158 lines
17 KiB
XML
158 lines
17 KiB
XML
<document id="5EF38A5105FF78F75029C4E22A9A6544" ID-DOI="0.1649/0010-065X-69.mo4.27" ID-ISSN="1938-4394" ID-Zenodo-Dep="10086147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095935056" checkinUser="carolina" docAuthor="Johnston, M. Andrew, Fleming, David, Franz, Nico M. & Smith, Aaron D." docDate="2015" docId="B31FCE62BB67FFB4FD22FF7BFC630854" docLanguage="en" docName="TheColeopBull.69.mo14.27-54.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Trogloderus costatus LeConte 1879" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="4F26B61ABB7EFFADFFC8FFB9FFAD0C40" masterDocTitle="Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts" masterLastPageNumber="54" masterPageNumber="27" pageNumber="52" updateTime="1699918030632" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="3A482EC6E9EEC38230B8DF4CB9A40418" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="27146B4C963736B25C46297C59C0E38E">
|
||
<mods:title id="FD2C7C45FBA39BEB3CACB9B33AE15DE3">Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="B15E164E30AC38F5CBB8439692C28EB2" type="personal">
|
||
<mods:role id="AF648E9A4343D7DBAECD398110E8D6E2">
|
||
<mods:roleTerm id="65F1830204077E7CC8DE7C622E1BCAEE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FF571067F7A230813253C7644E4EB5D6">Johnston, M. Andrew</mods:namePart>
|
||
<mods:affiliation id="5D792D14B15F0F83269ABFB9E6052821">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
|
||
<mods:nameIdentifier id="D3F1EC4678200E78DF96744516AFBFB2" type="email">ajohnston@asu.edu,</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="C7A9470D3D5C36D3871924017F847270" type="personal">
|
||
<mods:role id="4F4A861C9BBB2ED85F18554B52D12AEA">
|
||
<mods:roleTerm id="2C428F58C97B4A1A015EA38C19DB7383">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F846291928C3EE1C1FCD2BD6F3AA41D5">Fleming, David</mods:namePart>
|
||
<mods:affiliation id="2F91BC8566757E987E8E2C70856BB14A">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
|
||
<mods:nameIdentifier id="D43E3F75E63955D9F9AE3E6D454BA121" type="email">david.fleming.1@asu.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="C380B5BB90CB872848088BC942E7B760" type="personal">
|
||
<mods:role id="79A828D8F7C64A4CD58E879E12B33F5F">
|
||
<mods:roleTerm id="EE1E0C5B2875F88D7F36F9780BFFB7FE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="64CEC481756474BB33319FD49B7CAAA0">Franz, Nico M.</mods:namePart>
|
||
<mods:affiliation id="8EB36BDDF4BBB12878294950065C9185">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
|
||
<mods:nameIdentifier id="177804106FA0C7A8A6AB071F9C772925" type="email">nico.franz@asu.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="CA56F4DD0526C0ED2FFB06BB020F8596" type="personal">
|
||
<mods:role id="CCAD1C1AD184B9DDA5FDB3F5EC23AA29">
|
||
<mods:roleTerm id="0905B85E80A18EBD57ECFFD1645D2EC6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C968275BD12895463FFF0BDE6C12B3B6">Smith, Aaron D.</mods:namePart>
|
||
<mods:affiliation id="AC45C841B3B73807AE4D04E29FED81DE">Department of Biological Sciences, Northern Arizona University Flagstaff, AZ 86011 - 5640, U. S. A.</mods:affiliation>
|
||
<mods:nameIdentifier id="935951ADDFDF76A39581AB1D5025BA98" type="email">aaron.smith@nau.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E83F91EE6EF967C99E29E1755B61E7B5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="96AADFAB4B15DB0B2DBFA048FCB69157" type="host">
|
||
<mods:titleInfo id="686B7833068421D95EB3D3BFA74DBC9C">
|
||
<mods:title id="03C7A805FEBFD84A7F0C00D8FCA7DB58">The Coleopterists Bulletin</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="A8B7E15583AC3F8B0B780787E495C25C">
|
||
<mods:date id="EE26D3951A835D0744CBFBA5D04AB0FC">2015</mods:date>
|
||
<mods:detail id="2C342E59D9D29AA76E59D9823C2C51CD" type="series">
|
||
<mods:title id="F51BF9144A7115C97BA4CE5F963AE1E3">mo 14</mods:title>
|
||
</mods:detail>
|
||
<mods:detail id="FA47507C2F5023B67442E821C93B58F8" type="pubDate">
|
||
<mods:number id="FFA8E99A1472D3CB1D0342F8CF0C823F">2015-12-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0B62F1E28B97609A85AB36D9D6AC72DB" type="volume">
|
||
<mods:number id="D457C2BE8B4A92A5F2D699FEBCF3B037">69</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8E3A66787B68108B3BD6E8D3D111A5B2" unit="page">
|
||
<mods:start id="96151B73184C79C81C9406F12034BE61">27</mods:start>
|
||
<mods:end id="2E86DA0C7C36F2562BDC581FCDFB94DC">54</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="6F0F60A04CA776D01E86238151943051">journal article</mods:classification>
|
||
<mods:identifier id="C64170953FAAE9F8C0748E4FC9AD3357" type="DOI">0.1649/0010-065X-69.mo4.27</mods:identifier>
|
||
<mods:identifier id="F29BB659EB614DA69451FED1FB96363C" type="ISSN">1938-4394</mods:identifier>
|
||
<mods:identifier id="226E4DB2184702935B816388FC3343C8" type="Zenodo-Dep">10086147</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B31FCE62BB67FFB4FD22FF7BFC630854" LSID="urn:lsid:plazi:treatment:B31FCE62BB67FFB4FD22FF7BFC630854" httpUri="http://treatment.plazi.org/id/B31FCE62BB67FFB4FD22FF7BFC630854" lastPageNumber="52" pageId="25" pageNumber="52">
|
||
<subSubSection id="73AC2CFFBB67FFB4FD22FF7BFBFA0C97" box="[746,1111,194,215]" pageId="25" pageNumber="52" type="nomenclature">
|
||
<paragraph id="3B097F74BB67FFB4FD22FF7BFBFA0C97" blockId="25.[746,1111,194,242]" box="[746,1111,194,215]" pageId="25" pageNumber="52">
|
||
<heading id="6041C818BB67FFB4FD22FF7BFBFA0C97" box="[746,1111,194,215]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="52" reason="2">
|
||
<taxonomicName id="FCB604F7BB67FFB4FD22FF7BFBFA0C97" authority="LeConte, 1879" authorityName="LeConte" authorityYear="1879" box="[746,1111,194,215]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="09C2A366BB67FFB4FD22FF7BFBFA0C97" bold="true" box="[746,1111,194,215]" pageId="25" pageNumber="52">
|
||
<emphasis id="09C2A366BB67FFB4FD22FF7BFC140C97" bold="true" box="[746,953,194,215]" italics="true" pageId="25" pageNumber="52">Trogloderus costatus</emphasis>
|
||
LeConte, 1879
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC2CFFBB67FFB4FCADFF64FC710CB2" box="[869,988,220,242]" pageId="25" pageNumber="52" type="description">
|
||
<paragraph id="3B097F74BB67FFB4FCADFF64FC710CB2" blockId="25.[746,1111,194,242]" box="[869,988,220,242]" pageId="25" pageNumber="52">
|
||
(
|
||
<figureCitation id="A38D63F1BB67FFB4FCA4FF64FC780CB2" box="[876,981,220,242]" captionStart="Fig" captionStartId="2.[197,229,1597,1616]" captionTargetBox="[173,1193,193,1574]" captionTargetId="figure-1@2.[172,1194,192,1574]" captionTargetPageId="2" captionText="Fig. 1. Amphidorini species, dorsal habitus. A) Trogloderus costatus, western form, B) T. costatus, northern form, C) Embaphion depressum, D) Embaphion contusum. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120670" httpUri="https://zenodo.org/record/10120670/files/figure.png" pageId="25" pageNumber="52">Fig. 1A–B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC2CFFBB67FFB4FD65FEBEFBDC0D11" pageId="25" pageNumber="52" type="reference_group">
|
||
<paragraph id="3B097F74BB67FFB4FD65FEBEFBDC0D11" blockId="25.[685,1166,263,338]" pageId="25" pageNumber="52">
|
||
<taxonomicName id="FCB604F7BB67FFB4FD65FEBEFB230D5C" authority="Blaisdell, 1909" authorityName="Blaisdell" authorityYear="1909" box="[685,1166,263,284]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="tuberculatus">
|
||
<emphasis id="09C2A366BB67FFB4FD65FEBEFC590D5C" box="[685,1012,263,284]" italics="true" pageId="25" pageNumber="52">Trogloderus costatus tuberculatus</emphasis>
|
||
<bibRefCitation id="5F270285BB67FFB4FC34FEBEFB230D5C" author="Blaisdell" box="[1020,1166,263,284]" pageId="25" pageNumber="52" pagination="1 - 524" refId="ref12543" refString="Blaisdell, F. E. 1909. A monographic revision of the Coleoptera belonging to the Tenebrionidae tribe Eleodiini inhabiting the United States, Lower California, and adjacent islands. Bulletin of the United States National Museum 63: 1 - 524." type="journal article" year="1909">Blaisdell, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicName id="FCB604F7BB67FFB4FD65FE9BFBD50D77" authority="La Rivers, 1946" authorityName="La Rivers" authorityYear="1946" box="[685,1144,290,311]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="vandykei">
|
||
<emphasis id="09C2A366BB67FFB4FD65FE9BFC7E0D77" box="[685,979,290,311]" italics="true" pageId="25" pageNumber="52">Trogloderus costatus vandykei</emphasis>
|
||
<bibRefCitation id="5F270285BB67FFB4FC13FE9BFBD50D77" author="La Rivers" box="[987,1144,290,311]" pageId="25" pageNumber="52" pagination="35 - 44" refId="ref12990" refString="La Rivers, I. 1946. On the genus Trogloderus LeConte (Coleoptera: Tenebrionidae). Entomological News 57: 35 - 44." type="journal article" year="1946">La Rivers, 1946</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicName id="FCB604F7BB67FFB4FD65FE84FBDC0D11" authority="La Rivers, 1942" authorityName="La Rivers" authorityYear="1942" box="[685,1137,316,338]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="nevadus">
|
||
<emphasis id="09C2A366BB67FFB4FD65FE84FC600D12" box="[685,973,317,338]" italics="true" pageId="25" pageNumber="52">Trogloderus costatus nevadus</emphasis>
|
||
La Rivers, 1942
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC2CFFBB67FFB4FD0DFEDEFBA20E76" pageId="25" pageNumber="52" type="diagnosis">
|
||
<paragraph id="3B097F74BB67FFB4FD0DFEDEFBA20E76" blockId="25.[685,1172,359,1044]" pageId="25" pageNumber="52">
|
||
<emphasis id="09C2A366BB67FFB4FD0DFEDEFC830D3C" bold="true" box="[709,814,359,380]" pageId="25" pageNumber="52">Diagnosis.</emphasis>
|
||
Body elongate. Elytral disc with four sharp, elevated costae, not counting the sutural costa which may be present or absent. Pronotum coarsely sculptured, either evenly tuberculate or grossly punctured with a tendency towards tubercles laterally. Anterior angles prominent, posterior margin strongly constricted. Profemora spined in males, sinuate to dentate in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC2CFFBB67FFB4FD0DFD82FC5F0EC5" pageId="25" pageNumber="52" type="distribution">
|
||
<paragraph id="3B097F74BB67FFB4FD0DFD82FC5F0EC5" blockId="25.[685,1172,359,1044]" pageId="25" pageNumber="52">
|
||
<emphasis id="09C2A366BB67FFB4FD0DFD82FCE50E10" bold="true" box="[709,840,571,592]" pageId="25" pageNumber="52">Distribution.</emphasis>
|
||
Mohave, Coconino, La Paz, and Yuma Counties,
|
||
<collectingRegion id="F972B196BB67FFB4FC90FDEFFC060E2B" box="[856,939,598,619]" country="United States of America" name="Arizona" pageId="25" pageNumber="52">Arizona</collectingRegion>
|
||
.
|
||
<collectingRegion id="F972B196BB67FFB4FC7FFDEFFB970E2B" box="[951,1082,598,619]" country="United States of America" name="New Mexico" pageId="25" pageNumber="52">New Mexico</collectingRegion>
|
||
north to
|
||
<collectingRegion id="F972B196BB67FFB4FD65FDC9FCA60EC5" box="[685,779,624,645]" country="United States of America" name="Wyoming" pageId="25" pageNumber="52">Wyoming</collectingRegion>
|
||
and west to
|
||
<collectingRegion id="F972B196BB67FFB4FC43FDC9FC430EC5" box="[907,1006,624,645]" country="United States of America" name="California" pageId="25" pageNumber="52">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC2CFFBB67FFB4FD0DFD32FC630854" pageId="25" pageNumber="52" type="discussion">
|
||
<paragraph id="3B097F74BB67FFB4FD0DFD32FC630854" blockId="25.[685,1172,359,1044]" pageId="25" pageNumber="52">
|
||
<emphasis id="09C2A366BB67FFB4FD0DFD32FC840EE0" bold="true" box="[709,809,651,672]" pageId="25" pageNumber="52">Remarks.</emphasis>
|
||
The genus
|
||
<taxonomicName id="FCB604F7BB67FFB4FC66FD32FB810EE0" box="[942,1068,651,672]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09C2A366BB67FFB4FC66FD32FB810EE0" box="[942,1068,651,672]" italics="true" pageId="25" pageNumber="52">Trogloderus</emphasis>
|
||
</taxonomicName>
|
||
is monotypic, with six subspecies, following the revision of
|
||
<bibRefCitation id="5F270285BB67FFB4FD01FD79FCDF0E95" author="La Rivers" box="[713,882,704,725]" pageId="25" pageNumber="52" pagination="35 - 44" refId="ref12990" refString="La Rivers, I. 1946. On the genus Trogloderus LeConte (Coleoptera: Tenebrionidae). Entomological News 57: 35 - 44." type="journal article" year="1946">La Rivers (1946)</bibRefCitation>
|
||
, and is found in aeolian sand formations across the western
|
||
<collectingCountry id="43A13FE4BB67FFB4FC14FD62FBCF0EB0" box="[988,1122,731,752]" name="United States of America" pageId="25" pageNumber="52">United States</collectingCountry>
|
||
. The two forms found in
|
||
<collectingRegion id="F972B196BB67FFB4FCBFFD4CFC6E0F4A" box="[887,963,757,778]" country="United States of America" name="Arizona" pageId="25" pageNumber="52">Arizona</collectingRegion>
|
||
are quite distinct. In the west, following the
|
||
<collectingRegion id="F972B196BB67FFB4FC4CFCA9FC740F65" box="[900,985,784,805]" country="United States of America" name="Colorado" pageId="25" pageNumber="52">Colorado</collectingRegion>
|
||
River, is
|
||
<taxonomicName id="FCB604F7BB67FFB4FBF9FCA9FCAD0F00" authorityName="La Rivers" authorityYear="1946" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="vandykei">
|
||
<emphasis id="09C2A366BB67FFB4FBF9FCA9FCAD0F00" italics="true" pageId="25" pageNumber="52">T. costatus vandykei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="A38D63F1BB67FFB4FCC6FC93FCF40F7F" box="[782,857,810,831]" captionStart="Fig" captionStartId="2.[197,229,1597,1616]" captionTargetBox="[173,1193,193,1574]" captionTargetId="figure-1@2.[172,1194,192,1574]" captionTargetPageId="2" captionText="Fig. 1. Amphidorini species, dorsal habitus. A) Trogloderus costatus, western form, B) T. costatus, northern form, C) Embaphion depressum, D) Embaphion contusum. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120670" httpUri="https://zenodo.org/record/10120670/files/figure.png" pageId="25" pageNumber="52">Fig. 1A</figureCitation>
|
||
), which is quite robust, lacks the sutural costa, has smooth elytral intervals, and a heavily punctate pronotum. The northern form (
|
||
<figureCitation id="A38D63F1BB67FFB4FD7CFCC3FD510FCF" box="[692,764,890,911]" captionStart="Fig" captionStartId="2.[197,229,1597,1616]" captionTargetBox="[173,1193,193,1574]" captionTargetId="figure-1@2.[172,1194,192,1574]" captionTargetPageId="2" captionText="Fig. 1. Amphidorini species, dorsal habitus. A) Trogloderus costatus, western form, B) T. costatus, northern form, C) Embaphion depressum, D) Embaphion contusum. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120670" httpUri="https://zenodo.org/record/10120670/files/figure.png" pageId="25" pageNumber="52">Fig. 1B</figureCitation>
|
||
) from the
|
||
<collectingRegion id="F972B196BB67FFB4FCACFCC3FC020FCF" box="[868,943,890,911]" country="United States of America" name="Arizona" pageId="25" pageNumber="52">Arizona</collectingRegion>
|
||
Strip is variously determined as
|
||
<taxonomicName id="FCB604F7BB67FFB4FCC6FC2CFC5B0FEA" authorityName="Blaisdell" authorityYear="1909" box="[782,1014,917,938]" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="tuberculatus">
|
||
<emphasis id="09C2A366BB67FFB4FCC6FC2CFC5B0FEA" box="[782,1014,917,938]" italics="true" pageId="25" pageNumber="52">T. costatus tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCB604F7BB67FFB4FBE2FC2CFD560F84" authorityName="La Rivers" authorityYear="1942" class="Insecta" family="Tenebrionidae" genus="Trogloderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="costatus" subSpecies="nevadus">
|
||
<emphasis id="09C2A366BB67FFB4FBE2FC2CFD560F84" italics="true" pageId="25" pageNumber="52">T. costatus nevadus</emphasis>
|
||
</taxonomicName>
|
||
and is recognizable by the presence of a sutural costa, generally tuberculate elytral intervals, and evenly tuberculate pronotum. A revision of the genus is currently in progress.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |