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<document id="681CBB8C23358ADE02A472728CD524F2" ID-DOI="0.1649/0010-065X-69.mo4.27" ID-ISSN="1938-4394" ID-Zenodo-Dep="10086147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095935056" checkinUser="carolina" docAuthor="Johnston, M. Andrew, Fleming, David, Franz, Nico M. &amp; Smith, Aaron D." docDate="2015" docId="B31FCE62BB6BFFBBFD1FFA24FDB40EFE" docLanguage="en" docName="TheColeopBull.69.mo14.27-54.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Eleodes (Eleodes) obscurus" docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="4F26B61ABB7EFFADFFC8FFB9FFAD0C40" masterDocTitle="Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts" masterLastPageNumber="54" masterPageNumber="27" pageNumber="48" updateTime="1699918030632" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E5F2EA0730184E7F6472034E63615FD4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AB86D1C366A37AE8DC6B2AB1C0661786">Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts</mods:title>
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<mods:role id="DE69C575377627D14E227C50812FFB66">
<mods:roleTerm id="2CAFCCF25F117D37A62DE7FC8FACC5AF">Author</mods:roleTerm>
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<mods:namePart id="861BA6DBBA8C0C347D3A9CF010C12CD8">Johnston, M. Andrew</mods:namePart>
<mods:affiliation id="27006029011B842DD1772A35DFC71BA1">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="38EC911F5EBF4D03DDB4889CFB2F755A" type="email">ajohnston@asu.edu,</mods:nameIdentifier>
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<mods:namePart id="D74AACCC9A8CF00BAB5DF1E7514B8F23">Fleming, David</mods:namePart>
<mods:affiliation id="D555D6FAE9F707C8E3112420DFF1F798">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="5D1F065378EC532684E9B13F1C0EC15E" type="email">david.fleming.1@asu.edu</mods:nameIdentifier>
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<mods:namePart id="2241D8F8AA21AA004182EA7B208A0AB1">Franz, Nico M.</mods:namePart>
<mods:affiliation id="F5FBAE504530CFE0F9B218FA4D9C75F0">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="5AFACD7335858FCBB2F3020683A0D18D" type="email">nico.franz@asu.edu</mods:nameIdentifier>
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<mods:namePart id="4CA7D4011ED1AED6B661E3C471B1DF97">Smith, Aaron D.</mods:namePart>
<mods:affiliation id="90B3FCABA9E221F66275889E8A6AEFFF">Department of Biological Sciences, Northern Arizona University Flagstaff, AZ 86011 - 5640, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="F0AC4BB1E884CB363B6EC909C3B4B54A" type="email">aaron.smith@nau.edu</mods:nameIdentifier>
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<mods:titleInfo id="FCD5178540AA25DB870EF1BE2B0C5E6D">
<mods:title id="2A15BAB27ABCF458D73B74E0C3A612E1">The Coleopterists Bulletin</mods:title>
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<mods:part id="397AF5468B625FA0CABB4114B3154A82">
<mods:date id="D08E91DBF551A2F16FB685E1FCC3893B">2015</mods:date>
<mods:detail id="D117AC9F4EE225F22D707CA38102EBEC" type="series">
<mods:title id="73A94FF69D0E3E09182879998F74A535">mo 14</mods:title>
</mods:detail>
<mods:detail id="AE5D092E282CEE9AF8FEBC94FF157A90" type="pubDate">
<mods:number id="A594C13C26C43C4F7500C3FE6A4ABD78">2015-12-18</mods:number>
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<mods:detail id="F102DCD820D950BC4EB1D00AD78B3995" type="volume">
