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<document id="76990AC5569B23366F677DDB1F6F5DB8" ID-CLB-Dataset="6170" ID-DOI="10.11646/zootaxa.4975.2.1" ID-GBIF-Dataset="b98c5a41-914b-4398-8dca-2cf9b36d1817" ID-ISSN="1175-5326" ID-Zenodo-Dep="4806677" ID-ZooBank="6EB83A89-CC46-4F4E-99D5-B180A4677B7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622019802693" checkinUser="plazi" docAuthor="Parnaby, Harry &amp; Gill, Anthony C." docDate="2021" docId="B42F87F7FF85151FFF7EFAE29BFF8DA6" docLanguage="en" docName="zootaxa.4975.2.1.pdf" docOrigin="Zootaxa 4975 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macropus jukesii Miklouho-Maclay 1885" docType="treatment" docVersion="9" lastPageNumber="234" masterDocId="4816FF8FFF9A153EFFE9FF84996E8A20" masterDocTitle="Mammal type specimens in the Macleay Collections, University of Sydney" masterLastPageNumber="252" masterPageNumber="201" pageNumber="232" updateTime="1698991120771" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8724FAD332223B7AC4391868BDE8CF35">Mammal type specimens in the Macleay Collections, University of Sydney</mods:title>
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<mods:namePart id="77662C181F01E83A5F9F4E3DA2360DF3">Parnaby, Harry</mods:namePart>
<mods:affiliation id="1D0D379F30E8EC554C776478A902A214">Mammalogy, Australian Museum Research Institute, Australian Museum, 1 William St, Sydney, NSW 2010, Australia.</mods:affiliation>
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<mods:namePart id="65918124AAD516B4B2208D8C47345495">Gill, Anthony C.</mods:namePart>
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<mods:affiliation id="693799EF611CDB14B9981435339F9944">Macleay Collections, Chau Chak Wing Museum, University Avenue, The University of Sydney, NSW 2006, Australia. Anthony. c. gill @ sydney. edu. au; https: // orcid. org / 0000 - 0002 - 8990 - 3657 &amp; Ichthyology, Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney, NSW 2010.</mods:affiliation>
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<treatment id="B42F87F7FF85151FFF7EFAE29BFF8DA6" ID-DOI="http://doi.org/10.5281/zenodo.4806761" ID-GBIF-Taxon="182645612" ID-Zenodo-Dep="4806761" LSID="urn:lsid:plazi:treatment:B42F87F7FF85151FFF7EFAE29BFF8DA6" httpUri="http://treatment.plazi.org/id/B42F87F7FF85151FFF7EFAE29BFF8DA6" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232">
<subSubSection id="749C656AFF851521FF7EFAE29AD38FE2" pageId="31" pageNumber="232" type="nomenclature">
<paragraph id="3C3936E1FF851521FF7EFAE29BC78FA0" blockId="31.[151,681,1382,1409]" box="[151,681,1382,1409]" pageId="31" pageNumber="232">
<heading id="6771818DFF851521FF7EFAE29BC78FA0" bold="true" box="[151,681,1382,1409]" fontSize="11" level="1" pageId="31" pageNumber="232" reason="1">
<taxonomicName id="FB864D62FF851521FF7EFAE29BC78FA0" authority="Miklouho-Maclay, 1885 b" authorityName="Miklouho-Maclay" authorityYear="1885" box="[151,681,1382,1409]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FF851521FF7EFAE29BC78FA0" bold="true" box="[151,681,1382,1409]" pageId="31" pageNumber="232">
<emphasis id="0EF2EAF3FF851521FF7EFAE298078FA1" bold="true" box="[151,361,1382,1409]" italics="true" pageId="31" pageNumber="232">Macropus jukesii</emphasis>
<bibRefCitation id="58174B10FF851521FE99FAE29BC78FA0" author="Miklouho-Maclay, N." box="[368,681,1382,1408]" pageId="31" pageNumber="232" pagination="890 - 895" refId="ref38519" refString="Miklouho-Maclay, N. (1885 b) On two new species of Macropus from the south coast of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (4), 890 - 895, pl. 39." type="journal article" year="1885">Miklouho-Maclay, 1885b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3C3936E1FF851521FF7EFA2F9AD38FE2" blockId="31.[151,957,1450,1475]" box="[151,957,1450,1475]" pageId="31" pageNumber="232">
<emphasis id="0EF2EAF3FF851521FF7EFA2F98EB8FE2" box="[151,389,1451,1474]" italics="true" pageId="31" pageNumber="232">Proc. Linn. Soc. N.S.W.</emphasis>
(ser. 1) 9 (4), 891, plate 39, figs 16. (
<date id="48381021FF851521FCE3FA2F9ADF8FE2" box="[778,945,1450,1475]" pageId="31" pageNumber="232" value="1885-03-04">4th March 1885</date>
).
</paragraph>
</subSubSection>
<subSubSection id="749C656AFF85151EFF7EFA759D4388A5" lastPageId="32" lastPageNumber="233" pageId="31" pageNumber="232" type="description">
<paragraph id="3C3936E1FF851521FF7EFA759ACE8C2B" blockId="31.[151,928,1521,1547]" box="[151,928,1521,1547]" pageId="31" pageNumber="232">
Description read at the
<date id="48381021FF851521FE75FA759BE28C2B" box="[412,652,1521,1547]" pageId="31" pageNumber="232" value="1884-09-24">24th September 1884</date>
meeting of the LSNSW.
