376 lines
51 KiB
XML
376 lines
51 KiB
XML
<document ID-CLB-Dataset="6387" ID-DOI="http://dx.doi.org/10.3897/zookeys.1040.66209" ID-GBIF-Dataset="48cd698d-e51b-4bb2-bd4d-1794c8c5955f" ID-PMC="PMC8178291" ID-Pensoft-Pub="1313-2970-1040-167" ID-Pensoft-UUID="1F71B6B55BCB516A825666EA04FC0431" ID-PubMed="34135659" ID-ZooBank="910903D2476A439FAC73292D5B9A18C1" ModsDocID="1313-2970-1040-167" checkinTime="1622233190126" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Srisonchai, Ruttapon & Panha, Somsak" docDate="2021" docId="B4DFAF1674885A80AFDE56D716EED2C4" docLanguage="en" docName="ZooKeys 1040: 167-185" docOrigin="ZooKeys 1040" docPubDate="2021-05-28" docSource="http://dx.doi.org/10.3897/zookeys.1040.66209" docTitle="Tylopus sutchariti Likhitrakarn & Golovatch & Srisonchai & Panha 2021, sp. nov." docType="treatment" docUuid="5385A3A0-C129-41F3-9A8C-C3910EF9178C" docUuidSource="ZooBank" docVersion="6" id="1F71B6B55BCB516A825666EA04FC0431" lastPageNumber="167" masterDocId="1F71B6B55BCB516A825666EA04FC0431" masterDocTitle="Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="185" masterPageNumber="167" pageNumber="167" updateTime="1732754193849" updateUser="ExternalLinkService">
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<mods:titleInfo id="894C41AE55DAD7B9269C6B4D7DD180E1">
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<mods:title id="B188CEE10C0B560F673667633BD85AF1">Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="57E1B3087651782AE84E4DAF8B456835" type="personal">
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<mods:namePart id="656478F6C43732C6AE65ABC2BC0BC288">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="3BE36682F9B9E39397F247249A6279E5">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="3C713D7480888F315E5177E149F84841" type="personal">
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<mods:namePart id="B8BC58B1CB1ECEE38A65CB5412CB8EB6">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="7BE173051717EDEE2DE421A96C2DE2A0">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="2C7E3DBD6B18EF13C4121B2C4CAEE0B2" type="personal">
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<mods:roleTerm id="2129B06530D0E87DBEC6D559553BC99A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B3334BA7410CBF75A94973326602C8FA">Srisonchai, Ruttapon</mods:namePart>
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<mods:nameIdentifier id="9D95ABFC0FFE3A32D40B2CD90BA72658" type="ORCID">https://orcid.org/0000-0002-7142-0999</mods:nameIdentifier>
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<mods:affiliation id="BD48E0C2ADF5F495B41361FB0D7A1361">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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</mods:role>
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<mods:namePart id="A7DAB7A1A9816BB9D1B87CD06B2414FB">Panha, Somsak</mods:namePart>
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<mods:affiliation id="0D46CE63DE05CA27E43A92038BA6297F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="CE8CA982982576B76657A0F7467DAB29" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:date id="6D4090A117902BC708366015DB77FE20">2021</mods:date>
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<mods:number id="2710B48E603F4C2226878F27AEBAFBD7">2021-05-28</mods:number>
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<mods:number id="3A02FC69FA8414E12061EC39D73AC58F">1040</mods:number>
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<mods:url id="1CC7C3652F451AA234D5D9C64A8FBA44">http://dx.doi.org/10.3897/zookeys.1040.66209</mods:url>
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<mods:classification id="61E6DB034F6BB12812BF6A9EC8ADD4D6">journal article</mods:classification>
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<mods:identifier id="EAAE78726C4AC41AF43540D93831D54C" type="DOI">http://dx.doi.org/10.3897/zookeys.1040.66209</mods:identifier>
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<mods:identifier id="B5732283CB251DA698168EDB8C395CB2" type="Pensoft-Pub">1313-2970-1040-167</mods:identifier>
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<treatment id="B4DFAF1674885A80AFDE56D716EED2C4" ID-GBIF-Taxon="182408860" LSID="urn:lsid:zoobank.org:act:5385A3A0-C129-41F3-9A8C-C3910EF9178C" httpUri="http://treatment.plazi.org/id/B4DFAF1674885A80AFDE56D716EED2C4" lastPageNumber="167" pageId="0" pageNumber="167" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida" scope_subFamily="Paradoxosomatinae" scope_tribe="Sulciferini">
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<subSubSection id="0A04A1C38020317F9724D9BA3CF37F89" pageId="0" pageNumber="167" type="nomenclature">
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<paragraph id="8B6E66FFE253D71FC7BFC37C22235A25" pageId="0" pageNumber="167">
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<taxonomicName id="1D32AC7B56EB685E422D335C8F0B0A7D" ID-CoL="8KR4M" LSID="http://zoobank.org/5385A3A0-C129-41F3-9A8C-C3910EF9178C" authority="Likhitrakarn & Golovatch & Srisonchai & Panha, 2021" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti" status="sp. nov.">Tylopus sutchariti</taxonomicName>
