treatments-xml/data/B2/6A/C8/B26AC8752565FF81FF44FF01FC50FF4A.xml
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<document id="50F7700B44BDE0FB2309DE86F26DBAE9" ID-CLB-Dataset="30287" ID-DOI="10.11646/zootaxa.4407.2.1" ID-GBIF-Dataset="9cf2f92d-0c60-41da-891a-0452fe205926" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216693" ID-ZooBank="0409E956-3B2B-4D8B-A96C-26B70842FC28" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523435638400" checkinUser="plazi" docAuthor="Ramírez-Reyes, Tonatiuh &amp; Flores-Villela, Oscar" docDate="2018" docId="B26AC8752565FF81FF44FF01FC50FF4A" docLanguage="en" docName="zootaxa.4407.2.1.pdf" docOrigin="Zootaxa 4407 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllodactylus benedettii Ramírez-Reyes &amp; Flores-Villela 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="174" masterDocId="4E53B00D2574FF99FFD3FFDFFFE0FFE5" masterDocTitle="Taxonomic changes and description of two new species for the Phyllodactylus lanei complex (Gekkota: Phyllodactylidae) in Mexico" masterLastPageNumber="190" masterPageNumber="151" pageNumber="168" updateTime="1698536865782" updateUser="ExternalLinkService">
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<mods:title id="9D25FBADD9DAD6B49D0079E919CF66A3">Taxonomic changes and description of two new species for the Phyllodactylus lanei complex (Gekkota: Phyllodactylidae) in Mexico</mods:title>
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<mods:namePart id="859AF9B02E5381674B52F22CD19E5F96">Ramírez-Reyes, Tonatiuh</mods:namePart>
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<treatment id="B26AC8752565FF81FF44FF01FC50FF4A" ID-DOI="http://doi.org/10.5281/zenodo.5987595" ID-GBIF-Taxon="143536763" ID-Zenodo-Dep="5987595" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A" httpUri="http://treatment.plazi.org/id/B26AC8752565FF81FF44FF01FC50FF4A" lastPageId="24" lastPageNumber="174" pageId="17" pageNumber="168">
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<emphasis id="08B7A5712565FF88FF44FF01FDC9FF1D" bold="true" box="[151,553,222,248]" pageId="17" pageNumber="168">
<taxonomicName id="FDC302E02565FF88FF44FF01FE25FF1D" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[151,453,222,248]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712565FF88FF44FF01FE25FF1D" bold="true" box="[151,453,222,248]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1384180A2565FF88FE1FFF01FDC9FF1D" box="[460,553,222,248]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="72D92AE82565FF88FF44FEFAFC1AFD67" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="3A7C79632565FF88FF44FEFAFB05FE87" blockId="17.[151,1437,293,2010]" pageId="17" pageNumber="168">
<materialsCitation id="8AAB733E2565FF88FF44FEFAFB05FE87" ID-GBIF-Occurrence="1837804195" collectingDate="2014-09-15" collectionCode="MZFC" collectorName="La Huerta" latitude="19.5" location="Chamela Biological Station" longLatPrecision="763" longitude="-105.44" pageId="17" pageNumber="168" specimenCode="MZFC 28774" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="08B7A5712565FF88FF44FEFAFEEBFEDB" bold="true" box="[151,267,293,318]" pageId="17" pageNumber="168">
<typeStatus id="E578C7C12565FF88FF44FEFAFEE6FEDB" box="[151,262,293,318]" pageId="17" pageNumber="168" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male (
<specimenCode id="6A65D1182565FF88FE71FEF9FDA0FEDB" box="[418,576,294,318]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC 28774</specimenCode>
) collected at
<location id="3F1C2FB82565FF88FD09FEFAFBF6FEDB" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A:3F1C2FB82565FF88FD09FEFAFBF6FEDB" box="[730,1046,293,318]" latitude="19.5" longLatPrecision="763" longitude="-105.44" name="Chamela Biological Station" pageId="17" pageNumber="168" stateProvince="Jalisco">Chamela Biological Station</location>
,
<collectorName id="97361CB52565FF88FBF1FEF9FB77FED8" box="[1058,1175,294,317]" pageId="17" pageNumber="168">La Huerta</collectorName>
, state of
<collectingRegion id="F807B7812565FF88FB2CFEFAFAABFEDB" box="[1279,1355,293,318]" country="Mexico" name="Jalisco" pageId="17" pageNumber="168">Jalisco</collectingRegion>
(
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, -
<geoCoordinate id="5FF71FA42565FF88FF13FE96FECAFE87" box="[192,298,329,354]" direction="west" orientation="longitude" pageId="17" pageNumber="168" precision="555" value="-105.44">105.44W</geoCoordinate>
WGS84) on
<date id="4E7D5FA32565FF88FE6DFE96FD7FFE84" box="[446,671,329,354]" pageId="17" pageNumber="168" value="2014-09-15">
<collectingDate id="5E39A64B2565FF88FE6DFE96FD7FFE84" box="[446,671,329,354]" pageId="17" pageNumber="168" value="2014-09-15">September 15, 2014</collectingDate>
</date>
by Tonatiuh Ramírez Reyes and Alfredo Villarruel.
</materialsCitation>
</paragraph>
<paragraph id="3A7C79632565FF88FF14FEB2FC1AFD67" blockId="17.[151,1437,293,2010]" pageId="17" pageNumber="168">
<emphasis id="08B7A5712565FF88FF14FEB2FEDFFE63" bold="true" box="[199,319,365,390]" pageId="17" pageNumber="168">
<typeStatus id="E578C7C12565FF88FF14FEB2FEDFFE63" box="[199,319,365,390]" pageId="17" pageNumber="168" type="paratype">Paratypes</typeStatus>
</emphasis>
(14). An adult female (
<collectionCode id="5CD2E1A62565FF88FD91FEB1FD70FE63" box="[578,656,366,390]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
28773) collected at Chamela Biology Station, La Huerta municipality (
<geoCoordinate id="5FF71FA42565FF88FF4CFE4EFF1FFE4C" box="[159,255,401,426]" direction="north" orientation="latitude" pageId="17" pageNumber="168" precision="555" value="19.5">19.50 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42565FF88FEC0FE4EFE9EFE4F" box="[275,382,401,426]" direction="west" orientation="longitude" pageId="17" pageNumber="168" precision="555" value="-105.44">105.44W</geoCoordinate>
) on
<date id="4E7D5FA32565FF88FE61FE4EFD76FE4C" box="[434,662,401,426]" pageId="17" pageNumber="168" value="2014-09-15">September 15, 2014</date>
by Tonatiuh Ramírez Reyes and Alfredo Villarruel; (
<collectionCode id="5CD2E1A62565FF88FB25FE4EFAA4FE4C" box="[1270,1348,401,425]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21817- male,
<collectionCode id="5CD2E1A62565FF88FF0FFE69FEC8FE2B" box="[220,296,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21818-male,
<collectionCode id="5CD2E1A62565FF88FE17FE69FDF0FE2B" box="[452,528,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21877,
<collectionCode id="5CD2E1A62565FF88FDBFFE69FD58FE2B" box="[620,696,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21878-male,
<collectionCode id="5CD2E1A62565FF88FC80FE69FC7FFE2B" box="[851,927,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21879,
<collectionCode id="5CD2E1A62565FF88FC28FE69FBA7FE2B" box="[1019,1095,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21880-female,
<collectionCode id="5CD2E1A62565FF88FB2AFE69FAA5FE2B" box="[1273,1349,438,462]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="17" pageNumber="168">MZFC</collectionCode>
21885- male) collected at the Chamela Biology Station, La Huerta municipality (
<geoCoordinate id="5FF71FA42565FF88FC3BFE06FBA9FE14" box="[1000,1097,473,498]" direction="north" orientation="latitude" pageId="17" pageNumber="168" precision="555" value="19.5">19.50 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42565FF88FB8CFE06FB34FE14" box="[1119,1236,473,498]" direction="west" orientation="longitude" pageId="17" pageNumber="168" precision="555" value="-105.44">105.44 W</geoCoordinate>
) by Oscar
<collectingCountry id="42D439F32565FF88FA85FE06FA7CFE17" box="[1366,1436,473,498]" name="Indonesia" pageId="17" pageNumber="168">Flores</collectingCountry>
Villela in 2007; an adult female (
<collectionCode id="5CD2E1A62565FF88FDDAFE21FDDBFDF3" box="[521,571,510,534]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="17" pageNumber="168">IBH</collectionCode>
2133-2) and five adult males (
<collectionCode id="5CD2E1A62565FF88FC47FE21FC26FDF3" box="[916,966,510,534]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="17" pageNumber="168">IBH</collectionCode>
2133-3, 2133-4, 2133-5, 2133-7, 2133-9) collected at
<quantity id="FD3BD4862565FF88FEF3FDFEFEBDFDDF" box="[288,349,545,570]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="168" unit="km" value="2.0">2 km</quantity>
. SE, of the Station of Biology, Chamela (
<geoCoordinate id="5FF71FA42565FF88FCE2FDFEFC70FDDC" box="[817,912,545,570]" direction="north" orientation="latitude" pageId="17" pageNumber="168" precision="555" value="19.51">19.51 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42565FF88FC77FDFEFBF6FDDC" box="[932,1046,545,569]" direction="west" orientation="longitude" pageId="17" pageNumber="168" precision="555" value="-104.94">104.94 W</geoCoordinate>
) on
<date id="4E7D5FA32565FF88FB99FDFEFAEFFDDC" box="[1098,1295,545,570]" pageId="17" pageNumber="168" value="1971-10-27">October 27, 1971</date>
by Cornelio Sánchez; an adult female (
<collectionCode id="5CD2E1A62565FF88FE11FD99FE14FDBB" box="[450,500,582,606]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="17" pageNumber="168">IBH</collectionCode>
2138-4) collected at
<quantity id="FD3BD4862565FF88FD35FD9AFCC3FDBB" box="[742,803,581,606]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="km" value="5.0">5 km</quantity>
. S., de Chamela, Biology Station,
<collectionCode id="5CD2E1A62565FF88FB79FD99FAE2FDBB" box="[1194,1282,582,606]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="168">UNAM</collectionCode>
, municipality La Huerta (
<geoCoordinate id="5FF71FA42565FF88FEC5FDB5FE95FD64" box="[278,373,618,642]" direction="north" orientation="latitude" pageId="17" pageNumber="168" precision="555" value="19.49">19.49 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42565FF88FE5BFDB6FE1AFD67" box="[392,506,617,642]" direction="west" orientation="longitude" pageId="17" pageNumber="168" precision="555" value="-104.95">104.95 W</geoCoordinate>
) on
<date id="4E7D5FA32565FF88FDF8FDB5FD27FD67" box="[555,711,617,642]" pageId="17" pageNumber="168" value="1974-05-25">May 25, 1974</date>
by Gustavo Casas Andreu.
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82565FF88FF14FD52FE80FD0B" pageId="17" pageNumber="168" type="etymology">
<paragraph id="3A7C79632565FF88FF14FD52FE80FD0B" blockId="17.[151,1437,293,2010]" pageId="17" pageNumber="168">
<emphasis id="08B7A5712565FF88FF14FD52FEAEFD43" bold="true" box="[199,334,653,678]" pageId="17" pageNumber="168">Etymology.</emphasis>
The species is dedicated to the memory of the great Uruguayan writer Mario Benedetti in recognition of his prolific literary production and critical thinking of great importance in the political and social life of Latin America.
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82565FF8BFF14FD26FB9DFE81" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="diagnosis">
<paragraph id="3A7C79632565FF88FF14FD26FB1CFC23" blockId="17.[151,1437,293,2010]" pageId="17" pageNumber="168">
<emphasis id="08B7A5712565FF88FF14FD26FEA1FCF7" bold="true" box="[199,321,761,786]" pageId="17" pageNumber="168">Diagnosis.</emphasis>
<taxonomicName id="FDC302E02565FF88FE9FFD26FD8BFCF7" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[332,619,761,786]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712565FF88FE9FFD26FD8BFCF7" box="[332,619,761,786]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<emphasis id="08B7A5712565FF88FDA5FD26FD34FCF7" bold="true" box="[630,724,761,786]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="1384180A2565FF88FDA5FD26FD34FCF7" box="[630,724,761,786]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a species with the largest average size within the
<taxonomicName id="FDC302E02565FF88FA97FD25FA7CFCF7" authorityName="Smith" authorityYear="1935" box="[1348,1436,761,786]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FA97FD25FA7CFCF7" box="[1348,1436,761,786]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
complex, with a maximum SVL of
<quantity id="FD3BD4862565FF88FDF0FCC2FD69FCD0" box="[547,649,797,822]" metricMagnitude="-2" metricUnit="m" metricValue="7.52" pageId="17" pageNumber="168" unit="mm" value="75.2">75.2 mm</quantity>
.
<bibRefCitation id="5E5204922565FF88FD47FCC2FCCBFCD3" box="[660,811,797,822]" pageId="17" pageNumber="168" refId="ref16514" refString="Dixon, J. R. (1964) The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America. New Mexico State University Research Center, Scientific Bulletin, 1964, 1 - 139." type="journal article">Dixon (1964)</bibRefCitation>
established three fundamental characteristics to include species and subspecies within the
<taxonomicName id="FDC302E02565FF88FDCEFC9DFD90FCBF" authorityName="Smith" authorityYear="1935" box="[541,624,833,858]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FDCEFC9DFD90FCBF" box="[541,624,833,858]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
complex and differentiate them from
<taxonomicName id="FDC302E02565FF88FBF4FC9DFB32FCBF" authorityName="Wiegmann" authorityYear="1834" box="[1063,1234,833,858]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FBF4FC9DFB32FCBF" box="[1063,1234,833,858]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
: species of the
<taxonomicName id="FDC302E02565FF88FA54FC9DFF2FFC9B" authorityName="Smith" authorityYear="1935" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FA54FC9DFF2FFC9B" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
complex have a low number of interorbital scales (1219), white venter and low number of scales across the snout between the third labials (does not specify range of values).
<taxonomicName id="FDC302E02565FF88FC41FC56FB50FC47" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[914,1200,905,930]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FC41FC56FB50FC47" box="[914,1200,905,930]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
shows white-yellow venter, 14.4 interorbital scales on average, and 22.3 scales across the snout between the third labials.
