box="[199,661,188,213]" pageId="29" pageNumber="30">
<materialsCitation id="8DB3C27AFFAC791AFF6C25FD60928C97" ID-GBIF-Occurrence="1288043597" box="[199,661,188,213]" collectionCode="ZMUM" country="Russia" location="Russia" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<emphasis id="0FAF1435FFAC791AFF6C25FD63728C97" bold="true" box="[199,373,188,213]" pageId="29" pageNumber="30">
<typeStatus id="E2607685FFAC791AFF6C25FD63058C97" box="[199,258,188,213]" pageId="29" pageNumber="30">Type</typeStatus>
material.
</emphasis>
<collectionCode id="5BCA50E2FFAC791AFED725FC63D48C96" box="[380,467,189,212]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="29" pageNumber="30">ZMUM</collectionCode>
(W.266),
<collectingCountry id="45CC88B7FFAC791AFDE825FD60958C97" box="[579,658,188,213]" name="Russia" pageId="29" pageNumber="30">Russia</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FFAC791AFF6C259E675C8CBA" blockId="29.[151,1436,151,429]" box="[199,1371,223,248]" pageId="29" pageNumber="30">
<materialsCitation id="8DB3C27AFFAC791AFF6C259E675C8CBA" ID-GBIF-Occurrence="1288043593" box="[199,1371,223,248]" collectingDate="1995-08-18" collectorName="Cuc Phuong NP" location="Ninh Binh" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Ninh Binh">
<emphasis id="0FAF1435FFAC791AFF6C259E63B48CBA" bold="true" box="[199,435,223,248]" pageId="29" pageNumber="30">Examined material.</emphasis>
1 C (
<emphasis id="0FAF1435FFAC791AFE59259E60A98CBA" bold="true" box="[498,686,223,248]" pageId="29" pageNumber="30">SORC-V.050.01</emphasis>
) Cave &quot;Nguoi Xua&quot;,
<collectorName id="902EADF1FFAC791AFC3425A166548CB5" box="[927,1107,223,248]" pageId="29" pageNumber="30">Cuc Phuong NP</collectorName>
,
<collectingRegion id="FF1F06C5FFAC791AFBF4259E66D08CBA" box="[1119,1239,223,248]" country="Vietnam" name="Ninh Binh" pageId="29" pageNumber="30">Ninh Binh</collectingRegion>
,
<collectingDate id="5921170FFFAC791AFB49259E67508CBA" box="[1250,1367,223,248]" pageId="29" pageNumber="30" value="1995-08-18">
<date id="4965EEE7FFAC791AFB49259E67508CBA" box="[1250,1367,223,248]" pageId="29" pageNumber="30" value="1995-08-18">18/8/1995</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="3D64C827FFAC791AFF6C244561C58D5E" blockId="29.[151,1436,151,429]" box="[199,962,260,285]" pageId="29" pageNumber="30">
<materialsCitation id="8DB3C27AFFAC791AFF6C244561C58D5E" ID-GBIF-Occurrence="1288043561" box="[199,962,260,285]" collectorName="Cuc Phuong NP" location="Vietnam. Ninh Binh" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Vietnam. Ninh Binh">
<emphasis id="0FAF1435FFAC791AFF6C244560678D5F" bold="true" box="[199,608,260,285]" pageId="29" pageNumber="30">
Records from
<collectingRegion id="FF1F06C5FFAC791AFEDA244560678D5F" box="[369,608,260,285]" inferredBySuffix="true" pageId="29" pageNumber="30">
Vietnam.
<collectingRegion id="FF1F06C5FFAC791AFE4E244560678D5F" box="[485,608,260,285]" country="Vietnam" name="Ninh Binh" pageId="29" pageNumber="30">Ninh Binh</collectingRegion>
</collectingRegion>
</emphasis>
(
<collectorName id="902EADF1FFAC791AFDC4244561208D5E" box="[623,807,260,285]" pageId="29" pageNumber="30">Cuc Phuong NP</collectorName>
) (
<bibRefCitation id="594AB5D6FFAC791AFC96244561B28D5E" author="Thai" box="[829,949,260,285]" pageId="29" pageNumber="30" refString="Thai, T. B. (1982) The fauna of earthworms f the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Jurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai 1982</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFAC791AFF6C246660558D27" pageId="29" pageNumber="30" type="distribution">
<paragraph id="3D64C827FFAC791AFF6C2466609A8D02" blockId="29.[151,1436,151,429]" box="[199,669,295,320]" pageId="29" pageNumber="30">
<emphasis id="0FAF1435FFAC791AFF6C246663678D02" bold="true" box="[199,352,295,320]" pageId="29" pageNumber="30">Distribution.</emphasis>
Only known from
<collectingCountry id="45CC88B7FFAC791AFD9C2466609D8D02" box="[567,666,295,320]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="3D64C827FFAC791AFF6C240D60558D27" blockId="29.[151,1436,151,429]" box="[199,594,332,357]" pageId="29" pageNumber="30">
<emphasis id="0FAF1435FFAC791AFF6C240D63A78D27" bold="true" box="[199,416,332,357]" pageId="29" pageNumber="30">Vietnamese name.</emphasis>
Giun
<taxonomicName id="FADBB3A4FFAC791AFE4E240D60498D27" box="[485,590,332,357]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="ghilarovi">ghilarovi</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFAC791AFF6C242E67248DEF" pageId="29" pageNumber="30" type="discussion">
<paragraph id="3D64C827FFAC791AFF6C242E67248DEF" blockId="29.[151,1436,151,429]" pageId="29" pageNumber="30">
<emphasis id="0FAF1435FFAC791AFF6C242E633C8DCA" bold="true" box="[199,315,367,392]" pageId="29" pageNumber="30">Remarks.</emphasis>
<bibRefCitation id="594AB5D6FFAC791AFEEE242E60108DC5" author="Blakemore" box="[325,535,367,392]" pageId="29" pageNumber="30" refString="Blakemore, R. J. (2007 a) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. www. annelida. net / earthworm / Pheretimoids. pdf (accessed 6 Jun 2016)" type="journal article" year="2007" yearSuffix="a">Blakemore (2007a</bibRefCitation>
,
<bibRefCitation id="594AB5D6FFAC791AFD82243160688DCA" author="Blakemore" box="[553,623,367,392]" pageId="29" pageNumber="30" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">2008b</bibRefCitation>
) considered it as &quot;
<emphasis id="0FAF1435FFAC791AFCE6243066258DCA" box="[845,1058,367,392]" italics="true" pageId="29" pageNumber="30">species inquirenda</emphasis>
&quot;. Material was re-examined and absence of male copulatory pouches and hence placement of the species in
<taxonomicName id="FADBB3A4FFAC791AFC4924D566578DEF" box="[994,1104,404,429]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="29" pageNumber="30" phylum="Annelida" rank="genus">
<emphasis id="0FAF1435FFAC791AFC4924D566578DEF" box="[994,1104,404,429]" italics="true" pageId="29" pageNumber="30">Amynthas</emphasis>
</taxonomicName>
is confirmed here.
</paragraph>
</subSubSection>
</treatment>
</document>
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