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<document ID-CLB-Dataset="7610" ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="B471A3682E38576C82998670D6AE89EA" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina foja Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="464A09DF-E5D6-4D1D-9455-59676F42C60D" docUuidSource="ZooBank" docVersion="8" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1732748175969" updateUser="ExternalLinkService">
<mods:mods id="8D7A866C964FBC528F457D9F2DAC3969" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BA6A0C2D44F839D8269E315C61C30A35">
<mods:title id="17088EEC4FE16FA1BDF4F9B20687460D">Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="F2FFACDFCCB8765D776AB4C9A96F5FE9" type="personal">
<mods:role id="D6E99A3D570035DE5516C110E13A439A">
<mods:roleTerm id="91AEDA44650BBC05AED2EA7E3F459A4E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC9619EC9517DCA8EDFBDD4902AF1221">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="0C131896D805428AB66DB30B707B4C8B">Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="9E6E49A44164199AA9FC6FBDB8B262B0" type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="F70AB69A95531F4219FA4129A14FD0A2" type="personal">
<mods:role id="9DEE5A02CA92D314B278371A64FDCA68">
<mods:roleTerm id="68BB0F669E490B26D8E9FB2C41FBF398">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="499E387BBC35185CDBAC89876A4C4B46">Surbakti, Suriani</mods:namePart>
<mods:affiliation id="9BAF1A26DE8CB7BBEF8D8A658D7AEFF0">Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="431FC061CABADF709C9306E80D34581C" type="personal">
<mods:role id="099C62C7AC8243CE2A10C81F2A8C6C8C">
<mods:roleTerm id="2461C752B4F3F9D58677EC63287DD877">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFF5C3B2C05D6F133F4877DAD277D37F">Sumoked, Bob</mods:namePart>
<mods:affiliation id="5B7BCDDACFCD0779F2C26C81ED03D382">Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="A225EE6DB70A56F5618649DDD67A1443" type="personal">
<mods:role id="A03F68D108D88CEC18ED806AEB0557D8">
<mods:roleTerm id="0B126F89E17ECF5CF77F7D1D2C04A7AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B735976117E50C17064A015FEECBDFBD">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="AD157703A950E29C04EB74F4269B7F9E" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="048152F2E0107A21BBA66C6FAB6AA0A5">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="56CA1D9C7DF09447CB0AA1D7072768CB">text</mods:typeOfResource>
<mods:relatedItem id="FD5C0B083B82761F10461A7C9F9B05E5" type="host">
<mods:titleInfo id="9C2B2C9C4DB16A9E66F61B9227A1E512">
<mods:title id="3BBF031CC35CF27F13D46BEB8D1D3534">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="FA72A93B187139D56F85F8A69B5A3E1B">
<mods:date id="EE910EA954ACAF4EAF4736BEC4318BA8">2021</mods:date>
<mods:detail id="ED68C07C1674CE11431B37EAF0C94B5E" type="pubDate">
<mods:number id="670C00948339B71B3F5EBBB304B21B0A">2021-03-25</mods:number>
</mods:detail>
<mods:detail id="A2792F7219FAF0C36BDD1407315475A9" type="volume">
<mods:number id="394708B42BF9F88EA8D2FCAE109F26C3">1026</mods:number>
</mods:detail>
<mods:extent id="DAF41F78AD2727DCB1710B58069A549F" unit="page">
<mods:start id="B6174C7941190E8E77BD57A5677EE7B8">45</mods:start>
<mods:end id="5E74197FA9B654780EB78C4AA0E9D9A3">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3D383B8127B0AF961824314A54464B8A">
<mods:url id="1EF1F16FEB9183CEF6B978E080A31238">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification id="1DC6F6206592989C86773A63B259DBA3">journal article</mods:classification>
<mods:identifier id="CDCBB55D5F5854E4E2BA1C868F4B6316" type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier id="A4447CA055505E31CB1C07A55E21A3DC" type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier id="70C3559E87F11ACB64BB8CEFFB84EB2F" type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier id="D6B577143D3B91D3EFA21A343F49FD27" type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment id="B471A3682E38576C82998670D6AE89EA" ID-GBIF-Taxon="180680993" LSID="urn:lsid:zoobank.org:act:464A09DF-E5D6-4D1D-9455-59676F42C60D" httpUri="http://treatment.plazi.org/id/B471A3682E38576C82998670D6AE89EA" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="56CA6537C25327882A95E6DD1F906403" pageId="0" pageNumber="45" type="nomenclature">
<paragraph id="AD0B0A6904202ECB9626EED2E99F85B8" pageId="0" pageNumber="45">
<taxonomicName id="EECA8D6666C4FF086F16ECB31288EABC" ID-CoL="8LZKB" LSID="http://zoobank.org464A09DF-E5D6-4D1D-9455-59676F42C60D/" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja" status="sp. nov.">Exocelina foja Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel id="12726E84EB67159216660867F67A22A3" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation id="B12463533352DA2ABD3091B0B11080B6" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 1</figureCitation>
<figureCitation id="475843021FBCFD5EFC087F9A7BF19BCE" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">, 5</figureCitation>
<figureCitation id="CA8FFA72057BDF221E435FF825568594" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9C5F574F43AD12890A29672E10A72DB6" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="F16A6A80DC0D07F4266E831A9F68FA84" pageId="0" pageNumber="45">
<taxonomicName id="59F07D23CDAE48BC07C1CB72B17A7031" authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudosoppi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pseudosoppi" status="sp. nov.">Exocelina nr. pseudosoppi</taxonomicName>
#7286:
<bibRefCitation id="A3FDF6EC570A758B8532CC30A930445E" DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection id="85E81420AE43947EF67843F831E02DBC" pageId="0" pageNumber="45" type="type locality">
<paragraph id="DFF501A5F6067E77247CC60E86797476" pageId="0" pageNumber="45">
<typeStatus id="F0046E60D7CCD1965E7AE5A59D0ED329">Type</typeStatus>
locality.
</paragraph>
<paragraph id="BF84FFEAD689D0E2A816E481204DD342" pageId="0" pageNumber="45">
<materialsCitation id="115ED8A2A1F25442F18FE84F994648DF" ID-GBIF-Occurrence="3062138332" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="1700" latitude="-2.5718334" location="Foja Mts" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry id="19C982D55F30A33BA096811A26CB21A4" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="FC10DE397353E1DED00FC9ADADB1022D" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
,
<collectingMunicipality id="DBFAC4AA6812C8B8467AC885695E9E95">Sarmi Regency</collectingMunicipality>
,
<location id="21D3633A7BAC2CF23C47FAC8065D398A" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:21D3633A7BAC2CF23C47FAC8065D398A" country="Indonesia" latitude="-2.5718334" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" name="Foja Mts" stateProvince="Papua Barat">Foja Mts</location>
,
<geoCoordinate id="27B52B4174DB1584C119C76A8DF21DA6" degrees="02" direction="south" minutes="34" orientation="latitude" precision="1" seconds="18.6" value="-2.5718334">02°34'18.6&quot;S</geoCoordinate>
,
<geoCoordinate id="4433CD0A82FB8403D61918C563929565" degrees="138" direction="east" minutes="43" orientation="longitude" precision="1" seconds="02.1" value="138.71724">138°43'02.1&quot;E</geoCoordinate>
,
<elevation id="28AF3EAFD34F4BFB14CA36AAD9F6E2D1" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="312FB9904C9EFE573C091320DEB3C71A" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1CF10D2EE0CA054CA20F5E9284654B0" pageId="0" pageNumber="45" type="type material">
<paragraph id="664C6E773224477B650DC0A40DDB857E" pageId="0" pageNumber="45">
<typeStatus id="F2E8236F54A64B81C68F66FD19EA29ED">Type</typeStatus>
material.