</paragraph>
<paragraph id="3A7C79632565FF88FF14FC0EFD15FA7F" blockId="17.[151,1437,293,2010]" pageId="17" pageNumber="168">
Within the
<taxonomicName id="FDC302E02565FF88FE97FC0DFE74FC0F" authorityName="Smith" authorityYear="1935" box="[324,404,977,1002]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FE97FC0DFE74FC0F" box="[324,404,977,1002]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
complex (including insular and continental species), the only species that exceed
<quantity id="FD3BD4862565FF88FAFCFC0DFA9EFC0C" box="[1327,1406,978,1002]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="168" unit="mm" value="70.0">70 mm</quantity>
of SVL are
<taxonomicName id="FDC302E02565FF88FED3FC29FE8BFBEB" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[256,363,1013,1038]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FED3FC29FE8BFBEB" box="[256,363,1013,1038]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02565FF88FE75FC29FE18FBEB" authorityName="Smith" authorityYear="1935" box="[422,504,1013,1038]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FE75FC29FE18FBEB" box="[422,504,1013,1038]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
, two species that are very isolated geographically. It is clearly different from the nearest species
<taxonomicName id="FDC302E02565FF88FE82FBC5FE29FBD4" box="[337,457,1050,1073]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FE82FBC5FE29FBD4" box="[337,457,1050,1073]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
, which has a maximum SVL of
<quantity id="FD3BD4862565FF88FCB3FBC6FC2CFBD4" box="[864,972,1049,1074]" metricMagnitude="-2" metricUnit="m" metricValue="6.94" pageId="17" pageNumber="168" unit="mm" value="69.4">69.4 mm</quantity>
, considered as a medium size species. According to the statistical analyzes previously carried out, the combination of diagnostic morphological characters are the following (in addition to the SVL): It differs clearly from all species of
<taxonomicName id="FDC302E02565FF88FB69FBBEFA80FB9F" authorityName=", Dixon" authorityYear="1964" box="[1210,1376,1121,1146]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
<emphasis id="08B7A5712565FF88FB69FBBEFA80FB9F" box="[1210,1376,1121,1146]" italics="true" pageId="17" pageNumber="168">Phyllodactylus</emphasis>
</taxonomicName>
as it presents 62.6 (6069) longitudinal scales, 27 (2430) tubercles from head to tail, 14 (1113) interorbital scales, 29 (2532) scales across venter, 22 (2223) third labialsnout scales, 13 (1314) rows of tubercles across dorsum. Although
<taxonomicName id="FDC302E02565FF88FED8FB11FE79FB03" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[267,409,1229,1254]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FED8FB11FE79FB03" box="[267,409,1229,1254]" italics="true" pageId="17" pageNumber="168">P. benedettii</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="FDC302E02565FF88FDA5FB11FD03FB03" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[630,739,1229,1254]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FDA5FB11FD03FB03" box="[630,739,1229,1254]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
in some meristic characters, they present differences in the morphometric space (
<figureCitation id="A2F865E62565FF88FE58FB2EFE2FFAEF" box="[395,463,1265,1290]" captionStart="FIGURE 7" captionStartId="16.[151,250,1148,1170]" captionTargetBox="[161,1373,517,1099]" captionTargetId="figure@16.[151,1436,444,1127]" captionTargetPageId="16" captionText="FIGURE 7. Scatter plot of 11 morphometric variables for six species of the P.lanei complex:P.benedettii (black, group 1), P. kropotkini (aqua blue, group 2), P. rupinus (brown, group 3), P. lanei (blue, group 4), P. lupitae (red, group 5), P. isabelae (green, group 6). The first two components are shown with the percentage of variance on the axes." httpUri="https://zenodo.org/record/1216707/files/figure.png" pageId="17" pageNumber="168">Fig. 7</figureCitation>
). Some measures allow us to differentiate these species are the head-length and axillagroin length:
<taxonomicName id="FDC302E02565FF88FEEBFAC9FE26FACB" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[312,454,1301,1326]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FEEBFAC9FE26FACB" box="[312,454,1301,1326]" italics="true" pageId="17" pageNumber="168">P. benedettii</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862565FF88FE08FAC9FDA5FAC8" box="[475,581,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="17" pageNumber="168" unit="mm" value="17.9">17.9 mm</quantity>
and
<quantity id="FD3BD4862565FF88FD56FAC9FD0FFACB" box="[645,751,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="17" pageNumber="168" unit="mm" value="28.4">28.4 mm</quantity>
) and
<taxonomicName id="FDC302E02565FF88FCE4FAC9FC45FACB" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[823,933,1301,1326]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FCE4FAC9FC45FACB" box="[823,933,1301,1326]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862565FF88FC6AFACAFBD1FAC8" box="[953,1073,1301,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.686" pageId="17" pageNumber="168" unit="mm" value="16.86">16.86 mm</quantity>
and
<quantity id="FD3BD4862565FF88FBA2FAC9FB0CFACB" box="[1137,1260,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.793" pageId="17" pageNumber="168" unit="mm" value="27.93">27.93 mm</quantity>
). According to
<bibRefCitation id="5E5204922565FF88FF44FAE6FE72FAB7" box="[151,402,1337,1362]" pageId="17" pageNumber="168" refId="ref16092" refString="Castiglia, R, Aguayo, A. G., Bezerra, A. M. R., Flores-Villela, O. &amp; Gournung, E. (2009) Karyotypic diversification due to Robertsonian rearrangements in Phyllodactylus lanei Smith, 1935 (Squamata, Gekkonidae) from Mexico. Atti della Accademia Nazionale dei Lincei, Rendiconti Lincei, Scienze Fisiche e Naturali, 20, 77 - 82. https: // doi. org / 10.1007 / s 12210 - 009 - 0005 - 4" type="journal article">Castiglia et al. (2009</bibRefCitation>
;
<bibRefCitation id="5E5204922565FF88FE72FAE5FE3DFAB7" box="[417,477,1338,1362]" pageId="17" pageNumber="168" refId="ref16182" refString="Castiglia, R., Annesi, F., Bezerra, A. M. R., Garcia, A. &amp; Flores-Villela, O. (2010) Cytotaxonomy and DNA taxonomy of lizards (Squamata, Sauria) from a tropical dry forest in the Chamela-Cuixmala Biosphere Reserve on the coast of Jalisco, Mexico. Zootaxa, 2508, 1 - 29." type="journal article">2010</bibRefCitation>
), the karyotypes of
<emphasis id="08B7A5712565FF88FD1FFAE5FCDFFAB7" box="[716,831,1337,1362]" italics="true" pageId="17" pageNumber="168">
P. l.
<taxonomicName id="FDC302E02565FF88FCD4FAE6FCDFFAB7" authorityName="Smith" authorityYear="1935" box="[775,831,1337,1362]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">lanei</taxonomicName>
</emphasis>
(Tierra Colorada,
<collectingRegion id="F807B7812565FF88FBCFFAE5FB63FAB7" box="[1052,1155,1338,1362]" country="Mexico" name="Guerrero" pageId="17" pageNumber="168">Guerrero</collectingRegion>
) and
<emphasis id="08B7A5712565FF88FB19FAE5FABAFAB4" box="[1226,1370,1337,1362]" italics="true" pageId="17" pageNumber="168">
P. l.
<taxonomicName id="FDC302E02565FF88FAD7FAE5FABAFAB4" box="[1284,1370,1338,1361]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">rupinus</taxonomicName>
</emphasis>
from Chamela,
<collectingRegion id="F807B7812565FF88FEDBFA82FEB4FA93" box="[264,340,1373,1398]" country="Mexico" name="Jalisco" pageId="17" pageNumber="168">Jalisco</collectingRegion>
(here named
<taxonomicName id="FDC302E02565FF88FE3FFA81FD8FFA93" box="[492,623,1373,1398]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedetti">
<emphasis id="08B7A5712565FF88FE3FFA81FD8FFA93" box="[492,623,1373,1398]" italics="true" pageId="17" pageNumber="168">P. benedetti</emphasis>
</taxonomicName>
) are different. According to their studies
<taxonomicName id="FDC302E02565FF88FBEEFA81FB6CFA93" authorityName="Smith" authorityYear="1935" box="[1085,1164,1373,1398]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FBEEFA81FB6CFA93" box="[1085,1164,1373,1398]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
has 2n = 3334 and FN = 4041, while
<taxonomicName id="FDC302E02565FF88FE94FA5DFE30FA7F" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[327,464,1409,1434]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FE94FA5DFE30FA7F" box="[327,464,1409,1434]" italics="true" pageId="17" pageNumber="168">P. benedettii</emphasis>
</taxonomicName>
has 2n = 38 and FN = 38.
</paragraph>
<paragraph id="3A7C79632565FF8BFF14FA7AFB9DFE81" blockId="17.[151,1437,293,2010]" lastBlockId="18.[151,1437,151,933]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<taxonomicName id="FDC302E02565FF88FF14FA7AFE03FA5B" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[199,483,1445,1470]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FF14FA7AFE03FA5B" box="[199,483,1445,1470]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
has fewer than 30 tubercles from head to tail (2430, 27.4 avg.) similar to
<taxonomicName id="FDC302E02565FF88FAE1FA79FA7CFA5B" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[1330,1436,1445,1470]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FAE1FA79FA7CFA5B" box="[1330,1436,1445,1470]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(2731, 28.8 avg.),
<taxonomicName id="FDC302E02565FF88FEABFA15FDBEFA07" authorityName="Dixon" authorityYear="1960" box="[376,606,1481,1506]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="paucituberculatus">
<emphasis id="08B7A5712565FF88FEABFA15FDBEFA07" box="[376,606,1481,1506]" italics="true" pageId="17" pageNumber="168">P. paucituberculatus</emphasis>
</taxonomicName>
(28.7),
<taxonomicName id="FDC302E02565FF88FD69FA15FCA9FA07" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[698,841,1481,1506]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712565FF88FD69FA15FCA9FA07" box="[698,841,1481,1506]" italics="true" pageId="17" pageNumber="168">P. kropotkini</emphasis>
</taxonomicName>
(2531, 28.4 avg.) and
<taxonomicName id="FDC302E02565FF88FB8FFA15FB2FFA04" box="[1116,1231,1482,1505]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FB8FFA15FB2FFA04" box="[1116,1231,1482,1505]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
(2630, 28 avg.); other species have more than 30 tubercles from head to tail on average:
<taxonomicName id="FDC302E02565FF88FC66FA31FBE5F9E3" authorityName="Smith" authorityYear="1935" box="[949,1029,1517,1542]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FC66FA31FBE5F9E3" box="[949,1029,1517,1542]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
(3234, 33.2 avg.),
<taxonomicName id="FDC302E02565FF88FB39FA31FA82F9E3" box="[1258,1378,1517,1542]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712565FF88FB39FA31FA82F9E3" box="[1258,1378,1517,1542]" italics="true" pageId="17" pageNumber="168">P. isabelae</emphasis>
</taxonomicName>
(30 35, 32.2 avg.),
<taxonomicName id="FDC302E02565FF88FE97F9CDFE2EF9CC" authorityName="Taylor" authorityYear="1942" box="[324,462,1554,1577]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712565FF88FE97F9CDFE2EF9CC" box="[324,462,1554,1577]" italics="true" pageId="17" pageNumber="168">P. t. magnus</emphasis>
</taxonomicName>
(3540, 37 avg.),
<taxonomicName id="FDC302E02565FF88FD71F9CDFCF4F9CF" authorityName="Taylor" authorityYear="1940" box="[674,788,1553,1578]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712565FF88FD71F9CDFCF4F9CF" box="[674,788,1553,1578]" italics="true" pageId="17" pageNumber="168">P. muralis</emphasis>
</taxonomicName>
(3040, 33 avg.) and
<taxonomicName id="FDC302E02565FF88FBC2F9CDFB5CF9CF" authorityName="Wiegmann" authorityYear="1834" box="[1041,1212,1553,1578]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FBC2F9CDFB5CF9CF" box="[1041,1212,1553,1578]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
(3341, 36.4 avg.).
<taxonomicName id="FDC302E02565FF88FF44F9EAFE55F9AB" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[151,437,1589,1614]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FF44F9EAFE55F9AB" box="[151,437,1589,1614]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
showed 62 longitudinal ventral scales (LVS) on average (6069), other values were
<taxonomicName id="FDC302E02565FF88FA54F9E9FF14F997" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712565FF88FA54F9E9FF14F997" italics="true" pageId="17" pageNumber="168">P. isabelae</emphasis>
</taxonomicName>
(5258, 56.2 avg.),
<taxonomicName id="FDC302E02565FF88FE0BF985FDA2F997" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[472,578,1625,1650]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FE0BF985FDA2F997" box="[472,578,1625,1650]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(6064, 61.8 avg.),
<taxonomicName id="FDC302E02565FF88FCF6F985FC99F997" authorityName="Smith" authorityYear="1935" box="[805,889,1625,1650]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FCF6F985FC99F997" box="[805,889,1625,1650]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
(6074, 66 avg.),
<taxonomicName id="FDC302E02565FF88FB94F985FB58F994" box="[1095,1208,1626,1649]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FB94F985FB58F994" box="[1095,1208,1626,1649]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
(5667, 63 avg.),
<taxonomicName id="FDC302E02565FF88FA54F985FEE9F973" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712565FF88FA54F985FEE9F973" italics="true" pageId="17" pageNumber="168">P. kropotkini</emphasis>
</taxonomicName>
(6373, 67 avg.),
<taxonomicName id="FDC302E02565FF88FE04F9A1FDBFF970" authorityName="Taylor" authorityYear="1942" box="[471,607,1662,1685]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712565FF88FE04F9A1FDBFF970" box="[471,607,1662,1685]" italics="true" pageId="17" pageNumber="168">P. t. magnus</emphasis>
</taxonomicName>
(5258, 54 avg.),
<taxonomicName id="FDC302E02565FF88FCFEF9A1FC7EF973" authorityName="Taylor" authorityYear="1940" box="[813,926,1661,1686]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712565FF88FCFEF9A1FC7EF973" box="[813,926,1661,1686]" italics="true" pageId="17" pageNumber="168">P. muralis</emphasis>
</taxonomicName>
(5761, 59 avg.) and
<taxonomicName id="FDC302E02565FF88FB46F9A1FADEF973" authorityName="Wiegmann" authorityYear="1834" box="[1173,1342,1661,1686]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FB46F9A1FADEF973" box="[1173,1342,1661,1686]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
(5164, 57 avg.).
<taxonomicName id="FDC302E02565FF88FED1F97EFDFFF95F" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[258,543,1697,1722]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FED1F97EFDFFF95F" box="[258,543,1697,1722]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
presents 14.4 interorbital scales similar to
<taxonomicName id="FDC302E02565FF88FBD6F97DFB74F95F" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1029,1172,1697,1722]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712565FF88FBD6F97DFB74F95F" box="[1029,1172,1697,1722]" italics="true" pageId="17" pageNumber="168">P. kropotkini</emphasis>
</taxonomicName>
(1416, 14.6 avg.) and differs from
<taxonomicName id="FDC302E02565FF88FEFBF919FE43F93B" box="[296,419,1733,1758]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712565FF88FEFBF919FE43F93B" box="[296,419,1733,1758]" italics="true" pageId="17" pageNumber="168">P. isabelae</emphasis>
</taxonomicName>
(1420, 15.1 avg.),
<taxonomicName id="FDC302E02565FF88FD5CF919FD1BF93B" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[655,763,1733,1758]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FD5CF919FD1BF93B" box="[655,763,1733,1758]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(1420, 16.8 avg.),
<taxonomicName id="FDC302E02565FF88FC34F919FBD9F93B" authorityName="Smith" authorityYear="1935" box="[999,1081,1733,1758]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FC34F919FBD9F93B" box="[999,1081,1733,1758]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
(1517, 15.5 avg.),
<taxonomicName id="FDC302E02565FF88FAFAF919FA7BF938" box="[1321,1435,1734,1757]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FAFAF919FA7BF938" box="[1321,1435,1734,1757]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
(1417, 15.5 avg.),
<taxonomicName id="FDC302E02565FF88FEA1F935FE1AF8E4" authorityName="Taylor" authorityYear="1942" box="[370,506,1770,1793]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712565FF88FEA1F935FE1AF8E4" box="[370,506,1770,1793]" italics="true" pageId="17" pageNumber="168">P. t. magnus</emphasis>
</taxonomicName>
(2123, 23.3 avg.),
<taxonomicName id="FDC302E02565FF88FD0FF935FCADF8E7" authorityName="Taylor" authorityYear="1940" box="[732,845,1769,1794]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712565FF88FD0FF935FCADF8E7" box="[732,845,1769,1794]" italics="true" pageId="17" pageNumber="168">P. muralis</emphasis>
</taxonomicName>
(2127, 23.2 avg.), and
<taxonomicName id="FDC302E02565FF88FB8DF935FAE7F8E7" authorityName="Wiegmann" authorityYear="1834" box="[1118,1287,1769,1794]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FB8DF935FAE7F8E7" box="[1118,1287,1769,1794]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
(1622, 19.2 avg.).
<taxonomicName id="FDC302E02565FF88FF0DF8D2FE1AF8C3" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[222,506,1805,1830]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FF0DF8D2FE1AF8C3" box="[222,506,1805,1830]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
presents 29 scales across venter and differs from
<taxonomicName id="FDC302E02565FF88FBF9F8D1FB43F8C3" box="[1066,1187,1805,1830]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712565FF88FBF9F8D1FB43F8C3" box="[1066,1187,1805,1830]" italics="true" pageId="17" pageNumber="168">P. isabelae</emphasis>
</taxonomicName>
(2629, 27.8 avg.),
<taxonomicName id="FDC302E02565FF88FA54F8D1FF05F8AF" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FA54F8D1FF05F8AF" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(2629, 24.6 avg.),
<taxonomicName id="FDC302E02565FF88FE16F8EDFDF8F8AF" authorityName="Smith" authorityYear="1935" box="[453,536,1841,1866]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FE16F8EDFDF8F8AF" box="[453,536,1841,1866]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
(2932, 30 avg.),
<taxonomicName id="FDC302E02565FF88FD37F8EDFCB4F8AC" box="[740,852,1842,1865]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FD37F8EDFCB4F8AC" box="[740,852,1842,1865]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
(2428, 26.5 avg.),
<taxonomicName id="FDC302E02565FF88FBE6F8EDFB22F8AF" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1077,1218,1841,1866]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712565FF88FBE6F8EDFB22F8AF" box="[1077,1218,1841,1866]" italics="true" pageId="17" pageNumber="168">P. kropotkini</emphasis>
</taxonomicName>
(2933, 30.6 avg.),
<taxonomicName id="FDC302E02565FF88FF44F889FEC2F888" authorityName="Taylor" authorityYear="1942" box="[151,290,1878,1901]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712565FF88FF44F889FEC2F888" box="[151,290,1878,1901]" italics="true" pageId="17" pageNumber="168">P. t. magnus</emphasis>
</taxonomicName>
(2629, 27.3 avg.),
<taxonomicName id="FDC302E02565FF88FDD9F889FD99F88B" authorityName="Taylor" authorityYear="1940" box="[522,633,1877,1902]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712565FF88FDD9F889FD99F88B" box="[522,633,1877,1902]" italics="true" pageId="17" pageNumber="168">P. muralis</emphasis>
</taxonomicName>
(33) and
<taxonomicName id="FDC302E02565FF88FD3FF889FC77F88B" authorityName="Wiegmann" authorityYear="1834" box="[748,919,1877,1902]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FD3FF889FC77F88B" box="[748,919,1877,1902]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
(2733, 30.2 avg.).