</paragraph>
<paragraph id="3C3936E1FF851521FF7EF9BD9C6C8C73" blockId="31.[151,1437,1593,2015]" box="[151,1282,1593,1619]" pageId="31" pageNumber="232">
<emphasis id="0EF2EAF3FF851521FF7EF9BD98268C73" bold="true" box="[151,328,1593,1619]" pageId="31" pageNumber="232">Current name.</emphasis>
Dusky Pademelon
<taxonomicName id="FB864D62FF851521FDCAF9BD9ACE8C73" authority="(Schreber, 1778)" baseAuthorityName="Schreber" baseAuthorityYear="1778" box="[547,928,1593,1619]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="brunii">
<emphasis id="0EF2EAF3FF851521FDCAF9BD9BB08C73" box="[547,734,1593,1619]" italics="true" pageId="31" pageNumber="232">Thylogale brunii</emphasis>
(
<bibRefCitation id="58174B10FF851521FD05F9BD9AF68C73" author="Schreber, J. C. D. von" box="[748,920,1593,1619]" pageId="31" pageNumber="232" refId="ref40096" refString="Schreber, J. C. D. von (1778) Die Saugethiere in Abbildungen nach der Natur, mit Beschreibungen. Vol. 3. Walfgang Walther, Erlangen, Germany, pp. 551, pl. cliii. [as Der Filander, Didelphys Brunii] https: // doi. org / 10.5962 / bhl. title. 67399" type="book" year="1778">Schreber, 1778</bibRefCitation>
)
</taxonomicName>
following
<bibRefCitation id="58174B10FF851521FBF2F9BD9D908C73" author="Flannery, T. F." box="[1051,1278,1593,1619]" pageId="31" pageNumber="232" pagination="7 - 23" refId="ref35378" refString="Flannery, T. F. (1992) Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy, 15, 7 - 23." type="journal article" year="1992">Flannery (1992: 12)</bibRefCitation>
.
</paragraph>
<paragraph id="3C3936E1FF851521FF2EF9D99A558CC3" blockId="31.[151,1437,1593,2015]" pageId="31" pageNumber="232">
<emphasis id="0EF2EAF3FF851521FF2EF9D998CF8C57" bold="true" box="[199,417,1629,1655]" pageId="31" pageNumber="232">Taxonomic status.</emphasis>
First relegated to synonymy by
<bibRefCitation id="58174B10FF851521FCF5F9D99AA38C57" author="Thomas, O." box="[796,973,1629,1655]" pageId="31" pageNumber="232" refId="ref41096" refString="Thomas, O. (1888) Catalogue of the Marsupialia and Monotremata in the Collection of the British Museum (Natural History). British Museum (Natural History), London, xii + 401 pp. https: // doi. org / 10.5962 / bhl. title. 56181" type="book" year="1888">Thomas (1888)</bibRefCitation>
, who tentatively concluded, without being able to examine the
<typeStatus id="E33D8843FF851521FE4EF905988B8CBB" box="[423,485,1665,1691]" pageId="31" pageNumber="232">types</typeStatus>
, that it was a young example of
<taxonomicName id="FB864D62FF851521FCBCF9069D8D8CBC" authority="(Ramsay, 1877 d)" baseAuthorityName="Ramsay" baseAuthorityYear="1877" box="[853,1251,1665,1692]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="browni">
<emphasis id="0EF2EAF3FF851521FCBCF9069D758CBB" box="[853,1051,1665,1691]" italics="true" pageId="31" pageNumber="232">Macropus browni</emphasis>
(
<bibRefCitation id="58174B10FF851521FBC2F9059DB58CBC" author="Ramsay, E. P." box="[1067,1243,1665,1692]" pageId="31" pageNumber="232" pagination="307 - 309" refId="ref39605" refString="Ramsay, E. P. (1877 d) Description of a new species of Halmaturus, from New Ireland. Proceedings of the Linnean Society of New South Wales, Series 1, 1 (4), 307 - 309. https: // doi. org / 10.5962 / bhl. part. 12417" type="journal article" year="1877">Ramsay, 1877d</bibRefCitation>
)
</taxonomicName>
(now
<taxonomicName id="FB864D62FF851521FAC5F905998B8C9F" baseAuthorityName="Ramsay" baseAuthorityYear="1877" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="browni">
<emphasis id="0EF2EAF3FF851521FAC5F905998B8C9F" italics="true" pageId="31" pageNumber="232">Thylogale browni</emphasis>
</taxonomicName>
). Not usually recognised as a species by subsequent taxonomists. Prior to
<bibRefCitation id="58174B10FF851521FBCAF9219DB48C9F" author="Flannery, T. F." box="[1059,1242,1701,1727]" pageId="31" pageNumber="232" pagination="7 - 23" refId="ref35378" refString="Flannery, T. F. (1992) Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy, 15, 7 - 23." type="journal article" year="1992">Flannery (1992)</bibRefCitation>
,
<taxonomicName id="FB864D62FF851521FB0CF9219C408C9F" authorityName="Miklouho-Maclay" authorityYear="1885" box="[1253,1326,1701,1727]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FF851521FB0CF9219C408C9F" box="[1253,1326,1701,1727]" italics="true" pageId="31" pageNumber="232">jukesii</emphasis>
</taxonomicName>
was variously synonymised with
<taxonomicName id="FB864D62FF851521FE47F94E9B378CC3" authority="brunii" box="[430,601,1737,1763]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="subSpecies" species="brunii" subSpecies="brunii">
<emphasis id="0EF2EAF3FF851521FE47F94E9B378CC3" box="[430,601,1737,1763]" italics="true" pageId="31" pageNumber="232">T. brunii brunii</emphasis>
</taxonomicName>
or
<taxonomicName id="FB864D62FF851521FD96F94E9A5A8CC3" box="[639,820,1737,1763]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="subSpecies" species="brunii" subSpecies="browni">
<emphasis id="0EF2EAF3FF851521FD96F94E9A5A8CC3" box="[639,820,1737,1763]" italics="true" pageId="31" pageNumber="232">T. brunii browni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3C3936E1FF85151EFF2EF96898DF8B3D" blockId="31.[151,1437,1593,2015]" lastBlockId="32.[151,1437,151,2013]" lastPageId="32" lastPageNumber="233" pageId="31" pageNumber="232">
<emphasis id="0EF2EAF3FF851521FF2EF96898548D26" bold="true" box="[199,314,1772,1798]" pageId="31" pageNumber="232">
<typeStatus id="E33D8843FF851521FF2EF968985A8D26" box="[199,308,1772,1798]" pageId="31" pageNumber="232" type="syntype">Syntypes</typeStatus>
.