</paragraph>
<paragraph id="FB4104E968B2C6ADF2AE552650530B43" pageId="0" pageNumber="45">
<emphasis id="E8C810694B2F745872B99390DA735A0F" bold="true" pageId="0" pageNumber="45">
<emphasis id="DC9931497D1D7DF8A392284B517B3227" italics="true" pageId="0" pageNumber="45">
<typeStatus id="7C540E756DC7AD52A8A44A498BA30126">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation id="A9DD9342C128149F419243F76EBE44D7" ID-GBIF-Occurrence="3062138314" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot;
<collectingCountry id="ECDB54AB8E0C5E7D6344A9B3EDD63324" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="82185BB7B9F4BDE78EA669BA5FD46EEF" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="563E0FA5FD394E70DADF78CFAEE8E80E" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:563E0FA5FD394E70DADF78CFAEE8E80E" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="2FC21EB9B612D2BC864DBB621687923A" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="FDC82762F716ACA660B4BCF24AE4353C" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="5FE50324E41B1029732B96E828F70F37" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="5F72AF6DB2E4E1C2602324B5A61C7A22" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="C4542B121E7CF7ECBE1F9E6668CD2920" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="57B2944ECCB8061DC4245C224F3C4345" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode id="A1E1BD00569D456CBA6EDFCE6DDA0D1D" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="135DAC302903E193A2574B2E6C40CC27" pageId="0" pageNumber="45">
<materialsCitation id="22094F89018E77792F710D5A31FF0F7C" ID-GBIF-Occurrence="3062138330" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="45" specimenCount-female="19" specimenCount-male="26" typeStatus="paratype">
<emphasis id="B6C96E5A7E5F8ED6AEF81A7A559C128B" bold="true" pageId="0" pageNumber="45">
<emphasis id="A59E79124269AE6B81BD6939B4C29BB3" italics="true" pageId="0" pageNumber="45">
<typeStatus id="BE41A87D41EF07AC0923D7AE6FC74812">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount id="2F9722C59FEE59BF3893C45E301C18A8" type="male">26 males</specimenCount>
,
<specimenCount id="1019005D6541BE8312575E4C67B33A18" type="female">19 females</specimenCount>
with the same label as the
<typeStatus id="24D51E542549DE310B075A40601433FA">holotype</typeStatus>
, three males with additional handwritten labels &quot;creek A&quot;, &quot;creek C&quot; and &quot;creek D&quot; (
<collectionCode id="51162D1496678162D8769877B8690011" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="78E7965B087FCA33C61F76F1BA441DCC" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3DFC5E60310DF093C30C05940C6B974D" ID-GBIF-Occurrence="3062138333" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="659716E5103413B74995ECE8BF0E6C3D" type="male">1 male</specimenCount>
,
<specimenCount id="98F419C9DAC09B1405A92BF4A2BD11B1" type="female">6 females</specimenCount>
&quot;
<collectingCountry id="9570067CA3FDBE419DA0597599B2F595" name="Indonesia">Indonesia</collectingCountry>
(1700A):
<collectingRegion id="9E20D29B0ACFC0395753A7603A88A547" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="6838DEAF27438B31517C99228A9E9BA1" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:6838DEAF27438B31517C99228A9E9BA1" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="DBD064E87843431A311D690E7C58A671" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="AE58B047408AFFCA1D907B9EF0482FB5" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="64768C2AE7D2ABE5375322FD594B85F4" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="105CAD6BAF3834E0222973C9F713374E" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="AEFC9280EECC361C2B69209CB2E7F308" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="1AD410417784663DE29E31B3141E1CFC" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode id="939A5D66C7FC1E109372DC641EB2BAFD" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="6AA0A44CEF4F5BC32DAA24CDFD650A51" ID-GBIF-Occurrence="3062138324" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="ED1C7ECF8E40C9A60AC1E932DD292B23" type="male">4 males</specimenCount>
,
<specimenCount id="C9EE6AED756FFC01E011A65ECB932672" type="female">5 females</specimenCount>
&quot;
<collectingCountry id="68423CB730AE2F5381AEFF4C6A56884B" name="Indonesia">Indonesia</collectingCountry>
(1700B):