<mods:number id="D36D0679FC963CFE158D5751C99CF626">69</mods:number>
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<mods:start id="6AFCE9490B89EFB54FE55DCA55E457B8">27</mods:start>
<mods:end id="54656F367D86150D41AABDF11CA6D180">54</mods:end>
</mods:extent>
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<mods:classification id="2671F80850B3ED2D1F10023ED8008FEB">journal article</mods:classification>
<mods:identifier id="C9113E788D637DD925F6A1A177D0E428" type="DOI">0.1649/0010-065X-69.mo4.27</mods:identifier>
<mods:identifier id="072ABE7557115DBB1C73ECBCC46009BE" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="72C2FC762C4A61A62B8B5DC5C0F685FE" type="Zenodo-Dep">10086147</mods:identifier>
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<treatment id="B31FCE62BB6BFFBBFD1FFA24FDB40EFE" LSID="urn:lsid:plazi:treatment:B31FCE62BB6BFFBBFD1FFA24FDB40EFE" httpUri="http://treatment.plazi.org/id/B31FCE62BB6BFFBBFD1FFA24FDB40EFE" lastPageId="22" lastPageNumber="49" pageId="21" pageNumber="48">
<subSubSection id="73AC2CFFBB6BFFB8FD1FFA24FBC709F2" box="[727,1130,1437,1458]" pageId="21" pageNumber="48" type="nomenclature">
<paragraph id="3B097F74BB6BFFB8FD1FFA24FBC709F2" blockId="21.[727,1130,1437,1485]" box="[727,1130,1437,1458]" pageId="21" pageNumber="48">
<heading id="6041C818BB6BFFB8FD1FFA24FBC709F2" box="[727,1130,1437,1458]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="48" reason="2">
<taxonomicName id="FCB604F7BB6BFFB8FD1FFA24FBC709F2" authority="(Say, 1823)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[727,1130,1437,1458]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="48" phylum="Arthropoda" rank="species" species="obscurus" subGenus="Eleodes">
<emphasis id="09C2A366BB6BFFB8FD1FFA24FBC709F2" bold="true" box="[727,1130,1437,1458]" pageId="21" pageNumber="48">
<emphasis id="09C2A366BB6BFFB8FD1FFA24FC8E09F2" bold="true" box="[727,803,1437,1458]" italics="true" pageId="21" pageNumber="48">Eleodes</emphasis>
(
<emphasis id="09C2A366BB6BFFB8FCFDFA24FC2C09F2" bold="true" box="[821,897,1437,1458]" italics="true" pageId="21" pageNumber="48">Eleodes</emphasis>
)
<emphasis id="09C2A366BB6BFFB8FC5CFA24FC4109F2" bold="true" box="[916,1004,1437,1458]" italics="true" pageId="21" pageNumber="48">obscurus</emphasis>
(Say, 1823)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73AC2CFFBB6BFFB8FC97FA01FC810A6D" pageId="21" pageNumber="48" type="description">
<paragraph id="3B097F74BB6BFFB8FC97FA01FC4F098D" blockId="21.[727,1130,1437,1485]" box="[863,994,1463,1485]" pageId="21" pageNumber="48">
(
<figureCitation id="A38D63F1BB6BFFB8FCAEFA01FC76098D" box="[870,987,1463,1485]" captionStart="Fig" captionStartId="15.[175,207,1644,1663]" captionTargetBox="[151,1171,193,1621]" captionTargetId="figure-1@15.[151,1173,192,1621]" captionTargetPageId="15" captionText="Fig. 12. Eleodes species, dorsal habitus. A) E. dissimilis, female, B) E. nigrinus, male, C) E. obscurus sulcipennis, female, D) E. obscurus dispersus, male. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120694" httpUri="https://zenodo.org/record/10120694/files/figure.png" pageId="21" pageNumber="48">Fig. 12C–D</figureCitation>
)
</paragraph>
<paragraph id="3B097F74BB6BFFB8FD65FA5AFB390A52" blockId="21.[685,1172,1506,1581]" pageId="21" pageNumber="48">
