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<document id="B0E8586FDDCC41D6A06158460D5C0312" ID-CLB-Dataset="26014" ID-DOI="10.11646/zootaxa.4656.3.11" ID-GBIF-Dataset="99739ca8-be58-4843-aa6f-27a35bafe27a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3368943" ID-ZooBank="C465C40A-47AB-40D3-A7E6-E7B49FCAB131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565860060553" checkinUser="plazi" docAuthor="Quah, Evan S. H., Grismer, L. Lee, Wood, Perry L., Thura, Myint Kyaw, Oaks, Jamie R. &amp; Lin, Aung" docDate="2019" docId="B44887B9FFD5FFDCFF38FD40AFDFFD2B" docLanguage="en" docName="zootaxa.4656.3.11.pdf" docOrigin="Zootaxa 4656 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ansonia kyaiktiyoensis Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin 2019, sp. nov." docType="treatment" docUuid="34555D1C-97E2-41BD-8552-68347BFCCC3F" docUuidSource="ZooBank" docVersion="10" lastPageNumber="561" masterDocId="4871FFC1FFDFFFCCFFAFFFC4AE18FFD8" masterDocTitle="Discovery of the westernmost population of the genus Ansonia Stoliczka (Anura, Bufonidae) with the description of a new species from the Shan Plateau of eastern Myanmar" masterLastPageNumber="571" masterPageNumber="545" pageNumber="555" updateTime="1698751169668" updateUser="ExternalLinkService">
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<mods:title id="BE7F5E870878B90C9D819B80DC4A3A63">Discovery of the westernmost population of the genus Ansonia Stoliczka (Anura, Bufonidae) with the description of a new species from the Shan Plateau of eastern Myanmar</mods:title>
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<mods:namePart id="3E39687309FB26E89D90331666AEB38E">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="4C4AA5AEDD0CDD97D51E8988A9F49CCC">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="6C57F1203370DA8CC0827EB8C7682D1C">Thura, Myint Kyaw</mods:namePart>
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<mods:namePart id="FA8807E56B9746AB32C42C5584321A51">Oaks, Jamie R.</mods:namePart>
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<mods:namePart id="A9B0C33A26431E16E2D074EFA7A802E5">Lin, Aung</mods:namePart>
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<emphasis id="0E95EABDFFD5FFC6FF38FD40AFB5FD46" bold="true" box="[151,429,643,670]" italics="true" pageId="10" pageNumber="555">Ansonia kyaiktiyoensis</emphasis>
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<paragraph id="3C5E36AFFFD5FFC6FF38FD6CAD3FFD1A" blockId="10.[151,963,643,771]" box="[151,807,680,706]" pageId="10" pageNumber="555">Suggested common name: The Golden Rock Slender Toad</paragraph>
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, 
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&amp; 
<figureCitation id="A4DA2A2AFFD5FFC6FE82FD08AF48FD3E" box="[301,336,716,742]" captionStart="FIGURE 4" captionStartId="13.[151,250,1871,1896]" captionTargetBox="[151,1435,345,1852]" captionTargetId="figure@13.[151,1435,345,1852]" captionTargetPageId="13" captionText="FIGURE 4. (A) Calling adult male Ansonia inthanon from Doi Inthanon, Chiang Mai, Thailand. (B) Close-up of head of para- type of Ansonia kyaiktiyoensis sp. nov. (LSUHC 14067). Note the markings on the snout and under the eye as indicated by the arrows. (C) Close-up of head of A. inthanon from Doi Inthanon, Chiang Mai, Thailand. Note the absence of markings on the head. (Photographs by Ian Dugdale [A &amp; C] and Evan Quah [B])." httpUri="https://zenodo.org/record/3368953/files/figure.png" pageId="10" pageNumber="555">4B</figureCitation>
.
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.
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female (
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) collected on 
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4 November 
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in proximity of a stream below the 
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, 
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, 
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(
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, 
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, 
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in elevation) by 
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, 
<collectorName id="91145379FFD5FFC6FAF7FC97AF52FC49" pageId="10" pageNumber="555">Myint Kyaw Thura, L.</collectorName>
Lee Grismer, 
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, 
<collectorName id="91145379FFD5FFC6FD34FCB3AD4BFC48" box="[667,851,886,913]" pageId="10" pageNumber="555">Evan S.H. Quah</collectorName>
, and 
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</paragraph>
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<materialsCitation id="8C893CF2FFD5FFC6FF68FC5EAACBFC6C" ID-GBIF-Occurrence="2332606851" box="[199,1235,922,949]" collectingDate="2018-11-04" collectionCode="LSUHC" collectorName="Perry L. Wood, Jr. &amp; Myint Kyaw Thura, L. &amp; Jamie R. Oaks &amp; Evan S. H. Quah &amp; Aung Lin." country="Myanmar" county="Mon State" elevation="1033" latitude="97.09938" location="Adult" longLatPrecision="1" longitude="17.478863" municipality="Kyaiktiyo Pagoda" pageId="10" pageNumber="555" specimenCode="LSUHC 14067" specimenCount="1" typeStatus="paratype">
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<typeStatus id="E35A880DFFD5FFC6FF68FC5EAF2CFC6C" box="[199,308,922,948]" pageId="10" pageNumber="555" type="paratype">Paratype</typeStatus>
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. Sub-adult male (
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) bearing the same collection data as the holotype
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.
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<emphasis id="0E95EABDFFD5FFC6FF68FC78AF5AFC0F" bold="true" box="[199,322,956,983]" pageId="10" pageNumber="555">Diagnosis.</emphasis>
<taxonomicName id="FBE14D2CFFD5FFC6FEE6FC79AC51FC0F" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[329,585,957,983]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="555" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD5FFC6FEE6FC79AC51FC0F" box="[329,585,957,983]" italics="true" pageId="10" pageNumber="555">Ansonia kyaiktiyoensis</emphasis>
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<taxonomicNameLabel id="15A657C6FFD5FFC6FDFEFC79ACB3FC0F" box="[593,683,957,983]" pageId="10" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
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is recognised as a member of the genus 
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<emphasis id="0E95EABDFFD5FFC6FBD8FC79AACBFC0F" box="[1143,1235,957,983]" italics="true" pageId="10" pageNumber="555">Ansonia</emphasis>
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based on the molecular phylogenetic analyses (
<figureCitation id="A4DA2A2AFFD5FFC6FE42FC24AC34FC22" box="[493,556,992,1018]" captionStart="FIGURE 1" captionStartId="9.[151,250,1895,1920]" captionTargetBox="[341,1232,174,1873]" captionTargetId="figure@9.[336,1251,166,1875]" captionTargetPageId="9" captionText="FIGURE 1. Maximum likelihood (ML) phylogram for species of the genus Ansonia. Nodal support is indicated by Bayesian posterior probability (BPP) / Ultrafast bootstrap (UFBOOT). Black circles represent nodes supported by a BPP and UFB of 1.00 and 100, respectively. Blue node indicates the Thai-Malay Peninsula clade and red node indicates the Thai-Burmese clade. Species outside of the Thai-Malay Peninsula clade are represented by single tip labels for their lineages." httpUri="https://zenodo.org/record/3368945/files/figure.png" pageId="10" pageNumber="555">Fig. 1</figureCitation>
) and a combination of the following morphological characters: small body size (maximum 
<collectionCode id="5AF0AE6AFFD5FFC6FEB5FBC7AF56FBC5" box="[282,334,1027,1053]" pageId="10" pageNumber="555">SVL</collectionCode>
24.0 mm in males and females); long slender limbs bearing long slender digits with bulbous tips; absence of parotoid glands; weak subarticular tubercles; and membranous foot webbing (
<bibRefCitation id="58704B5EFFD5FFC6FB23FBE2AB57FB99" author="Chan, K. O. &amp; Wood, Jr., P. L. &amp; Anuar, S. &amp; Muin, M. A. &amp; Quah, E. S. H. &amp; Sumarli, A. X. Y. &amp; Grismer, L. L." box="[1164,1359,1062,1089]" pageId="10" pageNumber="555" pagination="427 - 440" refId="ref14958" refString="Chan, K. O., Wood, Jr., P. L., Anuar, S., Muin, M. A., Quah, E. S. H., Sumarli, A. X. Y. &amp; Grismer, L. L. (2014) A new species of upland Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from northeastern Peninsular Malaysia. Zootaxa, 3764 (4), 427 - 440. http: // doi. org / 10.11646 / zootaxa. 3764.4. 3" type="journal article" year="2014">
Chan 
<emphasis id="0E95EABDFFD5FFC6FB7EFBE3AB1CFB99" box="[1233,1284,1063,1089]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2014
</bibRefCitation>
; 
<bibRefCitation id="58704B5EFFD5FFC6FAF4FBE2AF0DFBBC" author="Davis, H. R. &amp; Grismer, L. L. &amp; Klabacka, R. L. &amp; Muin, M. A. M. &amp; Quah, E. S. H. &amp; Anuar, S. &amp; Wood, Jr., P. L. &amp; Sites, Jr., J. W." pageId="10" pageNumber="555" pagination="137 - 153" refId="ref15122" refString="Davis, H. R., Grismer, L. L., Klabacka, R. L., Muin, M. A. M., Quah, E. S. H., Anuar, S., Wood, Jr., P. L. &amp; Sites, Jr., J. W. (2016) The phylogenetic relationships of a new Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from a montane region in Peninsular Malaysia. Zootaxa, 4103 (2), 137 - 153. https: // doi. org / 10.11646 / zootaxa. 4103.2. 4" type="journal article" year="2016">
Davis 
<emphasis id="0E95EABDFFD5FFC6FF38FB8FAED2FBBC" box="[151,202,1098,1124]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2016
</bibRefCitation>
; 
<bibRefCitation id="58704B5EFFD5FFC6FE8EFB8EAC1BFBBC" author="Grismer, L. L. &amp; Wood, Jr., P. L. &amp; Aowphol, A. &amp; Cota, M. &amp; Grismer, M. S. &amp; Murdoch, M. &amp; Aguilar, C. &amp; Grismer, J. L." box="[289,515,1098,1124]" pageId="10" pageNumber="555" pagination="371 - 395" refId="ref15709" refString="Grismer, L. L., Wood, Jr., P. L., Aowphol, A., Cota, M., Grismer, M. S., Murdoch, M., Aguilar, C. &amp; Grismer, J. L. (2016) Out of Borneo, again and again: biogeography of the Stream Toad genus Ansonia Stoliczka (Anura: Bufonidae) and the discovery of the first limestone cave-dwelling species. Biological Journal of the Linnean Society, 120, 371 - 395. https: // doi. org / 10.1111 / bij. 1288 6" type="journal article" year="2016">
Grismer 
<emphasis id="0E95EABDFFD5FFC6FE2AFB8FAFA0FBBC" box="[389,440,1098,1124]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2016
</bibRefCitation>
; 
<bibRefCitation id="58704B5EFFD5FFC6FDA0FB8EAC96FBBC" author="Inger, R. F." box="[527,654,1098,1124]" pageId="10" pageNumber="555" pagination="473 - 503" refId="ref17010" refString="Inger, R. F. (1960) A review of the oriental toads of the genus Ansonia Stoliczka. Fieldiana. Zoology, 39, 473 - 503. http: // doi. org / 10.5962 / bhl. title. 269 7" type="journal article" year="1960">Inger 1960</bibRefCitation>
, 
<bibRefCitation id="58704B5EFFD5FFC6FD35FB8EACCDFBBC" author="Inger, R. F." box="[666,725,1098,1124]" pageId="10" pageNumber="555" pagination="1 - 402" refId="ref17056" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana. Zoology, 52, 1 - 402. https: // doi. org / 10.5962 / bhl. title. 314 7" type="journal article" year="1966">1966</bibRefCitation>
, 
<bibRefCitation id="58704B5EFFD5FFC6FD4EFB8EAD04FBBC" author="Inger, R. F." box="[737,796,1098,1124]" pageId="10" pageNumber="555" pagination="225 - 237" refId="ref17100" refString="Inger, R. F. (1992) Variation of apomorphic characters in stream-dwelling tadpoles of the bufonid genus Ansonia (Amphibia: Anura). Zoological Journal of the Linnaean Society, 105, 225 - 237. http: // doi. org / 10.1111 / j. 1096 - 3642.1992. tb 01230. x" type="journal article" year="1992">1992</bibRefCitation>
; 
<bibRefCitation id="58704B5EFFD5FFC6FC86FB8EAA3CFBBC" author="Wilkinson, J. A. &amp; Sellas A. B. &amp; Vindum, J." box="[809,1060,1098,1124]" pageId="10" pageNumber="555" pagination="54 - 68" refId="ref19225" refString="Wilkinson, J. A., Sellas A. B. &amp; Vindum, J. (2012) A new species of Ansonia (Anura: Bufonidae) from northern Tanintharyi Division, Myanmar. Zootaxa, 3163, 54 - 68. https: // doi. org / 10.11646 / zootaxa. 3163.1. 3" type="journal article" year="2012">
Wilkinson 
<emphasis id="0E95EABDFFD5FFC6FC0AFB8FADCFFBBC" box="[933,983,1098,1124]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2012
</bibRefCitation>
). It can be differentiated from all congeners by the following combination of characters: 
<collectionCode id="5AF0AE6AFFD5FFC6FD57FBA9AD34FB5F" box="[760,812,1133,1159]" pageId="10" pageNumber="555">SVL</collectionCode>
24.0 mm in the single male and female; snout projecting beyond lower jaw; tympanum visible; absence of interorbital and tarsal ridges; first finger shorter than second; finger and toe tips bulbous; approximately 1.5 phalanges free of web on fifth toe; presence of a yellow rictal tubercle at the angle of jaw; distinct dorsal tubercles; abdomen coarsely granular; absence of oblique flaps of skin on each side of vent; absence of wide, light-coloured patch below eye; presence of light-coloured interscapular spot; presence of large, discrete, bright-yellow submandibular spots; presence of markings on the snout consisting of streaks below the eye to the lip, and on the canthus rostralis to the nostril; dorsum grey-brown with a dark-brown, X-shaped marking on the back surrounding the interscapular spot; dark coloured markings on the rump; dorsum and flanks covered in orange-beige spots; a thin, dark dorsolateral stripe; iris yellow-gold; fore and hind limbs bearing orange-beige crossbars; and venter light-grey with yellow spots, especially near the flanks and the underside of the hind limbs.
</paragraph>
<paragraph id="3C5E36AFFFD5FFC6FF68FA08AB41F8D8" blockId="10.[151,1437,816,2039]" pageId="10" pageNumber="555">
<emphasis id="0E95EABDFFD5FFC6FF68FA08AFF9FA3F" bold="true" box="[199,481,1484,1511]" pageId="10" pageNumber="555">
Description of 
<typeStatus id="E35A880DFFD5FFC6FED5FA09AFF9FA3F" box="[378,481,1485,1511]" pageId="10" pageNumber="555" type="holotype">holotype</typeStatus>
</emphasis>
. Adult female, 
<collectionCode id="5AF0AE6AFFD5FFC6FD3CFA09ACDFFA3F" box="[659,711,1485,1511]" pageId="10" pageNumber="555">SVL</collectionCode>
24.0 mm; head longer than wide (HL/ 
<collectionCode id="5AF0AE6AFFD5FFC6FB2DFA09AAACFA3F" box="[1154,1204,1485,1511]" pageId="10" pageNumber="555">HW</collectionCode>
=1.04); snout shorter than wide (SL/ 
<collectionCode id="5AF0AE6AFFD5FFC6FE93FA34AF7FF9D2" box="[316,359,1520,1546]" pageId="10" pageNumber="555">SW</collectionCode>
=0.89), projecting beyond lower jaw, dorsally convex with a midline depression, truncate in dorsal view, truncate and sloping in lateral view; canthus rostralis distinct, lores vertical, slightly concave; nares open laterally just below canthus, nearly terminal on snout; distance between nares smaller than snout length (
<collectionCode id="5AF0AE6AFFD5FFC6FB66F9F2AAE3F989" box="[1225,1275,1590,1617]" country="USA" lsid="urn:lsid:biocol.org:col:15485" name="Indiana University" pageId="10" pageNumber="555" type="Herbarium">IND</collectionCode>
/SL=0.62) and snout width (
<collectionCode id="5AF0AE6AFFD5FFC6FE85F99DAF45F9AC" box="[298,349,1625,1652]" country="USA" lsid="urn:lsid:biocol.org:col:15485" name="Indiana University" pageId="10" pageNumber="555" type="Herbarium">IND</collectionCode>
/ 
<collectionCode id="5AF0AE6AFFD5FFC6FEC9F99DAF89F9AC" box="[358,401,1625,1652]" pageId="10" pageNumber="555">SW</collectionCode>
=0.55); eyes large, slightly protruding beyond labials in dorsal view, diameter less than snout length (
<collectionCode id="5AF0AE6AFFD5FFC6FF41F9B9AF0BF940" box="[238,275,1661,1688]" pageId="10" pageNumber="555">ED</collectionCode>
/SL=0.64) and interorbital distance (
<collectionCode id="5AF0AE6AFFD5FFC6FD1EF9B9ACCFF940" box="[689,727,1661,1688]" pageId="10" pageNumber="555">ED</collectionCode>
/ 
<collectionCode id="5AF0AE6AFFD5FFC6FD70F9B9AD0AF940" box="[735,786,1661,1688]" pageId="10" pageNumber="555">IOD</collectionCode>
=0.76); pupils horizontal; interorbital region flat, distance smaller than snout width (
<collectionCode id="5AF0AE6AFFD5FFC6FE1DF95BAFFCF962" box="[434,484,1695,1722]" pageId="10" pageNumber="555">IOD</collectionCode>
/ 
<collectionCode id="5AF0AE6AFFD5FFC6FE42F95BAC0FF962" box="[493,535,1695,1722]" pageId="10" pageNumber="555">SW</collectionCode>
=0.75) and snout length (
<collectionCode id="5AF0AE6AFFD5FFC6FC89F95BAD41F962" box="[806,857,1695,1722]" pageId="10" pageNumber="555">IOD</collectionCode>
/SL=0.85); tympanum distinct, oval, taller than wide, with horizontal axis less than eye diameter (
<collectionCode id="5AF0AE6AFFD5FFC6FD2AF907ACB3F905" box="[645,683,1731,1757]" pageId="10" pageNumber="555">TD</collectionCode>
/ 
<collectionCode id="5AF0AE6AFFD5FFC6FD1BF907ACC2F905" box="[692,730,1731,1757]" pageId="10" pageNumber="555">ED</collectionCode>
=0.39); choanae subcircular, separated by a distance larger than their diameter; vomerine ridge and teeth absent; tongue narrow, ending in median point, posterior one-half free.