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<taxonomicNameLabel id="FB808611FB28ADC82D29FC0BBF07909B" pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
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<figureCitation id="ED01B2AE3DF9EEBED97D5B9199808F2F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">Figs 1B</figureCitation>
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<figureCitation id="77F8123157F2DAE2A9CBB859504CD1AA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">, 5</figureCitation>
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<figureCitation id="E3848A07DACD0F11D75E0A309E2EF760" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">, 6</figureCitation>
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<figureCitation id="E4E1FBB401661A3CDE7259043E3E2CDC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D0F22ADF52AEB52E53889021E9E79093" pageId="0" pageNumber="167" type="materials_examined">
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<paragraph id="4468275E9A271737A583941316B85828" pageId="0" pageNumber="167">Material examined.</paragraph>
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<paragraph id="D761DDD7CD981CCBAB5D8D61B2B953F0" pageId="0" pageNumber="167">
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<materialsCitation id="7CB14CA9F682A14E7FBCB717E3D06178" collectingDate="2016-09-21" collectorName="R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="♂" stateProvince="Shan State" typeStatus="Holotype">
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<emphasis id="876F6AD817A9B09D16B4A7A482419DDA" bold="true" pageId="0" pageNumber="167">
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<emphasis id="2154AD7DA795CAC0267739B26E7417F3" italics="true" pageId="0" pageNumber="167">
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<typeStatus id="A9E228101460F4DD6C5A0B9FBEE2E098">Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry id="835F7CB0001AF177046803FE32BA851D" name="Myanmar">Myanmar</collectingCountry>
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-
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<emphasis id="19EE5FF8163764D23F5D68CD07E3DE1E" bold="true" pageId="0" pageNumber="167">
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<collectingRegion id="3A2EFD7829E98FB4F699AC99961D8AE9" country="Myanmar" name="Shan">Shan State</collectingRegion>
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</emphasis>
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•
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<specimenCount id="788152CFE9172D4B641067718E1006B5">♂</specimenCount>
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;
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<collectingMunicipality id="B4D1789D5579CCCA31377631F1AECDDB">Taunggyi District</collectingMunicipality>
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, near
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<location id="F14960500377E41C184A68EC84C37107" LSID="urn:lsid:plazi:treatment:B4DFAF1674885A80AFDE56D716EED2C4:F14960500377E41C184A68EC84C37107" country="Myanmar" latitude="20.754417" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" name="Montawa Cave" stateProvince="Shan State">Montawa Cave</location>
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; elev.
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<quantity id="726D052239D9A7A0B34727D4A036FC00" metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">
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<elevation id="EF7EBA80A06168C29A77ED4721A13BFE" metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">1204 m</elevation>
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</quantity>
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;
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<geoCoordinate id="8C0302D0EB0B169B39775FBC833FF2C9" degrees="20" direction="north" minutes="45" orientation="latitude" precision="1" seconds="15.9" value="20.754417">20°45'15.9"N</geoCoordinate>
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,
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<geoCoordinate id="CE550E55813AF9AEF6981B3632E7D965" degrees="97" direction="east" minutes="01" orientation="longitude" precision="1" seconds="03.4" value="97.01762">97°01'03.4"E</geoCoordinate>
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;
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<collectingDate id="56A037E7D91F1179A55729782FB4CF55" value="2016-09-21">21 Sep. 2016</collectingDate>
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;
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<collectorName id="079583498ECE6E0330F03E3EF844BD27">R. Srisonchai</collectorName>
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leg.; CUMZ
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</materialsCitation>
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.
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<materialsCitation id="6EBB49B65E5EE028D71EDCED29F16E75" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Shan State" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="1" specimenCount-female="1" stateProvince="Shan State" typeStatus="Paratype">
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<emphasis id="8776D7587BBF5F6FD1291313C8135548" bold="true" italics="true" pageId="0" pageNumber="167">
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<typeStatus id="D06FAA6D979951E0568411B7D7EE8D17">Paratype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="2FFCA20856273FD8BE80095F57EFF1BE" name="Myanmar">Myanmar</collectingCountry>
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-
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<emphasis id="88D96749A2606D5728342577F798BBCB" bold="true" pageId="0" pageNumber="167">
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<collectingRegion id="F6D95AED02D2B18F6DA3CF74FA9F291A" country="Myanmar" name="Shan">Shan State</collectingRegion>
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</emphasis>
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•
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<specimenCount id="BE68FDB7EA0D0318C10FECA3CF02AB9D" type="female">1 ♀</specimenCount>
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; same collection data as holotype; CUMZ
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FA507E77C2D56A8F595A8A3F12E9572E" pageId="0" pageNumber="167" type="diagnosis">
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<paragraph id="0BF7D41183BE5C414986DA15237394E5" pageId="0" pageNumber="167">Diagnosis.</paragraph>
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<paragraph id="6202066C47E4E897388C01B5AFBF8CA0" pageId="0" pageNumber="167">
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This new species comes to a dead end in couplet 5 in the latest key to
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<taxonomicName id="650EF9CE561E6B18D745129C98323952" authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
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<emphasis id="06AD4E889E158AACBAF1055B86D5C0FE" italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
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</taxonomicName>
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species (
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<bibRefCitation id="12C5DB581D145CCA25D542AB7E248DED" DOI="https://doi.org/10.5852/ejt.2016.195" author="Likhitrakarn, N" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="167" pagination="1 - 47" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2016. The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina. European Journal of Taxonomy 195: 1 - 47, DOI: https://doi.org/10.5852/ejt.2016.195" title="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina." url="https://doi.org/10.5852/ejt.2016.195" volume="195" year="2016">Likhitrakarn et al. 2016</bibRefCitation>
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), but it seems to be particularly similar to the later described and grossly sympatric
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<taxonomicName id="1B9A844F99F4656F9A30D5E02FDE93B8" lsidName="T. brehieri" pageId="0" pageNumber="167" rank="species" species="brehieri">
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<emphasis id="A9E1CA7888890F1F726DCD5E518195DD" italics="true" pageId="0" pageNumber="167">T. brehieri</emphasis>
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</taxonomicName>
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Golovatch, VandenSpiegel & Semenyuk, 2016, especially in its gonopod conformation. Both species compared come from Shan State, Myanmar (
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<bibRefCitation id="EEA645FF4843EA93F8512E34DD1B48B5" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="335 - 354" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" refId="B16" refString="Golovatch, SI, VandenSpiegel, D, Semenyuk, II, 2016. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI. Arthropoda Selecta 25 (4): 335 - 354, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" volume="25" year="2016">Golovatch et al. 2016</bibRefCitation>
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), but
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<taxonomicName id="5A415D825AC2FDEA867AE2F65D6F09DF" lsidName="T. sutchariti" pageId="0" pageNumber="167" rank="species" species="sutchariti">
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<emphasis id="4C8D49BA2952A88121EBBFD00F6C5191" italics="true" pageId="0" pageNumber="167">T. sutchariti</emphasis>
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</taxonomicName>
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sp. nov. differs in the presence of a small and triangular gonopod process
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<emphasis id="F1D43DB6BBE4410B31DC3DD772BA5BB6" bold="true" pageId="0" pageNumber="167">h</emphasis>
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(vs. absent), and the large and subtrapeziform apicolateral lobe (
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<emphasis id="C7AF698B86558A745AB504F8CDF82586" bold="true" pageId="0" pageNumber="167">l</emphasis>
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) with a smooth apical margin (Figs
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<figureCitation id="32F743866532B4FF3364C012F6B719AD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">6B, D</figureCitation>
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,
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<figureCitation id="25C7B8BD6AC1E3EA8866689CBC380DE8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7B</figureCitation>
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) (vs. a subtriangular
|
|
<emphasis id="7DB291B62FDED4050C1EF87C30F76FB4" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
with an apically rugose and denticulate margin), as well as the pleurosternal carinae being complete crests with a caudal tooth clearly visible until segments 16 (♂) or 13 (♀), thereafter missing (Fig.