</emphasis>
M.380, female, skin mount without stand, by subsequent determination by
<bibRefCitation id="58174B10FF851521FB46F9699C078D28" author="Stanbury, P. J." box="[1199,1385,1773,1800]" pageId="31" pageNumber="232" pagination="462 - 463" refId="ref40404" refString="Stanbury, P. J. (1969) Type specimens in the Macleay Museum, University of Sydney. IV. Mammals. Proceedings of the Linnean Society of New South Wales, 93 (3), 462 - 463." type="journal article" year="1969">Stanbury (1969)</bibRefCitation>
and
<bibRefCitation id="58174B10FF851521FF7EF895983C8D0B" author="Flannery, T. F." box="[151,338,1809,1835]" pageId="31" pageNumber="232" pagination="7 - 23" refId="ref35378" refString="Flannery, T. F. (1992) Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy, 15, 7 - 23." type="journal article" year="1992">Flannery (1992)</bibRefCitation>
. The skin mount equates to
<taxonomicName id="FB864D62FF851521FD75F8969A0E8D0B" authorityName="Miklouho-Maclay" authorityYear="1885" box="[668,864,1809,1835]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="31" pageNumber="232" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FF851521FD75F8969A0E8D0B" box="[668,864,1809,1835]" italics="true" pageId="31" pageNumber="232">Macropus jukesii</emphasis>
</taxonomicName>
adult stuffed female specimen (a) of the MAMU 1890s Catalogue. M.380.1, a broken cranium, in three main pieces (
<figureCitation id="A4BD2A64FF851521FC65F8B19AB08D6F" box="[908,990,1845,1871]" captionStart="FIGURE 13" captionStartId="33.[151,250,1506,1530]" captionTargetBox="[169,1419,891,1477]" captionTargetId="figure-376@33.[159,1427,885,1483]" captionTargetPageId="33" captionText="FIGURE 13. Cranial fragments and dentary of M380.1, syntype of Macropus jukesii Miklouho-Maclay 1885b [= Thylogale brunii (Schreber, 1778)], extracted from the skin in 1960. A and B, cranium; C, right dentary; D, right maxillary fragment; and E, premaxillary fragment. (left dentary not shown). Scale bar = 1 cm. (© Chau Chak Wing Museum, photo Robert Blackburn 2016)." figureDoi="http://doi.org/10.5281/zenodo.4806954" httpUri="https://zenodo.org/record/4806954/files/figure.png" pageId="31" pageNumber="232">Fig. 13</figureCitation>
), and two dentaries. The MAMU Mammal Register indicates that the skull was removed from skin M.
<quantity id="FB7E9B04FF851521FCB1F8DD9ACF8D53" box="[856,929,1881,1907]" metricMagnitude="0" metricUnit="m" metricValue="9.652" pageId="31" pageNumber="232" unit="in" value="380.0">380 in</quantity>
1960 and given the same number as the skin. In 2016 there were no numbers or writing on the skull or mandibles but two associated numbers on unattached labels are “temp 141” and IRN 121934 (internal record number from EMu database). This specimen is also marked M.380 on a note inside the skull box, in handwriting that appears to be Jenny Andersons, possibly prior to assignment of temporary numbers. The skull and mandible are badly corroded, and salt crystals adhering to the dentition are clearly evident, along with bone scratch marks consistent with those made by a scalpel blade. This skull is presumably the one extracted from the skin marked M.
<quantity id="FB7E9B04FFBA151EFD42FF5B9B9A8AD9" box="[683,756,223,249]" metricMagnitude="0" metricUnit="m" metricValue="9.652" pageId="32" pageNumber="233" unit="in" value="380.0">380 in</quantity>
1960, and its association with the skin had been lost prior to our examination in 2016.