<collectingRegion id="2285F49E447DC88F5FBD553840EA1BA6" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="F1036E62FA48332BFC68FAFCBEFFF844" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:F1036E62FA48332BFC68FAFCBEFFF844" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="65EDD2C173C562D7F9652817C5F673BD" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="81C84CF020D576F27E602E3B4403A5E0" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="F65A6D2CD19A433E060F1137F335EA27" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="DDDCD48DA6C2ECD707F9C499DECC5535" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="EB2A93FF8094BE96D3A9F4F01326BFAB" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="2E42B78264559AD3E3AEB4CE35DCD9BF" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode id="21140AB958AC4EF0567850F161957727" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="ED93911DAAD6AEEE2B0F6C961F483E10" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="427260592E2CD294D4C6AB8E1EC51868" ID-GBIF-Occurrence="3062138308" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="318F4566D35B9949B3869256C0B1F5F3" type="male">2 males</specimenCount>
,
<specimenCount id="0DC16B3B6AB770B7A0A9B8FF257C2A39" type="female">3 females</specimenCount>
&quot;
<collectingCountry id="2335B19F6688FED19C0E2E434F6C4F8F" name="Indonesia">Indonesia</collectingCountry>
(1700D):
<collectingRegion id="1673E0A809B3F7054C72473F946CC53F" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="38026C8BB9DDE9397DA80D50519986DD" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:38026C8BB9DDE9397DA80D50519986DD" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="603AFDCCF704A1A884A4C8AF01ACE7F6" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="1CDC6F90D33C27C3DCDBB1D9B648FFEB" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="042BDC9E0C8061405115EA83343427E2" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="A2F112DB4108DD00CA98FFEF72821362" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="7C73A84A73ABD602B48BCB17F6B24291" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="2533DEF4ABF60B74F389D7169CFFC1EE" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode id="E529275B046DD42B240671C489FED132" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B1DCFB8EF004F05CCC0A1C414CA70D42" ID-GBIF-Occurrence="3062138311" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="23" specimenCount-female="10" specimenCount-male="13" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="55330222DD35444936B6E2C9A8FE2399" type="male">13 males</specimenCount>
,
<specimenCount id="62D6D7792EED42B7D82F1DE35831F442" type="female">10 females</specimenCount>
&quot;
<collectingCountry id="B28B3806B87E10C2FED332D4315D5CE6" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="60565B1D73BE05223F4C45BA41712B48" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="0DAE6C6B38184A1EFFEB2F94B837BA1B" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:0DAE6C6B38184A1EFFEB2F94B837BA1B" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity id="2722A8ED31A4357016782CB5B080203E" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="7EB1B6558A6C4C45D7D3CA022FF29FE6" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date id="1EA950F1CD8988C1A18894866379EE94" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="375D9E4BB15792C92FD9BB4912F85DCF" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,&quot;, &quot;
<geoCoordinate id="5247171CAB34CD4E0531F5AA9CF1B534" degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate id="BE26945302B8A66C2A6DE87E038A791C" degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot;, two females with additional green text labels
<normalizedToken id="B09D622823E1286562FD073917162FBC" originalValue="“7357”">&quot;7357&quot;</normalizedToken>
and
<normalizedToken id="B5AA6EC161B39AB4ADFCCFA97CE33C54" originalValue="“7358”">&quot;7358&quot;</normalizedToken>
(
<collectionCode id="E885D157529A130DF2C4C4B013BDA845" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="D143C71D79B585AC1E7E675DD89B06AF" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E6BD41180497CDE179B3F76BAE11D412" ID-GBIF-Occurrence="3062138327" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1600" latitude="-2.561006" location="Foja Mountains" longLatPrecision="1" longitude="138.71149" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="3BA98CF685EDA9F6D46A5488C74936A9" type="male">2 males</specimenCount>