|
|
<figureCitation id="3E1950D9EFEAE3AFC21C593F8C9D21C7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B, D, E</figureCitation>
|
|
) (vs. visible until segment 10), while the sternal lobe between ♂ coxae 4 is deeply notched (Fig.
|
|
<figureCitation id="A6D99AC421399EFC9AC55442B5A0E0C2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5H, I</figureCitation>
|
|
) (vs. prominent and subquadrate).
|
|
</paragraph>
|
|
<caption id="30BF1FCAE4C7F60BE5A6D1D8B29643D1" doi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167" start="Figure 5" startId="F5">
|
|
<paragraph id="75ED61DFD15CD9F2B3F9B8B99CF8ADA4" pageId="0" pageNumber="167">
|
|
<emphasis id="681D607FF9BDAB617B5E94941D9DCFB2" bold="true" pageId="0" pageNumber="167">Figure 5.</emphasis>
|
|
<taxonomicName id="EBBC66A37B13AC993D7D562F7D7F9CB0" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
|
<emphasis id="882BC43E167D91303E58A1B30F7FF233" italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ)
|
|
<emphasis id="EAFC563E3503D407B0C693F82B01BC62" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
anterior part of body, dorsal and lateral views, respectively
|
|
<emphasis id="C25BF6AAB7A4C41C82F75D31D2B65DDD" bold="true" pageId="0" pageNumber="167">C, D</emphasis>
|
|
segments 10 and 11, dorsal and lateral views, respectively
|
|
<emphasis id="B97CD1B46E2C5F63706162EFE2C0D471" bold="true" pageId="0" pageNumber="167">E-G</emphasis>
|
|
posterior part of body, lateral, subdorsal and subventral views, respectively
|
|
<emphasis id="842697CEDB037C5BA261E7DB04607757" bold="true" pageId="0" pageNumber="167">H, I</emphasis>
|
|
sternal cones between coxae 4, subcaudal and sublateral views, respectively.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="B41352B1DB95A4F81B46A247F754F96A" pageId="0" pageNumber="167" type="description">
|
|
<paragraph id="F432E054708BA64BF7862C1D684BDDDF" pageId="0" pageNumber="167">Description.</paragraph>
|
|
<paragraph id="5C1F1C7927CF66E07CEBB7182C025388" pageId="0" pageNumber="167">Length 24.1 (♂) or 22.5 mm (♀), width of midbody pro- and metazona 1.8 and 2.7 mm (♂) or 2.2 and 2.6 mm (♀), respectively.</paragraph>
|
|
<paragraph id="49EE5BA05B16534A03DFBBD82CEA12A8" pageId="0" pageNumber="167">
|
|
Colouration of live animals dark brown (Fig.
|
|
<figureCitation id="2E789B7E3E117374693FEF7A659690A3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1B</figureCitation>
|
|
); venter and legs brown; colouration of alcohol material after two years of preservation blackish, calluses of paraterga yellowish brown, head and antennae dark brown, venter and legs light yellowish, increasingly darker brown distally (Fig.
|
|
<figureCitation id="3EB2765D479926AC614EC8B996613C10" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="0AD877E5D9559793B6AB42C900F97AFB" pageId="0" pageNumber="167">
|
|
Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae short (Figs
|
|
<figureCitation id="925CEB4D29C834513A72596F58C1D462" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1B</figureCitation>
|
|
,
|
|
<figureCitation id="89EE9BE45CBFDACCECE9EF03434D052C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
|
|
), reaching body segment 3 (♂) or surpassing body segment 2 (♀) when stretched dorsally. In width, head <segment 3 <4 <5 <collum <segment 2 <6-17 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 3+3 anterior, 1+1 intermediate, and 3+3 posterior; a small lateral incision near midway; caudal corner of paraterga rounded, slightly declined ventrad, not surpassing rear tergal margin (Fig.