</paragraph>
<paragraph id="3C5E36AFFFD5FFC6FF68F8CEAA9CF82F" blockId="10.[151,1437,816,2039]" pageId="10" pageNumber="555">
Forelimbs and fingers long and slender; finger length from shortest to longest: I&lt;II&lt;IV&lt;III; basal webbing not extending beyond proximal subarticular tubercle; fingertips bulbous, slightly dilated but not forming discs; subarticular tubercles indistinct; inner and outer metacarpal tubercles distinct, oval, raised, inner smaller than outer; supernumerary tubercles absent. Hind limbs and toes long and slender (CL/ 
<collectionCode id="5AF0AE6AFFD5FFC6FC49F8B7AA04F855" box="[998,1052,1907,1933]" pageId="10" pageNumber="555">SVL</collectionCode>
=0.42), foot shorter than tibia (
<collectionCode id="5AF0AE6AFFD5FFC6FADDF8B7AB8CF855" box="[1394,1428,1907,1933]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="555" type="Museum">PL</collectionCode>
/ CL=0.75); toe length from shortest to longest: I&lt;II&lt;V&lt;III&lt;IV; webbing formula: I ½–1.5 II ½– 
<date id="485F106FFFD5FFC6FB10F852AAF5F869" box="[1215,1261,1942,1969]" pageId="10" pageNumber="555">3 III</date>
1.5– 
<date id="485F106FFFD5FFC6FA89F852AB4EF869" box="[1318,1366,1942,1969]" pageId="10" pageNumber="555">3 IV</date>
3–1.5 V; toe tips bulbous, slightly dilated but not forming discs; subarticular tubercles indistinct; inner metatarsal tubercle oval, slightly raised; outer metatarsal tubercle raised, rounded, slightly smaller than inner.
</paragraph>
<paragraph id="3C5E36AFFFD4FFC7FF68FF5DAFF7FDBB" blockId="11.[151,1437,153,611]" pageId="11" pageNumber="556">
Upper eyelid, interorbital region, dorsal part of snout and canthus covered with small, flat tubercles bearing brown, keratinized tips; interorbital ridges absent; few randomly dispersed tubercles present on lores; single row of small spinules on upper lip and outer margin of upper eyelid; rictal tubercle at the corner of the jaw; supratympanic fold and parotoid gland absent; back, flanks and dorsal part of limbs bearing irregularly spaced large and small tubercles bearing brown keratinized spinules, larger tubercles often have more than one spinule; concentration of larger tubercles above tympanum and in scapular region forming an indistinct dorsolateral row posterior to eye and tympanum, extending to insertion of hind limbs; a series of brown or white conical, keratinized tubercles along edges of underside of mandible that are most dense near snout and corners of jaw, becoming randomly distributed in gular region; abdomen coarsely granular; all ventral surfaces except for manus and pes covered with coarse, evenly spaced tubercles, and the tubercles are largest and most dense on abdominal region of flanks and underside of thigh, becoming more finely granular in pectoral and gular regions; ventral and lateral surfaces of limbs bearing randomly distributed large and small conical tubercles, most terminating in keratinized tips. All measurements of the 
<typeStatus id="E35A880DFFD4FFC7FAC3FDE1AB84FDE7" box="[1388,1436,549,575]" pageId="11" pageNumber="556">type</typeStatus>
series are presented in 
<tableCitation id="71630314FFD4FFC7FE39FD8DAFF3FDBB" box="[406,491,585,611]" captionStart="TABLE 4" captionStartId="14.[151,239,154,178]" captionTargetPageId="14" captionText="TABLE 4. Selected measurements of the type series of Ansonia kyaiktiyoensis sp. nov. / = no data available." httpUri="http://table.plazi.org/id/689E6627FFD1FFC2FF38FF5EAB38FF6A" pageId="11" pageNumber="556" tableUuid="689E6627FFD1FFC2FF38FF5EAB38FF6A">Table 4</tableCitation>
.
</paragraph>
<caption id="689E6627FFD4FFC7FF38F854AC4FF816" httpUri="https://zenodo.org/record/3368949/files/figure.png" pageId="11" pageNumber="556" startId="11.[151,250,1936,1961]" targetBox="[171,1415,633,1917]" targetPageId="11">
<paragraph id="3C5E36AFFFD4FFC7FF38F854AC4FF816" blockId="11.[151,1436,1936,1998]" pageId="11" pageNumber="556">
<emphasis id="0E95EABDFFD4FFC7FF38F854AF15F871" bold="true" box="[151,269,1936,1961]" pageId="11" pageNumber="556">FIGURE 2</emphasis>
. (A) Paratype of 
<taxonomicName id="FBE14D2CFFD4FFC7FE12F856ACBCF871" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[445,676,1937,1961]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="556" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD4FFC7FE12F856ACBCF871" box="[445,676,1937,1961]" italics="true" pageId="11" pageNumber="556">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD4FFC7FD04F855ACE3F871" bold="true" box="[683,763,1937,1961]" pageId="11" pageNumber="556">
<taxonomicNameLabel id="15A657C6FFD4FFC7FD04F855ACE3F871" box="[683,763,1937,1961]" pageId="11" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(LSUHC 14067). (B) Hill forest habitat of 
<taxonomicName id="FBE14D2CFFD4FFC7FB1AF856AB84F871" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[1205,1436,1937,1961]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="556" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD4FFC7FB1AF856AB84F871" box="[1205,1436,1937,1961]" italics="true" pageId="11" pageNumber="556">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD4FFC7FF38F871AEF0F815" bold="true" box="[151,232,1973,1997]" pageId="11" pageNumber="556">
<taxonomicNameLabel id="15A657C6FFD4FFC7FF38F871AEF0F815" box="[151,232,1973,1997]" pageId="11" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at Kyaiktiyo, Mon State, Myanmar.
</paragraph>
</caption>
<paragraph id="3C5E36AFFFD3FFC0FF68FF5CAB4BFDC3" blockId="12.[151,1437,152,647]" pageId="12" pageNumber="557">
<emphasis id="0E95EABDFFD3FFC0FF68FF5CAC40FF6B" bold="true" box="[199,600,152,179]" pageId="12" pageNumber="557">
Colouration in life (
<figureCitation id="A4DA2A2AFFD3FFC0FE18FF5CAC08FF6B" box="[439,528,152,179]" captionStart="FIGURE 2" captionStartId="11.[151,250,1936,1961]" captionTargetBox="[171,1415,633,1917]" captionTargetId="figure@11.[171,1415,633,1917]" captionTargetPageId="11" captionText="FIGURE 2. (A) Paratype of Ansonia kyaiktiyoensis sp. nov. (LSUHC 14067). (B) Hill forest habitat of Ansonia kyaiktiyoensis sp. nov. at Kyaiktiyo, Mon State, Myanmar." httpUri="https://zenodo.org/record/3368949/files/figure.png" pageId="12" pageNumber="557">Fig. 2A</figureCitation>
&amp; 
<figureCitation id="A4DA2A2AFFD3FFC0FD95FF5CAC52FF6B" box="[570,586,152,179]" captionStart="FIGURE 3" captionStartId="12.[151,250,1975,2000]" captionTargetBox="[173,1415,669,1956]" captionTargetId="figure@12.[173,1415,669,1956]" captionTargetPageId="12" captionText="FIGURE 3. Dorsum (A) and venter (B) of type series of Ansonia kyaiktiyoensis sp. nov. Paratype (LSUHC 14067) on the left and holotype (LSUHC 14066) on the right." httpUri="https://zenodo.org/record/3368951/files/figure.png" pageId="12" pageNumber="557">3</figureCitation>
).
</emphasis>
Dorsum and flanks grey-brown and peppered with orange-beige spots; a dark-brown, X-shaped marking on the back starting behind the eyes and the lower arms of the X bordering the light- coloured, orange-beige insterscapular spot; another dark-brown arched marking across the sacrum that joins with the dark, posterior border of the interscapular spot on the midline of the back; and another dark-coloured marking on the urostyle; a narrow dorsolateral stripes runs from the posterior corner of the eyes, above the tympanum and ap- proximately halfway down the body between the forelimb and hind limb insertions before arching along the flanks to the anterior margin of the hind limb at its insertion point on the body; tips of tubercles on the dorsum and flanks orange-beige; colour of the iris is yellow-gold; a dark-coloured streak extends from below the eye to the lips, and another from the canthus rostralis to the nostril; yellow rictal tubercle at the corner of the jaw present; fore and hind limbs with orange-beige crossbars; venter light-grey with yellow spots that are more prominent on the flanks and the underside of the thighs; large, discrete, bright-yellow submandibular spots along the underside of lower jaw.
</paragraph>
<paragraph id="3C5E36AFFFD3FFC0FF68FDE0AC31FD5F" blockId="12.[151,1437,152,647]" pageId="12" pageNumber="557">
<emphasis id="0E95EABDFFD3FFC0FF68FDE0AC05FDE7" bold="true" box="[199,541,548,575]" pageId="12" pageNumber="557">Colouration in preservative.</emphasis>
Dorsum and flanks grey with lighter grey interscapular spot and dull orange- brown tubercles; markings on back and flanks dark-grey; venter light-grey with the yellow spots of the venter and submandibular spots turning cream.
</paragraph>
<caption id="689E6627FFD3FFC0FF38F873AC55F82C" httpUri="https://zenodo.org/record/3368951/files/figure.png" pageId="12" pageNumber="557" startId="12.[151,250,1975,2000]" targetBox="[173,1415,669,1956]" targetPageId="12">
<paragraph id="3C5E36AFFFD3FFC0FF38F873AC55F82C" blockId="12.[151,1436,1975,2036]" pageId="12" pageNumber="557">
<emphasis id="0E95EABDFFD3FFC0FF38F873AF15F808" bold="true" box="[151,269,1975,2000]" pageId="12" pageNumber="557">FIGURE 3</emphasis>
. Dorsum (A) and venter (B) of type series of 
<taxonomicName id="FBE14D2CFFD3FFC0FD70F87CADDFF817" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[735,967,1975,1999]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="12" pageNumber="557" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD3FFC0FD70F87CADDFF817" box="[735,967,1975,1999]" italics="true" pageId="12" pageNumber="557">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD3FFC0FC62F87CAA06F808" bold="true" box="[973,1054,1976,2000]" pageId="12" pageNumber="557">
<taxonomicNameLabel id="15A657C6FFD3FFC0FC62F87CAA06F808" box="[973,1054,1976,2000]" pageId="12" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype (LSUHC 14067) on the left and holotype (LSUHC 14066) on the right.
</paragraph>
</caption>
<paragraph id="3C5E36AFFFD2FFC1FF68FF5CAFB1FE9B" blockId="13.[151,1437,152,323]" pageId="13" pageNumber="558">
<emphasis id="0E95EABDFFD2FFC1FF68FF5CAF82FF6B" bold="true" box="[199,410,152,179]" pageId="13" pageNumber="558">
Variation (
<figureCitation id="A4DA2A2AFFD2FFC1FEE9FF5CAF93FF6B" box="[326,395,152,179]" captionStart="FIGURE 3" captionStartId="12.[151,250,1975,2000]" captionTargetBox="[173,1415,669,1956]" captionTargetId="figure@12.[173,1415,669,1956]" captionTargetPageId="12" captionText="FIGURE 3. Dorsum (A) and venter (B) of type series of Ansonia kyaiktiyoensis sp. nov. Paratype (LSUHC 14067) on the left and holotype (LSUHC 14066) on the right." httpUri="https://zenodo.org/record/3368951/files/figure.png" pageId="13" pageNumber="558">Fig. 3</figureCitation>
).
</emphasis>
The 
<typeStatus id="E35A880DFFD2FFC1FE7FFF5DAC28FF6B" box="[464,560,153,179]" pageId="13" pageNumber="558" type="paratype">paratype</typeStatus>
closely resembles the 
<typeStatus id="E35A880DFFD2FFC1FC84FF5DAD95FF6B" box="[811,909,153,179]" pageId="13" pageNumber="558" type="holotype">holotype</typeStatus>
in most aspects of colouration and morphology except for having more prominent yellow spotting on the underside of the jaws and venter that forms a reticulated pattern on the flanks and underside of the thighs. Similar to the 
<typeStatus id="E35A880DFFD2FFC1FCFAFF25ADA2FF23" box="[853,954,225,251]" pageId="13" pageNumber="558" type="holotype">holotype</typeStatus>
, it has the prominent markings on the snout that consist of streaks below the eye to the lip, and on the canthus rostralis to the nostril (
<figureCitation id="A4DA2A2AFFD2FFC1FB3EFEC1AAFCFEC7" box="[1169,1252,261,287]" captionStart="FIGURE 4" captionStartId="13.[151,250,1871,1896]" captionTargetBox="[151,1435,345,1852]" captionTargetId="figure@13.[151,1435,345,1852]" captionTargetPageId="13" captionText="FIGURE 4. (A) Calling adult male Ansonia inthanon from Doi Inthanon, Chiang Mai, Thailand. (B) Close-up of head of para- type of Ansonia kyaiktiyoensis sp. nov. (LSUHC 14067). Note the markings on the snout and under the eye as indicated by the arrows. (C) Close-up of head of A. inthanon from Doi Inthanon, Chiang Mai, Thailand. Note the absence of markings on the head. (Photographs by Ian Dugdale [A &amp; C] and Evan Quah [B])." httpUri="https://zenodo.org/record/3368953/files/figure.png" pageId="13" pageNumber="558">Fig 4B</figureCitation>
). Measurements are presented in 
<tableCitation id="71630314FFD2FFC1FEE0FEEDAFBDFE9B" box="[335,421,297,323]" captionStart="TABLE 4" captionStartId="14.[151,239,154,178]" captionTargetPageId="14" captionText="TABLE 4. Selected measurements of the type series of Ansonia kyaiktiyoensis sp. nov. / = no data available." httpUri="http://table.plazi.org/id/689E6627FFD1FFC2FF38FF5EAB38FF6A" pageId="13" pageNumber="558" tableUuid="689E6627FFD1FFC2FF38FF5EAB38FF6A">Table 4</tableCitation>
.
</paragraph>
<caption id="689E6627FFD2FFC1FF38F88BAD28F80B" httpUri="https://zenodo.org/record/3368953/files/figure.png" pageId="13" pageNumber="558" startId="13.[151,250,1871,1896]" targetBox="[151,1435,345,1852]" targetPageId="13">
<paragraph id="3C5E36AFFFD2FFC1FF38F88BAD28F80B" blockId="13.[151,1436,1871,2004]" pageId="13" pageNumber="558">
<emphasis id="0E95EABDFFD2FFC1FF38F88BAF15F8B0" bold="true" box="[151,269,1871,1896]" pageId="13" pageNumber="558">FIGURE 4</emphasis>
. (A) Calling adult male 
<taxonomicName id="FBE14D2CFFD2FFC1FDACF894ACAFF8BF" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[515,695,1871,1895]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="13" pageNumber="558" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="0E95EABDFFD2FFC1FDACF894ACAFF8BF" box="[515,695,1871,1895]" italics="true" pageId="13" pageNumber="558">Ansonia inthanon</emphasis>
</taxonomicName>
from Doi Inthanon, Chiang Mai, Thailand. (B) Close-up of head of para- type of 
<taxonomicName id="FBE14D2CFFD2FFC1FF4AF8B0AFD4F853" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[229,460,1907,1931]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="13" pageNumber="558" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD2FFC1FF4AF8B0AFD4F853" box="[229,460,1907,1931]" italics="true" pageId="13" pageNumber="558">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD2FFC1FE7CF8B7AC3BF853" bold="true" box="[467,547,1907,1931]" pageId="13" pageNumber="558">
<taxonomicNameLabel id="15A657C6FFD2FFC1FE7CF8B7AC3BF853" box="[467,547,1907,1931]" pageId="13" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(LSUHC 14067). Note the markings on the snout and under the eye as indicated by the arrows. (C) Close-up of head of 
<taxonomicName id="FBE14D2CFFD2FFC1FE48F85CAC46F877" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[487,606,1943,1967]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="13" pageNumber="558" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="0E95EABDFFD2FFC1FE48F85CAC46F877" box="[487,606,1943,1967]" italics="true" pageId="13" pageNumber="558">A. inthanon</emphasis>
</taxonomicName>
from Doi Inthanon, Chiang Mai, Thailand. Note the absence of markings on the head. (Photographs by Ian Dugdale [A &amp; C] and Evan Quah [B]).
</paragraph>
</caption>
<caption id="689E6627FFD1FFC2FF38FF5EAB38FF6A" ID-Table-UUID="689E6627FFD1FFC2FF38FF5EAB38FF6A" box="[151,1312,154,178]" httpUri="http://table.plazi.org/id/689E6627FFD1FFC2FF38FF5EAB38FF6A" pageId="14" pageNumber="559" targetBox="[166,1379,190,1964]" targetIsTable="true" targetPageId="14">
<paragraph id="3C5E36AFFFD1FFC2FF38FF5EAB38FF6A" blockId="14.[151,1312,154,178]" box="[151,1312,154,178]" pageId="14" pageNumber="559">
<emphasis id="0E95EABDFFD1FFC2FF38FF5EAF1BFF6A" bold="true" box="[151,259,154,178]" pageId="14" pageNumber="559">TABLE 4</emphasis>
. Selected measurements of the type series of 
<taxonomicName id="FBE14D2CFFD1FFC2FD45FF5EADC6FF6A" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[746,990,154,178]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="559" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD1FFC2FD45FF5EADC6FF6A" box="[746,990,154,178]" italics="true" pageId="14" pageNumber="559">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD1FFC2FC4BFF5EAA22FF6A" bold="true" box="[996,1082,154,178]" pageId="14" pageNumber="559">
<taxonomicNameLabel id="15A657C6FFD1FFC2FC4BFF5EAA22FF6A" box="[996,1082,154,178]" pageId="14" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ = no data available.
</paragraph>
</caption>
<paragraph id="3C5E36AFFFD1FFC2FF09FF7AAB0EF874" pageId="14" pageNumber="559">
<table id="4EE1C40FFFD10033FF09FF7AAB7BF874" box="[166,1379,190,1964]" gridcols="3" gridrows="45" pageId="14" pageNumber="559">
<tr id="82D134EDFFD10033FF09FF7AAB7BFF0F" box="[166,1379,190,215]" gridrow="0" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FF7AAA08FF0F" box="[166,1040,190,215]" gridcol="0" gridrow="0" pageId="14" pageNumber="559">Specimen</th>
<th id="C1005D91FFD10033FB80FF7AAAD0FF0F" box="[1071,1224,190,215]" gridcol="1" gridrow="0" pageId="14" pageNumber="559">LSUHC 14066</th>
<th id="C1005D91FFD10033FB45FF7AAB7BFF0F" box="[1258,1379,190,215]" gridcol="2" gridrow="0" pageId="14" pageNumber="559">LSUHC</th>
</tr>
<tr id="82D134EDFFD10033FF09FF26AB7BFF23" box="[166,1379,226,251]" gridrow="1" pageId="14" pageNumber="559" rowspan-0="1">
<td id="C1005D91FFD10033FB80FF26AAD0FF23" box="[1071,1224,226,251]" gridcol="1" gridrow="1" pageId="14" pageNumber="559">(holotype)</td>
<td id="C1005D91FFD10033FB45FF26AB7BFF23" box="[1258,1379,226,251]" gridcol="2" gridrow="1" pageId="14" pageNumber="559">(paratype)</td>
</tr>
<tr id="82D134EDFFD10033FF09FEC7AB7BFEC4" box="[166,1379,259,284]" gridrow="2" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FEC7AA08FEC4" box="[166,1040,259,284]" gridcol="0" gridrow="2" pageId="14" pageNumber="559">Sex</th>
<td id="C1005D91FFD10033FB80FEC7AAD0FEC4" box="[1071,1224,259,284]" gridcol="1" gridrow="2" pageId="14" pageNumber="559">Female</td>
<td id="C1005D91FFD10033FB45FEC7AB7BFEC4" box="[1258,1379,259,284]" gridcol="2" gridrow="2" pageId="14" pageNumber="559">Male</td>
</tr>
<tr id="82D134EDFFD10033FF09FEEDAB7BFE9A" box="[166,1379,297,322]" gridrow="3" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FEEDAA08FE9A" box="[166,1040,297,322]" gridcol="0" gridrow="3" pageId="14" pageNumber="559">Snout projecting beyond lower jaw (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FEEDAAD0FE9A" box="[1071,1224,297,322]" gridcol="1" gridrow="3" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FEEDAB7BFE9A" box="[1258,1379,297,322]" gridcol="2" gridrow="3" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FE8AAB7BFEBF" box="[166,1379,334,359]" gridrow="4" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FE8AAA08FEBF" box="[166,1040,334,359]" gridcol="0" gridrow="4" pageId="14" pageNumber="559">Tympanum visible (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FE8AAAD0FEBF" box="[1071,1224,334,359]" gridcol="1" gridrow="4" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FE8AAB7BFEBF" box="[1258,1379,334,359]" gridcol="2" gridrow="4" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FEB0AB7BFE55" box="[166,1379,372,397]" gridrow="5" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FEB0AA08FE55" box="[166,1040,372,397]" gridcol="0" gridrow="5" pageId="14" pageNumber="559">Light coloured rictal tubercle(s) at the corner of the jaw present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FEB0AAD0FE55" box="[1071,1224,372,397]" gridcol="1" gridrow="5" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FEB0AB7BFE55" box="[1258,1379,372,397]" gridcol="2" gridrow="5" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FE5EAB7BFE6B" box="[166,1379,410,435]" gridrow="6" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FE5EAA08FE6B" box="[166,1040,410,435]" gridcol="0" gridrow="6" pageId="14" pageNumber="559">Interorbital tubercle ridges present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FE5EAAD0FE6B" box="[1071,1224,410,435]" gridcol="1" gridrow="6" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FE5EAB7BFE6B" box="[1258,1379,410,435]" gridcol="2" gridrow="6" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FE04AB7BFE01" box="[166,1379,448,473]" gridrow="7" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FE04AA08FE01" box="[166,1040,448,473]" gridcol="0" gridrow="7" pageId="14" pageNumber="559">Opening of vocal sac on right (1) or left (0)</th>
<td id="C1005D91FFD10033FB80FE04AAD0FE01" box="[1071,1224,448,473]" gridcol="1" gridrow="7" pageId="14" pageNumber="559">/</td>
<td id="C1005D91FFD10033FB45FE04AB7BFE01" box="[1258,1379,448,473]" gridcol="2" gridrow="7" pageId="14" pageNumber="559">/</td>
</tr>
<tr id="82D134EDFFD10033FF09FE22AB7BFE27" box="[166,1379,486,511]" gridrow="8" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FE22AA08FE27" box="[166,1040,486,511]" gridcol="0" gridrow="8" pageId="14" pageNumber="559">Finger tips rounded or forming small discs (1) or expanded and estapulate (0) or</th>
<td id="C1005D91FFD10033FB80FE22AAD0FE27" box="[1071,1224,486,511]" gridcol="1" gridrow="8" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FE22AB7BFE27" box="[1258,1379,486,511]" gridcol="2" gridrow="8" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FDCEAB7BFD9B" box="[166,1379,522,579]" gridrow="9" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FDCEAA08FD9B" box="[166,1040,522,579]" gridcol="0" gridrow="9" pageId="14" pageNumber="559">neither (2) Toe tips rounded or forming small discs (1) or expanded and espatulate (0) or neither</th>
<td id="C1005D91FFD10033FB80FDCEAAD0FD9B" box="[1071,1224,522,579]" gridcol="1" gridrow="9" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FDCEAB7BFD9B" box="[1258,1379,522,579]" gridcol="2" gridrow="9" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FD8BAB7BFD50" box="[166,1379,591,648]" gridrow="10" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FD8BAA08FD50" box="[166,1040,591,648]" gridcol="0" gridrow="10" pageId="14" pageNumber="559">(2) 1st finger reaching the disc of the 2nd (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FD8BAAD0FD50" box="[1071,1224,591,648]" gridcol="1" gridrow="10" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FD8BAB7BFD50" box="[1258,1379,591,648]" gridcol="2" gridrow="10" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FD51AB7BFD76" box="[166,1379,661,686]" gridrow="11" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FD51AA08FD76" box="[166,1040,661,686]" gridcol="0" gridrow="11" pageId="14" pageNumber="559">No. of fingers with nuptial pads</th>
<td id="C1005D91FFD10033FB80FD51AAD0FD76" box="[1071,1224,661,686]" gridcol="1" gridrow="11" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FD51AB7BFD76" box="[1258,1379,661,686]" gridcol="2" gridrow="11" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FD7EAB7BFD0B" box="[166,1379,698,723]" gridrow="12" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FD7EAA08FD0B" box="[166,1040,698,723]" gridcol="0" gridrow="12" pageId="14" pageNumber="559">No. of free phalanges of V toe</th>
<td id="C1005D91FFD10033FB80FD7EAAD0FD0B" box="[1071,1224,698,723]" gridcol="1" gridrow="12" pageId="14" pageNumber="559">1.5</td>
<td id="C1005D91FFD10033FB45FD7EAB7BFD0B" box="[1258,1379,698,723]" gridcol="2" gridrow="12" pageId="14" pageNumber="559">1.5</td>
</tr>
<tr id="82D134EDFFD10033FF09FD24AB7BFD21" box="[166,1379,736,761]" gridrow="13" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FD24AA08FD21" box="[166,1040,736,761]" gridcol="0" gridrow="13" pageId="14" pageNumber="559">No. of free phalanges of IV toe</th>
<td id="C1005D91FFD10033FB80FD24AAD0FD21" box="[1071,1224,736,761]" gridcol="1" gridrow="13" pageId="14" pageNumber="559">3</td>
<td id="C1005D91FFD10033FB45FD24AB7BFD21" box="[1258,1379,736,761]" gridcol="2" gridrow="13" pageId="14" pageNumber="559">3</td>
</tr>
<tr id="82D134EDFFD10033FF09FCC2AB7BFCC7" box="[166,1379,774,799]" gridrow="14" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FCC2AA08FCC7" box="[166,1040,774,799]" gridcol="0" gridrow="14" pageId="14" pageNumber="559">No. of free phalanges of III toe</th>
<td id="C1005D91FFD10033FB80FCC2AAD0FCC7" box="[1071,1224,774,799]" gridcol="1" gridrow="14" pageId="14" pageNumber="559">1.5–3</td>
<td id="C1005D91FFD10033FB45FCC2AB7BFCC7" box="[1258,1379,774,799]" gridcol="2" gridrow="14" pageId="14" pageNumber="559">1.5–2.5</td>
</tr>
<tr id="82D134EDFFD10033FF09FCE8AB7BFC9D" box="[166,1379,812,837]" gridrow="15" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FCE8AA08FC9D" box="[166,1040,812,837]" gridcol="0" gridrow="15" pageId="14" pageNumber="559">No. of free phalanges of II toe</th>
<td id="C1005D91FFD10033FB80FCE8AAD0FC9D" box="[1071,1224,812,837]" gridcol="1" gridrow="15" pageId="14" pageNumber="559">0.5–1.5</td>
<td id="C1005D91FFD10033FB45FCE8AB7BFC9D" box="[1258,1379,812,837]" gridcol="2" gridrow="15" pageId="14" pageNumber="559">0.5–1.5</td>
</tr>
<tr id="82D134EDFFD10033FF09FC96AB7BFCB3" box="[166,1379,850,875]" gridrow="16" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FC96AA08FCB3" box="[166,1040,850,875]" gridcol="0" gridrow="16" pageId="14" pageNumber="559">No. of free phalanges of I toe</th>
<td id="C1005D91FFD10033FB80FC96AAD0FCB3" box="[1071,1224,850,875]" gridcol="1" gridrow="16" pageId="14" pageNumber="559">0.5</td>
<td id="C1005D91FFD10033FB45FC96AB7BFCB3" box="[1258,1379,850,875]" gridcol="2" gridrow="16" pageId="14" pageNumber="559">0.5</td>
</tr>
<tr id="82D134EDFFD10033FF09FCB3AB7BFC48" box="[166,1379,887,912]" gridrow="17" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FCB3AA08FC48" box="[166,1040,887,912]" gridcol="0" gridrow="17" pageId="14" pageNumber="559">Tarsal ridge present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FCB3AAD0FC48" box="[1071,1224,887,912]" gridcol="1" gridrow="17" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FCB3AB7BFC48" box="[1258,1379,887,912]" gridcol="2" gridrow="17" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FC59AB7BFC6E" box="[166,1379,925,950]" gridrow="18" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FC59AA08FC6E" box="[166,1040,925,950]" gridcol="0" gridrow="18" pageId="14" pageNumber="559">Inner metatarsal tubercle present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FC59AAD0FC6E" box="[1071,1224,925,950]" gridcol="1" gridrow="18" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FC59AB7BFC6E" box="[1258,1379,925,950]" gridcol="2" gridrow="18" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FC07AB7BFC04" box="[166,1379,963,988]" gridrow="19" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FC07AA08FC04" box="[166,1040,963,988]" gridcol="0" gridrow="19" pageId="14" pageNumber="559">Outer metatarsal tubercle present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FC07AAD0FC04" box="[1071,1224,963,988]" gridcol="1" gridrow="19" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FC07AB7BFC04" box="[1258,1379,963,988]" gridcol="2" gridrow="19" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FC2DAB7BFBDA" box="[166,1379,1001,1026]" gridrow="20" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FC2DAA08FBDA" box="[166,1040,1001,1026]" gridcol="0" gridrow="20" pageId="14" pageNumber="559">Submandibular tubercles in males present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FC2DAAD0FBDA" box="[1071,1224,1001,1026]" gridcol="1" gridrow="20" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FC2DAB7BFBDA" box="[1258,1379,1001,1026]" gridcol="2" gridrow="20" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FBCBAB7BFBF0" box="[166,1379,1039,1064]" gridrow="21" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FBCBAA08FBF0" box="[166,1040,1039,1064]" gridcol="0" gridrow="21" pageId="14" pageNumber="559">Dorsal tubercles present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FBCBAAD0FBF0" box="[1071,1224,1039,1064]" gridcol="1" gridrow="21" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FBCBAB7BFBF0" box="[1258,1379,1039,1064]" gridcol="2" gridrow="21" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FBF0AB7BFB95" box="[166,1379,1076,1101]" gridrow="22" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FBF0AA08FB95" box="[166,1040,1076,1101]" gridcol="0" gridrow="22" pageId="14" pageNumber="559">Dorsolateral row of enlarged tubercles present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FBF0AAD0FB95" box="[1071,1224,1076,1101]" gridcol="1" gridrow="22" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FBF0AB7BFB95" box="[1258,1379,1076,1101]" gridcol="2" gridrow="22" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FB9EAB7BFBAB" box="[166,1379,1114,1139]" gridrow="23" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FB9EAA08FBAB" box="[166,1040,1114,1139]" gridcol="0" gridrow="23" pageId="14" pageNumber="559">Rows of tubercles on back (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FB9EAAD0FBAB" box="[1071,1224,1114,1139]" gridcol="1" gridrow="23" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FB9EAB7BFBAB" box="[1258,1379,1114,1139]" gridcol="2" gridrow="23" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FB44AB7BFB41" box="[166,1379,1152,1177]" gridrow="24" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FB44AA08FB41" box="[166,1040,1152,1177]" gridcol="0" gridrow="24" pageId="14" pageNumber="559">Oblique flap of skin on each side of vent (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FB44AAD0FB41" box="[1071,1224,1152,1177]" gridcol="1" gridrow="24" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FB44AB7BFB41" box="[1258,1379,1152,1177]" gridcol="2" gridrow="24" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FB62AB7BFB67" box="[166,1379,1190,1215]" gridrow="25" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FB62AA08FB67" box="[166,1040,1190,1215]" gridcol="0" gridrow="25" pageId="14" pageNumber="559">Abdomen coarsely grannular (1), finely grannular (2), or tuberculate (0)</th>
<td id="C1005D91FFD10033FB80FB62AAD0FB67" box="[1071,1224,1190,1215]" gridcol="1" gridrow="25" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FB62AB7BFB67" box="[1258,1379,1190,1215]" gridcol="2" gridrow="25" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FB08AB7BFB3D" box="[166,1379,1228,1253]" gridrow="26" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FB08AA08FB3D" box="[166,1040,1228,1253]" gridcol="0" gridrow="26" pageId="14" pageNumber="559">Colour of iris</th>
<td id="C1005D91FFD10033FB80FB08AAD0FB3D" box="[1071,1224,1228,1253]" gridcol="1" gridrow="26" pageId="14" pageNumber="559">Yellow</td>
<td id="C1005D91FFD10033FB45FB08AB7BFB3D" box="[1258,1379,1228,1253]" gridcol="2" gridrow="26" pageId="14" pageNumber="559">Yellow</td>
</tr>
<tr id="82D134EDFFD10033FF09FB35AB7BFAD2" box="[166,1379,1265,1290]" gridrow="27" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FB35AA08FAD2" box="[166,1040,1265,1290]" gridcol="0" gridrow="27" pageId="14" pageNumber="559">Gular spotting present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FB35AAD0FAD2" box="[1071,1224,1265,1290]" gridcol="1" gridrow="27" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FB35AB7BFAD2" box="[1258,1379,1265,1290]" gridcol="2" gridrow="27" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FAD3AB7BFAE8" box="[166,1379,1303,1328]" gridrow="28" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FAD3AA08FAE8" box="[166,1040,1303,1328]" gridcol="0" gridrow="28" pageId="14" pageNumber="559">Wide, light patch below eye (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FAD3AAD0FAE8" box="[1071,1224,1303,1328]" gridcol="1" gridrow="28" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FAD3AB7BFAE8" box="[1258,1379,1303,1328]" gridcol="2" gridrow="28" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FAF9AB7BFA8E" box="[166,1379,1341,1366]" gridrow="29" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FAF9AA08FA8E" box="[166,1040,1341,1366]" gridcol="0" gridrow="29" pageId="14" pageNumber="559">White postorbital patch present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FAF9AAD0FA8E" box="[1071,1224,1341,1366]" gridcol="1" gridrow="29" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FAF9AB7BFA8E" box="[1258,1379,1341,1366]" gridcol="2" gridrow="29" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FAA7AB7BFAA4" box="[166,1379,1379,1404]" gridrow="30" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FAA7AA08FAA4" box="[166,1040,1379,1404]" gridcol="0" gridrow="30" pageId="14" pageNumber="559">Light spot between the scapulae present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FAA7AAD0FAA4" box="[1071,1224,1379,1404]" gridcol="1" gridrow="30" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FAA7AB7BFAA4" box="[1258,1379,1379,1404]" gridcol="2" gridrow="30" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FA4DAB7BFA7A" box="[166,1379,1417,1442]" gridrow="31" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FA4DAA08FA7A" box="[166,1040,1417,1442]" gridcol="0" gridrow="31" pageId="14" pageNumber="559">Light crossbar on hind limbs present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FA4DAAD0FA7A" box="[1071,1224,1417,1442]" gridcol="1" gridrow="31" pageId="14" pageNumber="559">1</td>
<td id="C1005D91FFD10033FB45FA4DAB7BFA7A" box="[1258,1379,1417,1442]" gridcol="2" gridrow="31" pageId="14" pageNumber="559">1</td>
</tr>
<tr id="82D134EDFFD10033FF09FA6AAB7BFA1F" box="[166,1379,1454,1479]" gridrow="32" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FA6AAA08FA1F" box="[166,1040,1454,1479]" gridcol="0" gridrow="32" pageId="14" pageNumber="559">Vertebral stripe present (1) or not (0)</th>
<td id="C1005D91FFD10033FB80FA6AAAD0FA1F" box="[1071,1224,1454,1479]" gridcol="1" gridrow="32" pageId="14" pageNumber="559">0</td>
<td id="C1005D91FFD10033FB45FA6AAB7BFA1F" box="[1258,1379,1454,1479]" gridcol="2" gridrow="32" pageId="14" pageNumber="559">0</td>
</tr>
<tr id="82D134EDFFD10033FF09FA10AB7BFA35" box="[166,1379,1492,1517]" gridrow="33" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FA10AA08FA35" box="[166,1040,1492,1517]" gridcol="0" gridrow="33" pageId="14" pageNumber="559">Discrete white or bright yellow spots along the underside of lower jaw large (2), small</th>
<td id="C1005D91FFD10033FB80FA10AAD0FA35" box="[1071,1224,1492,1517]" gridcol="1" gridrow="33" pageId="14" pageNumber="559">2</td>
<td id="C1005D91FFD10033FB45FA10AB7BFA35" box="[1258,1379,1492,1517]" gridcol="2" gridrow="33" pageId="14" pageNumber="559">2</td>
</tr>
<tr id="82D134EDFFD10033FF09FA3CAB7BF9EA" box="[166,1379,1528,1586]" gridrow="34" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09FA3CAA08F9EA" box="[166,1040,1528,1586]" gridcol="0" gridrow="34" pageId="14" pageNumber="559">(1) or absent (0) Finger length from shortest to longest</th>
<td id="C1005D91FFD10033FB80FA3CAAD0F9EA" box="[1071,1224,1528,1586]" gridcol="1" gridrow="34" pageId="14" pageNumber="559">1&lt;2&lt;4&lt;3</td>
<td id="C1005D91FFD10033FB45FA3CAB7BF9EA" box="[1258,1379,1528,1586]" gridcol="2" gridrow="34" pageId="14" pageNumber="559">1&lt;2&lt;4&lt;3</td>
</tr>
<tr id="82D134EDFFD10033FF09F9FBAB7BF980" box="[166,1379,1599,1624]" gridrow="35" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F9FBAA08F980" box="[166,1040,1599,1624]" gridcol="0" gridrow="35" pageId="14" pageNumber="559">Toe length from shortest to longest</th>
<td id="C1005D91FFD10033FB80F9FBAAD0F980" box="[1071,1224,1599,1624]" gridcol="1" gridrow="35" pageId="14" pageNumber="559">1&lt;2&lt;5&lt;3&lt;4</td>
<td id="C1005D91FFD10033FB45F9FBAB7BF980" box="[1258,1379,1599,1624]" gridcol="2" gridrow="35" pageId="14" pageNumber="559">1&lt;2&lt;5&lt;3&lt;4</td>
</tr>
<tr id="82D134EDFFD10033FF09F9A1AB7BF9A5" box="[166,1379,1637,1661]" gridrow="36" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F9A1AA08F9A5" box="[166,1040,1637,1661]" gridcol="0" gridrow="36" pageId="14" pageNumber="559">SVL (mm)</th>
<td id="C1005D91FFD10033FB80F9A1AAD0F9A5" box="[1071,1224,1637,1661]" gridcol="1" gridrow="36" pageId="14" pageNumber="559">24.0</td>
<td id="C1005D91FFD10033FB45F9A1AB7BF9A5" box="[1258,1379,1637,1661]" gridcol="2" gridrow="36" pageId="14" pageNumber="559">24.0</td>
</tr>
<tr id="82D134EDFFD10033FF09F94FAB7BF97B" box="[166,1379,1675,1699]" gridrow="37" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F94FAA08F97B" box="[166,1040,1675,1699]" gridcol="0" gridrow="37" pageId="14" pageNumber="559">HL (mm)</th>
<td id="C1005D91FFD10033FB80F94FAAD0F97B" box="[1071,1224,1675,1699]" gridcol="1" gridrow="37" pageId="14" pageNumber="559">7.20</td>
<td id="C1005D91FFD10033FB45F94FAB7BF97B" box="[1258,1379,1675,1699]" gridcol="2" gridrow="37" pageId="14" pageNumber="559">7.10</td>
</tr>
<tr id="82D134EDFFD10033FF09F974AB7BF911" box="[166,1379,1712,1737]" gridrow="38" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F974AA08F911" box="[166,1040,1712,1737]" gridcol="0" gridrow="38" pageId="14" pageNumber="559">HW (mm)</th>
<td id="C1005D91FFD10033FB80F974AAD0F911" box="[1071,1224,1712,1737]" gridcol="1" gridrow="38" pageId="14" pageNumber="559">6.91</td>
<td id="C1005D91FFD10033FB45F974AB7BF911" box="[1258,1379,1712,1737]" gridcol="2" gridrow="38" pageId="14" pageNumber="559">6.42</td>
</tr>
<tr id="82D134EDFFD10033FF09F912AB7BF937" box="[166,1379,1750,1775]" gridrow="39" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F912AA08F937" box="[166,1040,1750,1775]" gridcol="0" gridrow="39" pageId="14" pageNumber="559">SL (mm)</th>
<td id="C1005D91FFD10033FB80F912AAD0F937" box="[1071,1224,1750,1775]" gridcol="1" gridrow="39" pageId="14" pageNumber="559">3.24</td>
<td id="C1005D91FFD10033FB45F912AB7BF937" box="[1258,1379,1750,1775]" gridcol="2" gridrow="39" pageId="14" pageNumber="559">3.25</td>
</tr>
<tr id="82D134EDFFD10033FF09F938AB7BF8CD" box="[166,1379,1788,1813]" gridrow="40" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F938AA08F8CD" box="[166,1040,1788,1813]" gridcol="0" gridrow="40" pageId="14" pageNumber="559">SW (mm)</th>
<td id="C1005D91FFD10033FB80F938AAD0F8CD" box="[1071,1224,1788,1813]" gridcol="1" gridrow="40" pageId="14" pageNumber="559">3.66</td>
<td id="C1005D91FFD10033FB45F938AB7BF8CD" box="[1258,1379,1788,1813]" gridcol="2" gridrow="40" pageId="14" pageNumber="559">3.77</td>
</tr>
<tr id="82D134EDFFD10033FF09F8E6AB7BF8E2" box="[166,1379,1826,1850]" gridrow="41" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F8E6AA08F8E2" box="[166,1040,1826,1850]" gridcol="0" gridrow="41" pageId="14" pageNumber="559">IND (mm)</th>
<td id="C1005D91FFD10033FB80F8E6AAD0F8E2" box="[1071,1224,1826,1850]" gridcol="1" gridrow="41" pageId="14" pageNumber="559">2.00</td>
<td id="C1005D91FFD10033FB45F8E6AB7BF8E2" box="[1258,1379,1826,1850]" gridcol="2" gridrow="41" pageId="14" pageNumber="559">1.78</td>
</tr>
<tr id="82D134EDFFD10033FF09F883AB7BF8B8" box="[166,1379,1863,1888]" gridrow="42" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F883AA08F8B8" box="[166,1040,1863,1888]" gridcol="0" gridrow="42" pageId="14" pageNumber="559">IOD (mm)</th>
<td id="C1005D91FFD10033FB80F883AAD0F8B8" box="[1071,1224,1863,1888]" gridcol="1" gridrow="42" pageId="14" pageNumber="559">2.75</td>
<td id="C1005D91FFD10033FB45F883AB7BF8B8" box="[1258,1379,1863,1888]" gridcol="2" gridrow="42" pageId="14" pageNumber="559">2.62</td>
</tr>
<tr id="82D134EDFFD10033FF09F8AAAB7BF85E" box="[166,1379,1902,1926]" gridrow="43" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F8AAAA08F85E" box="[166,1040,1902,1926]" gridcol="0" gridrow="43" pageId="14" pageNumber="559">ED (mm)</th>
<td id="C1005D91FFD10033FB80F8AAAAD0F85E" box="[1071,1224,1902,1926]" gridcol="1" gridrow="43" pageId="14" pageNumber="559">2.08</td>
<td id="C1005D91FFD10033FB45F8AAAB7BF85E" box="[1258,1379,1902,1926]" gridcol="2" gridrow="43" pageId="14" pageNumber="559">1.92</td>
</tr>
<tr id="82D134EDFFD10033FF09F850AB7BF874" box="[166,1379,1940,1964]" gridrow="44" pageId="14" pageNumber="559">
<th id="C1005D91FFD10033FF09F850AA08F874" box="[166,1040,1940,1964]" gridcol="0" gridrow="44" pageId="14" pageNumber="559">TD (mm)</th>
<td id="C1005D91FFD10033FB80F850AAD0F874" box="[1071,1224,1940,1964]" gridcol="1" gridrow="44" pageId="14" pageNumber="559">0.81</td>
<td id="C1005D91FFD10033FB45F850AB7BF874" box="[1258,1379,1940,1964]" gridcol="2" gridrow="44" pageId="14" pageNumber="559">0.81</td>
</tr>
</table>
</paragraph>
<paragraph id="3C5E36AFFFD1FFC2FAEBFF7AAB9BFF0F" blockId="14.[1348,1411,190,215]" box="[1348,1411,190,215]" pageId="14" pageNumber="559">14067</paragraph>
<paragraph id="3C5E36AFFFD1FFC2FBF9F87FAB94F80A" blockId="14.[1110,1420,1979,2003]" box="[1110,1420,1979,2003]" pageId="14" pageNumber="559">
<tableNote id="C1073721FFD1FFC2FBF9F87FAB94F80A" box="[1110,1420,1979,2003]" pageId="14" pageNumber="559" targetBox="[166,1379,259,1964]" targetPageId="14">......continued on the next page</tableNote>
</paragraph>
<caption id="689E6627FFD0FFC3FF09FF64AF8CFF61" ID-Table-UUID="689E6627FFD0FFC3FF09FF64AF8CFF61" box="[166,404,160,185]" httpUri="http://table.plazi.org/id/689E6627FFD0FFC3FF09FF64AF8CFF61" pageId="15" pageNumber="560" startId="15.[166,250,160,185]" targetBox="[166,1411,197,784]" targetIsTable="true" targetPageId="15">
<paragraph id="3C5E36AFFFD0FFC3FF09FF64AF8CFF61" blockId="15.[166,404,160,185]" box="[166,404,160,185]" pageId="15" pageNumber="560">
<emphasis id="0E95EABDFFD0FFC3FF09FF64AF0BFF61" bold="true" box="[166,275,160,185]" pageId="15" pageNumber="560">TABLE 4.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="3C5E36AFFFD0FFC3FF09FF01AB0EFCC8" pageId="15" pageNumber="560">
<table id="4EE1C40FFFD00033FF09FF01AB9BFCC8" box="[166,1411,197,784]" gridcols="3" gridrows="16" pageId="15" pageNumber="560">
<tr id="82D134EDFFD00033FF09FF01AB9BFF06" box="[166,1411,197,222]" gridrow="0" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FF01AF12FF06" box="[166,266,197,222]" gridcol="0" gridrow="0" pageId="15" pageNumber="560">Specimen</th>
<th id="C1005D91FFD00033FB80FF01AAD0FF06" box="[1071,1224,197,222]" gridcol="1" gridrow="0" pageId="15" pageNumber="560">LSUHC 14066</th>
<th id="C1005D91FFD00033FB45FF01AB9BFF06" box="[1258,1411,197,222]" gridcol="2" gridrow="0" pageId="15" pageNumber="560">LSUHC 14067</th>
</tr>
<tr id="82D134EDFFD00033FF09FF2DAB9BFEDA" box="[166,1411,233,258]" gridrow="1" pageId="15" pageNumber="560" rowspan-0="1">
<td id="C1005D91FFD00033FB80FF2DAAD0FEDA" box="[1071,1224,233,258]" gridcol="1" gridrow="1" pageId="15" pageNumber="560">(holotype)</td>
<td id="C1005D91FFD00033FB45FF2DAB9BFEDA" box="[1258,1411,233,258]" gridcol="2" gridrow="1" pageId="15" pageNumber="560">(paratype)</td>
</tr>
<tr id="82D134EDFFD00033FF09FECFAB9BFEFC" box="[166,1411,267,292]" gridrow="2" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FECFAF12FEFC" box="[166,266,267,292]" gridcol="0" gridrow="2" pageId="15" pageNumber="560">ML (mm)</th>
<td id="C1005D91FFD00033FB80FECFAAD0FEFC" box="[1071,1224,267,292]" gridcol="1" gridrow="2" pageId="15" pageNumber="560">4.92</td>
<td id="C1005D91FFD00033FB45FECFAB9BFEFC" box="[1258,1411,267,292]" gridcol="2" gridrow="2" pageId="15" pageNumber="560">5.02</td>
</tr>
<tr id="82D134EDFFD00033FF09FEF5AB9BFE92" box="[166,1411,305,330]" gridrow="3" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FEF5AF12FE92" box="[166,266,305,330]" gridcol="0" gridrow="3" pageId="15" pageNumber="560">CL (mm)</th>
<td id="C1005D91FFD00033FB80FEF5AAD0FE92" box="[1071,1224,305,330]" gridcol="1" gridrow="3" pageId="15" pageNumber="560">9.97</td>
<td id="C1005D91FFD00033FB45FEF5AB9BFE92" box="[1258,1411,305,330]" gridcol="2" gridrow="3" pageId="15" pageNumber="560">9.62</td>
</tr>
<tr id="82D134EDFFD00033FF09FE9CAB9BFEA8" box="[166,1411,344,368]" gridrow="4" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FE9CAF12FEA8" box="[166,266,344,368]" gridcol="0" gridrow="4" pageId="15" pageNumber="560">PL (mm)</th>
<td id="C1005D91FFD00033FB80FE9CAAD0FEA8" box="[1071,1224,344,368]" gridcol="1" gridrow="4" pageId="15" pageNumber="560">7.44</td>
<td id="C1005D91FFD00033FB45FE9CAB9BFEA8" box="[1258,1411,344,368]" gridcol="2" gridrow="4" pageId="15" pageNumber="560">7.28</td>
</tr>
<tr id="82D134EDFFD00033FF09FEBAAB9BFE4E" box="[166,1411,382,406]" gridrow="5" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FEBAAF12FE4E" box="[166,266,382,406]" gridcol="0" gridrow="5" pageId="15" pageNumber="560">HL/HW</th>
<td id="C1005D91FFD00033FB80FEBAAAD0FE4E" box="[1071,1224,382,406]" gridcol="1" gridrow="5" pageId="15" pageNumber="560">1.04</td>
<td id="C1005D91FFD00033FB45FEBAAB9BFE4E" box="[1258,1411,382,406]" gridcol="2" gridrow="5" pageId="15" pageNumber="560">1.11</td>
</tr>
<tr id="82D134EDFFD00033FF09FE67AB9BFE64" box="[166,1411,419,444]" gridrow="6" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FE67AF12FE64" box="[166,266,419,444]" gridcol="0" gridrow="6" pageId="15" pageNumber="560">SL/SW</th>
<td id="C1005D91FFD00033FB80FE67AAD0FE64" box="[1071,1224,419,444]" gridcol="1" gridrow="6" pageId="15" pageNumber="560">0.89</td>
<td id="C1005D91FFD00033FB45FE67AB9BFE64" box="[1258,1411,419,444]" gridcol="2" gridrow="6" pageId="15" pageNumber="560">0.86</td>
</tr>
<tr id="82D134EDFFD00033FF09FE0CAB9BFE39" box="[166,1411,456,481]" gridrow="7" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FE0CAF12FE39" box="[166,266,456,481]" gridcol="0" gridrow="7" pageId="15" pageNumber="560">IND/SL</th>
<td id="C1005D91FFD00033FB80FE0CAAD0FE39" box="[1071,1224,456,481]" gridcol="1" gridrow="7" pageId="15" pageNumber="560">0.62</td>
<td id="C1005D91FFD00033FB45FE0CAB9BFE39" box="[1258,1411,456,481]" gridcol="2" gridrow="7" pageId="15" pageNumber="560">0.55</td>
</tr>
<tr id="82D134EDFFD00033FF09FE2AAB9BFDDF" box="[166,1411,494,519]" gridrow="8" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FE2AAF12FDDF" box="[166,266,494,519]" gridcol="0" gridrow="8" pageId="15" pageNumber="560">IND/SW</th>
<td id="C1005D91FFD00033FB80FE2AAAD0FDDF" box="[1071,1224,494,519]" gridcol="1" gridrow="8" pageId="15" pageNumber="560">0.55</td>
<td id="C1005D91FFD00033FB45FE2AAB9BFDDF" box="[1258,1411,494,519]" gridcol="2" gridrow="8" pageId="15" pageNumber="560">0.47</td>
</tr>
<tr id="82D134EDFFD00033FF09FDD0AB9BFDF5" box="[166,1411,532,557]" gridrow="9" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FDD0AF12FDF5" box="[166,266,532,557]" gridcol="0" gridrow="9" pageId="15" pageNumber="560">IOD/SW</th>
<td id="C1005D91FFD00033FB80FDD0AAD0FDF5" box="[1071,1224,532,557]" gridcol="1" gridrow="9" pageId="15" pageNumber="560">0.75</td>
<td id="C1005D91FFD00033FB45FDD0AB9BFDF5" box="[1258,1411,532,557]" gridcol="2" gridrow="9" pageId="15" pageNumber="560">0.69</td>
</tr>
<tr id="82D134EDFFD00033FF09FDFEAB9BFD8B" box="[166,1411,570,595]" gridrow="10" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FDFEAF12FD8B" box="[166,266,570,595]" gridcol="0" gridrow="10" pageId="15" pageNumber="560">IOD/SL</th>
<td id="C1005D91FFD00033FB80FDFEAAD0FD8B" box="[1071,1224,570,595]" gridcol="1" gridrow="10" pageId="15" pageNumber="560">0.85</td>
<td id="C1005D91FFD00033FB45FDFEAB9BFD8B" box="[1258,1411,570,595]" gridcol="2" gridrow="10" pageId="15" pageNumber="560">0.81</td>
</tr>
<tr id="82D134EDFFD00033FF09FDA4AB9BFDA1" box="[166,1411,608,633]" gridrow="11" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FDA4AF12FDA1" box="[166,266,608,633]" gridcol="0" gridrow="11" pageId="15" pageNumber="560">ED/SL</th>
<td id="C1005D91FFD00033FB80FDA4AAD0FDA1" box="[1071,1224,608,633]" gridcol="1" gridrow="11" pageId="15" pageNumber="560">0.64</td>
<td id="C1005D91FFD00033FB45FDA4AB9BFDA1" box="[1258,1411,608,633]" gridcol="2" gridrow="11" pageId="15" pageNumber="560">0.59</td>
</tr>
<tr id="82D134EDFFD00033FF09FD41AB9BFD46" box="[166,1411,645,670]" gridrow="12" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FD41AF12FD46" box="[166,266,645,670]" gridcol="0" gridrow="12" pageId="15" pageNumber="560">ED/IOD</th>
<td id="C1005D91FFD00033FB80FD41AAD0FD46" box="[1071,1224,645,670]" gridcol="1" gridrow="12" pageId="15" pageNumber="560">0.76</td>
<td id="C1005D91FFD00033FB45FD41AB9BFD46" box="[1258,1411,645,670]" gridcol="2" gridrow="12" pageId="15" pageNumber="560">0.73</td>
</tr>
<tr id="82D134EDFFD00033FF09FD68AB9BFD1C" box="[166,1411,684,708]" gridrow="13" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FD68AF12FD1C" box="[166,266,684,708]" gridcol="0" gridrow="13" pageId="15" pageNumber="560">TD/ED</th>
<td id="C1005D91FFD00033FB80FD68AAD0FD1C" box="[1071,1224,684,708]" gridcol="1" gridrow="13" pageId="15" pageNumber="560">0.39</td>
<td id="C1005D91FFD00033FB45FD68AB9BFD1C" box="[1258,1411,684,708]" gridcol="2" gridrow="13" pageId="15" pageNumber="560">0.42</td>
</tr>
<tr id="82D134EDFFD00033FF09FD16AB9BFD32" box="[166,1411,722,746]" gridrow="14" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FD16AF12FD32" box="[166,266,722,746]" gridcol="0" gridrow="14" pageId="15" pageNumber="560">CL/SVL</th>
<td id="C1005D91FFD00033FB80FD16AAD0FD32" box="[1071,1224,722,746]" gridcol="1" gridrow="14" pageId="15" pageNumber="560">0.42</td>
<td id="C1005D91FFD00033FB45FD16AB9BFD32" box="[1258,1411,722,746]" gridcol="2" gridrow="14" pageId="15" pageNumber="560">0.40</td>
</tr>
<tr id="82D134EDFFD00033FF09FD33AB9BFCC8" box="[166,1411,759,784]" gridrow="15" pageId="15" pageNumber="560">
<th id="C1005D91FFD00033FF09FD33AF12FCC8" box="[166,266,759,784]" gridcol="0" gridrow="15" pageId="15" pageNumber="560">PL/CL</th>
<td id="C1005D91FFD00033FB80FD33AAD0FCC8" box="[1071,1224,759,784]" gridcol="1" gridrow="15" pageId="15" pageNumber="560">0.75</td>
<td id="C1005D91FFD00033FB45FD33AB9BFCC8" box="[1258,1411,759,784]" gridcol="2" gridrow="15" pageId="15" pageNumber="560">0.76</td>
</tr>
</table>
</paragraph>
<paragraph id="3C5E36AFFFD0FFC3FF68FC84AC61FC58" blockId="15.[151,1437,832,2012]" pageId="15" pageNumber="560">
<emphasis id="0E95EABDFFD0FFC3FF68FC84AFAFFC83" bold="true" box="[199,439,832,859]" pageId="15" pageNumber="560">
Distribution (
<figureCitation id="A4DA2A2AFFD0FFC3FEC6FC84AFB6FC83" box="[361,430,832,859]" captionStart="FIGURE 5" captionStartId="17.[151,250,1967,1992]" captionTargetBox="[341,1242,166,1948]" captionTargetId="figure@17.[341,1246,166,1948]" captionTargetPageId="17" captionText="FIGURE 5. Distribution of Ansonia species along the Myanmar-Thai border and the type locality of Ansonia kyaiktiyoensis sp. nov. Star = type locality." httpUri="https://zenodo.org/record/3368955/files/figure.png" pageId="15" pageNumber="560">Fig. 5</figureCitation>
)
</emphasis>
. 
<taxonomicName id="FBE14D2CFFD0FFC3FE6AFC86ACDDFC83" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[453,709,833,860]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FE6AFC86ACDDFC83" box="[453,709,833,860]" italics="true" pageId="15" pageNumber="560">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD0FFC3FD63FC85AD3EFC83" bold="true" box="[716,806,833,859]" pageId="15" pageNumber="560">
<taxonomicNameLabel id="15A657C6FFD0FFC3FD63FC85AD3EFC83" box="[716,806,833,859]" pageId="15" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the forested hills (
<figureCitation id="A4DA2A2AFFD0FFC3FB72FC85AB2BFC83" box="[1245,1331,833,859]" captionStart="FIGURE 2" captionStartId="11.[151,250,1936,1961]" captionTargetBox="[171,1415,633,1917]" captionTargetId="figure@11.[171,1415,633,1917]" captionTargetPageId="11" captionText="FIGURE 2. (A) Paratype of Ansonia kyaiktiyoensis sp. nov. (LSUHC 14067). (B) Hill forest habitat of Ansonia kyaiktiyoensis sp. nov. at Kyaiktiyo, Mon State, Myanmar." httpUri="https://zenodo.org/record/3368949/files/figure.png" pageId="15" pageNumber="560">Fig. 2B</figureCitation>
) near the Kyaiktiyo Pagoda in Mon State, 
<collectingCountry id="44F6763FFFD0FFC3FDA9FCA2AC6DFC58" box="[518,629,870,896]" name="Myanmar" pageId="15" pageNumber="560">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="3C5E36AFFFD0FFC3FF68FC4DAFF5FBA4" blockId="15.[151,1437,832,2012]" pageId="15" pageNumber="560">
<emphasis id="0E95EABDFFD0FFC3FF68FC4DAF9AFC7B" bold="true" box="[199,386,904,931]" pageId="15" pageNumber="560">Natural history</emphasis>
. The 
<typeStatus id="E35A880DFFD0FFC3FE68FC4DAC31FC7B" box="[455,553,905,931]" pageId="15" pageNumber="560" type="holotype">holotype</typeStatus>
and 
<typeStatus id="E35A880DFFD0FFC3FDCBFC4EACDCFC7C" box="[612,708,906,932]" pageId="15" pageNumber="560" type="paratype">paratype</typeStatus>
were collected at night between 2000–2200 h in hill forest approximately 15–20 meters away from a stream. Both specimens were found sitting on top of leaf litter along the trail. The female 
<typeStatus id="E35A880DFFD0FFC3FEFBFC15AFAEFC33" box="[340,438,977,1003]" pageId="15" pageNumber="560" type="holotype">holotype</typeStatus>
was not gravid and the male 
<typeStatus id="E35A880DFFD0FFC3FD59FC16AD4EFC34" box="[758,854,978,1004]" pageId="15" pageNumber="560" type="paratype">paratype</typeStatus>
did not possess any secondary sexual characteristics (nuptial pads, distended vocal sacs, etc.), which indicates that the month of November may be outside the breeding season for the species. Other species of anurans found in the vicinity were 
<taxonomicName id="FBE14D2CFFD0FFC3FC45FBDDAA70FBEB" authorityName="Kuhl &amp; van Hasselt" authorityYear="1822" box="[1002,1128,1049,1075]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="genus">
<emphasis id="0E95EABDFFD0FFC3FC45FBDDAA70FBEB" box="[1002,1128,1049,1075]" italics="true" pageId="15" pageNumber="560">Megophrys</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="FBE14D2CFFD0FFC3FB30FBDDAB8DFBEB" baseAuthorityName="Blyth" baseAuthorityYear="1856" box="[1183,1429,1049,1075]" class="Amphibia" family="Ranidae" genus="Sylvirana" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="nigrovittata">
<emphasis id="0E95EABDFFD0FFC3FB30FBDDAB8DFBEB" box="[1183,1429,1049,1075]" italics="true" pageId="15" pageNumber="560">Sylvirana nigrovittata</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FF38FBFAAFC5FB8F" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[151,477,1085,1111]" class="Amphibia" family="Dicroglossidae" genus="Ingerana" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="tenasserimensis">
<emphasis id="0E95EABDFFD0FFC3FF38FBFAAEE6FB8F" box="[151,254,1086,1111]" italics="true" pageId="15" pageNumber="560">Ingerana</emphasis>
cf. 
<emphasis id="0E95EABDFFD0FFC3FE83FBFAAFC5FB8F" box="[300,477,1086,1111]" italics="true" pageId="15" pageNumber="560">tenasserimensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FE42FBFAAD10FB8F" baseAuthorityName="Sclater" baseAuthorityYear="1892" box="[493,776,1085,1111]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="limborgi">
<emphasis id="0E95EABDFFD0FFC3FE42FBFAAC63FB8F" box="[493,635,1086,1111]" italics="true" pageId="15" pageNumber="560">Limnonectes</emphasis>
cf. 
<emphasis id="0E95EABDFFD0FFC3FD06FBF9AD10FB8F" box="[681,776,1085,1111]" italics="true" pageId="15" pageNumber="560">limborgi</emphasis>
</taxonomicName>
, 
<emphasis id="0E95EABDFFD0FFC3FCB7FBFAAD30FB8F" box="[792,808,1086,1111]" italics="true" pageId="15" pageNumber="560">L</emphasis>
. cf. 
<emphasis id="0E95EABDFFD0FFC3FCF2FBF9ADB0FB8F" box="[861,936,1085,1111]" italics="true" pageId="15" pageNumber="560">taylori</emphasis>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC18FBF9AA3AFB8F" authorityName="Gistel" authorityYear="1848" box="[951,1058,1085,1111]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="genus">
<emphasis id="0E95EABDFFD0FFC3FC18FBF9AA3AFB8F" box="[951,1058,1085,1111]" italics="true" pageId="15" pageNumber="560">Philautus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="FBE14D2CFFD0FFC3FBF4FBF9AB84FB8F" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[1115,1436,1085,1111]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="0E95EABDFFD0FFC3FBF4FBF9AAFCFB8F" box="[1115,1252,1085,1111]" italics="true" pageId="15" pageNumber="560">Polypedates</emphasis>
cf. 
<emphasis id="0E95EABDFFD0FFC3FABDFBF9AB84FB8F" box="[1298,1436,1085,1111]" italics="true" pageId="15" pageNumber="560">leucomystax</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FF68FBA5AFFEFBA3" authorityName="Ahl" authorityYear="1927" box="[199,486,1121,1147]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="560" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="0E95EABDFFD0FFC3FF68FBA5AFFEFBA3" box="[199,486,1121,1147]" italics="true" pageId="15" pageNumber="560">Rhacophorus bipunctatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3C5E36AFFFD0FFC3FF68FB41AF2AFB1B" blockId="15.[151,1437,832,2012]" pageId="15" pageNumber="560">
<emphasis id="0E95EABDFFD0FFC3FF68FB41AF50FB47" bold="true" box="[199,328,1157,1183]" pageId="15" pageNumber="560">Etymology</emphasis>
. The specific epithet 
<taxonomicName id="FBE14D2CFFD0FFC3FD97FB41ACCDFB47" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[568,725,1157,1183]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis">
<emphasis id="0E95EABDFFD0FFC3FD97FB41ACCDFB47" box="[568,725,1157,1183]" italics="true" pageId="15" pageNumber="560">kyaiktiyoensis</emphasis>
</taxonomicName>
is in reference to the 
<typeStatus id="E35A880DFFD0FFC3FC69FB42ADEEFB78" box="[966,1014,1158,1184]" pageId="15" pageNumber="560">type</typeStatus>
locality of this species near the Kyaiktiyo Pagoda.
</paragraph>
<paragraph id="3C5E36AFFFD0FFC3FF68FB08AA54F8F0" blockId="15.[151,1437,832,2012]" pageId="15" pageNumber="560">
<emphasis id="0E95EABDFFD0FFC3FF68FB08AF7AFB3F" bold="true" box="[199,354,1228,1255]" pageId="15" pageNumber="560">Comparison.</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FEC2FB0AAC68FB3F" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[365,624,1229,1256]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FEC2FB0AAC68FB3F" box="[365,624,1229,1256]" italics="true" pageId="15" pageNumber="560">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD0FFC3FDD4FB09ACC0FB3F" bold="true" box="[635,728,1229,1255]" pageId="15" pageNumber="560">
<taxonomicNameLabel id="15A657C6FFD0FFC3FDD4FB09ACC0FB3F" box="[635,728,1229,1255]" pageId="15" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from 
<taxonomicName id="FBE14D2CFFD0FFC3FC4DFB0AAAC7FB30" authority="(Taylor)" authorityName="Taylor" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[994,1247,1229,1256]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="mcgregori">
<emphasis id="0E95EABDFFD0FFC3FC4DFB0AAA60FB3F" box="[994,1144,1230,1256]" italics="true" pageId="15" pageNumber="560">A. mcgregori</emphasis>
(Taylor)
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FAB3FB0AAF07FAD4" authority="(Boulenger)" authorityName="Boulenger" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="0E95EABDFFD0FFC3FAB3FB0AAB84FB3F" box="[1308,1436,1229,1256]" italics="true" pageId="15" pageNumber="560">A. muelleri</emphasis>
(Boulenger)
</taxonomicName>
by the presence of a visible tympanum. From 
<taxonomicName id="FBE14D2CFFD0FFC3FC8EFB36AA2BFAD4" authority="(Mocquard)" authorityName="Mocquard" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[801,1075,1265,1292]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="fuliginea">
<emphasis id="0E95EABDFFD0FFC3FC8EFB36ADBCFAD3" box="[801,932,1265,1292]" italics="true" pageId="15" pageNumber="560">A. fuliginea</emphasis>
(Mocquard)
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB92FB36AAFCFAD4" authority="Inger" authorityName="Inger" authorityYear="1966" box="[1085,1252,1265,1292]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="guibei">
<emphasis id="0E95EABDFFD0FFC3FB92FB36AAB9FAD3" box="[1085,1185,1265,1292]" italics="true" pageId="15" pageNumber="560">A. guibei</emphasis>
Inger
</taxonomicName>
, and 
<taxonomicName id="FBE14D2CFFD0FFC3FAB2FB36AECAFAE8" authority="Inger" authorityName="Inger" authorityYear="1966" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latidisca">
<emphasis id="0E95EABDFFD0FFC3FAB2FB36AB84FAD3" box="[1309,1436,1265,1292]" italics="true" pageId="15" pageNumber="560">A. latidisca</emphasis>
Inger
</taxonomicName>
it can be distinguished by its snout projecting beyond the lower jaw. 
<taxonomicName id="FBE14D2CFFD0FFC3FC48FAD2AAF0FAF7" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[999,1256,1301,1328]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FC48FAD2AAF0FAF7" box="[999,1256,1301,1328]" italics="true" pageId="15" pageNumber="560">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD0FFC3FB5FFAD1AB53FAF7" bold="true" box="[1264,1355,1301,1327]" pageId="15" pageNumber="560">
<taxonomicNameLabel id="15A657C6FFD0FFC3FB5FFAD1AB53FAF7" box="[1264,1355,1301,1327]" pageId="15" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from 
<taxonomicName id="FBE14D2CFFD0FFC3FEDEFAFEAD12FA8B" authority="Iskandar &amp; Mumpuni" authorityName="Iskandar &amp; Mumpuni" authorityYear="2004" box="[369,778,1337,1364]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="0E95EABDFFD0FFC3FEDEFAFEAC15FA8B" box="[369,525,1337,1364]" italics="true" pageId="15" pageNumber="560">A. glandulosa</emphasis>
Iskandar &amp; Mumpuni
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FCB9FAFEAD8DFA8B" authorityName="Inger" authorityYear="1966" box="[790,917,1337,1364]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latidisca">
<emphasis id="0E95EABDFFD0FFC3FCB9FAFEAD8DFA8B" box="[790,917,1337,1364]" italics="true" pageId="15" pageNumber="560">A. latidisca</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC0BFAFEAAAEFA8B" authority="Grismer, A." authorityName="Grismer, A." box="[932,1206,1337,1364]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latirostra">
<emphasis id="0E95EABDFFD0FFC3FC0BFAFEAA34FA8B" box="[932,1068,1337,1364]" italics="true" pageId="15" pageNumber="560">A. latirostra</emphasis>
Grismer, A.
</taxonomicName>
<taxonomicName id="FBE14D2CFFD0FFC3FB12FAFEAB29FA8B" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[1213,1329,1338,1363]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="mcgregori">
<emphasis id="0E95EABDFFD0FFC3FB12FAFEAB29FA8B" box="[1213,1329,1338,1363]" italics="true" pageId="15" pageNumber="560">mcgregori</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FA90FAFEAFA7FAA0" authority="Hertwig, Pui, Haas &amp; Das" authorityName="Hertwig, Pui, Haas &amp; Das" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="vidua">
<emphasis id="0E95EABDFFD0FFC3FA90FAFEAB84FA8B" box="[1343,1436,1337,1364]" italics="true" pageId="15" pageNumber="560">A. vidua</emphasis>
Hertwig, Pui, Haas &amp; Das
</taxonomicName>
and some specimens of 
<taxonomicName id="FBE14D2CFFD0FFC3FD7AFA9AAD4CFAAF" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[725,852,1373,1400]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="leptopus">
<emphasis id="0E95EABDFFD0FFC3FD7AFA9AAD4CFAAF" box="[725,852,1373,1400]" italics="true" pageId="15" pageNumber="560">A. leptopus</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FCCCFA9AADE6FAAF" authorityName="Inger" authorityYear="1960" box="[867,1022,1373,1400]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="longidigita">
<emphasis id="0E95EABDFFD0FFC3FCCCFA9AADE6FAAF" box="[867,1022,1373,1400]" italics="true" pageId="15" pageNumber="560">A. longidigita</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FB99FA9AAB56FAA0" authority="(Mocquard)" authorityName="Mocquard" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[1078,1358,1373,1400]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="spinulifer">
<emphasis id="0E95EABDFFD0FFC3FB99FA9AAAD9FAAF" box="[1078,1217,1373,1400]" italics="true" pageId="15" pageNumber="560">A. spinulifer</emphasis>
(Mocquard)
</taxonomicName>
by the absence of interorbital tubercle ridges. It can be further differentiated from 
<taxonomicName id="FBE14D2CFFD0FFC3FC79FA46AA4BFA43" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[982,1107,1409,1436]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="leptopus">
<emphasis id="0E95EABDFFD0FFC3FC79FA46AA4BFA43" box="[982,1107,1409,1436]" italics="true" pageId="15" pageNumber="560">A. leptopus</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FB28FA46AB38FA43" authorityName="Inger" authorityYear="1960" box="[1159,1312,1409,1436]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="longidigita">
<emphasis id="0E95EABDFFD0FFC3FB28FA46AB38FA43" box="[1159,1312,1409,1436]" italics="true" pageId="15" pageNumber="560">A. longidigita</emphasis>
</taxonomicName>
along with 
<taxonomicName id="FBE14D2CFFD0FFC3FF38FA62AF2AFA67" authorityName="Iskandar &amp; Mumpuni" authorityYear="2004" box="[151,306,1445,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="0E95EABDFFD0FFC3FF38FA62AF2AFA67" box="[151,306,1445,1472]" italics="true" pageId="15" pageNumber="560">A. glandulosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FE90FA62AFBCFA67" authorityName="Inger" authorityYear="1966" box="[319,420,1445,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="guibei">
<emphasis id="0E95EABDFFD0FFC3FE90FA62AFBCFA67" box="[319,420,1445,1472]" italics="true" pageId="15" pageNumber="560">A. guibei</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FE1EFA62AD89FA67" authority="Wood, Grismer, Ahmad &amp; Senawi" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[433,913,1445,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latiffi">
<emphasis id="0E95EABDFFD0FFC3FE1EFA62AC14FA67" box="[433,524,1445,1472]" italics="true" pageId="15" pageNumber="560">A. latiffi</emphasis>
Wood, Grismer, Ahmad &amp; Senawi
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC33FA62AA77FA18" authority="Inger" authorityName="Inger" authorityYear="1960" box="[924,1135,1445,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="platysoma">
<emphasis id="0E95EABDFFD0FFC3FC33FA62AA35FA67" box="[924,1069,1445,1472]" italics="true" pageId="15" pageNumber="560">A. platysoma</emphasis>
Inger
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FB0BFA62AB35FA67" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[1188,1325,1445,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="spinulifer">
<emphasis id="0E95EABDFFD0FFC3FB0BFA62AB35FA67" box="[1188,1325,1445,1472]" italics="true" pageId="15" pageNumber="560">A. spinulifer</emphasis>
</taxonomicName>
by the tip of its first finger not reaching the disc of the second finger. The absence of a tarsal ridge in 
<taxonomicName id="FBE14D2CFFD0FFC3FB2DFA0EAB26FA3B" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[1154,1342,1481,1508]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FB2DFA0EAB26FA3B" box="[1154,1342,1481,1508]" italics="true" pageId="15" pageNumber="560">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD0FFC3FAEBFA0DAB84FA3B" bold="true" box="[1348,1436,1481,1507]" pageId="15" pageNumber="560">
<taxonomicNameLabel id="15A657C6FFD0FFC3FAEBFA0DAB84FA3B" box="[1348,1436,1481,1507]" pageId="15" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differentiates it from 
<taxonomicName id="FBE14D2CFFD0FFC3FE24FA2AAC91F9D0" authority="Inger" authorityName="Inger" authorityYear="1960" box="[395,649,1517,1544]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="albomaculata">
<emphasis id="0E95EABDFFD0FFC3FE24FA2AAC5EF9DF" box="[395,582,1517,1544]" italics="true" pageId="15" pageNumber="560">A. albomaculata</emphasis>
Inger
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FD36FA2AAAEDF9DF" authority="Wood, Grismer, Ahmad &amp; Senawi" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[665,1269,1517,1544]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="0E95EABDFFD0FFC3FD36FA2AAD7FF9DF" box="[665,871,1517,1544]" italics="true" pageId="15" pageNumber="560">A. jeetsukumarani</emphasis>
Wood, Grismer, Ahmad &amp; Senawi
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FAAAFA2AAA15F9F3" authority="Grismer, Wood, Aowphol, Cota, Grismer, Murdoch, Aguilar &amp; Grismer" authorityName="Grismer, Wood, Aowphol, Cota, Grismer, Murdoch, Aguilar &amp; Grismer" authorityYear="2016" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="khaochangensis">
<emphasis id="0E95EABDFFD0FFC3FAAAFA2AAEC5F9F3" italics="true" pageId="15" pageNumber="560">A. khaochangensis</emphasis>
Grismer, Wood, Aowphol, Cota, Grismer, Murdoch, Aguilar &amp; Grismer
</taxonomicName>
, 
<emphasis id="0E95EABDFFD0FFC3FBB4F9D6AF0DF997" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FBB4F9D6AA63F9F3" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[1051,1147,1553,1580]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latiffi">A. latiffi</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB29F9D6AB04F9F3" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[1158,1308,1554,1580]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="mcgregori">A. mcgregori</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FA87F9D6AB80F9F3" authorityName="Inger" authorityYear="1960" box="[1320,1432,1554,1580]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="minuta">A. minuta</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FF38F9F2AF0DF997" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[151,277,1589,1616]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="muelleri">A. muelleri</taxonomicName>
</emphasis>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FEE1F9F2AFE0F997" authorityName="Stoliczka" authorityYear="1870" box="[334,504,1590,1616]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="penangensis">
<emphasis id="0E95EABDFFD0FFC3FEE1F9F2AFE0F997" box="[334,504,1590,1616]" italics="true" pageId="15" pageNumber="560">A. penangensis</emphasis>
</taxonomicName>
. Finger tips that are bulbous versus expanded and spatulate differentiate 
<taxonomicName id="FBE14D2CFFD0FFC3FA98F9F2AEE1F9AB" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FA98F9F2AEE1F9AB" italics="true" pageId="15" pageNumber="560">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFD0FFC3FEADF99DAF47F9AC" box="[258,351,1625,1652]" pageId="15" pageNumber="560" rank="species">
<emphasis id="0E95EABDFFD0FFC3FEADF99DAF40F9AB" bold="true" box="[258,344,1625,1651]" pageId="15" pageNumber="560">sp. nov</emphasis>
.
</taxonomicNameLabel>
from 
<emphasis id="0E95EABDFFD0FFC3FE09F99EACDDF9AB" box="[422,709,1625,1652]" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FE09F99EAC28F9AB" authorityName="Inger" authorityYear="1960" box="[422,560,1625,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="hanitschi">A. hanitschi</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FD92F99EACD9F9AB" authorityName="Inger" authorityYear="1966" box="[573,705,1625,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latidisca">A. latidisca</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FD62F99EAD98F9AC" authority="Inger" authorityName="Inger" authorityYear="1960" box="[717,896,1626,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="minuta">
<emphasis id="0E95EABDFFD0FFC3FD62F99EAD24F9AB" box="[717,828,1626,1652]" italics="true" pageId="15" pageNumber="560">A. minuta</emphasis>
Inger
</taxonomicName>
, 
<emphasis id="0E95EABDFFD0FFC3FC20F99EAA32F9AB" box="[911,1066,1625,1652]" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FC20F99EAA3EF9AB" authorityName="Inger" authorityYear="1960" box="[911,1062,1625,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="platysoma">A. platysoma</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FB9DF99EAAE4F9AB" authority="Dring" authorityName="Dring" authorityYear="1984" box="[1074,1276,1625,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="torrentis">
<emphasis id="0E95EABDFFD0FFC3FB9DF99EAAAAF9AB" box="[1074,1202,1626,1652]" italics="true" pageId="15" pageNumber="560">A. torrentis</emphasis>
Dring
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FA99F99EAB8DF9AB" box="[1334,1429,1625,1652]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="vidua">
<emphasis id="0E95EABDFFD0FFC3FA99F99EAB8DF9AB" box="[1334,1429,1625,1652]" italics="true" pageId="15" pageNumber="560">A. vidua</emphasis>
</taxonomicName>
. The absence of an enlarged row of dorsolateral tubercles in 
<taxonomicName id="FBE14D2CFFD0FFC3FC9FF9BAAA28F94F" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[816,1072,1661,1688]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FC9FF9BAAA28F94F" box="[816,1072,1661,1688]" italics="true" pageId="15" pageNumber="560">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFD0FFC3FB99F9B9AA89F940" box="[1078,1169,1661,1688]" pageId="15" pageNumber="560" rank="species">
<emphasis id="0E95EABDFFD0FFC3FB99F9B9AA92F94F" bold="true" box="[1078,1162,1661,1687]" pageId="15" pageNumber="560">sp. nov</emphasis>
.
</taxonomicNameLabel>
differentiates it from 
<taxonomicName id="FBE14D2CFFD0FFC3FA2BF9BAAFDDF963" authority="Inger &amp; Stuebing" authorityName="Inger &amp; Stuebing" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="echinata">
<emphasis id="0E95EABDFFD0FFC3FA2BF9BAAEEFF963" italics="true" pageId="15" pageNumber="560">A. echinata</emphasis>
Inger &amp; Stuebing
</taxonomicName>
, 
<emphasis id="0E95EABDFFD0FFC3FE7CF966AD17F963" box="[467,783,1697,1724]" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FE7CF966AC6BF963" authorityName="Iskandar &amp; Mumpuni" authorityYear="2004" box="[467,627,1697,1724]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="glandulosa">A. glandulosa</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FDD0F966AD12F963" box="[639,778,1697,1724]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latirostra">A. latirostra</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FCB8F966AAC2F963" authority="Matsui, Khonsue &amp; Panha" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" box="[791,1242,1697,1724]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="pilokensis">
<emphasis id="0E95EABDFFD0FFC3FCB8F966ADBFF963" box="[791,935,1697,1724]" italics="true" pageId="15" pageNumber="560">A. pilokensis</emphasis>
Matsui, Khonsue &amp; Panha
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB46F966AB6DF963" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[1257,1397,1697,1724]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="spinulifer">
<emphasis id="0E95EABDFFD0FFC3FB46F966AB6DF963" box="[1257,1397,1697,1724]" italics="true" pageId="15" pageNumber="560">A. spinulifer</emphasis>
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FA2BF966AF11F907" authorityName="Wilkinson, Sellas &amp; Vindum" authorityYear="2012" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="thinthinae">
<emphasis id="0E95EABDFFD0FFC3FA2BF966AF11F907" italics="true" pageId="15" pageNumber="560">A. thinthinae</emphasis>
</taxonomicName>
and some specimens of 
<taxonomicName id="FBE14D2CFFD0FFC3FDB8F902AC63F907" authorityName="Inger" authorityYear="1966" box="[535,635,1733,1760]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="guibei">
<emphasis id="0E95EABDFFD0FFC3FDB8F902AC63F907" box="[535,635,1733,1760]" italics="true" pageId="15" pageNumber="560">A. guibei</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FD1FF902AB85F907" authority="Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The" authorityName="Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The" authorityYear="2014" box="[688,1437,1733,1760]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="lumut">
<emphasis id="0E95EABDFFD0FFC3FD1FF902AD16F907" box="[688,782,1733,1760]" italics="true" pageId="15" pageNumber="560">A. lumut</emphasis>
Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The
</taxonomicName>
coarsely granular abdomen of 
<taxonomicName id="FBE14D2CFFD0FFC3FE42F92EACB2F8DB" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[493,682,1769,1796]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FE42F92EACB2F8DB" box="[493,682,1769,1796]" italics="true" pageId="15" pageNumber="560">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFD0FFC3FD1EF92DAD14F8DC" box="[689,780,1769,1796]" pageId="15" pageNumber="560" rank="species">
<emphasis id="0E95EABDFFD0FFC3FD1EF92DAD1EF8DB" bold="true" box="[689,774,1769,1795]" pageId="15" pageNumber="560">sp. nov</emphasis>
.
</taxonomicNameLabel>
differentiates it from 
<emphasis id="0E95EABDFFD0FFC3FBACF92EAEE5F8FF" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FBACF92EAA9EF8DB" box="[1027,1158,1769,1796]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="echinata">A. echinata</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB3EF92EAAE8F8DB" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[1169,1264,1769,1796]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latiffi">A. latiffi</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB54F92EAB80F8DB" authorityName="Inger" authorityYear="1960" box="[1275,1432,1769,1796]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="longidigita">A. longidigita</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FF38F8CAAEE1F8FF" authorityName="Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The" authorityYear="2014" box="[151,249,1805,1832]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="lumut">A. lumut</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FEABF8CAACCBF8FF" authority="Matsui, Khonsue &amp; Panha" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" box="[260,723,1805,1832]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="phuketensis">
<emphasis id="0E95EABDFFD0FFC3FEABF8CAAFBEF8FF" box="[260,422,1805,1832]" italics="true" pageId="15" pageNumber="560">A. phuketensis</emphasis>
Matsui, Khonsue &amp; Panha
</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FD4EF8CAADA9F8FF" authority="Kiew" authorityName="Kiew" box="[737,945,1805,1832]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="0E95EABDFFD0FFC3FD4EF8CAAD75F8FF" box="[737,877,1806,1832]" italics="true" pageId="15" pageNumber="560">A. siamensis</emphasis>
Kiew
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FC48F8CAAA5DF8FF" box="[999,1093,1805,1832]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="vidua">
<emphasis id="0E95EABDFFD0FFC3FC48F8CAAA5DF8FF" box="[999,1093,1805,1832]" italics="true" pageId="15" pageNumber="560">A. vidua</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3C5E36AFFFD0FFDCFF68F8F5AB64FE77" blockId="15.[151,1437,832,2012]" lastBlockId="16.[151,1437,153,755]" lastPageId="16" lastPageNumber="561" pageId="15" pageNumber="560">
The presence of large, discrete, bright-yellow, submandibular spots along the underside of the lower jaw (
<figureCitation id="A4DA2A2AFFD0FFC3FAEEF8F5AB8CF893" box="[1345,1428,1841,1867]" captionStart="FIGURE 3" captionStartId="12.[151,250,1975,2000]" captionTargetBox="[173,1415,669,1956]" captionTargetId="figure@12.[173,1415,669,1956]" captionTargetPageId="12" captionText="FIGURE 3. Dorsum (A) and venter (B) of type series of Ansonia kyaiktiyoensis sp. nov. Paratype (LSUHC 14067) on the left and holotype (LSUHC 14066) on the right." httpUri="https://zenodo.org/record/3368951/files/figure.png" pageId="15" pageNumber="560">Fig. 3B</figureCitation>
) distinguishes 
<taxonomicName id="FBE14D2CFFD0FFC3FE81F892AFFFF8B7" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[302,487,1877,1904]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FE81F892AFFFF8B7" box="[302,487,1877,1904]" italics="true" pageId="15" pageNumber="560">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFD0FFC3FE40F891AC50F8B7" bold="true" box="[495,584,1877,1903]" pageId="15" pageNumber="560">
<taxonomicNameLabel id="15A657C6FFD0FFC3FE40F891AC50F8B7" box="[495,584,1877,1903]" pageId="15" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from all its congeners except the Thai and Burmese species; 
<emphasis id="0E95EABDFFD0FFC3FB5AF892AC53F84B" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FB5AF892AB61F8B7" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[1269,1401,1877,1904]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="inthanon">A. inthanon</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FA2BF892AF1FF84B" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="pilokensis">A. pilokensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FEBCF8BEAFADF84B" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" box="[275,437,1913,1940]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="phuketensis">A. phuketensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FE6EF8BEAC53F84B" box="[449,587,1914,1940]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="siamensis">A. siamensis</taxonomicName>
</emphasis>
and 
<taxonomicName id="FBE14D2CFFD0FFC3FD2CF8BEAD0AF84B" authorityName="Wilkinson, Sellas &amp; Vindum" authorityYear="2012" box="[643,786,1913,1940]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="thinthinae">
<emphasis id="0E95EABDFFD0FFC3FD2CF8BEAD0AF84B" box="[643,786,1913,1940]" italics="true" pageId="15" pageNumber="560">A. thinthinae</emphasis>
</taxonomicName>
. The presence of a light spot between the scapulae further differentiates 
<taxonomicName id="FBE14D2CFFD0FFC3FE82F85AAFFDF86F" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[301,485,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFD0FFC3FE82F85AAFFDF86F" box="[301,485,1949,1976]" italics="true" pageId="15" pageNumber="560">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFD0FFC3FE43F859AC5EF860" box="[492,582,1949,1976]" pageId="15" pageNumber="560" rank="species">
<emphasis id="0E95EABDFFD0FFC3FE43F859AC27F86F" bold="true" box="[492,575,1949,1975]" pageId="15" pageNumber="560">sp. nov</emphasis>
.
</taxonomicNameLabel>
from 
<emphasis id="0E95EABDFFD0FFC3FD26F85AAAD3F86F" box="[649,1227,1949,1976]" italics="true" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FD26F85AAD58F86F" authorityName="Inger" authorityYear="1960" box="[649,832,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="albomaculata">A. albomaculata</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FCE4F85AADD3F86F" box="[843,971,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="echinata">A. echinata</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC7AF85AAA4FF86F" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[981,1111,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="fuliginea">A. fuliginea</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FBCDF85AAADFF86F" authorityName="Inger" authorityYear="1966" box="[1122,1223,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="guibei">A. guibei</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFD0FFC3FB7DF85AAB8EF860" authority="Inger" authorityName="Inger" authorityYear="1960" box="[1234,1430,1949,1976]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="hanitschi">
<emphasis id="0E95EABDFFD0FFC3FB7DF85AAB4CF86F" box="[1234,1364,1949,1976]" italics="true" pageId="15" pageNumber="560">A. hanitschi</emphasis>
Inger
</taxonomicName>
, 
<emphasis id="0E95EABDFFD0FFDCFF38F806AEF0FF6B" italics="true" lastPageId="16" lastPageNumber="561" pageId="15" pageNumber="560">
<taxonomicName id="FBE14D2CFFD0FFC3FF38F806AF7FF803" authorityName="Grismer, Wood, Aowphol, Cota, Grismer, Murdoch, Aguilar &amp; Grismer" authorityYear="2016" box="[151,359,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="khaochangensis">A. khaochangensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FEDEF806AFF7F803" authorityName="Inger" authorityYear="1966" box="[369,495,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="latidisca">A. latidisca</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FE56F806AC6EF803" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[505,630,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="leptopus">A. leptopus</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FD2FF806AD00F803" authorityName="Inger" authorityYear="1960" box="[640,792,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="longidigita">A. longidigita</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC8DF806ADACF803" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[802,948,1986,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="mcgregori">A. mcgregori</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FC11F806AA33F803" authorityName="Inger" authorityYear="1960" box="[958,1067,1986,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="minuta">A. minuta</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB9AF806AAA9F803" box="[1077,1201,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="meulleri">A. meulleri</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFD0FFC3FB15F806AB53F803" authorityName="Inger" authorityYear="1960" box="[1210,1355,1985,2012]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="15" pageNumber="560" phylum="Chordata" rank="species" species="platysoma">A. platysoma</taxonomicName>
, A. sia- mensis,
</emphasis>
<taxonomicName id="FBE14D2CFFCFFFDCFF40FF5DAC16FF6B" authority="Hendrickson" authorityName="Hendrickson" authorityYear="1966" box="[239,526,153,179]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="tiomanica">
<emphasis id="0E95EABDFFCFFFDCFF40FF5DAF62FF6B" box="[239,378,153,179]" italics="true" pageId="16" pageNumber="561">A. tiomanica</emphasis>
Hendrickson
</taxonomicName>
, 
<emphasis id="0E95EABDFFCFFFDCFDB4FF5DACE6FF6B" box="[539,766,153,179]" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFDB4FF5DAC80FF6B" authorityName="Dring" authorityYear="1984" box="[539,664,153,179]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="torrentis">A. torrentis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFD0CFF5DACE6FF6B" box="[675,766,153,179]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="vidua">A. vidua</taxonomicName>
</emphasis>
and some specimens of 
<taxonomicName id="FBE14D2CFFCFFFDCFBA4FF5DAB1CFF6B" authority="Grismer" authorityName="Grismer" authorityYear="2006" box="[1035,1284,153,179]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="endauensis">
<emphasis id="0E95EABDFFCFFFDCFBA4FF5DAABBFF6B" box="[1035,1187,153,179]" italics="true" pageId="16" pageNumber="561">A. endauensis</emphasis>
Grismer
</taxonomicName>
, 
<emphasis id="0E95EABDFFCFFFDCFABEFF5DAEE2FF0F" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFABEFF5DAB80FF6B" box="[1297,1432,153,179]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="latirostra">A. latirostra</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFF38FF79AEEEFF0F" authorityName="Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The" authorityYear="2014" box="[151,246,189,215]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="lumut">A. lumut</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFCFFFDCFEAFFF79AC0AFF0F" authority="Stoliczka" authorityName="Stoliczka" authorityYear="1870" box="[256,530,189,215]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="penangensis">
<emphasis id="0E95EABDFFCFFFDCFEAFFF79AFBEFF0F" box="[256,422,189,215]" italics="true" pageId="16" pageNumber="561">A. penangensis</emphasis>
Stoliczka
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFCFFFDCFDE9FF79AB84FF0F" authority="Davis, Grismer, Klabacka, Muin, Quah, Anuar, Wood &amp; Sites. The" authorityName="Davis, Grismer, Klabacka, Muin, Quah, Anuar, Wood &amp; Sites. The" box="[582,1436,189,215]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="smeagol">
<emphasis id="0E95EABDFFCFFFDCFDE9FF79ACD8FF0F" box="[582,704,189,215]" italics="true" pageId="16" pageNumber="561">A. smeagol</emphasis>
Davis, Grismer, Klabacka, Muin, Quah, Anuar, Wood &amp; Sites. The
</taxonomicName>
yellow-gold coloured iris further separates it from 
<emphasis id="0E95EABDFFCFFFDCFD6CFF25AA52FEC7" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFD6CFF25AD46FF23" authorityName="Grismer" authorityYear="2006" box="[707,862,225,251]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="endauensis">A. endauensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFCC6FF25AA2AFF23" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[873,1074,225,251]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="jeetsukumarani">A. jeetsukumarani</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFB92FF25AB16FF23" authorityName="Grismer, Wood, Aowphol, Cota, Grismer, Murdoch, Aguilar &amp; Grismer" authorityYear="2016" box="[1085,1294,225,251]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="khaochangensis">A. khaochangensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFAB6FF25AB80FF23" authorityName="Inger" authorityYear="1966" box="[1305,1432,225,251]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="latidisca">A. latidisca</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFF38FEC1AEEBFEC7" authorityName="Wood, Grismer, Ahmad &amp; Senawi" authorityYear="2008" box="[151,243,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="latiffi">A. latiffi</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFF52FEC1AF9BFEC7" box="[253,387,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="latirostra">A. latirostra</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFE22FEC1AFF4FEC7" authorityName="Chan, Wood, Anuar, Muin, Quah, Sumarli &amp; Grismer. The" authorityYear="2014" box="[397,492,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="lumut">A. lumut</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFE58FEC1AC91FEC7" baseAuthorityName="Taylor" baseAuthorityYear="1922" box="[503,649,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="mcgregori">A. mcgregori</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFD3DFEC1AD16FEC7" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[658,782,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="muelleri">A. muelleri</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFCB7FEC1ADD8FEC7" authorityName="Stoliczka" authorityYear="1870" box="[792,960,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFC65FEC1AA5EFEC7" box="[970,1094,261,287]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="smeagol">A. smeagol</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFCFFFDCFBFFFEC1AC2FFE9B" authority="Waser, Schweizer, Haas, Das, Jankowski, Pui &amp; Hertwig" authorityName="Waser, Schweizer, Haas, Das, Jankowski, Pui &amp; Hertwig" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="teneritas">
<emphasis id="0E95EABDFFCFFFDCFBFFFEC1AAD6FEC7" box="[1104,1230,261,287]" italics="true" pageId="16" pageNumber="561">A. teneritas</emphasis>
Waser, Schweizer, Haas, Das, Jankowski, Pui &amp; Hertwig
</taxonomicName>
, 
<emphasis id="0E95EABDFFCFFFDCFDEBFEEDAD4EFE9B" box="[580,854,297,323]" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFDEBFEEDACCBFE9B" authorityName="Hendrickson" authorityYear="1966" box="[580,723,297,323]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="tiomanica">A. tiomanica</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFD71FEEDAD4EFE9B" box="[734,854,297,323]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="torrestis">A. torrestis</taxonomicName>
</emphasis>
and 
<taxonomicName id="FBE14D2CFFCFFFDCFC24FEEDADFFFE9B" box="[907,999,297,323]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="vidua">
<emphasis id="0E95EABDFFCFFFDCFC24FEEDADFFFE9B" box="[907,999,297,323]" italics="true" pageId="16" pageNumber="561">A. vidua</emphasis>
</taxonomicName>
that have iris colours ranging from reddish gold to black. The absence of a wide, light-coloured patch below the eye separates 
<taxonomicName id="FBE14D2CFFCFFFDCFBE2FE89AB1CFEBF" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[1101,1284,333,359]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFCFFFDCFBE2FE89AB1CFEBF" box="[1101,1284,333,359]" italics="true" pageId="16" pageNumber="561">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFCFFFDCFAA5FE89AB79FEBF" bold="true" box="[1290,1377,333,359]" pageId="16" pageNumber="561">
<taxonomicNameLabel id="15A657C6FFCFFFDCFAA5FE89AB79FEBF" box="[1290,1377,333,359]" pageId="16" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from 
<emphasis id="0E95EABDFFCFFFDCFF38FEB5AFF2FE53" box="[151,490,369,395]" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFF38FEB5AF48FE53" authorityName="Inger" authorityYear="1960" box="[151,336,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="albomaculata">A. albomaculata</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFEF2FEB5AFF2FE53" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" box="[349,490,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="pilokensis">A. pilokensis</taxonomicName>
</emphasis>
, 
<emphasis id="0E95EABDFFCFFFDCFE55FEB5AD3FFE53" box="[506,807,369,395]" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFE55FEB5AC84FE53" authorityName="Matsui, Khonsue &amp; Panha" authorityYear="2018" box="[506,668,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="phuketensis">A. phuketensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFD07FEB5AD3FFE53" box="[680,807,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="teneritas">A. teneritas</taxonomicName>
</emphasis>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFC98FEB5ADDEFE53" authorityName="Wilkinson, Sellas &amp; Vindum" authorityYear="2012" box="[823,966,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="thinthinae">
<emphasis id="0E95EABDFFCFFFDCFC98FEB5ADDEFE53" box="[823,966,369,395]" italics="true" pageId="16" pageNumber="561">A. thinthinae</emphasis>
</taxonomicName>
and some specimens of 
<emphasis id="0E95EABDFFCFFFDCFB73FEB5AC01FE77" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFB73FEB5AB60FE53" authorityName="Grismer" authorityYear="2006" box="[1244,1400,369,395]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="endauensis">A. endauensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFA2BFEB5AEE6FE77" authorityName="Inger" authorityYear="1960" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="hanitschi">A. hanitschi</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFEA7FE51AF9DFE77" authorityName="Matsui, Khonsue &amp; Nabhitabhata" authorityYear="2005" box="[264,389,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kraensis">A. kraensis</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFE20FE51AC0DFE77" box="[399,533,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="latirostra">A. latirostra</taxonomicName>
,
</emphasis>
<taxonomicName id="FBE14D2CFFCFFFDCFD8FFE51ACF3FE77" authority="Inger" authorityName="Inger" authorityYear="1960" box="[544,747,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="malayana">
<emphasis id="0E95EABDFFCFFFDCFD8FFE51ACB2FE77" box="[544,682,405,431]" italics="true" pageId="16" pageNumber="561">A. malayana</emphasis>
Inger
</taxonomicName>
, 
<emphasis id="0E95EABDFFCFFFDCFD58FE51AA0CFE77" box="[759,1044,405,431]" italics="true" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFD58FE51AD7DFE77" authorityName="Inger" authorityYear="1960" box="[759,869,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="minuta">A. minuta</taxonomicName>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFCC0FE51AA0CFE77" authorityName="Stoliczka" authorityYear="1870" box="[879,1044,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
</emphasis>
, 
<taxonomicName id="FBE14D2CFFCFFFDCFB8DFE51AAADFE77" authorityName="Inger" authorityYear="1960" box="[1058,1205,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="platysoma">
<emphasis id="0E95EABDFFCFFFDCFB8DFE51AAADFE77" box="[1058,1205,405,431]" italics="true" pageId="16" pageNumber="561">A. platysoma</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFCFFFDCFB45FE51AB6DFE77" authorityName="Hendrickson" authorityYear="1966" box="[1258,1397,405,431]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="tiomanica">
<emphasis id="0E95EABDFFCFFFDCFB45FE51AB6DFE77" box="[1258,1397,405,431]" italics="true" pageId="16" pageNumber="561">A. tiomanica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="74FB6524FFCFFFDCFF68FE7DAFDFFD2B" pageId="16" pageNumber="561" type="reference_group">
<paragraph id="3C5E36AFFFCFFFDCFF68FE7DAFDFFD2B" blockId="16.[151,1437,153,755]" pageId="16" pageNumber="561">
<taxonomicName id="FBE14D2CFFCFFFDCFF68FE7DAFDFFE0B" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[199,455,441,467]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFCFFFDCFF68FE7DAFDFFE0B" box="[199,455,441,467]" italics="true" pageId="16" pageNumber="561">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFCFFFDCFE60FE7DAC32FE0B" box="[463,554,441,467]" pageId="16" pageNumber="561" rank="species">
<emphasis id="0E95EABDFFCFFFDCFE60FE7DAC3CFE0B" bold="true" box="[463,548,441,467]" pageId="16" pageNumber="561">sp. nov</emphasis>
.
</taxonomicNameLabel>
most closely resembles 
<taxonomicName id="FBE14D2CFFCFFFDCFC92FE7DADA1FE0B" ID-CoL="679WN" authorityName="Matsui, Khonsue &amp; Nabhitabhata" authorityYear="2005" box="[829,953,441,467]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kraensis">
<emphasis id="0E95EABDFFCFFFDCFC92FE7DADA1FE0B" box="[829,953,441,467]" italics="true" pageId="16" pageNumber="561">A. kraensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="FBE14D2CFFCFFFDCFC5EFE7DAA6CFE0B" ID-CoL="679WM" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[1009,1140,441,467]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="0E95EABDFFCFFFDCFC5EFE7DAA6CFE0B" box="[1009,1140,441,467]" italics="true" pageId="16" pageNumber="561">A. inthanon</emphasis>
</taxonomicName>
in morphology but can be distinguished from the former by having yellow spots and blotches on the venter that are especially prominent near the flanks (
<figureCitation id="A4DA2A2AFFCFFFDCFEB7FDC5AF77FDC3" box="[280,367,513,539]" captionStart="FIGURE 3" captionStartId="12.[151,250,1975,2000]" captionTargetBox="[173,1415,669,1956]" captionTargetId="figure@12.[173,1415,669,1956]" captionTargetPageId="12" captionText="FIGURE 3. Dorsum (A) and venter (B) of type series of Ansonia kyaiktiyoensis sp. nov. Paratype (LSUHC 14067) on the left and holotype (LSUHC 14066) on the right." httpUri="https://zenodo.org/record/3368951/files/figure.png" pageId="16" pageNumber="561">Fig. 3B</figureCitation>
) as opposed to the venter being silvery white and mottled in dark-brown (
<bibRefCitation id="58704B5EFFCFFFDCFB6DFDC5AB80FDC3" author="Matsui, M. &amp; Khonsue, W. &amp; Nabhitabhata, J." box="[1218,1432,512,539]" pageId="16" pageNumber="561" pagination="809 - 814" refId="ref17626" refString="Matsui, M., Khonsue, W. &amp; Nabhitabhata, J. (2005) A new Ansonia from the Isthmus of Kra, Thailand (Amphibia, Anura, Bufonidae). Zoological Science, 22, 809 - 814. http: // doi. org / 10.2108 / zsj. 22.80 9" type="journal article" year="2005">
Matsui 
<emphasis id="0E95EABDFFCFFFDCFAB5FDC6AB55FDC3" box="[1306,1357,513,539]" italics="true" pageId="16" pageNumber="561">et al</emphasis>
. 2005
</bibRefCitation>
: 
<figureCitation id="A4DA2A2AFFCFFFDCFF38FDE1AEF5FDE7" box="[151,237,549,575]" captionStart="FIGURE 1" captionStartId="9.[151,250,1895,1920]" captionTargetBox="[341,1232,174,1873]" captionTargetId="figure@9.[336,1251,166,1875]" captionTargetPageId="9" captionText="FIGURE 1. Maximum likelihood (ML) phylogram for species of the genus Ansonia. Nodal support is indicated by Bayesian posterior probability (BPP) / Ultrafast bootstrap (UFBOOT). Black circles represent nodes supported by a BPP and UFB of 1.00 and 100, respectively. Blue node indicates the Thai-Malay Peninsula clade and red node indicates the Thai-Burmese clade. Species outside of the Thai-Malay Peninsula clade are represented by single tip labels for their lineages." httpUri="https://zenodo.org/record/3368945/files/figure.png" pageId="16" pageNumber="561">Fig. 1B</figureCitation>
). From its sister species 
<taxonomicName id="FBE14D2CFFCFFFDCFDACFDE1AC9EFDE7" ID-CoL="679WM" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[515,646,549,575]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="0E95EABDFFCFFFDCFDACFDE1AC9EFDE7" box="[515,646,549,575]" italics="true" pageId="16" pageNumber="561">A. inthanon</emphasis>
</taxonomicName>
, it can be distinguished by the presence of dark markings on the snout and lips (
<figureCitation id="A4DA2A2AFFCFFFDCFEADFD8DAF50FDBB" box="[258,328,585,611]" captionStart="FIGURE 4" captionStartId="13.[151,250,1871,1896]" captionTargetBox="[151,1435,345,1852]" captionTargetId="figure@13.[151,1435,345,1852]" captionTargetPageId="13" captionText="FIGURE 4. (A) Calling adult male Ansonia inthanon from Doi Inthanon, Chiang Mai, Thailand. (B) Close-up of head of para- type of Ansonia kyaiktiyoensis sp. nov. (LSUHC 14067). Note the markings on the snout and under the eye as indicated by the arrows. (C) Close-up of head of A. inthanon from Doi Inthanon, Chiang Mai, Thailand. Note the absence of markings on the head. (Photographs by Ian Dugdale [A &amp; C] and Evan Quah [B])." httpUri="https://zenodo.org/record/3368953/files/figure.png" pageId="16" pageNumber="561">Fig. 4</figureCitation>
), a dark dorsolateral stripe, and less extensive webbing on the toes (
<tableCitation id="71630314FFCFFFDCFBFBFD8DAAB5FDBB" box="[1108,1197,585,611]" captionStart="TABLE 3" captionStartId="7.[151,227,479,501]" captionTargetPageId="7" captionText="TABLE 3. Uncorrected p-distances calculated for Ansonia using MEGA v6.06 (Tamura et al. 2013)." httpUri="http://table.plazi.org/id/689E6627FFD8FFCBFF38FE1BAA35FE2D" pageId="16" pageNumber="561" tableUuid="689E6627FFD8FFCBFF38FE1BAA35FE2D">Table 3</tableCitation>
; 
<bibRefCitation id="58704B5EFFCFFFDCFB14FD8DAB88FDBB" author="Matsui, M. &amp; Nabhitabhata, J. &amp; Panha, S." box="[1211,1424,585,611]" pageId="16" pageNumber="561" pagination="448 - 454" refId="ref17737" refString="Matsui, M., Nabhitabhata, J. &amp; Panha, S. (1998) A new Ansonia from northern Thailand (Anura, Bufonidae). Herpetologica, 54, 448 - 454." type="journal article" year="1998">
Matsui 
<emphasis id="0E95EABDFFCFFFDCFABDFD8EAB5CFDBB" box="[1298,1348,585,611]" italics="true" pageId="16" pageNumber="561">et al</emphasis>
. 1998
</bibRefCitation>
). 
<taxonomicName id="FBE14D2CFFCFFFDCFF38FDA9AF4CFD5F" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[151,340,621,647]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFCFFFDCFF38FDA9AF4CFD5F" box="[151,340,621,647]" italics="true" pageId="16" pageNumber="561">A. kyaiktiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A657C6FFCFFFDCFEF3FDA9AFA0FD5F" box="[348,440,621,647]" pageId="16" pageNumber="561" rank="species">
<emphasis id="0E95EABDFFCFFFDCFEF3FDA9AFA9FD5F" bold="true" box="[348,433,621,647]" pageId="16" pageNumber="561">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from 
<taxonomicName id="FBE14D2CFFCFFFDCFDE0FDA9ACCBFD5F" ID-CoL="679WM" authorityName="Matsui, Nabhitabhata &amp; Panha" authorityYear="1998" box="[591,723,621,647]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="0E95EABDFFCFFFDCFDE0FDA9ACCBFD5F" box="[591,723,621,647]" italics="true" pageId="16" pageNumber="561">A. inthanon</emphasis>
</taxonomicName>
in the number of free phalanges on the second toe (0.5–1.5 vs. 0.5–1); third toe (1.5–2.5 or 3.0 vs. 0.5–2.00); fourth toe (3 vs. 2.66–2.75); and fifth toe (1.5 vs. 0.5) (
<tableCitation id="71630314FFCFFFDCFAA9FD55AB42FD73" box="[1286,1370,657,683]" captionStart="TABLE 4" captionStartId="14.[151,239,154,178]" captionTargetPageId="14" captionText="TABLE 4. Selected measurements of the type series of Ansonia kyaiktiyoensis sp. nov. / = no data available." httpUri="http://table.plazi.org/id/689E6627FFD1FFC2FF38FF5EAB38FF6A" pageId="16" pageNumber="561" tableUuid="689E6627FFD1FFC2FF38FF5EAB38FF6A">Table 4</tableCitation>
&amp; 
<tableCitation id="71630314FFCFFFDCFAD1FD54AB96FD73" box="[1406,1422,656,683]" captionStart="TABLE 5" captionStartId="18.[151,225,499,521]" captionTargetPageId="18" captionText="TABLE 5. Diagnostic characters differentiating the species of Ansonia kyaiktiyoensis sp. nov. from all other nominal species of Ansonia. Bold character states are those that discretely separate A. kyaiktiyoensis sp. nov. / = no data available" httpUri="http://table.plazi.org/id/689E6627FFCDFFDEFF38FE37AF84FDFF" pageId="16" pageNumber="561" tableUuid="689E6627FFCDFFDEFF38FE37AF84FDFF">5</tableCitation>
). Other diagnostic characters that separate 
<taxonomicName id="FBE14D2CFFCFFFDCFDCFFD71AD47FD17" authority="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin, 2019" authorityName="Quah &amp; Grismer &amp; Wood &amp; Thura &amp; Oaks &amp; Lin" authorityYear="2019" box="[608,863,693,719]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="species" species="kyaiktiyoensis" status="sp. nov.">
<emphasis id="0E95EABDFFCFFFDCFDCFFD71AD47FD17" box="[608,863,693,719]" italics="true" pageId="16" pageNumber="561">Ansonia kyaiktiyoensis</emphasis>
</taxonomicName>
<emphasis id="0E95EABDFFCFFFDCFCC9FD71ADA7FD17" bold="true" box="[870,959,693,719]" pageId="16" pageNumber="561">
<taxonomicNameLabel id="15A657C6FFCFFFDCFCC9FD71ADA7FD17" box="[870,959,693,719]" pageId="16" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the other nominal species of 
<taxonomicName id="FBE14D2CFFCFFFDCFAEFFD71AB84FD17" ID-CoL="X5N" authorityName="Stoliczka" authorityYear="1870" box="[1344,1436,693,719]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="16" pageNumber="561" phylum="Chordata" rank="genus">
<emphasis id="0E95EABDFFCFFFDCFAEFFD71AB84FD17" box="[1344,1436,693,719]" italics="true" pageId="16" pageNumber="561">Ansonia</emphasis>
</taxonomicName>
are summarized in 
<tableCitation id="71630314FFCFFFDCFEC3FD1DAFD9FD2B" box="[364,449,728,755]" captionStart="TABLE 5" captionStartId="18.[151,225,499,521]" captionTargetPageId="18" captionText="TABLE 5. Diagnostic characters differentiating the species of Ansonia kyaiktiyoensis sp. nov. from all other nominal species of Ansonia. Bold character states are those that discretely separate A. kyaiktiyoensis sp. nov. / = no data available" httpUri="http://table.plazi.org/id/689E6627FFCDFFDEFF38FE37AF84FDFF" pageId="16" pageNumber="561" tableUuid="689E6627FFCDFFDEFF38FE37AF84FDFF">Table 5</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>