</paragraph>
<paragraph id="3C3936E1FFBA151EFF2EFEA39D4388A5" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
<materialsCitation id="8CEE3CBCFFBA151EFF2EFEA39D4788A5" ID-GBIF-Occurrence="3128590317" collectorName="P. Mpzesly" country="Guinea-Bissau" county="University of Sydney" location="South Coast" municipality="Museum of Geology" pageId="32" pageNumber="233" specimenCount="1" specimenCount-female="1">
M.380.2, by subsequent determination here; cranium and two dentaries (
<figureCitation id="A4BD2A64FFBA151EFC15FEA39D218B61" box="[1020,1103,295,321]" captionStart="FIGURE 14" captionStartId="34.[151,250,1767,1791]" captionTargetBox="[170,1428,206,1730]" captionTargetId="figure-25@34.[151,1436,181,1743]" captionTargetPageId="34" captionText="FIGURE 14. Skull and mandible of M.380.2, syntype of Macropus jukesii Miklouho-Maclay 1885b [= Thylogale brunii (Schreber, 1778)], the skull illustrated in the original account, and associated paper label written by Miklouho-Maclay. Writing on the braincase is likely to be Miklouho-Maclays. Scale bar = 2 cm. (© Chau Chak Wing Museum, photo Robert Blackburn 2016)." figureDoi="http://doi.org/10.5281/zenodo.4806956" httpUri="https://zenodo.org/record/4806956/files/figure.png" pageId="32" pageNumber="233">Fig. 14</figureCitation>
), unsexed, equates to
<taxonomicName id="FB864D62FFBA151EFAA0FEA398618B45" authorityName="Miklouho-Maclay" authorityYear="1885" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBA151EFAA0FEA398618B45" italics="true" pageId="32" pageNumber="233">Macropus jukesii</emphasis>
</taxonomicName>
“(b) skull” of MAMU 1890s Catalogue. An ink inscription of “
<taxonomicName id="FB864D62FFBA151EFC08FECF9DC78B45" authorityName="Miklouho-Maclay" authorityYear="1885" box="[993,1193,331,357]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="jukesii">Macropus jukesii</taxonomicName>
” across the braincase appears to have been written by Miklouho-Maclay (
<figureCitation id="A4BD2A64FFBA151EFD1BFEEB9A268BA9" box="[754,840,367,393]" captionStart="FIGURE 14" captionStartId="34.[151,250,1767,1791]" captionTargetBox="[170,1428,206,1730]" captionTargetId="figure-25@34.[151,1436,181,1743]" captionTargetPageId="34" captionText="FIGURE 14. Skull and mandible of M.380.2, syntype of Macropus jukesii Miklouho-Maclay 1885b [= Thylogale brunii (Schreber, 1778)], the skull illustrated in the original account, and associated paper label written by Miklouho-Maclay. Writing on the braincase is likely to be Miklouho-Maclays. Scale bar = 2 cm. (© Chau Chak Wing Museum, photo Robert Blackburn 2016)." figureDoi="http://doi.org/10.5281/zenodo.4806956" httpUri="https://zenodo.org/record/4806956/files/figure.png" pageId="32" pageNumber="233">Fig. 14</figureCitation>
). The skull was removed from a body preserved in “brine” received from Andrew Goldie (
<bibRefCitation id="58174B10FFBA151EFDBBFE179A1C8B8D" author="Miklouho-Maclay, N." box="[594,882,402,429]" pageId="32" pageNumber="233" pagination="890 - 895" refId="ref38519" refString="Miklouho-Maclay, N. (1885 b) On two new species of Macropus from the south coast of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (4), 890 - 895, pl. 39." type="journal article" year="1885">Miklouho-Maclay 1885b</bibRefCitation>
). The body, which could subsequently have been turned into a skin, has not been located but is possibly AM M.2033, an adult female study skin, received as a body in alcohol without a skull, from MAMU in 1907 (see Parnaby
<emphasis id="0EF2EAF3FFBA151EFCA2FE5F9AE48BD5" box="[843,906,475,501]" italics="true" pageId="32" pageNumber="233">et. al.</emphasis>
2017). A loose paper label in the box with skull M.380.02 (
<figureCitation id="A4BD2A64FFBA151EFEFDFE7B98088839" box="[276,358,511,537]" captionStart="FIGURE 14" captionStartId="34.[151,250,1767,1791]" captionTargetBox="[170,1428,206,1730]" captionTargetId="figure-25@34.[151,1436,181,1743]" captionTargetPageId="34" captionText="FIGURE 14. Skull and mandible of M.380.2, syntype of Macropus jukesii Miklouho-Maclay 1885b [= Thylogale brunii (Schreber, 1778)], the skull illustrated in the original account, and associated paper label written by Miklouho-Maclay. Writing on the braincase is likely to be Miklouho-Maclays. Scale bar = 2 cm. (© Chau Chak Wing Museum, photo Robert Blackburn 2016)." figureDoi="http://doi.org/10.5281/zenodo.4806956" httpUri="https://zenodo.org/record/4806956/files/figure.png" pageId="32" pageNumber="233">Fig. 14</figureCitation>
) is likely to have been written by Miklouho-Maclay and states “
<taxonomicName id="FB864D62FFBA151EFBD5FE7B9C668839" authority="Jukesly" authorityName="Jukesly" box="[1084,1288,511,537]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">Macropus Jukesly</taxonomicName>
[sic] Maclay from the
<location id="3959603AFFBA151EFEE8FDA798E5881D" LSID="urn:lsid:plazi:treatment:B42F87F7FF85151FFF7EFAE29BFF8DA6:3959603AFFBA151EFEE8FDA798E5881D" box="[257,395,547,573]" country="Guinea-Bissau" county="University of Sydney" municipality="Museum of Geology" name="South Coast" pageId="32" pageNumber="233">South Coast</location>
of N.