|
|
<figureCitation id="2F114A736680F3AEB0526A6B3846C03F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="330BCAF3CE476A86A9CF60314C948056" pageId="0" pageNumber="167">
|
|
Tegument rather smooth and shining, prozonae very finely shagreened, metaterga smooth and finely rugulose; surface below paraterga finely microgranulate (Fig.
|
|
<figureCitation id="FD22D789A4E94C4F4E75073C789A21CA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A-F</figureCitation>
|
|
). Postcollum metaterga with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points. Tergal setae long, strong, slender, about 1/3 metatergal length. Axial line visible only on metazonae.
|
|
</paragraph>
|
|
<paragraph id="C27CA6D00CA1D91CFCF00EEB35D1F17C" pageId="0" pageNumber="167">
|
|
Paraterga strongly developed (Fig.
|
|
<figureCitation id="B8123CD47F6C00CAEE5AA977E4217D78" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A-F</figureCitation>
|
|
), especially well so in ♂, set high, at upper 1/3 of midbody height, slightly upturned, but remaining below dorsum; anterior edge well-developed, mostly regularly rounded and narrowly bordered, continuous with callus; caudal corner narrowly rounded to fully pointed, extending increasingly past rear tergal margin, especially well so on segments 15-19; on segments 16-19, tips strongly curved mesad, posterior edge slightly oblique (Fig.
|
|
<figureCitation id="F1823F59997395E53B2942EF43B4BB18" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing segments (Fig.
|
|
<figureCitation id="55C1CF428D64FD26AB566A37743D05A6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5D</figureCitation>
|
|
). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Paraterga 2 broad, horizontal, anterior edge angular, lateral edge with three evident incisions, one in anterior 1/3, middle one at midway, caudal incision near tip; anterior incision particularly evident. Paraterga 3 and 4 with two small incisions at lateral edge (Fig.
|
|
<figureCitation id="70EE8A3890C4C3390B14E6C956C60FE1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A</figureCitation>
|
|
), one in anterior 1/3, the other at midway, anterior one also particularly evident. Following segments each with lateral edge showing an evident incision near front 1/3 (Fig.
|
|
<figureCitation id="D0C160E57E914DB4C7D4D334E3A521D0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5C</figureCitation>
|
|
). Ozopores evident, lateral, lying in an ovoid groove at about 1/3 metatergal length in front of posterior edge of metaterga (Fig.
|
|
<figureCitation id="07685F64A6BACF65AB54D991021BE3C4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5D</figureCitation>
|
|
). Transverse sulcus usually distinct (Fig.
|
|
<figureCitation id="66C5DEBA4157C364A4FB5C67B3B6868B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
), complete and visible on metaterga 5-18, deep, narrow, reaching bases of paraterga, line-shaped, clearly beaded at bottom. Stricture between pro- and metazonae wide, deep, clearly ribbed at bottom down to base of paraterga (Fig.
|
|
<figureCitation id="0452D77644937595B3D06BBDF1895015" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5A, C, F</figureCitation>
|
|
). Pleurosternal carinae complete crests on segment 2-4 (Fig.
|
|
<figureCitation id="9C3A36CD6CBB8A812DB6E3496DC7732B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B</figureCitation>
|
|
), with anteriorly bulged crests and a sharp denticle caudally on segments 5-8 (♂, ♀), thereafter increasingly reduced and broken, remaining only a small sharp caudal tooth until segment 16 (♂) or 13 (♀), thereafter missing (Fig.
|
|
<figureCitation id="785128D21DE70BB0D2CFF407885D9FEB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5B, D, E</figureCitation>
|
|
). Epiproct (Fig.
|
|
<figureCitation id="960743A4672538EF5623A8A0298BEA21" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5E-G</figureCitation>
|
|
) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed caudally, both pointed at tip; pre-apical papillae evident, lying close to tip. Hypoproct roundly subtrapeziform (Fig.
|
|
<figureCitation id="E47F60A3D72A422C46F5A969E1A179BF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5G</figureCitation>
|
|
), small setiferous knobs at caudal edge well-separated and evident.
|
|
</paragraph>
|
|
<paragraph id="9A68BA0F02E618D29062A1E7A16AF909" pageId="0" pageNumber="167">
|
|
Sterna densely setose, without modifications (Fig.
|
|
<figureCitation id="513994AC967ABAD017282F259AD4DB30" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5G</figureCitation>
|
|
); cross-impressions shallow; a deeply notched sternal lobe between ♂ coxae 4 (Fig.