,
<specimenCount id="E99C120E5279EACAE9DAA4E88DEEEF96" type="female">4 females</specimenCount>
&quot;
<collectingCountry id="7473F427E52336ACA14F7A3F2D0B00D1" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="8039FD662CFFB50C265A855E032B67EE" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="9BA00C217927AE911629CA60C6BF64CA" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:9BA00C217927AE911629CA60C6BF64CA" country="Indonesia" latitude="-2.561006" longLatPrecision="1" longitude="138.71149" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, river camp,
<quantity id="73310D89D641EB2335CE9E2FD6F7C453" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="ED4C847B86E86244B6D95BDFB2F52978" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="D0B413AE0D1BCA9318AFA4ED0C86DBAA" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="B90DEDC374D2727F1B970E3E3480EB30" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="6ADDEC22F310FA98CF7CB900C188A0D2" degrees="2.561006" direction="south" orientation="latitude" precision="1" value="-2.561006">-2.561006</geoCoordinate>
<geoCoordinate id="5F11E0418E007D54914DAAD27E26BC2C" degrees="138.711487" direction="east" orientation="longitude" precision="1" value="138.71149">138.711487</geoCoordinate>
, Sumoked (Pap059)&quot;, one male with an additional handwritten label &quot;forest near bog camp&quot;, the other male with an additional green text label
<normalizedToken id="E2B55879AB03361BBD7C3A9A6F1785C3" originalValue="“7286”">&quot;7286&quot;</normalizedToken>
(
<collectionCode id="662AFBD15052CCD3164BB0ADBCAA6B47" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="912D676E7C664483C27F9C8BDAC30AC0" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9CAB1C735FE10BF3DB064696BA9E4C92" pageId="0" pageNumber="45" type="description">
<paragraph id="23A1437E4081318FBF1627180E46535C" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="0B2473CAF38B267415970AE517BC730D" pageId="0" pageNumber="45">
<emphasis id="5D5F5C9EA1961749ED43BE189210127C" bold="true" pageId="0" pageNumber="45">
<emphasis id="560D59A52FAB039218E447CEC790C7F2" italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small: TL-H
<quantity id="963EB5FD7F29A26C748C7DD19B917178" metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.85" metricValueMin="3.3" unit="mm" value="3.575" valueMax="3.85" valueMin="3.3">3.30-3.85 mm</quantity>
, TL
<quantity id="CF3ECB56F1E039D779F687EC7627AABE" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.70-4.30 mm</quantity>
, MW
<quantity id="930495D7DB70295397D7DBBF604D8631" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.80-2.10 mm</quantity>
(
<typeStatus id="9B530C536CD0B98C04FAADD31241F347">holotype</typeStatus>
: TL-H
<quantity id="E0B93C4C2D0D152DF68C714EBC0B60D7" metricMagnitude="-3" metricUnit="m" metricValue="3.85" unit="mm" value="3.85">3.85 mm</quantity>
, TL
<quantity id="6C1727CDF5E9D520B29F2DF3AE17D112" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.30 mm</quantity>
,
<collectionCode id="01A2D6D1939F9B61593A845A7C2994FE" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity id="1BBD3891DD092DC09BBD5DB68DCCABBB" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.10 mm</quantity>
), with oblong-oval habitus (Fig.
<figureCitation id="95D162528844B0FC128D1C958C8255A4" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">1</figureCitation>
).
</paragraph>
<paragraph id="72B8A18C3645B0B291151E3800656C87" pageId="0" pageNumber="45">
<emphasis id="212623975A98AFBAAE438B6BC68BC41C" bold="true" pageId="0" pageNumber="45">
<emphasis id="C8D41B61B759B08B1868092EFCF08F18" italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally dark brown to piceous, with paler head and sides of pronotum (Fig.
<figureCitation id="B9A17B92459A9962AE549EDE56B3B5EC" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">1</figureCitation>
). Head more or less uniformly dark brown to reddish brown, darker around eyes, or slightly paler anteriorly; pronotum dark brown to piceous on disc and distinctly paler (to yellowish red) anteriorly, posteriorly, and especially laterally; dark area on disc sometimes represented just as median band; elytra dark brown to piceous, with reddish brown sutural lines; head appendages and legs yellowish red to reddish brown. Teneral specimens paler.
</paragraph>
<caption id="2B81681EC08578E7291228884C4BD28A" doi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45" start="Figures 1–4" startId="F1">
<paragraph id="DC4F031B8D42B80210CDD554659389A0" pageId="0" pageNumber="45">
<emphasis id="7980FF656EFA95186005A291C9C037C0" bold="true" pageId="0" pageNumber="45">Figures 1-4.</emphasis>
Habitus and colouration of
<typeStatus id="18C7D5480E1634D0727820F2D81A9786">holotype</typeStatus>
<emphasis id="07AA19C97E57672245F6C601CB46FD61" bold="true" pageId="0" pageNumber="45">1</emphasis>
<taxonomicName id="9616424B2ECF39461E62A13409B0AD2A" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja">
<emphasis id="7C90543AF941ECE8B74AE4EF7A84010E" italics="true" pageId="0" pageNumber="45">Exocelina foja</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="678C2D43D676D25742C0257B5D25938E" bold="true" pageId="0" pageNumber="45">2</emphasis>
<taxonomicName id="8B7F8EB4C69592E8ECCA910E0BD5A51D" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. waaf" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="waaf">
<emphasis id="F1F7384F8FEF5142B5CF3C1DDBDC76DB" italics="true" pageId="0" pageNumber="45">E. waaf</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="5A2D529DAF46A478E79667A2E4BA08C9" bold="true" pageId="0" pageNumber="45">3</emphasis>
<taxonomicName id="92062A03E06E01F5E475E49314B52A59" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai">
<emphasis id="7148732B214EEF21122A443ED7FFD838" italics="true" pageId="0" pageNumber="45">E. riberai</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="94824816B2C0FACD7CF550B1CAE566D0" bold="true" pageId="0" pageNumber="45">4</emphasis>
<taxonomicName id="28ABA64292942E52312BE8AA345A47F6" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis id="45FCE2484CF8A4373FFE47B2567B2BC7" italics="true" pageId="0" pageNumber="45">E. apistefti</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph id="670EC7DAB495F38E34D2717EDB024CFB" pageId="0" pageNumber="45">
<emphasis id="B9F2CF179402B835133E7F188AE63FCE" bold="true" pageId="0" pageNumber="45">
<emphasis id="E2E145131583C300CE3E49848467A26B" italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with fine punctation and microreticulation. Head with dense and coarse punctation (spaces between punctures 0-3 times size of punctures), distinctly finer and sparser anteriorly and posteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with distinctly finer and sparser punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Pronotum and elytra with weakly impressed microreticulation; head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and fine sparse punctation, coarser and denser on two terminal abdominal ventrites.
</paragraph>
<paragraph id="B692E7F6CE6BCE90089DA6437B131232" pageId="0" pageNumber="45">
<emphasis id="2BE2678E7E96389553246CAAEE2A0BE0" bold="true" pageId="0" pageNumber="45">
<emphasis id="3DD9BD546DB7700BA9BCA1A73F567BF1" italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex medially, with distinct bead and few setae. Abdominal ventrite 6 broadly rounded apically.
</paragraph>
<paragraph id="945F8BF05B8A21C5137D58A8BCF20B33" pageId="0" pageNumber="45">
<emphasis id="8F34EF5D85C9E96BE86ADB772ED6D778" bold="true" pageId="0" pageNumber="45">
<emphasis id="6F076219010D1A7723C25A069301A969" italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of eleven and posterior row of six short setae (Fig.
<figureCitation id="40B6BF0663F169E25623BA865D8801EC" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex broad, curved downwards, and sharply pointed at tip; in ventral view, with distinct submedian constriction, distal part narrower than proximal one, apex truncate (Fig.
<figureCitation id="DF0BF2F58E2DDE7D3EE42147BDA48AA4" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5B, C</figureCitation>
). Paramere with strong notch on dorsal side and subdistal part relatively large and elongate; subdistal setae very few, dense and flattened: three upper longer, thinner, curved at apex and four lower shorter, almost straight, thicker; proximal setae hair-like, numerous, dense, but distinctly more inconspicuous than subdistal ones (Fig.
<figureCitation id="D785A3EE276E7336D06A6F2D1FD18E6A" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with 5-9 lateral striae on each side.
</paragraph>
<caption id="8B866637C0D5F10B48B80F6DFF537D8F" doi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45" start="Figures 5, 6" startId="F2">
<paragraph id="3F0BEFB84C2F43B2675244CE63491FE7" pageId="0" pageNumber="45">
<emphasis id="57C88F93EDC829A110014479A33813B0" bold="true" pageId="0" pageNumber="45">Figures 5, 6.</emphasis>
<emphasis id="8F49A156D95CE43390DF373FB1D1365F" bold="true" pageId="0" pageNumber="45">5</emphasis>
<taxonomicName id="7543D4D4630BCF1A54F9A21DE349BD7D" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja">
<emphasis id="DF8D80DC4EB8EBC9D679E3EC57F54E64" italics="true" pageId="0" pageNumber="45">Exocelina foja</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus id="F89262ED9959FA20253CA7245303A0EB">holotype</typeStatus>
<emphasis id="BB4CFBD4D349C604E313C84AFAAED901" bold="true" pageId="0" pageNumber="45">6</emphasis>
<taxonomicName id="C971E49A91A7ED2425BA35DF9416A8A3" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. waaf" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="waaf">
<emphasis id="4537459B456697D7E70D1DF08C7D84D0" italics="true" pageId="0" pageNumber="45">E. waaf</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus id="453AB3A52023D9F2A17A00C79E278155">holotype</typeStatus>
<emphasis id="264952057F7E7A419A39525F6B3E31AA" bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and
<quantity id="536FFF18F13221B7B558E4DD0F87B1CB" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view
<emphasis id="0F473E97999ADF3883AB8883C802AA39" bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view
<emphasis id="3E0A53DAC0F5AEF39786734FF2CD6298" bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view
<emphasis id="D68E52C6B7DBA425C84CF7104C88A190" bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph id="2B53752C3F5B1680F04DBA908EE1B8D7" pageId="0" pageNumber="45">
<emphasis id="E526A0ADD4A66494605FEF305917D57D" bold="true" pageId="0" pageNumber="45">
<emphasis id="CDDBB53DEC615EA8AFF2C9D7FA810FE3" italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without striae.
</paragraph>
</subSubSection>
<subSubSection id="5FD909A8BF1563EEFB733BD15E17CA4B" pageId="0" pageNumber="45" type="affinities">
<paragraph id="E4744BBB3C10D8E5AA1D91DD607B765E" pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph id="5B8044FB426AFAD008C95EF9C84BCB47" pageId="0" pageNumber="45">
The species evidently belongs to the
<taxonomicName id="33152683808E6E140A80121A2497D86C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis id="9E9C7818D1FED987834D7FB68EB54F36" italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. Within the group, it can be placed close to
<taxonomicName id="A0B5F0EC962AAC8F7A4AEA153EDB2DB9" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. oceai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oceai">
<emphasis id="947AB7AF8112CA9F49EF42869C002F89" italics="true" pageId="0" pageNumber="45">E. oceai</emphasis>
</taxonomicName>
<bibRefCitation id="4895D37E29EA7A7860AE84536B031C7A" DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
due to presence of the pronotal lateral bead. Shape of its median lobe is similar to that of
<taxonomicName id="5C4703EC712038EE4489625E945E02E0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pseudosoppi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pseudosoppi">
<emphasis id="91BC795433006676250C14A8A2770C29" italics="true" pageId="0" pageNumber="45">E. pseudosoppi</emphasis>
</taxonomicName>
<bibRefCitation id="10CD21AC0FBAF1B70290ECBCAC40693A" DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
; setation of the paramere is very characteristic.
</paragraph>
</subSubSection>
<subSubSection id="727A7AC361344A4B511AF2A2BC8D6049" pageId="0" pageNumber="45" type="distribution">
<paragraph id="BF65FC35AD198AB97DD00DEB1FE69D17" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="AE9E903CC4E53BBE70F81C6E43702B64" pageId="0" pageNumber="45">
<materialsCitation id="CE7CE28B5BF60678C62239F449860E78" ID-GBIF-Occurrence="3062138325" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry id="89ACACE0DDD533FA4FC257E529FF667B" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="7CDD42FA161208A67A5B747B5AA0C745" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
.
<collectingMunicipality id="1E36D7912F80ADDC45479E495EB6D8A4">This</collectingMunicipality>
species is known only from
<location id="BB3F48F11C533ADABE8DABBB08F53A26" LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:BB3F48F11C533ADABE8DABBB08F53A26" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, from and near the
<typeStatus id="C6179684EED7877A9D9D0F2B8AFB7FF2">type</typeStatus>
locality (Fig.
<figureCitation id="C438C3162FFCC9D4F16A6D8499D57651" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="748E169AEB7763D8541DBABCE125900D" pageId="0" pageNumber="45" type="habitat">
<paragraph id="D396A720712AA4C40A0CDDA3F6A930A1" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="B7E1E6EDB4DE8ADB9B23F050B39B97A5" pageId="0" pageNumber="45">The specimens were collected from small, shallow forest creeks.</paragraph>
</subSubSection>
<subSubSection id="D917F08700C38B9B0124B5CAEC185E0E" pageId="0" pageNumber="45" type="etymology">
<paragraph id="D9F4D39FDC97494E01C0FE97CFBF0E5A" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="5E294B8375138C241B403CEE99C60AC2" pageId="0" pageNumber="45">The species is named after Foja Mountains. The name is a noun in the nominative singular standing in apposition.</paragraph>
</subSubSection>
</treatment>
</document>
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