<taxonomicName id="FCB604F7BB6BFFB8FD65FA5AFB8209B7" authority="LeConte, 1858" authorityName="LeConte" authorityYear="1858" box="[685,1071,1506,1528]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="48" phylum="Arthropoda" rank="subSpecies" species="obscurus" subSpecies="dispersus">
<emphasis id="09C2A366BB6BFFB8FD65FA5AFC0909B8" box="[685,932,1507,1528]" italics="true" pageId="21" pageNumber="48">Eleodes obscurus dispersus</emphasis>
LeConte, 1858
</taxonomicName>
(
<figureCitation id="A38D63F1BB6BFFB8FBF3FA5BFB2009B7" box="[1083,1165,1506,1527]" captionStart="Fig" captionStartId="15.[175,207,1644,1663]" captionTargetBox="[151,1171,193,1621]" captionTargetId="figure-1@15.[151,1173,192,1621]" captionTargetPageId="15" captionText="Fig. 12. Eleodes species, dorsal habitus. A) E. dissimilis, female, B) E. nigrinus, male, C) E. obscurus sulcipennis, female, D) E. obscurus dispersus, male. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120694" httpUri="https://zenodo.org/record/10120694/files/figure.png" pageId="21" pageNumber="48">Fig. 12D</figureCitation>
)
<taxonomicName id="FCB604F7BB6BFFB8FD65FA44FB390A52" authority="Mannerheim, 1843" authorityName="Mannerheim" authorityYear="1843" box="[685,1172,1533,1554]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="48" phylum="Arthropoda" rank="subSpecies" species="obscurus" subSpecies="sulcipennis">
<emphasis id="09C2A366BB6BFFB8FD65FA44FC7D0A52" box="[685,976,1533,1554]" italics="true" pageId="21" pageNumber="48">Eleodes obscurus sulcipennis</emphasis>
Mannerheim, 1843
</taxonomicName>
</paragraph>
<paragraph id="3B097F74BB6BFFB8FD0FF9A1FC810A6D" blockId="21.[685,1172,1506,1581]" box="[711,812,1560,1581]" pageId="21" pageNumber="48">
(
<figureCitation id="A38D63F1BB6BFFB8FD06F9A1FC880A6D" box="[718,805,1560,1581]" captionStart="Fig" captionStartId="15.[175,207,1644,1663]" captionTargetBox="[151,1171,193,1621]" captionTargetId="figure-1@15.[151,1173,192,1621]" captionTargetPageId="15" captionText="Fig. 12. Eleodes species, dorsal habitus. A) E. dissimilis, female, B) E. nigrinus, male, C) E. obscurus sulcipennis, female, D) E. obscurus dispersus, male. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120694" httpUri="https://zenodo.org/record/10120694/files/figure.png" pageId="21" pageNumber="48">Fig. 12C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73AC2CFFBB6BFFBBFD0DF9FBFE800D1F" lastPageId="22" lastPageNumber="49" pageId="21" pageNumber="48" type="diagnosis">
<paragraph id="3B097F74BB6BFFBBFD0DF9FBFE800D1F" blockId="21.[685,1173,1602,1729]" lastBlockId="22.[173,660,195,702]" lastPageId="22" lastPageNumber="49" pageId="21" pageNumber="48">
<emphasis id="09C2A366BB6BFFB8FD0DF9FBFC830A17" bold="true" box="[709,814,1602,1623]" pageId="21" pageNumber="48">Diagnosis.</emphasis>
Body elongate, large (&gt;
<quantity id="FC4ED291BB6BFFB8FBF1F9FBFB240A17" box="[1081,1161,1602,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="48" unit="mm" value="25.0">25 mm</quantity>
). Pronotum strongly arcuate laterally, narrowing posteriorly, anterior angles slightly produced, rounded. Prosternum extending posteriorly of procoxae, without distinct process. Profemora spined in males, unspined in females, probasitarsis thickened beneath, with bulbous lobe projecting anteriorly from plantar surface. Elytra parallel-sided to slightly inflated posteriorly in females, noticeably sculptured with either muricate to bluntly tuberclate or with deeply sulcate striae.
</paragraph>
</subSubSection>
<subSubSection id="73AC2CFFBB68FFBBFF0DFEDCFD3D0DA6" pageId="22" pageNumber="49" type="distribution">
<paragraph id="3B097F74BB68FFBBFF0DFEDCFD3D0DA6" blockId="22.[173,660,195,702]" pageId="22" pageNumber="49">
<emphasis id="09C2A366BB68FFBBFF0DFEDCFE920D3A" bold="true" box="[197,319,357,378]" pageId="22" pageNumber="49">Distribution.</emphasis>
Coconino, Navajo, Apache, Yavapai, Gila, Maricopa, Pinal, Pima, Santa Cruz, and Cochise Counties, Arizona. Broadly distributed from Texas north to
<collectingRegion id="F972B196BB68FFBBFECAFE0FFE0F0D8B" box="[258,418,438,459]" country="Canada" name="New Brunswick" pageId="22" pageNumber="49">New Brunswick</collectingRegion>
,
<collectingCountry id="43A13FE4BB68FFBBFE65FE0FFE5A0D8B" box="[429,503,438,459]" name="Canada" pageId="22" pageNumber="49">Canada</collectingCountry>
, west to
<collectingRegion id="F972B196BB68FFBBFD9AFE0FFEA60DA6" country="Canada" name="British Columbia" pageId="22" pageNumber="49">British Columbia</collectingRegion>
,
<collectingCountry id="43A13FE4BB68FFBBFEDDFE68FEF00DA6" box="[277,349,465,486]" name="Canada" pageId="22" pageNumber="49">Canada</collectingCountry>
, and south to
<collectingRegion id="F972B196BB68FFBBFE2BFE68FD9A0DA6" box="[483,567,465,486]" country="Mexico" name="Durango" pageId="22" pageNumber="49">Durango</collectingRegion>
,
<collectingCountry id="43A13FE4BB68FFBBFD8AFE68FD210DA6" box="[578,652,465,486]" name="Mexico" pageId="22" pageNumber="49">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="73AC2CFFBB68FFBBFF0DFE55FDB40EFE" pageId="22" pageNumber="49" type="discussion">
<paragraph id="3B097F74BB68FFBBFF0DFE55FDB40EFE" blockId="22.[173,660,195,702]" pageId="22" pageNumber="49">
<emphasis id="09C2A366BB68FFBBFF0DFE55FE850E41" bold="true" box="[197,296,492,513]" pageId="22" pageNumber="49">Remarks.</emphasis>
This large species is easily recognizable by its size and
<emphasis id="09C2A366BB68FFBBFEBFFDBEFE1B0E5C" box="[375,438,519,540]" italics="true" pageId="22" pageNumber="49">gestalt</emphasis>
, though there are two subspecies present in the state with quite different elytral sculpturing.
<taxonomicName id="FCB604F7BB68FFBBFEADFD84FDC10E12" authorityName="LeConte" authorityYear="1858" box="[357,620,573,594]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="obscurus" subSpecies="dispersus">
<emphasis id="09C2A366BB68FFBBFEADFD84FDC10E12" box="[357,620,573,594]" italics="true" pageId="22" pageNumber="49">Eleodes obscurus dispersus</emphasis>
</taxonomicName>
can be found in northeastern
<collectingRegion id="F972B196BB68FFBBFE52FDE1FE4B0E2D" box="[410,486,600,621]" country="United States of America" name="Arizona" pageId="22" pageNumber="49">Arizona</collectingRegion>
and has the elytra muricate to bluntly tuberculate.
<taxonomicName id="FCB604F7BB68FFBBFE20FDCAFEB40EE3" authorityName="Mannerheim" authorityYear="1843" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="obscurus" subSpecies="sulcipennis">
<emphasis id="09C2A366BB68FFBBFE20FDCAFEB40EE3" italics="true" pageId="22" pageNumber="49">Eleodes obscurus sulcipennis</emphasis>
</taxonomicName>
is broadly distributed in
<collectingRegion id="F972B196BB68FFBBFDD2FD37FDCA0EE3" box="[538,615,654,675]" country="United States of America" name="Arizona" pageId="22" pageNumber="49">Arizona</collectingRegion>
and can be separated by its sulcate elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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