|
|
<figureCitation id="C9F4563B8984B4AF57C92B3F5DEA6572" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure5" httpUri="https://binary.pensoft.net/fig/549198" pageId="0" pageNumber="167">5H, I</figureCitation>
|
|
). Legs long and slender, midbody ones ca 1.4-1.5 (♂) or 1.1-1.2 (♀) as long as body height; ♂ legs of segments 4-17 with prefemora distinctly swollen laterally; ♂ legs of segments 2-16 each with femur, postfemur, tibia and tarsus with particularly dense setae and carrying ventral microgranulations (Fig.
|
|
<figureCitation id="FD966AEE97A71B6F49537FED1237DC04" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7C</figureCitation>
|
|
), tarsal brushes absent.
|
|
</paragraph>
|
|
<paragraph id="61C7C5266EC9E9FF33826701C1155352" pageId="0" pageNumber="167">
|
|
Gonopods (Figs
|
|
<figureCitation id="1884141F1C8381969917251501916D5A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167">6</figureCitation>
|
|
,
|
|
<figureCitation id="17CA075E24940737D184D7DCFAE0BA34" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tylopus sutchariti sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and triangular process of femorite, l apicolateral lobe of femorite, m dorsoapical lobe of femorite, sl tip of solenomere, sph solenophore, z short and simple process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167">7A, B</figureCitation>
|
|
) complex; coxa slightly curved caudad, sparsely setose distoventrally. Prefemorite as usual, densely setose, about 1/2 as long as femorite +
|
|
<normalizedToken id="F2BBB886745551D80BAE6E15628A6E5D" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part. Femorite rather stout, suberect, expanded distad, showing a distinct mesal groove/hollow (
|
|
<emphasis id="348215F8527A7C2B6F55E6DF9E828A0D" bold="true" pageId="0" pageNumber="167">g</emphasis>
|
|
) and a prominent, rounded, dorso-apical lobe (
|
|
<emphasis id="62284A1AC86FBA401AF302892FA2BE1A" bold="true" pageId="0" pageNumber="167">m</emphasis>
|
|
), apicolateral lobe (
|
|
<emphasis id="670582560669F0061BA462D7A94F8D56" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
) large, subtrapeziform, with a smooth apical margin, mostly delimited at base by a transverse sulcus, with process
|
|
<emphasis id="BADE73A6A3ECD46D62EA24105984410F" bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
short and triangular; process (
|
|
<emphasis id="447C782E3FBB8C7690A73547133CA491" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
) rather short and simple, narrowly rounded at tip. Solenophore (
|
|
<emphasis id="3073E01FB8C328D610A049A7ADF54270" bold="true" pageId="0" pageNumber="167">sph</emphasis>
|
|
) typically coiled, lamellar, expanded apically into a subtruncate lobe, almost fully sheathing a similarly long, flagelliform solenomere, with only its tip (
|
|
<emphasis id="29FC0B561BC8B4C12448A2B925629147" bold="true" pageId="0" pageNumber="167">sl</emphasis>
|
|
) being exposed.