<taxonomicName id="FDC302E02565FF88FBACF88AFA7BF88B" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1151,1435,1877,1902]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF88FBACF88AFA7BF88B" box="[1151,1435,1877,1902]" italics="true" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
presents 22 third labialsnout scales on average (2223) and differs from
<taxonomicName id="FDC302E02565FF88FC07F8A5FBADF877" box="[980,1101,1913,1938]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712565FF88FC07F8A5FBADF877" box="[980,1101,1913,1938]" italics="true" pageId="17" pageNumber="168">P. isabelae</emphasis>
</taxonomicName>
(1924, 21.4 avg.),
<taxonomicName id="FDC302E02565FF88FAE1F8A5FA7CF877" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[1330,1436,1913,1938]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712565FF88FAE1F8A5FA7CF877" box="[1330,1436,1913,1938]" italics="true" pageId="17" pageNumber="168">P. lupitae</emphasis>
</taxonomicName>
(2328, 25.5 avg.),
<taxonomicName id="FDC302E02565FF88FEA0F841FE27F853" authorityName="Smith" authorityYear="1935" box="[371,455,1949,1974]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712565FF88FEA0F841FE27F853" box="[371,455,1949,1974]" italics="true" pageId="17" pageNumber="168">P. lanei</emphasis>
</taxonomicName>
(2023, 21.2 avg.),
<taxonomicName id="FDC302E02565FF88FD78F841FCFCF850" box="[683,796,1950,1973]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712565FF88FD78F841FCFCF850" box="[683,796,1950,1973]" italics="true" pageId="17" pageNumber="168">P. rupinus</emphasis>
</taxonomicName>
(1822, 20.6 avg.),
<taxonomicName id="FDC302E02565FF88FBD3F841FB6DF853" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1024,1165,1949,1974]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712565FF88FBD3F841FB6DF853" box="[1024,1165,1949,1974]" italics="true" pageId="17" pageNumber="168">P. kropotkini</emphasis>
</taxonomicName>
(1921, 20.2 avg.),
<taxonomicName id="FDC302E02565FF88FAA2F841FF0EF83C" authorityName="Taylor" authorityYear="1942" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712565FF88FAA2F841FF0EF83C" italics="true" pageId="17" pageNumber="168">P. t. magnus</emphasis>
</taxonomicName>
(2426, 24.6 avg.),
<taxonomicName id="FDC302E02565FF88FE0BF81DFDA7F83F" authorityName="Taylor" authorityYear="1940" box="[472,583,1985,2010]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712565FF88FE0BF81DFDA7F83F" box="[472,583,1985,2010]" italics="true" pageId="17" pageNumber="168">P. muralis</emphasis>
</taxonomicName>
(2428, 25.2 avg.) and
<taxonomicName id="FDC302E02565FF88FCB3F81DFBEBF83F" authorityName="Wiegmann" authorityYear="1834" box="[864,1035,1985,2010]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712565FF88FCB3F81DFBEBF83F" box="[864,1035,1985,2010]" italics="true" pageId="17" pageNumber="168">P. tuberculosus</emphasis>
</taxonomicName>
(2126, 24.2 avg.).
<taxonomicName id="FDC302E02565FF8BFB26F81EFEE5FF55" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="18" lastPageNumber="169" order="Squamata" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712565FF8BFB26F81EFEE5FF55" italics="true" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">Phyllodactylus benedettii</emphasis>
</taxonomicName>
presents 13.8 rows of tubercles across dorsum on average (1314) and differs from
<taxonomicName id="FDC302E02566FF8BFB17FF46FADDFF55" box="[1220,1341,151,176]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712566FF8BFB17FF46FADDFF55" box="[1220,1341,151,176]" italics="true" pageId="18" pageNumber="169">P. isabelae</emphasis>
</taxonomicName>
(1518, 16.7 avg.),
<taxonomicName id="FDC302E02566FF8BFEC6FF62FE9BFF30" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[277,379,188,213]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712566FF8BFEC6FF62FE9BFF30" box="[277,379,188,213]" italics="true" pageId="18" pageNumber="169">P. lupitae</emphasis>
</taxonomicName>
(1415, 14.8 avg.),
<taxonomicName id="FDC302E02566FF8BFDB2FF62FD54FF30" authorityName="Smith" authorityYear="1935" box="[609,692,188,213]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712566FF8BFDB2FF62FD54FF30" box="[609,692,188,213]" italics="true" pageId="18" pageNumber="169">P. lanei</emphasis>
</taxonomicName>
(1416, 15.6 avg.),
<taxonomicName id="FDC302E02566FF8BFC45FF62FBE4FF31" box="[918,1028,189,212]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712566FF8BFC45FF62FBE4FF31" box="[918,1028,189,212]" italics="true" pageId="18" pageNumber="169">P. rupinus</emphasis>
</taxonomicName>
(1315, 14 avg.),
<taxonomicName id="FDC302E02566FF8BFB07FF62FA82FF30" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1236,1378,188,213]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712566FF8BFB07FF62FA82FF30" box="[1236,1378,188,213]" italics="true" pageId="18" pageNumber="169">P. kropotkini</emphasis>
</taxonomicName>
(12 14, 13.4 avg.),
<taxonomicName id="FDC302E02566FF8BFE93FF3EFE29FF1D" authorityName="Taylor" authorityYear="1942" box="[320,457,225,248]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712566FF8BFE93FF3EFE29FF1D" box="[320,457,225,248]" italics="true" pageId="18" pageNumber="169">P. t. magnus</emphasis>
</taxonomicName>
(1315, 14.3 avg.),
<taxonomicName id="FDC302E02566FF8BFD7EFF3EFCFEFF1D" authorityName="Taylor" authorityYear="1940" box="[685,798,223,248]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712566FF8BFD7EFF3EFCFEFF1D" box="[685,798,223,248]" italics="true" pageId="18" pageNumber="169">P. muralis</emphasis>
</taxonomicName>
(1213, 12.2 avg.) and
<taxonomicName id="FDC302E02566FF8BFBF8FF3EFB34FF1D" authorityName="Wiegmann" authorityYear="1834" box="[1067,1236,223,248]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712566FF8BFBF8FF3EFB34FF1D" box="[1067,1236,223,248]" italics="true" pageId="18" pageNumber="169">P. tuberculosus</emphasis>
</taxonomicName>
(1217, 14 avg.). Finally,
<taxonomicName id="FDC302E02566FF8BFF21FEDAFE9BFEF8" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[242,379,260,285]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712566FF8BFF21FEDAFE9BFEF8" box="[242,379,260,285]" italics="true" pageId="18" pageNumber="169">P. benedettii</emphasis>
</taxonomicName>
presents the largest values in the following measures:
<quantity id="FD3BD4862566FF8BFC0EFEDBFBA1FEF9" box="[989,1089,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="169" unit="mm" value="5.5">5.50 mm</quantity>
in length of the 4th toepad and
<quantity id="FD3BD4862566FF8BFF44FEF8FF1AFEDA" box="[151,250,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="18" pageNumber="169" unit="mm" value="35.8">35.8 mm</quantity>
in axilla-groin length (LAG); compared to
<taxonomicName id="FDC302E02566FF8BFD33FEF6FCB5FEA5" box="[736,853,295,320]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712566FF8BFD33FEF6FCB5FEA5" box="[736,853,295,320]" italics="true" pageId="18" pageNumber="169">P. isabelae</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862566FF8BFCB4FEF7FC20FEDA" box="[871,960,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="169" unit="mm" value="4.1">4.1 mm</quantity>
,
<quantity id="FD3BD4862566FF8BFC1FFEF7FBD3FEA5" box="[972,1075,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.2199999999999998" pageId="18" pageNumber="169" unit="mm" value="22.2">22.2 mm</quantity>
),
<taxonomicName id="FDC302E02566FF8BFB95FEF6FB4FFEA5" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[1094,1199,295,320]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712566FF8BFB95FEF6FB4FFEA5" box="[1094,1199,295,320]" italics="true" pageId="18" pageNumber="169">P. lupitae</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862566FF8BFB6EFEF7FAF6FEDA" box="[1213,1302,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="18" pageNumber="169" unit="mm" value="4.9">4.9 mm</quantity>
,
<quantity id="FD3BD4862566FF8BFAF1FEF7FA95FEA5" box="[1314,1397,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="169" unit="mm" value="31.0">31 mm</quantity>
),
<taxonomicName id="FDC302E02566FF8BFA54FEF6FF2FFE80" authorityName="Smith" authorityYear="1935" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712566FF8BFA54FEF6FF2FFE80" italics="true" pageId="18" pageNumber="169">P. lanei</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862566FF8BFF0DFE93FED7FE81" box="[222,311,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="18" pageNumber="169" unit="mm" value="4.6">4.6 mm</quantity>
,
<quantity id="FD3BD4862566FF8BFE90FE93FE4BFE81" box="[323,427,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="18" pageNumber="169" unit="mm" value="21.1">21.1 mm</quantity>
),
<taxonomicName id="FDC302E02566FF8BFE6EFE92FDCFFE81" box="[445,559,333,356]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712566FF8BFE6EFE92FDCFFE81" box="[445,559,333,356]" italics="true" pageId="18" pageNumber="169">P. rupinus</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862566FF8BFDEDFE93FD62FE81" box="[574,642,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="169" unit="mm" value="5.0">5 mm</quantity>
,
<quantity id="FD3BD4862566FF8BFD5EFE93FD12FE81" box="[653,754,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="18" pageNumber="169" unit="mm" value="25.6">25.6 mm</quantity>
) and
<taxonomicName id="FDC302E02566FF8BFCE2FE92FC5EFE80" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[817,958,332,357]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712566FF8BFCE2FE92FC5EFE80" box="[817,958,332,357]" italics="true" pageId="18" pageNumber="169">P. kropotkini</emphasis>
</taxonomicName>
(
<quantity id="FD3BD4862566FF8BFC1DFE93FBF2FE81" box="[974,1042,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="169" unit="mm" value="5.0">5 mm</quantity>
,
<quantity id="FD3BD4862566FF8BFBCEFE93FB91FE81" box="[1053,1137,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="169" unit="mm" value="23.0">23 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82566FF8AFF14FEB0FC95F863" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" type="description">
<paragraph id="3A7C79632566FF8BFF14FEB0FC05FC40" blockId="18.[151,1437,151,933]" pageId="18" pageNumber="169">
<emphasis id="08B7A5712566FF8BFF14FEB0FDA6FE6D" bold="true" box="[199,582,367,392]" pageId="18" pageNumber="169">
Description of
<typeStatus id="E578C7C12566FF8BFEAAFEB0FE00FE6D" box="[377,480,367,392]" pageId="18" pageNumber="169" type="holotype">holotype</typeStatus>
(
<figureCitation id="A2F865E62566FF8BFE23FEB0FDD7FE6D" box="[496,567,367,392]" captionStart="FIGURE 9" captionStartId="18.[151,250,1714,1736]" captionTargetBox="[151,1422,985,1693]" captionTargetId="figure@18.[151,1436,985,1693]" captionTargetPageId="18" captionText="FIGURE 9. Dorsal and ventral view of the holotype of Phyllodactylus benedettii sp. nov. (MZFC 28774). The black bars represent the scale (10 mm)." httpUri="https://zenodo.org/record/1216711/files/figure.png" pageId="18" pageNumber="169">Fig. 9</figureCitation>
).
</emphasis>
(All bilateral counts are given as right/left). Adult male, robust body, head not flattened, neck slightly differentiated from the head. The head scales are granular, most are slightly globose, and are interspersed with circular-oblique flat scales that are mainly located between the eyes towards the tip of the snout. The rostral scale presents an intermediate striation near to middle of the scale (a groove that extends longitudinally to the middle part of the rostral scale); nostril bordered by a simple supranasal (one supranasal scale only), first labial, rostral and two postnasals; two supranasal scales contact each other, bordered by 7 postnasals; 22 interorbital scales beginning at the anterior ocular border; 15 scales across the snout from the second right labial, 24 scales across the snout at the level of the third labial scale on the right side; 8/9 loreal scales from the first ocular scale to the nostril; 11/10 supralabial scales; posterior border of the first supralabial in contact with the nostril (right and left); 8/8 infralabial scales; elongated auricular opening smaller than ocular diameter; occipital scales different in size and shape to the interorbital scales, while the interorbital scales are more or less uniform in size and shape (oval shape), they are smaller than the occipital ones. The occipital scales are larger than the interorbital scales, they have irregular shapes and presents some circular tubercles intermixed; mental scale is equally wide as long (
<quantity id="FD3BD4862566FF8BFF4CFC9BFF1AFCB9" box="[159,250,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="169" unit="mm" value="3.6">3.6 mm</quantity>
), forms an irregular &quot;V&quot; towards the posterior section, two postmental scales closely in contact each other and with the first and second infralabials; postmental scales in contact with a row of 4 scales, the 2 intermediates larger than the extremes, 10 scales of uniform size border the row of scales.
</paragraph>
<caption id="6EBC29EB2566FF8BFF44F96DFE57F902" httpUri="https://zenodo.org/record/1216711/files/figure.png" pageId="18" pageNumber="169" startId="18.[151,250,1714,1736]" targetBox="[151,1422,985,1693]" targetPageId="18">
<paragraph id="3A7C79632566FF8BFF44F96DFE57F902" blockId="18.[151,1436,1714,1767]" pageId="18" pageNumber="169">
<emphasis id="08B7A5712566FF8BFF44F96DFEF5F922" bold="true" box="[151,277,1714,1736]" pageId="18" pageNumber="169">FIGURE 9.</emphasis>
Dorsal and ventral view of the holotype of
<taxonomicName id="FDC302E02566FF8BFD39F96DFC0DF92D" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[746,1005,1714,1736]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712566FF8BFD39F96DFC0DF92D" box="[746,1005,1714,1736]" italics="true" pageId="18" pageNumber="169">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<emphasis id="08B7A5712566FF8BFC25F96CFBA9F92D" bold="true" box="[1014,1097,1715,1736]" pageId="18" pageNumber="169">
<taxonomicNameLabel id="1384180A2566FF8BFC25F96CFBA9F92D" box="[1014,1097,1715,1736]" pageId="18" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZFC 28774). The black bars represent the scale (10 mm).
</paragraph>
</caption>
<paragraph id="3A7C79632566FF8AFF14F8D1FC4AFF12" blockId="18.[151,1436,1806,2011]" lastBlockId="19.[151,1436,151,861]" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169">
Body with granular scales; 13 rows of strongly keeled dorsal tubercles with variable size; 25 tubercles in a middle dorsal row from base of the head to base of the tail; 21/15 tubercles from axilla to groin; 34 rows of cycloid scales across the venter; 87 longitudinal ventral scales from the first differentiated scale on the gular region to the cloacal opening; ventral scales larger compared to the lateral scales of the body and the gular region; scales slightly imbricate on the anterior and posterior extremities, scales dorsally granular intercalated with strongly keeled tubercles of variable size. The tail of the specimen is original (not regenerated) and measures
<quantity id="FD3BD4862566FF8BFB62F81DFAF5F83F" box="[1201,1301,1986,2011]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" pageId="18" pageNumber="169" unit="mm" value="55.5">55.5 mm</quantity>
with the tip broken. Dorsal scales of the tail are imbricate with intercalated tubercles and juxtaposed ventral scales, keeled tubercles extend to more than half of the tail, as they extend to the tip of the tail the tubercles gradually flatten until they form keeled scales. Does not present femoral or precloacal pores.
</paragraph>
<paragraph id="3A7C79632567FF8AFF14FEDBFE90FE80" blockId="19.[151,1436,151,861]" pageId="19" pageNumber="170">
Digital lamella formulae: right posterior (
<date id="4E7D5FA32567FF8AFD44FEDBFCCFFEF9" box="[663,815,260,284]" pageId="19" pageNumber="170" value="2012-07-13" valueMax="2012-09-10" valueMin="2012-07-13">7-9-12-13-10</date>
), left posterior (
<date id="4E7D5FA32567FF8AFC30FEDBFB8CFEF8" box="[995,1132,260,285]" pageId="19" pageNumber="170" value="2011-06-13" valueMax="2011-08-09" valueMin="2011-06-13">6-8-11-13-9</date>
); right anterior (
<date id="4E7D5FA32567FF8AFAF0FEDBFF54FEA5" pageId="19" pageNumber="170" value="2012-06-11" valueMax="2012-09-11" valueMin="2012-06-11">6-9-12-11- 11</date>
), left anterior (
<date id="4E7D5FA32567FF8AFEB0FEF8FDE8FEA5" box="[355,520,295,320]" pageId="19" pageNumber="170" value="2013-11-06" valueMax="2013-12-10" valueMin="2013-11-06">6-10-13-11-12</date>
), fourth finger of the extremities longer than others; digital toepads longer than wide on all fingers.
</paragraph>
<paragraph id="3A7C79632567FF8AFF14FEAFFC94FE35" blockId="19.[151,1436,151,861]" pageId="19" pageNumber="170">Measurements in mm: snout-vent length 70.9; axilla-groin 35.8; head length 16.5; head width 13.2; snout length (to eye) 10.05; eye diameter 4.1; auricular opening (maximum) 1.6; length of fourth finger 5.5; length of fourth toe 5.7; width between eye supercilliaries 9; internaral 2.1.</paragraph>
<paragraph id="3A7C79632567FF8AFF14FE03FEAAFDBA" blockId="19.[151,1436,151,861]" pageId="19" pageNumber="170">
Coloration of
<typeStatus id="E578C7C12567FF8AFEA1FE03FE34FE10" box="[370,468,476,501]" pageId="19" pageNumber="170" type="holotype">holotype</typeStatus>
in ethanol (
<figureCitation id="A2F865E62567FF8AFDBCFE03FD5AFE11" box="[623,698,476,501]" captionStart="FIGURE 8" captionStartId="16.[151,250,1924,1946]" captionTargetBox="[161,1384,1304,1875]" captionTargetId="figure@16.[151,1436,1219,1902]" captionTargetPageId="16" captionText="FIGURE 8. Scatter plot of 19 bioclimatic variables to four species: P. lanei (green, group 1), P. rupinus (orange, group 2), P. benedettii (Black, group 3) and P. kropotkini (Red dot, group 4). The first two components are shown with the percentage of variance on the axes." httpUri="https://zenodo.org/record/1216709/files/figure.png" pageId="19" pageNumber="170">Fig. 8</figureCitation>
). The
<typeStatus id="E578C7C12567FF8AFCC3FE03FC92FE10" box="[784,882,476,501]" pageId="19" pageNumber="170" type="holotype">holotype</typeStatus>
exhibits relatively little coloration, the dorsal background color varies from brown to creamy yellow, with dark brown faint irregular spots (less evident compared to live coloration), the ventral surface varies yellowish and cream; half of the ventral color on the tail to the tip is cream.
</paragraph>
<paragraph id="3A7C79632567FF8AFF14FDB2FE78FCB8" blockId="19.[151,1436,151,861]" pageId="19" pageNumber="170">
Pattern and color in life (
<figureCitation id="A2F865E62567FF8AFE22FDB3FDA8FD61" box="[497,584,620,645]" captionStart="FIGURE 10" captionStartId="19.[151,250,1773,1795]" captionTargetBox="[151,1436,895,1752]" captionTargetId="figure@19.[151,1436,895,1752]" captionTargetPageId="19" captionText="FIGURE 10. Phyllodactylus benedettii sp. nov. from the type locality (Chamela-Cuixmala Biosphere Reserve, Jalisco) in life (Photo by Israel Solano Zavaleta)." httpUri="https://zenodo.org/record/1216713/files/figure.png" pageId="19" pageNumber="170">Fig. 10</figureCitation>
). Dorsally presents a clear background, a mixture of cream-pink, on this background, patterns of irregular patches dark brown color and some completely black spots are present, some of these spots are slightly surrounded by yellowish scales; on the middle dorsal part a longitudinal line of the background color runs from head to tail; scales around the eyes and some supralabials are slightly colored light yellow; ventrally presents a white-cream coloration in combination with light yellow, more evident towards the ventral region; the pupils are black on a bronze iris. Dorsal coloration of the tail is white-cream with brown stripes and some black points.
</paragraph>
<caption id="6EBC29EB2567FF8AFF44F932FE12F8C7" httpUri="https://zenodo.org/record/1216713/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,250,1773,1795]" targetBox="[151,1436,895,1752]" targetPageId="19">
<paragraph id="3A7C79632567FF8AFF44F932FE12F8C7" blockId="19.[151,1436,1773,1826]" pageId="19" pageNumber="170">
<emphasis id="08B7A5712567FF8AFF44F932FEC0F8E6" bold="true" box="[151,288,1773,1795]" pageId="19" pageNumber="170">FIGURE 10.</emphasis>
<taxonomicName id="FDC302E02567FF8AFEF4F932FDC8F8E6" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[295,552,1773,1795]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712567FF8AFEF4F932FDC8F8E6" box="[295,552,1773,1795]" italics="true" pageId="19" pageNumber="170">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<emphasis id="08B7A5712567FF8AFDFCF931FD60F8E6" bold="true" box="[559,640,1774,1795]" pageId="19" pageNumber="170">
<taxonomicNameLabel id="1384180A2567FF8AFDFCF931FD60F8E6" box="[559,640,1774,1795]" pageId="19" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the type locality (Chamela-Cuixmala Biosphere Reserve, Jalisco) in life (Photo by Israel Solano Zavaleta).
</paragraph>
</caption>
<paragraph id="3A7C79632567FF8AFF14F896FC95F863" blockId="19.[151,1436,1865,2034]" pageId="19" pageNumber="170">
<emphasis id="08B7A5712567FF8AFF14F896FEDCF887" bold="true" box="[199,316,1865,1890]" pageId="19" pageNumber="170">Variation.</emphasis>
All meristic and morphometric characters are presented with mean values, standard deviation and range of meristic (
<tableCitation id="77414CD82567FF8AFEB0F8B2FE59F863" box="[355,441,1901,1926]" captionStart="TABLE2" captionStartId="7.[125,204,659,680]" captionText="TABLE2. Mean, stanđarđ đeviationanđ rangeof meristic variables examineđ forspecies of the P.tuberculosus group." pageId="19" pageNumber="170">Table 2</tableCitation>
) and morphometric variables (
<tableCitation id="77414CD82567FF8AFCC3F8B2FC87F863" box="[784,871,1901,1926]" captionStart="TABLE 7" captionStartId="12.[120,199,575,596]" captionText="TABLE 7.Mean,stanđarđ đeviation anđrange ofmorphometric variables examineđforsix speciesofPhyllodactylus." pageId="19" pageNumber="170">Table 7</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82567FF8DFF14F84EFBD6FC65" lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170" type="distribution">
<paragraph id="3A7C79632567FF8DFF14F84EFBD6FC65" blockId="19.[151,1436,1865,2034]" lastBlockId="20.[151,1437,151,896]" lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170">
<emphasis id="08B7A5712567FF8AFF14F84EFE18F84F" bold="true" box="[199,504,1937,1962]" pageId="19" pageNumber="170">Distribution and habitat.</emphasis>
<taxonomicName id="FDC302E02567FF8AFDD0F84EFCC2F84F" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[515,802,1937,1962]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712567FF8AFDD0F84EFCC2F84F" box="[515,802,1937,1962]" italics="true" pageId="19" pageNumber="170">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<emphasis id="08B7A5712567FF8AFCFFF84EFC6AF84F" bold="true" box="[812,906,1937,1962]" pageId="19" pageNumber="170">
<taxonomicNameLabel id="1384180A2567FF8AFCFFF84EFC6AF84F" box="[812,906,1937,1962]" pageId="19" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is an endemic species to the state of
<collectingRegion id="F807B7812567FF8AFA9AF84EFA78F84F" box="[1353,1432,1937,1962]" country="Mexico" name="Jalisco" pageId="19" pageNumber="170">Jalisco</collectingRegion>
, restricted to the Chamela-Cuixmala Biosphere Reserve. According to the INEGI digital map of elevation (resolution of
<quantity id="FD3BD4862567FF8AFEE5F806FE93F817" box="[310,371,2009,2034]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="170" unit="m" value="15.0">15 m</quantity>
), the
<typeStatus id="E578C7C12567FF8AFE62F805FE01F817" box="[433,481,2010,2034]" pageId="19" pageNumber="170">type</typeStatus>
locality is located at
<quantity id="FD3BD4862567FF8AFD00F805FCDDF817" box="[723,829,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="170" unit="m" value="100.0">100 masl</quantity>
, located in the biogeographic province of the Western Pacific surrounded by deciduous tropical forests (
<figureCitation id="A2F865E62560FF8DFD14FF48FCFAFF55" box="[711,794,151,176]" captionStart="FIGURE 11" captionStartId="20.[151,250,1440,1462]" captionTargetBox="[151,1436,949,1419]" captionTargetId="figure@20.[151,1436,949,1419]" captionTargetPageId="20" captionText="FIGURE 11. Deciduos tropical forests from the type locality (Chamela-Cuixmala Biosphere Reserve, Jalisco) habitat of Phyllodactylus benedettii sp. nov. (Photos by Tonatiuh Ramírez Reyes)." httpUri="https://zenodo.org/record/1216715/files/figure.png" pageId="20" pageNumber="171">Fig. 11</figureCitation>
). The microhabitats reported for this species are diverse: soil, trees, shrubs and in habitats modified by man as houses or bridges or some other constructions (meteorological stations). They are particularly abundant in rocks and crevices and in people's houses or buildings where it feeds on insects and other invertebrates (
<bibRefCitation id="5E5204922560FF8DFD1BFEDBFC3FFEF9" box="[712,991,260,285]" pageId="20" pageNumber="171" refId="ref16620" refString="Garcia, A. &amp; Ceballos, G. (1994) Guia de campo de los reptiles y anfibios de la costa de Jalisco, Mexico. Fundacion Ecologica de Cuixmala, A. C., Instituto de Biologia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 176 pp." type="book">García &amp; Ceballos 1994</bibRefCitation>
;
<bibRefCitation id="5E5204922560FF8DFC38FEDBFAA9FEF9" box="[1003,1353,260,284]" pageId="20" pageNumber="171" refId="ref17263" refString="Ponce-Campos, P. &amp; Garcia-Aguayo, A. (2007) Phyllodactylus lanei. The IUCN Red List of Threatened Species. Version 2014.2. Available from: http: // www. iucnredlist. org (accessed 27 October 2014)" type="journal article">Ponce-Campos &amp; García 2007</bibRefCitation>
;
<bibRefCitation id="5E5204922560FF8DFA87FEDBFEA0FEA5" pageId="20" pageNumber="171" refId="ref18396" refString="Vitt, L. J. &amp; Caldwell, J. P. (2014) Herpetology: An Introductory Biology of Amphibians and Reptiles. 4 th Edition. Academic Press, San Diego, 757 pp. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 386919 - 7.00002 - 2" type="book">Vitt &amp; Caldwell 2014</bibRefCitation>
; pers. obs.). Males of
<taxonomicName id="FDC302E02560FF8DFDEDFEF6FD29FEA5" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[574,713,295,320]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712560FF8DFDEDFEF6FD29FEA5" box="[574,713,295,320]" italics="true" pageId="20" pageNumber="171">P. benedettii</emphasis>
</taxonomicName>
have an extended period of reproduction (AugustMarch) with two peaks, one in rainy season (AugustOctober) and another one in dry season (NovemberMarch). Reproductive behavior (courtship and mating) begins in August which coincides with the beginning of rainy season. The females produce clutches of two eggs, with up to three clutches during the breeding season, with a peak of egg production from December to March coinciding with the dry season (
<bibRefCitation id="5E5204922560FF8DFCF8FE68FB64FE35" box="[811,1156,439,464]" pageId="20" pageNumber="171" refId="ref17554" refString="Ramirez-Sandoval, E., Ramirez-Bautista, A. &amp; Vitt, J. L. (2006) Reproduction in the lizard Phyllodactylus lanei (Squamata: Gekkonidae) from the Pacific Coast of Mexico. Copeia, 1, 1 - 9. https: // doi. org / 10.1643 / 0045 - 8511 (2006) 006 [0001: RITLPL] 2.0. CO; 2" type="journal article">
Ramírez-Sandoval
<emphasis id="08B7A5712560FF8DFBD5FE66FBDFFE35" box="[1030,1087,439,464]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
2006
</bibRefCitation>
). This species presents a distinct karyotype from
<taxonomicName id="FDC302E02560FF8DFE67FE02FDE9FE10" authorityName="Smith" authorityYear="1935" box="[436,521,476,501]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712560FF8DFE67FE02FDE9FE10" box="[436,521,476,501]" italics="true" pageId="20" pageNumber="171">P. lanei</emphasis>
</taxonomicName>
presumably due to Robertsonian fusions/fissions with 19 pairs of telocentric chromosomes (2n = 38, FN = 38) (
<bibRefCitation id="5E5204922560FF8DFDF0FE20FCEEFDFD" box="[547,782,511,536]" pageId="20" pageNumber="171" refId="ref16092" refString="Castiglia, R, Aguayo, A. G., Bezerra, A. M. R., Flores-Villela, O. &amp; Gournung, E. (2009) Karyotypic diversification due to Robertsonian rearrangements in Phyllodactylus lanei Smith, 1935 (Squamata, Gekkonidae) from Mexico. Atti della Accademia Nazionale dei Lincei, Rendiconti Lincei, Scienze Fisiche e Naturali, 20, 77 - 82. https: // doi. org / 10.1007 / s 12210 - 009 - 0005 - 4" type="journal article">
Castiglia
<emphasis id="08B7A5712560FF8DFD43FDDEFD2AFDFD" box="[656,714,511,536]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
2009
</bibRefCitation>
). They are lizards of nocturnal habits and feed on insects, although to date there have been no studies on the composition of their diet (
<bibRefCitation id="5E5204922560FF8DFBF0FDFBFAA2FDD9" box="[1059,1346,548,573]" pageId="20" pageNumber="171" refId="ref16620" refString="Garcia, A. &amp; Ceballos, G. (1994) Guia de campo de los reptiles y anfibios de la costa de Jalisco, Mexico. Fundacion Ecologica de Cuixmala, A. C., Instituto de Biologia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 176 pp." type="book">García &amp; Ceballos 1994</bibRefCitation>
;
<bibRefCitation id="5E5204922560FF8DFA81FDFBFEA7FD85" pageId="20" pageNumber="171" refId="ref18396" refString="Vitt, L. J. &amp; Caldwell, J. P. (2014) Herpetology: An Introductory Biology of Amphibians and Reptiles. 4 th Edition. Academic Press, San Diego, 757 pp. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 386919 - 7.00002 - 2" type="book">Vitt &amp; Caldwell 2014</bibRefCitation>
). Representative species of tropical dry forest of Chamela-Cuixmala Biosphere Reserve are:
<emphasis id="08B7A5712560FF8DFF44FDB3FC79FD60" box="[151,921,620,645]" italics="true" pageId="20" pageNumber="171">Sciadodendrom excelsum, Brosimum alicastrum, Orbignya cohune</emphasis>
and
<emphasis id="08B7A5712560FF8DFC04FDB3FB3EFD60" box="[983,1246,620,645]" italics="true" pageId="20" pageNumber="171">Tabebuia donell-smithi</emphasis>
(
<bibRefCitation id="5E5204922560FF8DFB22FDB3FF33FD4D" pageId="20" pageNumber="171" refId="ref16295" refString="Ceballos, G., Szekely, A., Garcia, A., Rodriguez, P. &amp; Noguera, F. (1999) Programa de Manejo de la Reserva de la Biosfera Chamela-Cuixmala. Instituto Nacional de Ecologia, SEMARNAP, Mexico, D. F., 133 pp." type="book">
Ceballos
<emphasis id="08B7A5712560FF8DFAB3FDB2FA7CFD60" box="[1376,1436,620,645]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
1999
</bibRefCitation>
). The deciduous tropical forests is near the mouth of the Cuixmala River and is characterized by the abundance of water in the rainy season, the climate is tropical (warm-humid), has an annual average temperature of 25 °C and the annual rainfall varies from
<quantity id="FD3BD4862560FF8DFDAFFD07FCC1FD0A" box="[636,801,728,752]" metricMagnitude="-1" metricUnit="m" metricValue="8.74" metricValueMax="10.0" metricValueMin="7.4799999999999995" pageId="20" pageNumber="171" unit="mm" value="874.0" valueMax="1000.0" valueMin="748.0">7481000 mm</quantity>
(
<bibRefCitation id="5E5204922560FF8DFCE2FD07FBADFD15" box="[817,1101,727,752]" pageId="20" pageNumber="171" refId="ref16620" refString="Garcia, A. &amp; Ceballos, G. (1994) Guia de campo de los reptiles y anfibios de la costa de Jalisco, Mexico. Fundacion Ecologica de Cuixmala, A. C., Instituto de Biologia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 176 pp." type="book">García &amp; Ceballos 1994</bibRefCitation>
;
<bibRefCitation id="5E5204922560FF8DFB88FD08FAA6FD15" box="[1115,1350,727,752]" pageId="20" pageNumber="171" refId="ref16295" refString="Ceballos, G., Szekely, A., Garcia, A., Rodriguez, P. &amp; Noguera, F. (1999) Programa de Manejo de la Reserva de la Biosfera Chamela-Cuixmala. Instituto Nacional de Ecologia, SEMARNAP, Mexico, D. F., 133 pp." type="book">
Ceballos
<emphasis id="08B7A5712560FF8DFB15FD06FAE1FD15" box="[1222,1281,727,752]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
1999
</bibRefCitation>
). Other species of reptiles with nocturnal activity that inhabit the deciduous tropical forests are:
<emphasis id="08B7A5712560FF8DFB45FD23FE7BFCB8" italics="true" pageId="20" pageNumber="171">
<taxonomicName id="FDC302E02560FF8DFB45FD23FF24FCDD" authority=", Boa" authorityName="Boa" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="frenatus">Hemidactylus frenatus, Boa</taxonomicName>
constrictor, Lampropeltis triangulum, Leptodeira maculata, Pseudoficimia frontalis, Senticolis triaspis, Tropidodipsas philippii
</emphasis>
(
<taxonomicName id="FDC302E02560FF8DFE78FC9BFDA8FCB8" authorityName="Jan" authorityYear="1863" box="[427,584,836,861]" class="Reptilia" family="Colubridae" genus="Sibon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="philippi">
<emphasis id="08B7A5712560FF8DFE78FC9BFDA8FCB8" box="[427,584,836,861]" italics="true" pageId="20" pageNumber="171">Sibon philippi</emphasis>
</taxonomicName>
),
<emphasis id="08B7A5712560FF8DFD8CFC9BFE18FC65" italics="true" pageId="20" pageNumber="171">
<taxonomicName id="FDC302E02560FF8DFD8CFC9BFC53FCB8" authority=", Thamnophis" authorityName="Thamnophis" box="[607,947,836,861]" class="Reptilia" family="Colubridae" genus="Tantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="bocourti">Tantilla bocourti, Thamnophis</taxonomicName>
valida,
<taxonomicName id="FDC302E02560FF8DFBDFFC9BFA7CFCB9" authority=", Micrurus" authorityName="Micrurus" box="[1036,1436,836,861]" class="Reptilia" family="Colubridae" genus="Trimorphodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="biscutatus">Trimorphodon biscutatus, Micrurus</taxonomicName>
distans, Agkistrodon bilineatus,
</emphasis>
and
<taxonomicName id="FDC302E02560FF8DFDFCFCB8FBD1FC65" authority="(Garcia &amp; Ceballos 1994)" authorityName="Garcia &amp; Ceballos" authorityYear="1994" baseAuthorityName="Garcia &amp; Ceballos" baseAuthorityYear="1994" box="[559,1073,871,896]" class="Reptilia" family="Viperidae" genus="Crotalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="basiliscus">
<emphasis id="08B7A5712560FF8DFDFCFCB8FCE5FC65" box="[559,773,871,896]" italics="true" pageId="20" pageNumber="171">Crotalus basiliscus</emphasis>
(
<bibRefCitation id="5E5204922560FF8DFCC7FCB7FBCAFC65" box="[788,1066,871,896]" pageId="20" pageNumber="171" refId="ref16620" refString="Garcia, A. &amp; Ceballos, G. (1994) Guia de campo de los reptiles y anfibios de la costa de Jalisco, Mexico. Fundacion Ecologica de Cuixmala, A. C., Instituto de Biologia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 176 pp." type="book">García &amp; Ceballos 1994</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="6EBC29EB2560FF8DFF44FA7FFC92FA30" httpUri="https://zenodo.org/record/1216715/files/figure.png" pageId="20" pageNumber="171" startId="20.[151,250,1440,1462]" targetBox="[151,1436,949,1419]" targetPageId="20">
<paragraph id="3A7C79632560FF8DFF44FA7FFC92FA30" blockId="20.[151,1436,1440,1493]" pageId="20" pageNumber="171">
<emphasis id="08B7A5712560FF8DFF44FA7FFEC3FA50" bold="true" box="[151,291,1440,1462]" pageId="20" pageNumber="171">FIGURE 11.</emphasis>
Deciduos tropical forests from the type locality (Chamela-Cuixmala Biosphere Reserve, Jalisco) habitat of
<taxonomicName id="FDC302E02560FF8DFF44FA60FE78FA30" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[151,408,1471,1493]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="benedettii" status="sp. nov.">
<emphasis id="08B7A5712560FF8DFF44FA60FE78FA30" box="[151,408,1471,1493]" italics="true" pageId="20" pageNumber="171">Phyllodactylus benedettii</emphasis>
</taxonomicName>
<emphasis id="08B7A5712560FF8DFE4DFA60FE0EFA31" bold="true" box="[414,494,1471,1492]" pageId="20" pageNumber="171">
<taxonomicNameLabel id="1384180A2560FF8DFE4DFA60FE0EFA31" box="[414,494,1471,1492]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Photos by Tonatiuh Ramírez Reyes).
</paragraph>
</caption>
<subSubSection id="72D92AE82560FF8DFF14FA23FDD2F80F" pageId="20" pageNumber="171" type="discussion">
<paragraph id="3A7C79632560FF8DFF14FA23FD49F820" blockId="20.[151,1437,1532,2026]" pageId="20" pageNumber="171">
<emphasis id="08B7A5712560FF8DFF14FA23FD50F9F0" bold="true" box="[199,688,1532,1557]" pageId="20" pageNumber="171">Comments on conservation and threats.</emphasis>
This species inhabits the Chamela-Cuixmala Biosphere Reserve decreed in 1993. There is a minor threat posed by the localities bordering the reserve, and according to the observations made by the first author, apparently there is a large population of
<taxonomicName id="FDC302E02560FF8DFBD1F99AFB6AF9B8" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1026,1162,1604,1629]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712560FF8DFBD1F99AFB6AF9B8" box="[1026,1162,1604,1629]" italics="true" pageId="20" pageNumber="171">P. benedettii</emphasis>
</taxonomicName>
geckos inside Chamela- Cuixmala Biology Station (UNAM). There are two main threats: the false beliefs of some people who consider this species as poisonous (local inhabitants), and the possible negative effects due to the presence of invasive gecko
<taxonomicName id="FDC302E02560FF8DFF44F96FFE73F92C" box="[151,403,1712,1737]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="08B7A5712560FF8DFF44F96FFE73F92C" box="[151,403,1712,1737]" italics="true" pageId="20" pageNumber="171">Hemidactylus frenatus</emphasis>
</taxonomicName>
, a species with a very large population that may exploit the same ecological niche. To date there is no specific study on niche overlap between
<taxonomicName id="FDC302E02560FF8DFD08F90AFCB8F908" box="[731,856,1749,1773]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="08B7A5712560FF8DFD08F90AFCB8F908" box="[731,856,1749,1773]" italics="true" pageId="20" pageNumber="171">H. frenatus</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02560FF8DFC5DF90BFBD4F908" authorityName=", Dixon" authorityYear="1964" box="[910,1076,1748,1773]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="genus">
<emphasis id="08B7A5712560FF8DFC5DF90BFBD4F908" box="[910,1076,1748,1773]" italics="true" pageId="20" pageNumber="171">Phyllodactylus</emphasis>
</taxonomicName>
species in
<collectingCountry id="42D439F32560FF8DFB63F90BFAE8F908" box="[1200,1288,1748,1773]" name="Mexico" pageId="20" pageNumber="171">
<collectingRegion id="F807B7812560FF8DFB63F90BFAE8F908" box="[1200,1288,1748,1773]" country="Mexico" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingRegion>
</collectingCountry>
, however, its potential danger to local biodiversity has been documented, including interspecific negative interactions with native geckos from different areas of the world; one example of this is the displacement and reduction of six gecko species of the genus
<taxonomicName id="FDC302E02560FF8DFE57F89EFE32F8BD" authorityName="Kluge" authorityYear="1983" box="[388,466,1857,1880]" class="Reptilia" family="Gekkonidae" genus="Nactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="genus">
<emphasis id="08B7A5712560FF8DFE57F89EFE32F8BD" box="[388,466,1857,1880]" italics="true" pageId="20" pageNumber="171">Nactus</emphasis>
</taxonomicName>
on the Mascarene Islands and
<taxonomicName id="FDC302E02560FF8DFCE7F89FFBA5F8BC" baseAuthorityName="Duméril &amp; Bibron" baseAuthorityYear="1836" box="[820,1093,1856,1881]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="08B7A5712560FF8DFCE7F89FFBA5F8BC" box="[820,1093,1856,1881]" italics="true" pageId="20" pageNumber="171">Lepidodactylus lugubris</emphasis>
</taxonomicName>
throughout Pacific islands. In these cases
<taxonomicName id="FDC302E02560FF8DFECDF8BAFE7DF898" box="[286,413,1893,1917]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="08B7A5712560FF8DFECDF8BAFE7DF898" box="[286,413,1893,1917]" italics="true" pageId="20" pageNumber="171">H. frenatus</emphasis>
</taxonomicName>
has been observed stalking, lunging towards and biting at other geckos (Global Invasive Species Database 2015). Due to the above, and as a potential risk to local biodiversity in
<collectingCountry id="42D439F32560FF8DFBACF857FB38F844" box="[1151,1240,1928,1953]" name="Mexico" pageId="20" pageNumber="171">
<collectingRegion id="F807B7812560FF8DFBACF857FB38F844" box="[1151,1240,1928,1953]" country="Mexico" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingRegion>
</collectingCountry>
, it is classified as a high risk invasive species (
<bibRefCitation id="5E5204922560FF8DFE0BF872FD7DF820" box="[472,669,1964,1989]" pageId="20" pageNumber="171" refId="ref16350" refString="CONABIO (2016) Sistema de informacion sobre especies invasoras en Mexico. Comision Nacional para el Conocimiento y Uso de la Biodiversidad. Available from: http: // www. biodiversidad. gob. mx / invasoras (accessed 13 March 2018)" type="journal article">CONABIO 2016</bibRefCitation>
).
</paragraph>
<paragraph id="3A7C79632560FF8DFF44F80FFDD2F80F" blockId="20.[151,1437,1532,2026]" box="[151,562,2000,2026]" pageId="20" pageNumber="171">
<emphasis id="08B7A5712560FF8DFF44F80FFDD2F80F" bold="true" box="[151,562,2000,2026]" pageId="20" pageNumber="171">
<taxonomicName id="FDC302E02560FF8DFF44F80FFE2DF80F" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[151,461,2000,2026]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="174" phylum="Chordata" rank="species" species="kropotkini" status="sp. nov.">
<emphasis id="08B7A5712560FF8DFF44F80FFE2DF80F" bold="true" box="[151,461,2000,2026]" italics="true" pageId="20" pageNumber="171">Phyllodactylus kroPotkini</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82561FF8CFF44FF63FC62FE6D" pageId="21" pageNumber="172" type="materials_examined">
<paragraph id="3A7C79632561FF8CFF44FF63FEB9FEF8" blockId="21.[151,1437,188,2013]" pageId="21" pageNumber="172">
<materialsCitation id="8AAB733E2561FF8CFF44FF63FEB9FEF8" ID-GBIF-Occurrence="1837804194" collectingDate="2014-03-15" collectionCode="MZFC" county="Adult" latitude="18.29" location="Huerta Vieja" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" pageId="21" pageNumber="172" specimenCode="MZFC 28736" specimenCount="1" stateProvince="Guerrero" typeStatus="holotype">
<emphasis id="08B7A5712561FF8CFF44FF63FEEBFF30" bold="true" box="[151,267,188,213]" pageId="21" pageNumber="172">
<typeStatus id="E578C7C12561FF8CFF44FF63FEE6FF30" box="[151,262,188,213]" pageId="21" pageNumber="172" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="D31D01EF2561FF8CFEC9FF63FEBAFF30" box="[282,346,188,213]" pageId="21" pageNumber="172">Adult</collectingCounty>
male (
<specimenCode id="6A65D1182561FF8CFE66FF63FDBAFF30" box="[437,602,188,213]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="21" pageNumber="172">MZFC 28736</specimenCode>
) collected in
<collectingMunicipality id="DA18E3192561FF8CFCDAFF62FC2CFF30" box="[777,972,188,213]" pageId="21" pageNumber="172">Nueva Filadelfia</collectingMunicipality>
(
<location id="3F1C2FB82561FF8CFC31FF63FB9DFF30" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A:3F1C2FB82561FF8CFC31FF63FB9DFF30" box="[994,1149,188,213]" county="Adult" latitude="18.29" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" name="Huerta Vieja" pageId="21" pageNumber="172" stateProvince="Guerrero">Huerta Vieja</location>
) in the municipality of
<location id="3F1C2FB82561FF8CFF44FF00FEF6FF1D" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A:3F1C2FB82561FF8CFF44FF00FEF6FF1D" box="[151,278,223,248]" county="Adult" latitude="18.29" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" name="Tlapehuala" pageId="21" pageNumber="172" stateProvince="Guerrero">Tlapehuala</location>
,
<collectingRegion id="F807B7812561FF8CFEF2FF3FFE66FF1D" box="[289,390,224,248]" country="Mexico" name="Guerrero" pageId="21" pageNumber="172">Guerrero</collectingRegion>
(
<geoCoordinate id="5FF71FA42561FF8CFE46FF3FFE13FF12" box="[405,499,224,248]" direction="north" orientation="latitude" pageId="21" pageNumber="172" precision="555" value="18.29">18.29 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42561FF8CFDD5FF3FFD96FF12" box="[518,630,224,248]" direction="west" orientation="longitude" pageId="21" pageNumber="172" precision="555" value="-100.49">100.49 W</geoCoordinate>
WGS84), collected on
<date id="4E7D5FA32561FF8CFCAEFF00FBCFFF1D" box="[893,1071,223,248]" pageId="21" pageNumber="172" value="2014-03-15">
<collectingDate id="5E39A64B2561FF8CFCAEFF00FBCFFF1D" box="[893,1071,223,248]" pageId="21" pageNumber="172" value="2014-03-15">March 15, 2014</collectingDate>
</date>
by Tonatiuh Ramírez Reyes and Joel Rosas Avila.
</materialsCitation>
</paragraph>
<paragraph id="3A7C79632561FF8CFF14FEF8FC62FE6D" blockId="21.[151,1437,188,2013]" pageId="21" pageNumber="172">
<materialsCitation id="8AAB733E2561FF8CFF14FEF8FC62FE6D" ID-GBIF-Occurrence="1837804191" collectingDate="2014-03-16" collectionCode="MZFC" county="Four" latitude="18.29" location="Huerta Vieja" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" pageId="21" pageNumber="172" specimenCode="MZFC 28735, MZFC 28737, MZFC 28738, MZFC 28739" specimenCount="2" stateProvince="Guerrero" typeStatus="paratype">
<emphasis id="08B7A5712561FF8CFF14FEF8FEDFFEA5" bold="true" box="[199,319,295,320]" pageId="21" pageNumber="172">
<typeStatus id="E578C7C12561FF8CFF14FEF8FEDFFEA5" box="[199,319,295,320]" pageId="21" pageNumber="172" type="paratype">Paratypes</typeStatus>
</emphasis>
(4).
<collectingCounty id="D31D01EF2561FF8CFEA4FEF7FE4CFEDA" box="[375,428,296,319]" pageId="21" pageNumber="172">Four</collectingCounty>
male adults (
<specimenCode id="6A65D1182561FF8CFD96FEF7FD04FEA5" box="[581,740,295,320]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="21" pageNumber="172">MZFC 28735</specimenCode>
,
<specimenCode id="6A65D1182561FF8CFD22FEF7FC6EFEA5" box="[753,910,296,320]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="21" pageNumber="172">MZFC 28737</specimenCode>
,
<specimenCode id="6A65D1182561FF8CFC49FEF7FBD8FEA5" box="[922,1080,296,320]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="21" pageNumber="172">MZFC 28738</specimenCode>
,
<specimenCode id="6A65D1182561FF8CFB97FEF7FB00FEA5" box="[1092,1248,296,320]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="21" pageNumber="172">MZFC 28739</specimenCode>
) collected at the
<typeStatus id="E578C7C12561FF8CFF44FE92FF27FE80" box="[151,199,333,357]" pageId="21" pageNumber="172">type</typeStatus>
locality,
<collectingMunicipality id="DA18E3192561FF8CFEE0FE92FE11FE80" box="[307,497,332,357]" pageId="21" pageNumber="172">Nueva Filadelfia</collectingMunicipality>
(
<location id="3F1C2FB82561FF8CFDD1FE93FD7AFE80" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A:3F1C2FB82561FF8CFDD1FE93FD7AFE80" box="[514,666,332,357]" county="Four" latitude="18.29" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" name="Huerta Vieja" pageId="21" pageNumber="172" stateProvince="Guerrero">Huerta Vieja</location>
),
<location id="3F1C2FB82561FF8CFD7CFE93FCCEFE80" LSID="urn:lsid:plazi:treatment:B26AC8752565FF81FF44FF01FC50FF4A:3F1C2FB82561FF8CFD7CFE93FCCEFE80" box="[687,814,332,357]" county="Four" latitude="18.29" longLatPrecision="765" longitude="-100.49" municipality="Nueva Filadelfia" name="Tlapehuala" pageId="21" pageNumber="172" stateProvince="Guerrero">Tlapehuala</location>
,
<collectingRegion id="F807B7812561FF8CFCE8FE93FC40FE81" box="[827,928,332,356]" country="Mexico" name="Guerrero" pageId="21" pageNumber="172">Guerrero</collectingRegion>
(
<geoCoordinate id="5FF71FA42561FF8CFC61FE93FBF3FE81" box="[946,1043,332,356]" direction="north" orientation="latitude" pageId="21" pageNumber="172" precision="555" value="18.29">18.29 N</geoCoordinate>
, -
<geoCoordinate id="5FF71FA42561FF8CFBFBFE93FB7CFE81" box="[1064,1180,332,356]" direction="west" orientation="longitude" pageId="21" pageNumber="172" precision="555" value="-100.49">100.49 W</geoCoordinate>
WGS84) collected on
<date id="4E7D5FA32561FF8CFF44FEB0FEA9FE6D" box="[151,329,367,392]" pageId="21" pageNumber="172" value="2014-03-16">
<collectingDate id="5E39A64B2561FF8CFF44FEB0FEA9FE6D" box="[151,329,367,392]" pageId="21" pageNumber="172" value="2014-03-16">March 16, 2014</collectingDate>
</date>
by Tonatiuh Ramírez Reyes and Joel Rosas Ávila.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82561FF8CFF14FE4BFC83FE10" pageId="21" pageNumber="172" type="etymology">
<paragraph id="3A7C79632561FF8CFF14FE4BFC83FE10" blockId="21.[151,1437,188,2013]" pageId="21" pageNumber="172">
<emphasis id="08B7A5712561FF8CFF14FE4BFEAEFE48" bold="true" box="[199,334,404,429]" pageId="21" pageNumber="172">Etymology.</emphasis>
The species is dedicated to the memory of the great Russian philosopher, geographer and naturalist Piotr Kropotkin, who made great scientific and theoretical contributions about mutual support and altruism in some animal populations (including human society).
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82561FF8CFF14FE20FB33F959" pageId="21" pageNumber="172" type="diagnosis">
<paragraph id="3A7C79632561FF8CFF14FE20FD18FBD1" blockId="21.[151,1437,188,2013]" pageId="21" pageNumber="172">
<emphasis id="08B7A5712561FF8CFF14FE20FEA1FDFD" bold="true" box="[199,321,511,536]" pageId="21" pageNumber="172">Diagnosis.</emphasis>
<taxonomicName id="FDC302E02561FF8CFE98FE20FD8DFDFD" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[331,621,511,536]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini" status="sp. nov.">
<emphasis id="08B7A5712561FF8CFE98FE20FD8DFDFD" box="[331,621,511,536]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
<emphasis id="08B7A5712561FF8CFDA4FE20FD34FDFD" bold="true" box="[631,724,511,536]" pageId="21" pageNumber="172">
<taxonomicNameLabel id="1384180A2561FF8CFDA4FE20FD34FDFD" box="[631,724,511,536]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a medium-sized gecko within the
<taxonomicName id="FDC302E02561FF8CFB57FDDEFB3AFDFD" authorityName="Smith" authorityYear="1935" box="[1156,1242,511,536]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFB57FDDEFB3AFDFD" box="[1156,1242,511,536]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
complex.
<bibRefCitation id="5E5204922561FF8CFA85FE20FF01FDD8" pageId="21" pageNumber="172" refId="ref16514" refString="Dixon, J. R. (1964) The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America. New Mexico State University Research Center, Scientific Bulletin, 1964, 1 - 139." type="journal article">Dixon (1964)</bibRefCitation>
established three fundamental characteristics to include species and subspecies within the
<taxonomicName id="FDC302E02561FF8CFB33FDFAFAD4FDD8" authorityName="Smith" authorityYear="1935" box="[1248,1332,548,573]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFB33FDFAFAD4FDD8" box="[1248,1332,548,573]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
complex and differentiate them from
<taxonomicName id="FDC302E02561FF8CFE0CFD96FD6CFD85" authorityName="Wiegmann" authorityYear="1834" box="[479,652,583,608]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFE0CFD96FD6CFD85" box="[479,652,583,608]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
, species of the
<taxonomicName id="FDC302E02561FF8CFC94FD96FC7BFD85" authorityName="Smith" authorityYear="1935" box="[839,923,583,608]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFC94FD96FC7BFD85" box="[839,923,583,608]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
complex have a low number of interorbital scales (1219), white venter and low number of scales across the snout between the third labials (does not specify range of values).
<taxonomicName id="FDC302E02561FF8CFE8FFD50FD9DFD4D" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[348,637,655,680]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFE8FFD50FD9DFD4D" box="[348,637,655,680]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
has 14.6 interorbital scales on average (1415), white venter and 20.2 third labialsnout scales on average (1921).
<taxonomicName id="FDC302E02561FF8CFD47FD6BFC53FD28" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[660,947,692,717]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFD47FD6BFC53FD28" box="[660,947,692,717]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
presents a maximum recorded SVL of
<quantity id="FD3BD4862561FF8CFAB8FD6BFF23FD0A" metricMagnitude="-2" metricUnit="m" metricValue="6.2299999999999995" pageId="21" pageNumber="172" unit="mm" value="62.3">62.3 mm</quantity>
that clearly differs from the nearest species
<taxonomicName id="FDC302E02561FF8CFD62FD06FCE4FD15" authorityName="Smith" authorityYear="1935" box="[689,772,727,752]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFD62FD06FCE4FD15" box="[689,772,727,752]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(max.
<quantity id="FD3BD4862561FF8CFC81FD07FC42FD15" box="[850,930,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="21" pageNumber="172" unit="mm" value="71.0">71 mm</quantity>
) and it differs from other species:
<taxonomicName id="FDC302E02561FF8CFAF0FD06FA7CFD15" box="[1315,1436,727,752]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFAF0FD06FA7CFD15" box="[1315,1436,727,752]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(max. 57.5),
<taxonomicName id="FDC302E02561FF8CFEFBFD22FE72FCF0" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[296,402,764,789]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFEFBFD22FE72FCF0" box="[296,402,764,789]" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(max.
<quantity id="FD3BD4862561FF8CFE31FD23FDABFCF1" box="[482,587,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="7.33" pageId="21" pageNumber="172" unit="mm" value="73.3">73.3 mm</quantity>
),
<taxonomicName id="FDC302E02561FF8CFD8DFD22FD2FFCF1" box="[606,719,765,788]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFD8DFD22FD2FFCF1" box="[606,719,765,788]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(max. 69.4),
<taxonomicName id="FDC302E02561FF8CFCBAFD22FC13FCF0" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[873,1011,764,789]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFCBAFD22FC13FCF0" box="[873,1011,764,789]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(max. 74.2) and
<taxonomicName id="FDC302E02561FF8CFB66FD22FAFBFCF0" authorityName="Taylor" authorityYear="1942" box="[1205,1307,764,789]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="bordai">
<emphasis id="08B7A5712561FF8CFB66FD22FAFBFCF0" box="[1205,1307,764,789]" italics="true" pageId="21" pageNumber="172">P. bordai</emphasis>
</taxonomicName>
(max.
<quantity id="FD3BD4862561FF8CFAB8FD23FF27FCDD" metricMagnitude="-2" metricUnit="m" metricValue="5.89" pageId="21" pageNumber="172" unit="mm" value="58.9">58.9 mm</quantity>
).
<taxonomicName id="FDC302E02561FF8CFF0AFCC0FE18FCDD" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[217,504,799,824]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini" status="sp. nov.">
<emphasis id="08B7A5712561FF8CFF0AFCC0FE18FCDD" box="[217,504,799,824]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
<emphasis id="08B7A5712561FF8CFE2CFCC0FDB8FCDD" bold="true" box="[511,600,799,824]" pageId="21" pageNumber="172">
<taxonomicNameLabel id="1384180A2561FF8CFE2CFCC0FDB8FCDD" box="[511,600,799,824]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows the following combination of characters: 28 (2531) tubercles from head to tail, 67 (6373) longitudinal scales, 13 (1214) rows of tubercles across dorsum, 30 (2933) scales across venter, 14 (1415) interorbital scales and 20 (1921) third labialsnout scales.
<taxonomicName id="FDC302E02561FF8CFBE2FCB8FAB5FC65" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1073,1365,871,896]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFBE2FCB8FAB5FC65" box="[1073,1365,871,896]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
has a partial overlap with
<taxonomicName id="FDC302E02561FF8CFE55FC52FE3DFC40" authorityName="Smith" authorityYear="1935" box="[390,477,908,933]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFE55FC52FE3DFC40" box="[390,477,908,933]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02561FF8CFDCFFC52FD76FC40" box="[540,662,908,933]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFDCFFC52FD76FC40" box="[540,662,908,933]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
on morphometric space (
<figureCitation id="A2F865E62561FF8CFC14FC53FBF0FC41" box="[967,1040,908,933]" captionStart="FIGURE 7" captionStartId="16.[151,250,1148,1170]" captionTargetBox="[161,1373,517,1099]" captionTargetId="figure@16.[151,1436,444,1127]" captionTargetPageId="16" captionText="FIGURE 7. Scatter plot of 11 morphometric variables for six species of the P.lanei complex:P.benedettii (black, group 1), P. kropotkini (aqua blue, group 2), P. rupinus (brown, group 3), P. lanei (blue, group 4), P. lupitae (red, group 5), P. isabelae (green, group 6). The first two components are shown with the percentage of variance on the axes." httpUri="https://zenodo.org/record/1216707/files/figure.png" pageId="21" pageNumber="172">Fig. 7</figureCitation>
), however
<taxonomicName id="FDC302E02561FF8CFB46FC52FAC6FC40" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1173,1318,908,933]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFB46FC52FAC6FC40" box="[1173,1318,908,933]" italics="true" pageId="21" pageNumber="172">P. kropotkini</emphasis>
</taxonomicName>
is clearly differentiated from
<taxonomicName id="FDC302E02561FF8CFEA7FC6EFE26FC2D" authorityName="Smith" authorityYear="1935" box="[372,454,943,968]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFEA7FC6EFE26FC2D" box="[372,454,943,968]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
by maximum length (SVL) (
<figureCitation id="A2F865E62561FF8CFCC5FC70FCBCFC2D" box="[790,860,943,968]" captionStart="FIGURE6" captionStartId="15.[151,250,799,821]" captionTargetBox="[193,1398,193,768]" captionTargetId="figure@15.[193,1398,193,768]" captionTargetPageId="15" captionText="FIGURE6. Box plot of SVLfor sixspecies ofPhyllodactylus, on thehorizontal axis:P.isabelae (1), P.lupitae (2),P.lanei(3), P. rupinus (4), P. benedettii (5), P. kropotkini (6), the vertical axis indicates the length in mm. The medians are indicated by horizontal lines inside the boxes, the black dots indicate outliers." httpUri="https://zenodo.org/record/1216705/files/figure.png" pageId="21" pageNumber="172">Fig. 6</figureCitation>
); it also presents the follow measures: axilla-groin length (
<quantity id="FD3BD4862561FF8CFF3FFC0BFE84FC09" box="[236,356,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.254" pageId="21" pageNumber="172" unit="mm" value="22.54">22.54 mm</quantity>
), snout length (
<quantity id="FD3BD4862561FF8CFDC2FC0BFD9BFC09" box="[529,635,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="9.34" pageId="21" pageNumber="172" unit="mm" value="9.34">9.34 mm</quantity>
), auricular opening (
<quantity id="FD3BD4862561FF8CFCB0FC0BFC2DFC09" box="[867,973,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="21" pageNumber="172" unit="mm" value="1.78">1.78 mm</quantity>
), while
<taxonomicName id="FDC302E02561FF8CFBF6FC0AFB9AFC08" authorityName="Smith" authorityYear="1935" box="[1061,1146,980,1005]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFBF6FC0AFB9AFC08" box="[1061,1146,980,1005]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
has
<quantity id="FD3BD4862561FF8CFB7DFC0BFAF2FC09" box="[1198,1298,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="21" pageNumber="172" unit="mm" value="24.7">24.7 mm</quantity>
axilla-groin length,
<quantity id="FD3BD4862561FF8CFF3FFC27FE86FBEA" box="[236,358,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.061" pageId="21" pageNumber="172" unit="mm" value="10.61">10. 61 mm</quantity>
snout length and auricular opening (
<quantity id="FD3BD4862561FF8CFCD6FC28FC8BFBF5" box="[773,875,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="21" pageNumber="172" unit="mm" value="1.95">1.95 mm</quantity>
) and
<taxonomicName id="FDC302E02561FF8CFC78FC26FBC1FBF5" box="[939,1057,1015,1040]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFC78FC26FBC1FBF5" box="[939,1057,1015,1040]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
has
<quantity id="FD3BD4862561FF8CFB8AFC27FB5DFBEA" box="[1113,1213,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="21" pageNumber="172" unit="mm" value="18.7">18.7 mm</quantity>
axilla-groin length,
<quantity id="FD3BD4862561FF8CFF44FBC3FF1BFBD1" box="[151,251,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="8.18" pageId="21" pageNumber="172" unit="mm" value="8.18">8.18 mm</quantity>
snout length and auricular opening
<quantity id="FD3BD4862561FF8CFD5EFBC3FD14FBD1" box="[653,756,1052,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="21" pageNumber="172" unit="mm" value="1.51">1.51 mm</quantity>
.
</paragraph>
<paragraph id="3A7C79632561FF8CFF14FB9EFB33F959" blockId="21.[151,1437,188,2013]" pageId="21" pageNumber="172">
<taxonomicName id="FDC302E02561FF8CFF14FB9EFEB6FBBD" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[199,342,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFF14FB9EFEB6FBBD" box="[199,342,1087,1112]" italics="true" pageId="21" pageNumber="172">P. kropotkini</emphasis>
</taxonomicName>
has a low number of interorbital scales (1415, 14.6 avg.) similar to
<taxonomicName id="FDC302E02561FF8CFBA4FB9EFAE2FBBD" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1143,1282,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFBA4FB9EFAE2FBBD" box="[1143,1282,1087,1112]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(1316, 14.4 avg.), all other species of
<taxonomicName id="FDC302E02561FF8CFE66FBBBFDBBFB98" authorityName=", Dixon" authorityYear="1964" box="[437,603,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="genus">
<emphasis id="08B7A5712561FF8CFE66FBBBFDBBFB98" box="[437,603,1124,1149]" italics="true" pageId="21" pageNumber="172">Phyllodactylus</emphasis>
</taxonomicName>
exceed 15 interorbital scales on average,
<taxonomicName id="FDC302E02561FF8CFBFEFBBAFB45FB98" box="[1069,1189,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFBFEFBBAFB45FB98" box="[1069,1189,1124,1149]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(1416, 15.1 avg.),
<taxonomicName id="FDC302E02561FF8CFA54FBBAFF05FB45" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFA54FBBAFF05FB45" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(1420, 16.83 avg.),
<taxonomicName id="FDC302E02561FF8CFE0FFB56FDCEFB45" authorityName="Smith" authorityYear="1935" box="[476,558,1159,1184]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFE0FFB56FDCEFB45" box="[476,558,1159,1184]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(1517, 15.5 avg.),
<taxonomicName id="FDC302E02561FF8CFCC8FB56FC6DFB45" box="[795,909,1161,1184]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFCC8FB56FC6DFB45" box="[795,909,1161,1184]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(1417, 15.5 avg.),
<taxonomicName id="FDC302E02561FF8CFBA4FB56FAE3FB45" authorityName="Taylor" authorityYear="1942" box="[1143,1283,1161,1184]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712561FF8CFBA4FB56FAE3FB45" box="[1143,1283,1161,1184]" italics="true" pageId="21" pageNumber="172">P. t. magnus</emphasis>
</taxonomicName>
(2123, 22.3 avg.),
<taxonomicName id="FDC302E02561FF8CFF0DFB72FEB0FB20" authorityName="Taylor" authorityYear="1940" box="[222,336,1196,1221]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712561FF8CFF0DFB72FEB0FB20" box="[222,336,1196,1221]" italics="true" pageId="21" pageNumber="172">P. muralis</emphasis>
</taxonomicName>
(2127, 23.2 avg.),
<taxonomicName id="FDC302E02561FF8CFDE7FB72FD3EFB20" authorityName="Wiegmann" authorityYear="1834" box="[564,734,1196,1221]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFDE7FB72FD3EFB20" box="[564,734,1196,1221]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
(1622, 19.2 avg.). There are an average of 30.6 scales across the venter (2933), unlike
<taxonomicName id="FDC302E02561FF8CFE13FB0EFDD8FB0D" box="[448,568,1231,1256]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFE13FB0EFDD8FB0D" box="[448,568,1231,1256]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(2629, 27.8 avg.),
<taxonomicName id="FDC302E02561FF8CFCC8FB0EFC64FB0D" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[795,900,1231,1256]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFCC8FB0EFC64FB0D" box="[795,900,1231,1256]" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(2326, 24.6 avg.),
<taxonomicName id="FDC302E02561FF8CFBB5FB0EFB5AFB0D" authorityName="Smith" authorityYear="1935" box="[1126,1210,1231,1256]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFBB5FB0EFB5AFB0D" box="[1126,1210,1231,1256]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(2932, 30 avg.),
<taxonomicName id="FDC302E02561FF8CFA54FB0EFF0DFAE9" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFA54FB0EFF0DFAE9" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(2428, 26.5 avg.) and
<taxonomicName id="FDC302E02561FF8CFDFBFB2AFD5AFAE8" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[552,698,1268,1293]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFDFBFB2AFD5AFAE8" box="[552,698,1268,1293]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(2532, 29.1 avg.).
<taxonomicName id="FDC302E02561FF8CFC16FB2BFB0EFAE8" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[965,1262,1268,1293]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFC16FB2BFB0EFAE8" box="[965,1262,1268,1293]" italics="true" pageId="21" pageNumber="172">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
presents 67.6 longitudinal scales from the gular region to the anus on average (6373), compared to
<taxonomicName id="FDC302E02561FF8CFB5FFAC6FAE0FAD5" box="[1164,1280,1305,1328]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFB5FFAC6FAE0FAD5" box="[1164,1280,1305,1328]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(6373, 63.6 avg.),
<taxonomicName id="FDC302E02561FF8CFF0EFAE2FED0FAB0" authorityName="Smith" authorityYear="1935" box="[221,304,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFF0EFAE2FED0FAB0" box="[221,304,1340,1365]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(6074, 66 avg.),
<taxonomicName id="FDC302E02561FF8CFE2FFAE2FD65FAB0" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[508,645,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFE2FFAE2FD65FAB0" box="[508,645,1340,1365]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(6069, 62.6 avg.),
<taxonomicName id="FDC302E02561FF8CFCB5FAE2FC0DFAB1" authorityName="Taylor" authorityYear="1942" box="[870,1005,1341,1364]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712561FF8CFCB5FAE2FC0DFAB1" box="[870,1005,1341,1364]" italics="true" pageId="21" pageNumber="172">P. t. magnus</emphasis>
</taxonomicName>
(5258, 54.6 avg.),
<taxonomicName id="FDC302E02561FF8CFB1DFAE2FADEFAB0" authorityName="Taylor" authorityYear="1940" box="[1230,1342,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712561FF8CFB1DFAE2FADEFAB0" box="[1230,1342,1340,1365]" italics="true" pageId="21" pageNumber="172">P. muralis</emphasis>
</taxonomicName>
(5761, 59 avg.) and
<taxonomicName id="FDC302E02561FF8CFEFFFABEFE37FA9D" authorityName="Wiegmann" authorityYear="1834" box="[300,471,1375,1400]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFEFFFABEFE37FA9D" box="[300,471,1375,1400]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
(5164, 57.7 avg.). This species also has 28.4 tubercles from head to tail on average (2531), different from
<taxonomicName id="FDC302E02561FF8CFE72FA5AFDF1FA79" box="[417,529,1413,1436]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFE72FA5AFDF1FA79" box="[417,529,1413,1436]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(2630; 28 avg.),
<taxonomicName id="FDC302E02561FF8CFD0CFA5AFCD0FA78" authorityName="Smith" authorityYear="1935" box="[735,816,1412,1437]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFD0CFA5AFCD0FA78" box="[735,816,1412,1437]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(3234, 33.2 avg.),
<taxonomicName id="FDC302E02561FF8CFBC6FA5AFB7EFA78" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1045,1182,1412,1437]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFBC6FA5AFB7EFA78" box="[1045,1182,1412,1437]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
, (2430, 27.4 avg.),
<taxonomicName id="FDC302E02561FF8CFA54FA5AFF15FA25" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFA54FA5AFF15FA25" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(3035, 32.2 avg.),
<taxonomicName id="FDC302E02561FF8CFE0EFA76FDA8FA25" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[477,584,1447,1472]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFE0EFA76FDA8FA25" box="[477,584,1447,1472]" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(2731, 28.8 avg.),
<emphasis id="08B7A5712561FF8CFCE2FA76FC52FA25" box="[817,946,1449,1472]" italics="true" pageId="21" pageNumber="172">P. t. magnu</emphasis>
s (3540, 38 avg.),
<taxonomicName id="FDC302E02561FF8CFB42FA76FAE3FA25" authorityName="Taylor" authorityYear="1940" box="[1169,1283,1447,1472]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712561FF8CFB42FA76FAE3FA25" box="[1169,1283,1447,1472]" italics="true" pageId="21" pageNumber="172">P. muralis</emphasis>
</taxonomicName>
(3040, 33.6 avg.) and
<taxonomicName id="FDC302E02561FF8CFED5FA12FE4FFA00" authorityName="Wiegmann" authorityYear="1834" box="[262,431,1484,1509]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFED5FA12FE4FFA00" box="[262,431,1484,1509]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
(3341, 36.4 avg.) tubercles. It also presents 13.4 rows of tubercles on average across the dorsum (1214), different from
<taxonomicName id="FDC302E02561FF8CFE29FA2EFD8AF9ED" box="[506,618,1521,1544]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFE29FA2EFD8AF9ED" box="[506,618,1521,1544]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(1315, 14 avg.),
<taxonomicName id="FDC302E02561FF8CFCE5FA2EFC5FF9ED" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[822,959,1519,1544]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFCE5FA2EFC5FF9ED" box="[822,959,1519,1544]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(1314, 14.4 avg.),
<taxonomicName id="FDC302E02561FF8CFB4CFA2EFAE8F9ED" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[1183,1288,1519,1544]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFB4CFA2EFAE8F9ED" box="[1183,1288,1519,1544]" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(1415, 14.8 avg.),
<taxonomicName id="FDC302E02561FF8CFF0DF9CAFECFF9C8" authorityName="Smith" authorityYear="1935" box="[222,303,1556,1581]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFF0DF9CAFECFF9C8" box="[222,303,1556,1581]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(1416, 15.6 avg.),
<taxonomicName id="FDC302E02561FF8CFDC5F9CAFD6FF9C8" box="[534,655,1556,1581]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFDC5F9CAFD6FF9C8" box="[534,655,1556,1581]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(1518, 16.7 avg.),
<taxonomicName id="FDC302E02561FF8CFCA7F9CAFC1DF9C9" authorityName="Taylor" authorityYear="1942" box="[884,1021,1557,1580]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712561FF8CFCA7F9CAFC1DF9C9" box="[884,1021,1557,1580]" italics="true" pageId="21" pageNumber="172">P. t. magnus</emphasis>
</taxonomicName>
(1315, 14.33 avg.),
<taxonomicName id="FDC302E02561FF8CFB23F9CAFABEF9C8" authorityName="Taylor" authorityYear="1940" box="[1264,1374,1556,1581]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712561FF8CFB23F9CAFABEF9C8" box="[1264,1374,1556,1581]" italics="true" pageId="21" pageNumber="172">P. muralis</emphasis>
</taxonomicName>
(12 13, 12.2 avg.) and
<taxonomicName id="FDC302E02561FF8CFEA1F9E6FDFEF9B5" authorityName="Wiegmann" authorityYear="1834" box="[370,542,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFEA1F9E6FDFEF9B5" box="[370,542,1591,1616]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
(1217, 14 avg.). Finally
<taxonomicName id="FDC302E02561FF8CFC9CF9E6FC00F9B5" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[847,992,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712561FF8CFC9CF9E6FC00F9B5" box="[847,992,1591,1616]" italics="true" pageId="21" pageNumber="172">P. kropotkini</emphasis>
</taxonomicName>
presents 20.2 third labialsnout scales (1921), similar to
<taxonomicName id="FDC302E02561FF8CFEBCF982FE3CF991" box="[367,476,1629,1652]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712561FF8CFEBCF982FE3CF991" box="[367,476,1629,1652]" italics="true" pageId="21" pageNumber="172">P. rupinus</emphasis>
</taxonomicName>
(1822, 20.6 avg.) and it differs from the other species of
<taxonomicName id="FDC302E02561FF8CFBBDF983FAF4F990" authorityName=", Dixon" authorityYear="1964" box="[1134,1300,1628,1653]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="genus">
<emphasis id="08B7A5712561FF8CFBBDF983FAF4F990" box="[1134,1300,1628,1653]" italics="true" pageId="21" pageNumber="172">Phyllodactylus</emphasis>
</taxonomicName>
:
<taxonomicName id="FDC302E02561FF8CFAF0F982FA7CF990" box="[1315,1436,1628,1653]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712561FF8CFAF0F982FA7CF990" box="[1315,1436,1628,1653]" italics="true" pageId="21" pageNumber="172">P. isabelae</emphasis>
</taxonomicName>
(1924, 21.4 avg.),
<taxonomicName id="FDC302E02561FF8CFEA6F95EFE00F97D" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[373,480,1663,1688]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712561FF8CFEA6F95EFE00F97D" box="[373,480,1663,1688]" italics="true" pageId="21" pageNumber="172">P. lupitae</emphasis>
</taxonomicName>
(2328, 25.5 avg.),
<taxonomicName id="FDC302E02561FF8CFD16F95EFCF7F97D" authorityName="Smith" authorityYear="1935" box="[709,791,1663,1688]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712561FF8CFD16F95EFCF7F97D" box="[709,791,1663,1688]" italics="true" pageId="21" pageNumber="172">P. lanei</emphasis>
</taxonomicName>
(2023, 21.2 avg.),
<taxonomicName id="FDC302E02561FF8CFBD3F95EFB6AF97D" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[1024,1162,1663,1688]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="benedettii">
<emphasis id="08B7A5712561FF8CFBD3F95EFB6AF97D" box="[1024,1162,1663,1688]" italics="true" pageId="21" pageNumber="172">P. benedettii</emphasis>
</taxonomicName>
(2223, 22.3 avg.),
<taxonomicName id="FDC302E02561FF8CFAA2F95EFF0EF959" authorityName="Taylor" authorityYear="1942" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="subSpecies" species="tuberculosus" subSpecies="magnus">
<emphasis id="08B7A5712561FF8CFAA2F95EFF0EF959" italics="true" pageId="21" pageNumber="172">P. t. magnus</emphasis>
</taxonomicName>
(2426, 24.6 avg.),
<taxonomicName id="FDC302E02561FF8CFE02F97AFDA1F958" authorityName="Taylor" authorityYear="1940" box="[465,577,1700,1725]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="muralis">
<emphasis id="08B7A5712561FF8CFE02F97AFDA1F958" box="[465,577,1700,1725]" italics="true" pageId="21" pageNumber="172">P. muralis</emphasis>
</taxonomicName>
(2428, 25.2 avg.) and
<taxonomicName id="FDC302E02561FF8CFC9EF97AFC17F958" authorityName="Wiegmann" authorityYear="1834" box="[845,1015,1700,1725]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="08B7A5712561FF8CFC9EF97AFC17F958" box="[845,1015,1700,1725]" italics="true" pageId="21" pageNumber="172">P. tuberculosus</emphasis>
</taxonomicName>
(2126, 24.2 avg.).
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82561FF8FFF14F918FDA4F8F2" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172" type="description">
<paragraph id="3A7C79632561FF8FFF14F918FCDCFD85" blockId="21.[151,1437,188,2013]" lastBlockId="22.[151,1436,151,608]" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172">
<emphasis id="08B7A5712561FF8CFF14F918FD83F905" bold="true" box="[199,611,1735,1760]" pageId="21" pageNumber="172">
Description of
<typeStatus id="E578C7C12561FF8CFE53F918FE07F905" box="[384,487,1735,1760]" pageId="21" pageNumber="172" type="holotype">holotype</typeStatus>
(
<figureCitation id="A2F865E62561FF8CFE28F918FDB4F905" box="[507,596,1735,1760]" captionStart="FIGURE 12" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[151,1384,661,1384]" captionTargetId="figure@22.[151,1436,661,1384]" captionTargetPageId="22" captionText="FIGURE 12. Dorsal and ventral view of the holotype of Phyllodactylus kropotkini sp. nov. (MZFC 28736). The black bars represent the scale (10 mm)." httpUri="https://zenodo.org/record/1216717/files/figure.png" pageId="21" pageNumber="172">Fig. 12</figureCitation>
).
</emphasis>
Adult male, medium body proportions (non-robust), short neck, head differentiated from the body. The head scales are granular, mostly flattened and some slightly globose towards the anterior part of the eye openings. Rostral scales have an intermediate longitudinal groove that reaches the middle part of the scale; nasal orifice bordered by a simple supranasal, rostral and 2 postnasals; nasal orifice in contact with the first supralabial, this has a fold that resembles a 3rd postnasal scale; 2 supranasal scales in contact with each other, bordered by 6 postmental scales; 25 interorbital scales from the anterior ocular border; 18 scales across snout starting with the 2nd labial; 26 scales across snout starting with 3rd labial scales; 12/12 loreal scales from the nostril to eye; 12/12 supralabials; edge of 1
<taxonomicName id="FDC302E02561FF8CFDA0F81BFCEBF838" box="[627,779,1988,2013]" pageId="21" pageNumber="172" rank="subSpecies" subSpecies="supralabial">st supralabial</taxonomicName>
in contact with the nostril (right and left) forming a groove near mental scale (right and left); 10/11 infralabial scales; auricular opening elongated and smaller than ocular diameter; occipital scales different in size and shape to interorbital with some interspersed circular tubercles; mental scale forms an irregular &quot;V&quot; towards the posterior region; two postmental scales in contact each other and with the 1
<taxonomicName id="FDC302E02562FF8FFEDAFEDBFEB3FEF8" box="[265,339,260,285]" pageId="22" pageNumber="173" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd infralabial scales (right and left); postmental scales in contact with a row of 5 scales, the central scale (pentagon shape) is larger than pair of adjacent scales (right and left). Body with granular scales, has 13 rows of strongly keeled dorsal tubercles of variable size; 30 tubercles in a dorsal row starting in the base of the head to base of the tail; 24/19 tubercles axilla-groin; 32 rows of cycloid scales across venter; 81 ventral longitudinal scales from the first differentiated scale in the gular region to the anus; ventral scales larger than the lateral sclaes of the body and the gular region; scales slightly imbricate on the four extremities. The tail of the specimen is original (not regenerated) and measures
<quantity id="FD3BD4862562FF8FFD2DFE03FC86FE11" box="[766,870,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="22" pageNumber="173" unit="mm" value="39.6">39.6 mm</quantity>
with the tip broken, dorsal scales of the tail are subimbricate and mostly flattened. Therefore, the tubercles are clearly defined. Does not present femoral or precloacal pores. Lamellar formulae: anterior right leg (
<date id="4E7D5FA32562FF8FFCDBFDFBFC4CFDD9" box="[776,940,548,572]" pageId="22" pageNumber="173" value="2012-08-13" valueMax="2012-10-11" valueMin="2012-08-13">8-10-12-13-11</date>
), anterior left leg (
<date id="4E7D5FA32562FF8FFBACFDFBFAC3FDD9" box="[1151,1315,548,572]" pageId="22" pageNumber="173" value="2013-09-13" valueMax="2013-10-11" valueMin="2013-09-13">9-10-13-13-11</date>
), posterior right leg (
<date id="4E7D5FA32562FF8FFED6FD98FE4AFD85" box="[261,426,583,608]" pageId="22" pageNumber="173" value="2014-08-15" valueMax="2014-11-13" valueMin="2014-08-15">8-11-14-15-13</date>
), posterior left leg (
<date id="4E7D5FA32562FF8FFD5BFD97FCCFFD85" box="[648,815,584,608]" pageId="22" pageNumber="173" value="2014-08-14" valueMax="2014-12-13" valueMin="2014-08-14">8-12-14-14-13</date>
).
</paragraph>
<caption id="6EBC29EB2562FF8FFF44FAA2FE57FA57" httpUri="https://zenodo.org/record/1216717/files/figure.png" pageId="22" pageNumber="173" startId="22.[151,250,1405,1427]" targetBox="[151,1384,661,1384]" targetPageId="22">
<paragraph id="3A7C79632562FF8FFF44FAA2FE57FA57" blockId="22.[151,1436,1405,1458]" pageId="22" pageNumber="173">
<emphasis id="08B7A5712562FF8FFF44FAA2FEC1FA77" bold="true" box="[151,289,1405,1427]" pageId="22" pageNumber="173">FIGURE 12.</emphasis>
Dorsal and ventral view of the holotype of
<taxonomicName id="FDC302E02562FF8FFD3EFAA2FC13FA76" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[749,1011,1405,1427]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="173" phylum="Chordata" rank="species" species="kropotkini" status="sp. nov.">
<emphasis id="08B7A5712562FF8FFD3EFAA2FC13FA76" box="[749,1011,1405,1427]" italics="true" pageId="22" pageNumber="173">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
<emphasis id="08B7A5712562FF8FFC2FFAA2FBAEFA77" bold="true" box="[1020,1102,1405,1426]" pageId="22" pageNumber="173">
<taxonomicNameLabel id="1384180A2562FF8FFC2FFAA2FBAEFA77" box="[1020,1102,1405,1426]" pageId="22" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZFC 28736). The black bars represent the scale (10 mm).
</paragraph>
</caption>
<paragraph id="3A7C79632562FF8FFF14FA00FD02F9DA" blockId="22.[151,1437,1502,2031]" pageId="22" pageNumber="173">Measurements in mm. Snout-vent length 62.2; axilla-groin 23; head length 16.9; head width 12.3; snout length (to eye) 7.5; eye diameter 3.3; auricular opening (maximum) 1.7; length of fourth finger 5; length of fourth toe 6; width between eye supercilliaries 8.1; internaral 1.5.</paragraph>
<paragraph id="3A7C79632562FF8FFF14F995FF14F92A" blockId="22.[151,1437,1502,2031]" pageId="22" pageNumber="173">
Coloration of
<typeStatus id="E578C7C12562FF8FFEB6F995FE27F986" box="[357,455,1610,1635]" pageId="22" pageNumber="173" type="holotype">holotype</typeStatus>
in ethanol (
<figureCitation id="A2F865E62562FF8FFD9CF995FD43F986" box="[591,675,1610,1635]" captionStart="FIGURE 12" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[151,1384,661,1384]" captionTargetId="figure@22.[151,1436,661,1384]" captionTargetPageId="22" captionText="FIGURE 12. Dorsal and ventral view of the holotype of Phyllodactylus kropotkini sp. nov. (MZFC 28736). The black bars represent the scale (10 mm)." httpUri="https://zenodo.org/record/1216717/files/figure.png" pageId="22" pageNumber="173">Fig. 12</figureCitation>
). The
<typeStatus id="E578C7C12562FF8FFD3FF995FCAEF986" box="[748,846,1610,1635]" pageId="22" pageNumber="173" type="holotype">holotype</typeStatus>
presents a coloration that mixes the cream and light gray coloration on dorsal background, with a few dorsal dark brown spots; the ventral coloration is white with a slight light yellowish on posterior legs. The ventral coloration of the tail tip is dark gray, as well as the tips of the toepads.
</paragraph>
<paragraph id="3A7C79632562FF8FFF14F905FDA4F8F2" blockId="22.[151,1437,1502,2031]" pageId="22" pageNumber="173">
<emphasis id="08B7A5712562FF8FFF14F905FEDCF916" bold="true" box="[199,316,1754,1779]" pageId="22" pageNumber="173">Variation.</emphasis>
All characters are presented with mean values, standard deviation and range of meristic (
<tableCitation id="77414CD82562FF8FFAEFF905FA74F916" box="[1340,1428,1754,1779]" captionStart="TABLE2" captionStartId="7.[125,204,659,680]" captionText="TABLE2. Mean, stanđarđ đeviationanđ rangeof meristic variables examineđ forspecies of the P.tuberculosus group." pageId="22" pageNumber="173">Table 2</tableCitation>
) and morphometric variables (
<tableCitation id="77414CD82562FF8FFE0CF921FDD6F8F2" box="[479,566,1790,1815]" captionStart="TABLE 7" captionStartId="12.[120,199,575,596]" captionText="TABLE 7.Mean,stanđarđ đeviation anđrange ofmorphometric variables examineđforsix speciesofPhyllodactylus." pageId="22" pageNumber="173">Table 7</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72D92AE82562FF8EFF14F8FDFDBAFE6D" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173" type="distribution">
<paragraph id="3A7C79632562FF8EFF14F8FDFDBAFE6D" blockId="22.[151,1437,1502,2031]" lastBlockId="23.[151,1437,151,392]" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173">
<emphasis id="08B7A5712562FF8FFF14F8FDFE15F8DE" bold="true" box="[199,501,1826,1851]" pageId="22" pageNumber="173">Distribution and habitat.</emphasis>
<taxonomicName id="FDC302E02562FF8FFE2DF8FDFCC0F8DE" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[510,800,1826,1851]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="173" phylum="Chordata" rank="species" species="kropotkini">
<emphasis id="08B7A5712562FF8FFE2DF8FDFCC0F8DE" box="[510,800,1826,1851]" italics="true" pageId="22" pageNumber="173">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
is endemic to the middle region of the Balsas Basin at locality Nueva Filadelfia (Huerta Vieja), municipality of Tlapehuala,
<collectingRegion id="F807B7812562FF8FFC48F898FBE0F8BA" box="[923,1024,1863,1887]" country="Mexico" name="Guerrero" pageId="22" pageNumber="173">Guerrero</collectingRegion>
(
<figureCitation id="A2F865E62562FF8FFBDCF899FB81F8BA" box="[1039,1121,1862,1887]" captionStart="FIGURE 13" captionStartId="23.[151,250,1430,1452]" captionTargetBox="[151,1436,445,1409]" captionTargetId="figure@23.[151,1436,445,1409]" captionTargetPageId="23" captionText="FIGURE 13. Residuals of Tropical dry Forest at the type locality (Nueva Filadelfia, Tlapehuala, Guerrero) habitat of Phyllodactylus kropotkini sp.nov. (Photo by Tonatiuh Ramírez Reyes)." httpUri="https://zenodo.org/record/1216719/files/figure.png" pageId="22" pageNumber="173">Fig. 13</figureCitation>
). This species was observed mainly on rocks and between cracks in caves near the
<typeStatus id="E578C7C12562FF8FFD25F8B4FCC6F866" box="[758,806,1899,1923]" pageId="22" pageNumber="173">type</typeStatus>
locality. The specimens were collected in the house of a person who allowed collection inside his home. Due to its presumably small distribution area and its new taxonomic status, no further information on its reproduction or ecology is known. According to the digital elevation model of INEGI (resolution
<quantity id="FD3BD4862562FF8FFD82F809FD70F80A" box="[593,656,2006,2031]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="173" unit="m" value="15.0">15 m</quantity>
), the locality is at
<quantity id="FD3BD4862562FF8FFCB8F809FC55F80B" box="[875,949,2006,2031]" metricMagnitude="2" metricUnit="m" metricValue="5.79" pageId="22" pageNumber="173" unit="m" value="579.0">579 m</quantity>
of elevation in relict lowland tropical dry forest in a habitat clearly disturbed by agricultural and livestock activity. Annual temperature for the locality is of 27 °C, with maximum temperatures of 38.6 °C and annual precipitation of
<quantity id="FD3BD4862563FF8EFC0CFF63FBABFF31" box="[991,1099,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.099" pageId="23" pageNumber="174" unit="mm" value="1099.0">1099 mm</quantity>
on average. The Balsas Basin contains a great diversity of species of the genus
<taxonomicName id="FDC302E02563FF8EFD07FF3EFCCEFF1D" authorityName="N.J. Jacquin ex Linnaeus" authorityYear="1762" box="[724,814,225,248]" class="Magnoliopsida" family="Burseraceae" genus="Bursera" higherTaxonomySource="CoL" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="174" phylum="Magnoliopsida" rank="genus">
<emphasis id="08B7A5712563FF8EFD07FF3EFCCEFF1D" box="[724,814,225,248]" italics="true" pageId="23" pageNumber="174">Bursera</emphasis>
</taxonomicName>
, in fact this biogeographic province is considered the likely center of diversification and endemism of
<taxonomicName id="FDC302E02563FF8EFD13FEDAFCFAFEF9" authorityName="N.J. Jacquin ex Linnaeus" authorityYear="1762" box="[704,794,261,284]" class="Magnoliopsida" family="Burseraceae" genus="Bursera" higherTaxonomySource="CoL" kingdom="Plantae" order="Sapindales" pageId="23" pageNumber="174" phylum="Magnoliopsida" rank="genus">
<emphasis id="08B7A5712563FF8EFD13FEDAFCFAFEF9" box="[704,794,261,284]" italics="true" pageId="23" pageNumber="174">Bursera</emphasis>
</taxonomicName>
(
<bibRefCitation id="5E5204922563FF8EFCF9FEDBFC5CFEF9" box="[810,956,260,285]" pageId="23" pageNumber="174" refId="ref18166" refString="Toledo, A. (2003) Rios, costas, mares. Hacia un analisis integrado de las regiones hidrologicas de Mexico. INE-Semarnat, Mexico, 117 pp." type="book">Toledo 2003</bibRefCitation>
;
<bibRefCitation id="5E5204922563FF8EFC1AFEDBFB10FEF9" box="[969,1264,260,285]" pageId="23" pageNumber="174" refId="ref17216" refString="Pineda-Garcia, F., Arredondo-Amezcua, L. &amp; Ibarra-Manriquez, G. (2007) Riqueza y diversidad de especies lenosas del bosque tropical caducifolio El Tarimo, Cuenca del Balsas, Guerrero. Revista Mexicana de Biodiversidad, 78, 129 - 139." type="journal article">
Pineda-García
<emphasis id="08B7A5712563FF8EFBA2FEDAFB4BFEF8" box="[1137,1195,260,285]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
2007
</bibRefCitation>
). Other species of reptiles that inhabit this province are:
<emphasis id="08B7A5712563FF8EFD5AFEF8FEF2FE6D" italics="true" pageId="23" pageNumber="174">
<taxonomicName id="FDC302E02563FF8EFD5AFEF8FBC2FEDA" authority=", Ctenosaura" authorityName="Ctenosaura" box="[649,1058,295,320]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="frenatus">Hemidactylus frenatus, Ctenosaura</taxonomicName>
pectinata, Sceloporus horridus, Urosaurus bicarinatus, Drymarchon corais, Leptodeira maculata, Salvadora mexicana,
<taxonomicName id="FDC302E02563FF8EFBAEFE93FA7CFE81" authority=", Micrurus" authorityName="Micrurus" box="[1149,1436,332,357]" class="Reptilia" family="Viperidae" genus="Crotalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="simus">Crotalus simus, Micrurus</taxonomicName>
laticollaris
</emphasis>
(
<bibRefCitation id="5E5204922563FF8EFEF3FEB0FDAFFE6D" box="[288,591,367,392]" pageId="23" pageNumber="174" refId="ref17666" refString="Reyna-Alvarez, J., Suazo-Ortuno, I. &amp; Alvarado-Diaz, J. (2010) Herpetofauna del Municipio de Huetamo, Michoacan, Mexico. Biologicas, 12 (1), 40 ¯ 45." type="journal article">
Reyna-Álvarez
<emphasis id="08B7A5712563FF8EFE00FEAEFDEBFE6D" box="[467,523,367,392]" italics="true" pageId="23" pageNumber="174">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="6EBC29EB2563FF8EFF44FA49FC8BFA2E" httpUri="https://zenodo.org/record/1216719/files/figure.png" pageId="23" pageNumber="174" startId="23.[151,250,1430,1452]" targetBox="[151,1436,445,1409]" targetPageId="23">
<paragraph id="3A7C79632563FF8EFF44FA49FC8BFA2E" blockId="23.[151,1436,1430,1483]" pageId="23" pageNumber="174">
<emphasis id="08B7A5712563FF8EFF44FA49FEC5FA4E" bold="true" box="[151,293,1430,1452]" pageId="23" pageNumber="174">FIGURE 13.</emphasis>
Residuals of Tropical dry Forest at the type locality (Nueva Filadelfia, Tlapehuala, Guerrero) habitat of
<taxonomicName id="FDC302E02563FF8EFF44FA6AFE7BFA2E" authority="Ramírez-Reyes &amp; Flores-Villela, 2018" authorityName="Ramírez-Reyes &amp; Flores-Villela" authorityYear="2018" box="[151,411,1461,1483]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="kropotkini" status="sp. nov.">
<emphasis id="08B7A5712563FF8EFF44FA6AFE7BFA2E" box="[151,411,1461,1483]" italics="true" pageId="23" pageNumber="174">Phyllodactylus kropotkini</emphasis>
</taxonomicName>
<emphasis id="08B7A5712563FF8EFE72FA6AFE11FA2F" bold="true" box="[417,497,1461,1482]" pageId="23" pageNumber="174">
<taxonomicNameLabel id="1384180A2563FF8EFE72FA6AFE11FA2F" box="[417,497,1461,1482]" pageId="23" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Photo by Tonatiuh Ramírez Reyes).
</paragraph>
</caption>
<subSubSection id="72D92AE82563FF81FF14FA28FC50FF4A" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="discussion">
<paragraph id="3A7C79632563FF8EFF14FA28FD44F902" blockId="23.[151,1437,1527,2021]" pageId="23" pageNumber="174">
<emphasis id="08B7A5712563FF8EFF14FA28FD29F9F5" bold="true" box="[199,713,1527,1552]" pageId="23" pageNumber="174">Comments about conservation and threats.</emphasis>
This species inhabits the middle region of Balsas Basin, and the
<typeStatus id="E578C7C12563FF8EFF44F9C3FF27F9D1" box="[151,199,1564,1588]" pageId="23" pageNumber="174">type</typeStatus>
locality is a remnant tropical dry forest; the species inhabits an area highly disturbed by agricultural and livestock activities, however, this species is able to adapt effectively to modified habitats, inhabiting houses or other
<typeStatus id="E578C7C12563FF8EFF04F9BBFEF2F999" box="[215,274,1636,1660]" pageId="23" pageNumber="174">types</typeStatus>
of buildings (bridges, fences, etc.). The main threat to this species (and in general to all species of the
<taxonomicName id="FDC302E02563FF8EFA54F9BAFF2FF945" authorityName="Smith" authorityYear="1935" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712563FF8EFA54F9BAFF2FF945" italics="true" pageId="23" pageNumber="174">P. lanei</emphasis>
</taxonomicName>
complex) are the myths and false beliefs of the people, who consider this species as dangerous or poisonous (
<bibRefCitation id="5E5204922563FF8EFF4DF973FE5EF921" box="[158,446,1707,1732]" pageId="23" pageNumber="174" refId="ref16620" refString="Garcia, A. &amp; Ceballos, G. (1994) Guia de campo de los reptiles y anfibios de la costa de Jalisco, Mexico. Fundacion Ecologica de Cuixmala, A. C., Instituto de Biologia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 176 pp." type="book">García &amp; Ceballos 1994</bibRefCitation>
). This false belief has led some people to implement &quot;eradication&quot; campaigns of this species in houses or wherever geckos are seen.
</paragraph>
<paragraph id="3A7C79632563FF8EFF14F92CFDA6F825" blockId="23.[151,1437,1527,2021]" pageId="23" pageNumber="174">
<emphasis id="08B7A5712563FF8EFF14F92CFE59F8E9" bold="true" box="[199,441,1779,1804]" pageId="23" pageNumber="174">Taxonomic changes.</emphasis>
Although the purpose of the present study was not to re-describe the species of the
<taxonomicName id="FDC302E02563FF8EFA54F92AFF2FF8D5" authorityName="Smith" authorityYear="1935" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712563FF8EFA54F92AFF2FF8D5" italics="true" pageId="23" pageNumber="174">P. lanei</emphasis>
</taxonomicName>
complex, we will mention some points that are important to consider for the previously described species
<taxonomicName id="FDC302E02563FF8EFA54F8C6FF2FF8B1" authorityName="Smith" authorityYear="1935" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712563FF8EFA54F8C6FF2FF8B1" italics="true" pageId="23" pageNumber="174">P. lanei</emphasis>
</taxonomicName>
,
<taxonomicName id="FDC302E02563FF8EFF0FF8E2FEACF8B1" box="[220,332,1853,1876]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712563FF8EFF0FF8E2FEACF8B1" box="[220,332,1853,1876]" italics="true" pageId="23" pageNumber="174">P. rupinus</emphasis>
</taxonomicName>
,
<taxonomicName id="FDC302E02563FF8EFE88F8E2FE32F8B1" box="[347,466,1851,1876]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712563FF8EFE88F8E2FE32F8B1" box="[347,466,1851,1876]" italics="true" pageId="23" pageNumber="174">P. isabelae</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02563FF8EFDDBF8E2FD92F8B1" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[520,626,1851,1876]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712563FF8EFDDBF8E2FD92F8B1" box="[520,626,1851,1876]" italics="true" pageId="23" pageNumber="174">P. lupitae</emphasis>
</taxonomicName>
. The original description of
<taxonomicName id="FDC302E02563FF8EFC7CF8E2FBE3F8B1" authorityName="Smith" authorityYear="1935" box="[943,1027,1851,1876]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712563FF8EFC7CF8E2FBE3F8B1" box="[943,1027,1851,1876]" italics="true" pageId="23" pageNumber="174">P. lanei</emphasis>
</taxonomicName>
can be consulted in
<bibRefCitation id="5E5204922563FF8EFB35F8E4FA9CF8B1" box="[1254,1404,1851,1876]" pageId="23" pageNumber="174" refId="ref18024" refString="Smith, H. M. (1935) Miscellaneous notes on Mexican lizards. The University of Kansas Science Bulletin, 36, 119 - 155." type="journal article">Smith (1935)</bibRefCitation>
,
<taxonomicName id="FDC302E02563FF8EFA54F8E2FF0DF89D" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712563FF8EFA54F8E2FF0DF89D" italics="true" pageId="23" pageNumber="174">P. rupinus</emphasis>
</taxonomicName>
can be consulted in
<bibRefCitation id="5E5204922563FF8EFE0FF880FD96F89D" box="[476,630,1887,1912]" pageId="23" pageNumber="174" refId="ref16514" refString="Dixon, J. R. (1964) The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America. New Mexico State University Research Center, Scientific Bulletin, 1964, 1 - 139." type="journal article">Dixon (1964)</bibRefCitation>
and
<taxonomicName id="FDC302E02563FF8EFD62F8BEFCC8F89D" box="[689,808,1887,1912]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712563FF8EFD62F8BEFCC8F89D" box="[689,808,1887,1912]" italics="true" pageId="23" pageNumber="174">P. isabelae</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02563FF8EFCB5F8BEFA78F89D" authority="in Castro-Franco &amp; Uribe-Pena (1992)" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[870,1432,1887,1912]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712563FF8EFCB5F8BEFC32F89D" box="[870,978,1887,1912]" italics="true" pageId="23" pageNumber="174">P. lupitae</emphasis>
in
<bibRefCitation id="5E5204922563FF8EFC29F8BFFA78F89D" box="[1018,1432,1887,1912]" pageId="23" pageNumber="174" refId="ref16250" refString="Castro-Franco, R. &amp; Uribe-Pena, Z. (1992) Dos subespecies nuevas de Phyllodactylus lanei (Sauria: Gekkonidae). Anales del Instituto de Biologia. Universidad Nacional Autonoma de Mexico, Serie Zoologia, 63, 113 - 123." type="journal article">Castro-Franco &amp; Uribe-Peña (1992)</bibRefCitation>
</taxonomicName>
. The last publication includes an identification key for
<taxonomicName id="FDC302E02563FF8EFCD8F85AFC81F879" authorityName="Smith" authorityYear="1935" box="[779,865,1923,1948]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lanei">
<emphasis id="08B7A5712563FF8EFCD8F85AFC81F879" box="[779,865,1923,1948]" italics="true" pageId="23" pageNumber="174">P. lanei</emphasis>
</taxonomicName>
,
<taxonomicName id="FDC302E02563FF8EFCA2F85AFC04F879" box="[881,996,1925,1948]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="rupinus">
<emphasis id="08B7A5712563FF8EFCA2F85AFC04F879" box="[881,996,1925,1948]" italics="true" pageId="23" pageNumber="174">P. rupinus</emphasis>
</taxonomicName>
,
<taxonomicName id="FDC302E02563FF8EFC26F85AFB8EF879" box="[1013,1134,1923,1948]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="08B7A5712563FF8EFC26F85AFB8EF879" box="[1013,1134,1923,1948]" italics="true" pageId="23" pageNumber="174">P. isabelae</emphasis>
</taxonomicName>
and
<taxonomicName id="FDC302E02563FF8EFB7EF85AFAF9F879" authorityName="in Castro-Franco &amp; Uribe-Pena" authorityYear="1992" box="[1197,1305,1923,1948]" class="Reptilia" family="Gekkonidae" genus="Phyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lupitae">
<emphasis id="08B7A5712563FF8EFB7EF85AFAF9F879" box="[1197,1305,1923,1948]" italics="true" pageId="23" pageNumber="174">P. lupitae</emphasis>
</taxonomicName>
and agrees mostly with the results presented here.
</paragraph>
<paragraph id="3A7C79632563FF81FF14F813FC50FF4A" blockId="23.[151,1437,1527,2021]" lastBlockId="24.[151,944,151,176]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">The original descriptions are extensive enough and can be reviewed for a detailed taxonomic identification, here we will only mention characters that can help us identify the taxa.</paragraph>
</subSubSection>
</treatment>
</document>