<collectingCountry id="44917671FFBA151EFE3FFDA79B49881D" box="[470,551,547,573]" name="Guinea-Bissau" pageId="32" pageNumber="233">Guinea</collectingCountry>
near Annabada (
<collectorName id="91735337FFBA151EFD00FDA79A00881E" box="[745,878,547,574]" pageId="32" pageNumber="233">P. Mpzesly</collectorName>
)”. A more recent label (a disused NSW Railways parcel office freight tag) has “The Curator,
<collectingMunicipality id="DC5DAC9BFFBA151EFD61FDC39A1A8841" box="[648,884,583,609]" pageId="32" pageNumber="233">Museum of Geology</collectingMunicipality>
,
<collectingCounty id="D5584E6DFFBA151EFC68FDC39D198841" box="[897,1143,583,609]" pageId="32" pageNumber="233">University of Sydney</collectingCounty>
” printed on one side, and “
<taxonomicName id="FB864D62FFBA151EFF48FDEF980788A5" authority="Jukesii" authorityName="Jukesii" box="[161,361,619,645]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">Macropus Jukesii</taxonomicName>
<typeStatus id="E33D8843FFBA151EFE86FDEF98DB88A5" box="[367,437,619,645]" pageId="32" pageNumber="233">TYPE</typeStatus>
<specimenCount id="2A80FD68FFBA151EFE55FDEF98BF88A5" box="[444,465,619,645]" pageId="32" pageNumber="233" type="female"></specimenCount>
Macleay Museum” scrawled in pen on the other side
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="749C656AFFBA151EFF2EFD0B9DDB88ED" pageId="32" pageNumber="233" type="materials_examined">
<paragraph id="3C3936E1FFBA151EFF2EFD0B9DDB88ED" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
<emphasis id="0EF2EAF3FFBA151EFF2EFD0B98088888" bold="true" box="[199,358,654,681]" pageId="32" pageNumber="233">Type locality.</emphasis>
<collectingRegion id="FE42F803FFBA151EFE87FD0B9B678889" box="[366,521,655,681]" country="Papua New Guinea" name="National Capital" pageId="32" pageNumber="233">Port Moresby</collectingRegion>
<materialsCitation id="8CEE3CBCFFBA151EFDF9FD0B9DDE88ED" ID-GBIF-Occurrence="3128590305" country="Papua New Guinea" location="Central Province" pageId="32" pageNumber="233" specimenCount="1" stateProvince="National Capital">
area,
<location id="3959603AFFBA151EFDA7FD0B9A608889" LSID="urn:lsid:plazi:treatment:B42F87F7FF85151FFF7EFAE29BFF8DA6:3959603AFFBA151EFDA7FD0B9A608889" box="[590,782,655,681]" country="Papua New Guinea" name="Central Province" pageId="32" pageNumber="233" stateProvince="National Capital">Central Province</location>
,
<collectingCountry id="44917671FFBA151EFCF2FD0B9A978889" box="[795,1017,655,681]" name="Papua New Guinea" pageId="32" pageNumber="233">Papua New Guinea</collectingCountry>
. Given as “hills near Anuabada (
<collectingRegion id="FE42F803FFBA151EFA85FD0B999188EE" country="Papua New Guinea" name="National Capital" pageId="32" pageNumber="233">Port Moresby</collectingRegion>
)” (
<bibRefCitation id="58174B10FFBA151EFEF6FD379B2E88ED" author="Miklouho-Maclay, N." box="[287,576,690,717]" pageId="32" pageNumber="233" pagination="890 - 895" refId="ref38519" refString="Miklouho-Maclay, N. (1885 b) On two new species of Macropus from the south coast of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (4), 890 - 895, pl. 39." type="journal article" year="1885">Miklouho-Maclay, 1885b</bibRefCitation>
) [= Hanuabada village, now a suburb of
<collectingRegion id="FE42F803FFBA151EFBE3FD379DC488ED" box="[1034,1194,690,717]" country="Papua New Guinea" name="National Capital" pageId="32" pageNumber="233">Port Moresby</collectingRegion>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="749C656AFFBA151FFF2EFD529BFF8DA6" lastPageId="33" lastPageNumber="234" pageId="32" pageNumber="233" type="discussion">
<paragraph id="3C3936E1FFBA151EFF2EFD529DF689CD" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
<emphasis id="0EF2EAF3FFBA151EFF2EFD52982188D0" bold="true" box="[199,335,726,752]" pageId="32" pageNumber="233">Comments.</emphasis>
The skull that we believe to be the one illustrated by Miklouho-Maclay was mistakenly associated with skin M.380 when registered in c. 1964. The broken skull extracted from skin M.
<quantity id="FB7E9B04FFBA151EFB88FD7F9DC28935" box="[1121,1196,763,789]" metricMagnitude="0" metricUnit="m" metricValue="9.652" pageId="32" pageNumber="233" unit="in" value="380.0">380 in</quantity>
1960 had been overlooked during registration and in subsequent decades was assigned temporary numbers as a skull without data. We are confident that we have correctly associated the damaged skull (now M.380.1) with the skin and that this is the skull extracted in 1960. Holes drilled in the jaw show green copper residue which match holes in appropriate places in the skin, indicating that the skull and skin were wired together. A second intact skull and dentaries now registered as M.380.2, is most likely the skull illustrated in his account. It resembles the one illustrated by Miklouho-Maclay and our measurements of the skull closely match those taken from his illustration (
<tableCitation id="7104035AFFBA151EFBDAFC579DE489CD" box="[1075,1162,979,1005]" captionStart="TABLE 7" captionStartId="35.[151,239,151,175]" captionTargetPageId="35" captionText="TABLE 7. Selected cranial and dental measurements (mm) of M.380.2, likely syntype of Macropus jukesii Miklouho- Maclay, 1885b [= Thylogale brunii (Schreber, 1778)] compared to measurements from his plate 39 from an original, unbound copy of the journal." httpUri="http://table.plazi.org/id/68F96669FFB9151DFF7EFF1398A38AD7" pageId="32" pageNumber="233" tableUuid="68F96669FFB9151DFF7EFF1398A38AD7">Table 7</tableCitation>
).
</paragraph>
<paragraph id="3C3936E1FFBA151EFF2EFC739C038E15" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
Miklouho-Maclay did not explicitly state the number of specimens used in his description of
<taxonomicName id="FB864D62FFBA151EFB5AFC739C008E31" authorityName="Miklouho-Maclay" authorityYear="1885" box="[1203,1390,1015,1041]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBA151EFB5AFC739C008E31" box="[1203,1390,1015,1041]" italics="true" pageId="32" pageNumber="233">Macropus jukesii</emphasis>
</taxonomicName>
and it is unclear from his ambiguous introductory text and subsequent remarks whether he had one or
<specimenCount id="2A80FD68FFBA151EFB2AFB9F9C078E15" box="[1219,1385,1051,1077]" pageId="32" pageNumber="233" type="generic">two specimens</specimenCount>
.
</paragraph>
<paragraph id="3C3936E1FFBA151EFF2EFBBB9B7B8C72" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
We have established that Miklouho-Maclay examined
<specimenCount id="2A80FD68FFBA151EFCC0FBBB9ABC8E79" box="[809,978,1087,1113]" pageId="32" pageNumber="233" type="generic">two specimens</specimenCount>
, both in Macleays Collection. Our interpretation of his account is that he initially examined a female skin mount, without access to a skull, which was in the skin. He subsequently became aware of a consignment of spirit-preserved wallaby skins obtained by Macleay from Andrew Goldie, one of which Miklouho-Maclay believed belonged to his proposed species
<taxonomicName id="FB864D62FFBA151EFB18FB2F9C0F8EE5" authorityName="Miklouho-Maclay" authorityYear="1885" box="[1265,1377,1195,1221]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBA151EFB18FB2F9C0F8EE5" box="[1265,1377,1195,1221]" italics="true" pageId="32" pageNumber="233">M. jukesii</emphasis>
</taxonomicName>
. The latter specimen contained a skull, which Miklouho-Maclay remarked had greatly facilitated his descriptive account. Miklouho-Maclays alternate use of the terms “stuffed specimen” and a spirit specimen implies
<specimenCount id="2A80FD68FFBA151EFB2CFB779C008F2D" box="[1221,1390,1267,1293]" pageId="32" pageNumber="233" type="generic">two specimens</specimenCount>
. We reject an alternative interpretation that a single spirit specimen was turned into a dried skin and skull. His statement on page 891 implies that he was already aware of the existence of his new species (the stuffed skin) when he located the spirit preserved specimen: “Having been informed by Mr Masters ...that a few skins....were preserved in spirits, I examined them at the first opportunity and had the good chance of finding amongst them a skin, with the skull, of one of the new species,
<emphasis id="0EF2EAF3FFBA151EFE4CFA239B768FC5" italics="true" pageId="32" pageNumber="233">which discovery put me in the position of examining the dentition and making the description of this species more complete.</emphasis>
[our emphasis]”. A subsequent statement on page 894, in which he discussed the visibility of a lateral fold of enamel on the upper third incisor, implies that he had
<specimenCount id="2A80FD68FFBA151EFBC7FA6B9DB68C29" box="[1070,1240,1519,1545]" pageId="32" pageNumber="233" type="generic">two specimens</specimenCount>
: “(Examining the incisors of the stuffed specimen, I was not at all sure about the existence of the fold and convinced myself of it only after having examined the skull)”.
</paragraph>
<paragraph id="3C3936E1FFBA151EFF2EF9DF99918D99" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
The MAMU 1890s Catalogue has only two entries under “
<taxonomicName id="FB864D62FFBA151EFC8BF9DF9D448C55" authority="Jukesi" authorityName="Jukesi" box="[866,1066,1627,1653]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">Macropus Jukesi</taxonomicName>
”: “a, female. adult, stuffed.” and on a separate line: “b, skull.”, with both entered as “
<collectingRegion id="FE42F803FFBA151EFD36F9FB9AEC8CB9" box="[735,898,1663,1689]" country="Papua New Guinea" name="National Capital" pageId="32" pageNumber="233">Port Moresby</collectingRegion>
”. Our interpretation of the procedure adopted in the Catalogue is that a separate alphabetical letter was assigned to each specimen of a species, where “specimen” equates to an individual animal. Consequently, a skin would be assigned a letter but without indication of whether the skull was extracted, but a specimen consisting of a skull only (without skin or skeleton), would be assigned a separate alphabetical designation. If our interpretation is correct, then two individuals are listed under
<taxonomicName id="FB864D62FFBA151EFAC5F88B998E8D6D" authorityName="Miklouho-Maclay" authorityYear="1885" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBA151EFAC5F88B998E8D6D" italics="true" pageId="32" pageNumber="233">Macropus jukesii</emphasis>
</taxonomicName>
in the Catalogue. It is possible that a skin in alcohol associated with the second skull was overlooked when the 1890s Catalogue was compiled, particularly if its association with the skull had been lost. This is likely, as there were at least half a dozen alcohol skins of
<taxonomicName id="FB864D62FFBA151EFD6CF8FF9B9B8DB5" baseAuthorityName="Flannery" baseAuthorityYear="1992" box="[645,757,1915,1941]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis id="0EF2EAF3FFBA151EFD6CF8FF9B9B8DB5" box="[645,757,1915,1941]" italics="true" pageId="32" pageNumber="233">Thylogale</emphasis>
</taxonomicName>
and
<taxonomicName id="FB864D62FFBA151EFCD9F8FF9ACC8DB4" baseAuthorityName="Groves &amp; Flannery" baseAuthorityYear="1989" box="[816,930,1915,1940]" class="Mammalia" family="Macropodidae" genus="Dorcopsis" kingdom="Animalia" order="Diprotodontia" pageId="32" pageNumber="233" phylum="Chordata" rank="genus">
<emphasis id="0EF2EAF3FFBA151EFCD9F8FF9ACC8DB4" box="[816,930,1915,1940]" italics="true" pageId="32" pageNumber="233">Dorcopsis</emphasis>
</taxonomicName>
at that time, none of which had registration numbers.
</paragraph>
<paragraph id="3C3936E1FFBA151FFF2EF8479AA588A5" blockId="32.[151,1437,151,2013]" lastBlockId="33.[151,1437,151,861]" lastPageId="33" lastPageNumber="234" pageId="32" pageNumber="233">
It seems that extracting skulls from mounted specimens in Macleays Collection was not an option because it would degrade the presentation of the specimen. This provides further support for our interpretation of the original account.
<bibRefCitation id="58174B10FFBB151FFF14FF3F9B448AF5" author="Miklouho-Maclay, N." box="[253,554,186,213]" pageId="33" pageNumber="234" pagination="890 - 895" refId="ref38519" refString="Miklouho-Maclay, N. (1885 b) On two new species of Macropus from the south coast of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (4), 890 - 895, pl. 39." type="journal article" year="1885">Miklouho-Maclay (1885b)</bibRefCitation>
described
<taxonomicName id="FB864D62FFBB151FFD4CFF3F9A1A8AF5" authorityName="Miklouho-Maclay" authorityYear="1885" box="[677,884,187,213]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0EF2EAF3FFBB151FFD4CFF3F9A1A8AF5" box="[677,884,187,213]" italics="true" pageId="33" pageNumber="234">Macropus gracilis</emphasis>
</taxonomicName>
in the same paper in which he named
<taxonomicName id="FB864D62FFBB151FFAC5FF3F998E8AD9" authorityName="Miklouho-Maclay" authorityYear="1885" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBB151FFAC5FF3F998E8AD9" italics="true" pageId="33" pageNumber="234">Macropus jukesii</emphasis>
</taxonomicName>
. He lamented on page 894 that, as he did not have a skull, he was unable to describe the skull and dentition of
<taxonomicName id="FB864D62FFBB151FFF5CFE8798EC8B3D" authorityName="Miklouho-Maclay" authorityYear="1885" box="[181,386,259,285]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0EF2EAF3FFBB151FFF5CFE8798EC8B3D" box="[181,386,259,285]" italics="true" pageId="33" pageNumber="234">Macropus gracilis</emphasis>
</taxonomicName>
. His illustration of the lateral view of the incisors must have been drawn with the skull
<emphasis id="0EF2EAF3FFBB151FFAB9FE879CFB8B3C" box="[1360,1429,259,284]" italics="true" pageId="33" pageNumber="234">in situ</emphasis>
. This suggests that skull extraction from mounted skins was not an option available to Miklouho-Maclay. Further support for the preference of not damaging Macleays important skin mounts by skull removal comes from Ramsay. In his description of what he thought at the time was a single specimen of
<taxonomicName id="FB864D62FFBB151FFBE2FEEB9D8A8BA9" authorityName="Ramsay" authorityYear="1877" box="[1035,1252,367,393]" class="Mammalia" family="Macropodidae" genus="Petrogale" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="assimilis">
<emphasis id="0EF2EAF3FFBB151FFBE2FEEB9D8A8BA9" box="[1035,1252,367,393]" italics="true" pageId="33" pageNumber="234">Petrogale assimilis</emphasis>
</taxonomicName>
in the Macleay Collection,
<bibRefCitation id="58174B10FFBB151FFEF0FE179B648B8E" author="Ramsay, E. P." box="[281,522,403,430]" pageId="33" pageNumber="234" pagination="359 - 361" refId="ref39663" refString="Ramsay, E. P. (1877 e) Description of a supposed new species of Rock Wallaby from the Palm Islands, on the North-east Coast of Australia, proposed to be called Petrogale assimilis. Proceedings of the Linnean Society of New South Wales, Series 1, 1 (4), 359 - 361. https: // doi. org / 10.5962 / bhl. part. 12421" type="journal article" year="1877">Ramsay (1877e: 360)</bibRefCitation>
stated that he could not properly describe skull and dentition because this “cannot easily be examined without risk of deterioration [to the skin].” If so, this supports the view that Miklouho-Maclays description and illustrations of the skull of
<taxonomicName id="FB864D62FFBB151FFD92FE5F9A528BD5" authorityName="Miklouho-Maclay" authorityYear="1885" box="[635,828,475,501]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBB151FFD92FE5F9A528BD5" box="[635,828,475,501]" italics="true" pageId="33" pageNumber="234">Macropus jukesii</emphasis>
</taxonomicName>
were not based on the skull removed from the stuffed female specimen, but that he examined a second specimen. Considering his unresolved conundrum about whether the differences in incisor morphology between the male
<taxonomicName id="FB864D62FFBB151FFCF2FDA79AF9881D" authorityName="Miklouho-Maclay" authorityYear="1885" box="[795,919,547,573]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="0EF2EAF3FFBB151FFCF2FDA79AF9881D" box="[795,919,547,573]" italics="true" pageId="33" pageNumber="234">M. gracilis</emphasis>
</taxonomicName>
and the female
<taxonomicName id="FB864D62FFBB151FFBB9FDA79DAE881D" authorityName="Miklouho-Maclay" authorityYear="1885" box="[1104,1216,547,573]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBB151FFBB9FDA79DAE881D" box="[1104,1216,547,573]" italics="true" pageId="33" pageNumber="234">M. jukesii</emphasis>
</taxonomicName>
were due to sex or species differences, a second skull of
<taxonomicName id="FB864D62FFBB151FFDA9FDC39BC08841" authorityName="Miklouho-Maclay" authorityYear="1885" box="[576,686,583,609]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBB151FFDA9FDC39BC08841" box="[576,686,583,609]" italics="true" pageId="33" pageNumber="234">M. jukesii</emphasis>
</taxonomicName>
had to be either a female, or an incomplete skin of indeterminable sex. The skin (AM M.2033) thought to belong with this skull is a female.
</paragraph>
<paragraph id="3C3936E1FFBB151FFF2EFD0B9893897D" blockId="33.[151,1437,151,861]" pageId="33" pageNumber="234">
We are confident that skin mount M.380 is the specimen examined by Miklouho-Maclay. The sex of skin M.380 is not indicated in the MAMU Mammal Register but it has a conspicuously enlarged pouch. Miklouho-Maclay did not illustrate the stuffed skin. We took two measurements of the skin and both are a close approximation to those given by Miklouho-Maclay: his measurement of
<quantity id="FB7E9B04FFBB151FFD2BFD7E9A718935" box="[706,799,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="33" pageNumber="234" unit="mm" value="635.0">635 mm</quantity>
for “tip of nose to base of tail” versus our measurement of c.
<quantity id="FB7E9B04FFBB151FFF39FC9B98438919" box="[208,301,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="6.3" pageId="33" pageNumber="234" unit="mm" value="630.0">630 mm</quantity>
taken along the spine following the body contours, and his “tail tip to base of tail” of
<quantity id="FB7E9B04FFBB151FFB18FC9B9C208919" box="[1265,1358,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="33" pageNumber="234" unit="mm" value="370.0">370 mm</quantity>
versus our measurement of c.
<quantity id="FB7E9B04FFBB151FFE70FCC79897897D" box="[409,505,835,861]" metricMagnitude="-1" metricUnit="m" metricValue="3.6799999999999997" pageId="33" pageNumber="234" unit="mm" value="368.0">368 mm</quantity>
.
</paragraph>
<caption id="68F96669FFBB151FFF7EFA6699B78C46" ID-DOI="http://doi.org/10.5281/zenodo.4806954" ID-Zenodo-Dep="4806954" httpUri="https://zenodo.org/record/4806954/files/figure.png" pageId="33" pageNumber="234" startId="33.[151,250,1506,1530]" targetBox="[169,1419,891,1477]" targetPageId="33">
<paragraph id="3C3936E1FFBB151FFF7EFA6699B78C46" blockId="33.[151,1437,1506,1638]" pageId="33" pageNumber="234">
<emphasis id="0EF2EAF3FFBB151FFF7EFA6698758FDA" bold="true" box="[151,283,1506,1530]" pageId="33" pageNumber="234">FIGURE 13</emphasis>
. Cranial fragments and dentary of M380.1, syntype of
<taxonomicName id="FB864D62FFBB151FFCB0FA679C7E8FDA" authority="Miklouho-Maclay 1885 b" authorityName="Miklouho-Maclay" authorityYear="1885" box="[857,1296,1506,1530]" class="Mammalia" family="Macropodidae" genus="Macropus" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="jukesii">
<emphasis id="0EF2EAF3FFBB151FFCB0FA679D668FDA" box="[857,1032,1506,1530]" italics="true" pageId="33" pageNumber="234">Macropus jukesii</emphasis>
<bibRefCitation id="58174B10FFBB151FFBF8FA669C7E8FDA" author="Miklouho-Maclay, N." box="[1041,1296,1506,1530]" pageId="33" pageNumber="234" pagination="890 - 895" refId="ref38519" refString="Miklouho-Maclay, N. (1885 b) On two new species of Macropus from the south coast of New Guinea. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (4), 890 - 895, pl. 39." type="journal article" year="1885">Miklouho-Maclay 1885b</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="FB864D62FFBB151FFADEFA6698E78C3E" authority="(Schreber, 1778)" baseAuthorityName="Schreber" baseAuthorityYear="1778" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="33" pageNumber="234" phylum="Chordata" rank="species" species="brunii">
<emphasis id="0EF2EAF3FFBB151FFADEFA6699BB8C3E" italics="true" pageId="33" pageNumber="234">Thylogale brunii</emphasis>
(
<bibRefCitation id="58174B10FFBB151FFF0BF98298EC8C3E" author="Schreber, J. C. D. von" box="[226,386,1542,1566]" pageId="33" pageNumber="234" refId="ref40096" refString="Schreber, J. C. D. von (1778) Die Saugethiere in Abbildungen nach der Natur, mit Beschreibungen. Vol. 3. Walfgang Walther, Erlangen, Germany, pp. 551, pl. cliii. [as Der Filander, Didelphys Brunii] https: // doi. org / 10.5962 / bhl. title. 67399" type="book" year="1778">Schreber, 1778</bibRefCitation>
)
</taxonomicName>
], extracted from the skin in 1960. A and B, cranium; C, right dentary; D, right maxillary fragment; and E, premaxillary fragment. (left dentary not shown). Scale bar = 1 cm. (© Chau Chak Wing Museum, photo Robert Blackburn 2016).
</paragraph>
</caption>
<paragraph id="3C3936E1FFBB151FFF2EF9109BFF8DA6" blockId="33.[151,1437,1684,1926]" pageId="33" pageNumber="234">
The inadequately defined type material of this taxon has generated considerable confusion prior to our study.
<bibRefCitation id="58174B10FFBB151FFF7EF93D985A8CF3" author="Tate, G. H. H." box="[151,308,1720,1747]" pageId="33" pageNumber="234" pagination="1 - 10" refId="ref40535" refString="Tate, G. H. H. (1940) Notes on the types of certain early described species of monotremes, marsupials, Muridae, and bats from the Indo-Australian region. American Museum Novitates, 1061, 1 - 10." type="journal article" year="1940">Tate (1940: 5)</bibRefCitation>
examined the material during a visit to MAMU in 1937, which at that time consisted of a skin mount and one skull. We concur with his conclusion that the skull (now M.380.2) was the “type”, and that it was dubiously associated with the skin mount. Tate probably recognised that the skin mount had a skull
<emphasis id="0EF2EAF3FFBB151FFB6CF8859DA48D3A" box="[1157,1226,1793,1818]" italics="true" pageId="33" pageNumber="234">in situ</emphasis>
but he was unsure of the type status of the skin mount. Skull M.380.2 would also have been incorrectly associated with the
<typeStatus id="E33D8843FFBB151FFAD3F8A09CF28D1E" box="[1338,1436,1828,1854]" pageId="33" pageNumber="234" type="holotype">holotype</typeStatus>
skin when examined by
<bibRefCitation id="58174B10FFBB151FFE4DF8CC9B348D43" author="Flannery, T. F." box="[420,602,1864,1891]" pageId="33" pageNumber="234" pagination="7 - 23" refId="ref35378" refString="Flannery, T. F. (1992) Taxonomic revision of the Thylogale brunii complex (Macropodidae: Marsupialia) in Melanesia, with description of a new species. Australian Mammalogy, 15, 7 - 23." type="journal article" year="1992">Flannery (1992)</bibRefCitation>
, but his conclusion that the
<typeStatus id="E33D8843FFBB151FFC67F8CC9A9E8D42" box="[910,1008,1864,1890]" pageId="33" pageNumber="234" type="holotype">holotype</typeStatus>
is a young animal stands because both skulls have incompletely erupted last molars.
</paragraph>
</subSubSection>
</treatment>
</document>