|
|
</paragraph>
|
|
<caption id="858DF022827B039614D6EE9651123061" doi="10.3897/zookeys.1040.66209.figure6" httpUri="https://binary.pensoft.net/fig/549199" pageId="0" pageNumber="167" start="Figure 6" startId="F6">
|
|
<paragraph id="D4D90B88D1B20A3865F45722902EE0EB" pageId="0" pageNumber="167">
|
|
<emphasis id="DB66262ED1252D394364C8F559A91428" bold="true" pageId="0" pageNumber="167">Figure 6.</emphasis>
|
|
<taxonomicName id="76FFD06636A5F42A0A5B3ECC81A97337" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
|
<emphasis id="0E1B87D58D39CD70D40CCBCF12AF7B90" italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ), left gonopod
|
|
<emphasis id="9384CE173D2A342BF9B13671DB97D4D8" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
mesal and lateral views, respectively
|
|
<emphasis id="746D26AFEB376C24CA01C8ED6D621536" bold="true" pageId="0" pageNumber="167">C-F</emphasis>
|
|
distal part, submesal, sublateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="112E5F651E7545EC8E35FEA5043E05DF" pageId="0" pageNumber="167" type="name">
|
|
<paragraph id="9F0187B42E4AD83BFCAF681ABE63B291" pageId="0" pageNumber="167">Name.</paragraph>
|
|
<paragraph id="4AD077C6D9AC89E5DCCCB5B25CED531A" pageId="0" pageNumber="167">To honour Dr. Chirasak Sutcharit, Professor at the Department of Biology of the Chulalongkorn University, Bangkok, who participated in collecting the type series.</paragraph>
|
|
<caption id="D46D3EBEBC8992E19199B529A2B80E86" doi="10.3897/zookeys.1040.66209.figure7" httpUri="https://binary.pensoft.net/fig/549200" pageId="0" pageNumber="167" start="Figure 7" startId="F7">
|
|
<paragraph id="7D60BCF6599AA808B64B0249809DE647" pageId="0" pageNumber="167">
|
|
<emphasis id="AF30255AE2355B46CD9C8065047E7ED2" bold="true" pageId="0" pageNumber="167">Figure 7.</emphasis>
|
|
<taxonomicName id="0371080F70D4245C1E8BFA3FE8A5B32E" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
|
<emphasis id="BB0C9CCB474A3DD741EDE5DAEBE9CDAD" italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ)
|
|
<emphasis id="70BF032DDCD5B1E527BC3FE13D16FE4D" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
|
left gonopod, mesal and lateral views, respectively
|
|
<emphasis id="66DAD425C8D02BFB0F5B0B85E6000E63" bold="true" pageId="0" pageNumber="167">C</emphasis>
|
|
leg of segment 10. Abbreviations:
|
|
<emphasis id="6B10BA9979E3CF45AE75EB8A81A3D826" bold="true" pageId="0" pageNumber="167">g</emphasis>
|
|
mesal groove/hollow of femorite,
|
|
<emphasis id="8E8EF2415FCEF2328D4A04D6711B900E" bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
short and triangular process of femorite,
|
|
<emphasis id="B5EA43E5F03ADC11BD2071EAAC478F28" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
apicolateral lobe of femorite,
|
|
<emphasis id="ECBB6AD637727CB82B0509E0609E5555" bold="true" pageId="0" pageNumber="167">m</emphasis>
|
|
dorsoapical lobe of femorite,
|
|
<emphasis id="68B475846A78484D0F81750B31F7BA3B" bold="true" pageId="0" pageNumber="167">sl</emphasis>
|
|
tip of solenomere,
|
|
<emphasis id="D5C9498684C74825EAF3A3373ACB427C" bold="true" pageId="0" pageNumber="167">sph</emphasis>
|
|
solenophore,
|
|
<emphasis id="730A64BA3455E4DC105E7CBC658AF5E4" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
short and simple process of femorite. Scale bars: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="6B02EBD11200FF554C1D39485E4A3154" doi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167" start="Figure 8" startId="F8">
|
|
<paragraph id="FE54CE5056A6890C5ADFD4E07D0E22B8" pageId="0" pageNumber="167">
|
|
<emphasis id="E62797942618599E7C63376909B50AB4" bold="true" pageId="0" pageNumber="167">Figure 8.</emphasis>
|
|
Gonopod structure of
|
|
<taxonomicName id="315189647B74D6C0D9B3CF9AB2AEADCF" authorityName="Golovatch & Enghoff" authorityYear="1993" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus rugosus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="rugosus">
|
|
<emphasis id="B91A671F67BF42B5F73B1639EF159BAB" italics="true" pageId="0" pageNumber="167">Tylopus rugosus</emphasis>
|
|
</taxonomicName>
|
|
Golovatch & Enghoff, 1993, ♂ holotype, left gonopod
|
|
<emphasis id="74974C06D4A2D262B01ED0DE943FD236" bold="true" pageId="0" pageNumber="167">A-C</emphasis>
|
|
mesal, lateral and dorsal views, respectively. Abbreviations:
|
|
<emphasis id="990394A89E11FA96171DBD60E2DF96B7" bold="true" pageId="0" pageNumber="167">h</emphasis>
|
|
strong hook-shaped process of femorite,
|
|
<emphasis id="0D27A17C512297C51423FBF083433A81" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
|
apicolateral lobe of femorite,
|
|
<emphasis id="EF0DD33AED862ACB3355FEEEA6D705BA" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
|
serrate process of femorite. Scale bar: 0.5 mm (after
|
|
<bibRefCitation id="C97F0CC0218B6C67BC358839316B15D5" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="167" pagination="85 - 125" refId="B14" refString="Golovatch, SI, Enghoff, H, 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125" title="Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="19" year="1993">Golovatch and Enghoff 1993</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="E0A7D089B2F2B0F2F3ED837FF1349FAE" pageId="0" pageNumber="167" type="remarks">
|
|
<paragraph id="F43D74F06C4A04D484AE63D40DED9EFA" pageId="0" pageNumber="167">Remark.</paragraph>
|
|
<paragraph id="B4E06B77DD1306B55DD12C8650821C97" pageId="0" pageNumber="167">
|
|
Both new species described here have been found to occur syntopically (Fig.
|
|
<figureCitation id="985508D2F33D777772939263E0FE1C7B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributions of Tylopus species recorded from Myanmar. Pink circle: T. brehieri Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk: T. punctus Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle: T. monticola sp. nov.; Blue square: T. sutchariti sp. nov. and T. monticola sp. nov.; Orange diamond: T. silvestris (Pocock, 1895); Green triangle: T. doriae (Pocock, 1895)." figureDoi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167">9</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="987AA7A6A9298C05E150DC5DB284FB51" doi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167" start="Figure 9" startId="F9">
|
|
<paragraph id="AEA9946DF4DE9386D78C4FF35CA7D35C" pageId="0" pageNumber="167">
|
|
<emphasis id="C4F10DFB8FF8E1E010CE31ED08B5FDEF" bold="true" pageId="0" pageNumber="167">Figure 9.</emphasis>
|
|
Distributions of
|
|
<taxonomicName id="D0E0D2C79FFEEBFC1E5BA52559CA9825" authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="AC2268D4DF81F7D7CC53367A095231C6" italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
|
|
</taxonomicName>
|
|
species recorded from Myanmar. Pink circle:
|
|
<taxonomicName id="1178F64830FA9BCBE9BCE81F4BA5600A" lsidName="T. brehieri" pageId="0" pageNumber="167" rank="species" species="brehieri">
|
|
<emphasis id="03381DD957884501AB5D9EB9CBEEEE4B" italics="true" pageId="0" pageNumber="167">T. brehieri</emphasis>
|
|
</taxonomicName>
|
|
Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk:
|
|
<taxonomicName id="1672E873FBF54500D061C7EBD40CE65C" lsidName="T. punctus" pageId="0" pageNumber="167" rank="species" species="punctus">
|
|
<emphasis id="CE623EBE4AF39324E5FF6DE2FD719F37" italics="true" pageId="0" pageNumber="167">T. punctus</emphasis>
|
|
</taxonomicName>
|
|
Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle:
|
|
<taxonomicName id="73B9263C043D8EF6E245E8E98A667B9A" lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
|
<emphasis id="528CDD83CCDC14D24F7F3E0073F68312" italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.; Blue square:
|
|
<taxonomicName id="1E857CFFF3EBD2698DF9DB15A44CDE75" lsidName="T. sutchariti" pageId="0" pageNumber="167" rank="species" species="sutchariti">
|
|
<emphasis id="650C0DCE41E6E042E2DB2E7FC75ACDD0" italics="true" pageId="0" pageNumber="167">T. sutchariti</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName id="439B4C6C633A25BDB7C2FAD11807327F" lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
|
<emphasis id="69303CA7784A1E7D3622E1E05F55A2CC" italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.; Orange diamond:
|
|
<taxonomicName id="EB5527603157DA645BBB0561707BAA2F" lsidName="T. silvestris" pageId="0" pageNumber="167" rank="species" species="silvestris">
|
|
<emphasis id="BDFD381640672D8BC25724EBC329EF8C" italics="true" pageId="0" pageNumber="167">T. silvestris</emphasis>
|
|
</taxonomicName>
|
|
(Pocock, 1895); Green triangle:
|
|
<taxonomicName id="C6A2E9BEAE36F74EE8F472658222FDEF" lsidName="T. doriae" pageId="0" pageNumber="167" rank="species" species="doriae">
|
|
<emphasis id="34F60BBAB6551F9BC455BAF2AA547390" italics="true" pageId="0" pageNumber="167">T. doriae</emphasis>
|
|
</taxonomicName>
|
|
(Pocock, 1895).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |