treatments-xml/data/B2/11/8B/B2118B04FFD5FF9CA6C30473FB1BB41F.xml

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<mods:title id="73D1240B5662C886A34E67EE45594A16">Three new species and a new record of the spider genus Mallinella Strand, 1906 from Brunei (Araneae, Zodariidae)</mods:title>
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<mods:namePart id="2BF61D42F8C6B27F21346833B08232FA">Dankittipakul, Pakawin</mods:namePart>
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<mods:namePart id="24C60EF6E897C06F3D5C954D5AEAC850">Koh, Joseph K. H.</mods:namePart>
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<mods:date id="6728984889F44E9427C7565E06761DE2">2014</mods:date>
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<treatment id="B2118B04FFD5FF9CA6C30473FB1BB41F" ID-DOI="http://doi.org/10.5281/zenodo.6132154" ID-GBIF-Taxon="119509615" ID-Zenodo-Dep="6132154" LSID="urn:lsid:plazi:treatment:B2118B04FFD5FF9CA6C30473FB1BB41F" httpUri="http://treatment.plazi.org/id/B2118B04FFD5FF9CA6C30473FB1BB41F" lastPageId="5" lastPageNumber="382" pageId="1" pageNumber="378">
<subSubSection id="72A26999FFD5FF98A6C30473FD92B0B9" pageId="1" pageNumber="378" type="nomenclature">
<paragraph id="3A073A12FFD5FF98A6C30473FC99B047" blockId="1.[151,834,1842,1902]" box="[151,834,1842,1869]" pageId="1" pageNumber="378">
<heading id="614F8D7EFFD5FF98A6C30473FC99B047" bold="true" box="[151,834,1842,1869]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
<emphasis id="08CCE600FFD5FF98A6C30473FC99B047" bold="true" box="[151,834,1842,1869]" pageId="1" pageNumber="378">
<taxonomicName id="FDB84191FFD5FF98A6C30473FD05B047" ID-CoL="3XNM2" authority="Koh &amp; Dankittipakul" authorityName="Koh &amp; Dankittipakul" box="[151,734,1842,1869]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD5FF98A6C30473FE11B046" bold="true" box="[151,458,1842,1868]" italics="true" pageId="1" pageNumber="378">Mallinella merimbunenis</emphasis>
Koh &amp; Dankittipakul
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD5FF98A4B10472FC99B047" box="[741,834,1843,1869]" pageId="1" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3A073A12FFD5FF98A6C30414FEA1B064" blockId="1.[151,834,1842,1902]" box="[151,378,1877,1902]" pageId="1" pageNumber="378">
(
<figureCitation id="A2832697FFD5FF98A6CB0414FEC4B064" box="[159,287,1877,1902]" captionStart-0="FIGURES 1 – 3. 1 – 3" captionStart-1="FIGURES 4 – 11. 4" captionStartId-0="2.[151,264,1049,1072]" captionStartId-1="2.[151,264,1887,1910]" captionTargetBox-0="[177,1418,195,1028]" captionTargetBox-1="[172,1415,1139,1867]" captionTargetId-0="figure@2.[175,1419,195,1029]" captionTargetId-1="figure@2.[172,1416,1139,1868]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 3. 1 – 3. Habitus. 1. Mallinella myrmecophaga sp. nov., female holotype. 2. M. platyrhyncha sp. nov., female holotype. 3. M. merimbunenis sp. nov., female paratype." captionText-1="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri-0="https://zenodo.org/record/225301/files/figure.png" httpUri-1="https://zenodo.org/record/225302/files/figure.png" pageId="1" pageNumber="378">Figs 3–4, 8</figureCitation>
, 
<figureCitation id="A2832697FFD5FF98A77F0414FEA9B064" box="[299,370,1877,1902]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="1" pageNumber="378">15–21</figureCitation>
)
</paragraph>
<paragraph id="3A073A12FFD5FF98A6C304DCFD92B0B9" blockId="1.[151,585,1949,1971]" box="[151,585,1949,1971]" pageId="1" pageNumber="378">
<treatmentCitationGroup id="1AA81D3CFFD5FF98A6C304DCFD92B0B9" box="[151,585,1949,1971]" pageId="1" pageNumber="378">
<taxonomicName id="FDB84191FFD5FF98A6C304DCFEDAB0B9" ID-CoL="5KLT" box="[151,257,1949,1971]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">
<emphasis id="08CCE600FFD5FF98A6C304DCFEDAB0B9" box="[151,257,1949,1971]" italics="true" pageId="1" pageNumber="378">Mallinella</emphasis>
</taxonomicName>
sp. A 
<treatmentCitation id="BB191C03FFD5FF98A71604DCFD9FB0B9" author="Koh" box="[322,580,1949,1971]" page="335" pageId="1" pageNumber="378" year="2013">
<bibRefCitation id="5E2947E3FFD5FF98A71604DCFDCAB0B9" author="Koh" box="[322,529,1949,1971]" pageId="1" pageNumber="378" refString="Koh, J. K. H. &amp; Leong, T. M. (2013) Biodiversity in the Heart of Borneo: Spiders of Brunei Darussalam. Natural History Publications (Borneo), Kota Kinabalu, 357 pp." type="book" year="2013">Koh &amp; Leong, 2013</bibRefCitation>
: 335
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.
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</paragraph>
</subSubSection>
<caption id="6EC76A9AFFD6FF9BA6C30758FD0FB345" httpUri="https://zenodo.org/record/225301/files/figure.png" pageId="2" pageNumber="379" targetBox="[177,1418,195,1028]" targetPageId="2">
<paragraph id="3A073A12FFD6FF9BA6C30758FD0FB345" blockId="2.[151,1436,1049,1103]" pageId="2" pageNumber="379">
<emphasis id="08CCE600FFD6FF9BA6C30758FEE7B325" bold="true" box="[151,316,1049,1072]" pageId="2" pageNumber="379">FIGURES 1–3.</emphasis>
1–3. Habitus. 1. 
<taxonomicName id="FDB84191FFD6FF9BA7A1075BFCDFB33A" box="[501,772,1050,1072]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="myrmecophaga" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA7A1075BFCDFB33A" box="[501,772,1050,1072]" italics="true" pageId="2" pageNumber="379">Mallinella myrmecophaga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA558075BFC84B325" box="[780,863,1050,1071]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA558075BFC84B325" bold="true" box="[780,863,1050,1071]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype. 2. 
<taxonomicName id="FDB84191FFD6FF9BA215075AFB30B33A" box="[1089,1259,1050,1072]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="platyrhyncha" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA215075AFB30B33A" box="[1089,1259,1050,1072]" italics="true" pageId="2" pageNumber="379">M. platyrhyncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA2A0075BFA9CB325" box="[1268,1351,1050,1071]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA2A0075BFA9CB325" bold="true" box="[1268,1351,1050,1071]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype. 3. 
<taxonomicName id="FDB84191FFD6FF9BA741077BFE1DB345" box="[277,454,1081,1103]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA741077BFE1DB345" box="[277,454,1081,1103]" italics="true" pageId="2" pageNumber="379">M. merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA7990778FDC6B344" box="[461,541,1081,1102]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA7990778FDC6B344" bold="true" box="[461,541,1081,1102]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype.
</paragraph>
</caption>
<caption id="6EC76A9AFFD6FF9BA6C3041EFE99B0FA" httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="2" pageNumber="379" targetBox="[172,1415,1139,1867]" targetPageId="2">
<paragraph id="3A073A12FFD6FF9BA6C3041EFE99B0FA" blockId="2.[151,1436,1887,2032]" pageId="2" pageNumber="379">
<emphasis id="08CCE600FFD6FF9BA6C3041EFE9DB07E" bold="true" box="[151,326,1887,1910]" pageId="2" pageNumber="379">FIGURES 4–11.</emphasis>
4. Leg I–III, showing tubercle (indicated by arrow). 5–9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. 
<taxonomicName id="FDB84191FFD6FF9BA758043FFDD7B09E" box="[268,524,1918,1940]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA758043FFDD7B09E" box="[268,524,1918,1940]" italics="true" pageId="2" pageNumber="379">Mallinella merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA440043EFDBEB09E" box="[532,613,1919,1940]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA440043EFDBEB09E" bold="true" box="[532,613,1919,1940]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype. 5. 
<taxonomicName id="FDB84191FFD6FF9BA57A043EFABDB09E" authority="Dankittipakul, Jocque &amp; Singtripop, 2012" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[814,1382,1918,1940]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="bigemina">
<emphasis id="08CCE600FFD6FF9BA57A043EFC74B09E" box="[814,943,1918,1940]" italics="true" pageId="2" pageNumber="379">M. bigemina</emphasis>
Dankittipakul, Jocqué &amp; Singtripop, 2012
</taxonomicName>
, new material from Brunei. 6. 
<taxonomicName id="FDB84191FFD6FF9BA7C204DFFDCCB0B9" box="[406,535,1949,1971]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="bigemina">
<emphasis id="08CCE600FFD6FF9BA7C204DFFDCCB0B9" box="[406,535,1949,1971]" italics="true" pageId="2" pageNumber="379">M. bigemina</emphasis>
</taxonomicName>
, female holotype. 7. 
<taxonomicName id="FDB84191FFD6FF9BA4A504DFFB17B0B9" authority="Bosmans &amp; Hillyard, 1990" authorityName="Bosmans &amp; Hillyard" authorityYear="1990" box="[753,1228,1949,1971]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="08CCE600FFD6FF9BA4A504DFFC7BB0B9" box="[753,928,1949,1971]" italics="true" pageId="2" pageNumber="379">M. albomaculata</emphasis>
(Bosmans &amp; Hillyard, 1990)
</taxonomicName>
, female paratype. 8. 
<taxonomicName id="FDB84191FFD6FF9BA6C304FCFE93B0D8" box="[151,328,1980,2002]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA6C304FCFE93B0D8" box="[151,328,1980,2002]" italics="true" pageId="2" pageNumber="379">M. merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA71B04FDFE7BB0DB" box="[335,416,1980,2001]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA71B04FDFE7BB0DB" bold="true" box="[335,416,1980,2001]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype. 9–10. 
<taxonomicName id="FDB84191FFD6FF9BA4C804FCFC80B0D8" box="[668,859,1980,2002]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="myrmecophaga" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA4C804FCFC80B0D8" box="[668,859,1980,2002]" italics="true" pageId="2" pageNumber="379">M. myrmecophaga</emphasis>
</taxonomicName>
<emphasis id="08CCE600FFD6FF9BA53604FDFC69B0DB" bold="true" box="[866,946,1980,2001]" pageId="2" pageNumber="379">
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA53604FDFC69B0DB" box="[866,946,1980,2001]" pageId="2" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype. 11. 
<taxonomicName id="FDB84191FFD6FF9BA2C204FCFAE6B0D8" box="[1174,1341,1980,2002]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="platyrhyncha" status="sp. nov.">
<emphasis id="08CCE600FFD6FF9BA2C204FCFAE6B0D8" box="[1174,1341,1980,2002]" italics="true" pageId="2" pageNumber="379">M. platyrhyncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD6FF9BA31004FDFA4EB0DB" box="[1348,1429,1980,2001]" pageId="2" pageNumber="379" rank="species">
<emphasis id="08CCE600FFD6FF9BA31004FDFA4EB0DB" bold="true" box="[1348,1429,1980,2001]" pageId="2" pageNumber="379">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype.
</paragraph>
</caption>
<caption id="6EC76A9AFFD7FF9AA6C30407FA40B0B2" httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="3" pageNumber="380" targetBox="[167,1417,236,1838]" targetPageId="3">
<paragraph id="3A073A12FFD7FF9AA6C30407FA40B0B2" blockId="3.[151,1437,1862,1976]" pageId="3" pageNumber="380">
<emphasis id="08CCE600FFD7FF9AA6C30407FE8CB056" bold="true" box="[151,343,1862,1885]" pageId="3" pageNumber="380">FIGURES 12–21.</emphasis>
12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12–14. 
<taxonomicName id="FDB84191FFD7FF9AA6B704C5FCBDB090" authority="Dankittipakul, Jocque &amp; Singtripop, 2012" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[227,870,1924,1946]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="bigemina">
<emphasis id="08CCE600FFD7FF9AA6B704C5FE69B090" box="[227,434,1924,1946]" italics="true" pageId="3" pageNumber="380">Mallinella bigemina</emphasis>
Dankittipakul, Jocqué &amp; Singtripop, 2012
</taxonomicName>
, new female from Brunei. 15–17. 
<taxonomicName id="FDB84191FFD7FF9AA29104C4FAADB090" box="[1221,1398,1924,1946]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD7FF9AA29104C4FAADB090" box="[1221,1398,1924,1946]" italics="true" pageId="3" pageNumber="380">M. merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD7FF9AA32904C5FF18B0B2" pageId="3" pageNumber="380" rank="species">
<emphasis id="08CCE600FFD7FF9AA32904C5FF18B0B2" bold="true" pageId="3" pageNumber="380">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype. 18–21. 
<taxonomicName id="FDB84191FFD7FF9AA7EC04E2FDB2B0B2" box="[440,617,1954,1976]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD7FF9AA7EC04E2FDB2B0B2" box="[440,617,1954,1976]" italics="true" pageId="3" pageNumber="380">M. merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD7FF9AA42404E2FD1BB0B2" box="[624,704,1955,1976]" pageId="3" pageNumber="380" rank="species">
<emphasis id="08CCE600FFD7FF9AA42404E2FD1BB0B2" bold="true" box="[624,704,1955,1976]" pageId="3" pageNumber="380">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype. Scale bar: 0.1 mm (15–17); 0.25 mm (12–14, 18–21).
</paragraph>
</caption>
<subSubSection id="72A26999FFD0FF9DA6C303D6FE25B560" pageId="4" pageNumber="381" type="materials_examined">
<paragraph id="3A073A12FFD0FF9DA6C303D6FE25B560" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA6C303D6FCF5B7BA" bold="true" box="[151,814,151,176]" pageId="4" pageNumber="381">
Material examined. 
<typeStatus id="E50384B0FFD0FF9DA7CE03D6FDD0B7BA" box="[410,523,151,176]" pageId="4" pageNumber="381" type="holotype">Holotype</typeStatus>
: ♂, 
<collectingCountry id="42AF7A82FFD0FF9DA41A03D6FD66B7BA" box="[590,701,151,176]" name="Brunei" pageId="4" pageNumber="381">BRUNEI</collectingCountry>
, 
<emphasis id="08CCE600FFD0FF9DA48603D9FCFFB7A5" bold="true" box="[722,804,152,175]" italics="true" pageId="4" pageNumber="381">Tutong</emphasis>
:
</emphasis>
Tasek Merimbun Heritage Park, Zone C3, mixed dipterocarp forest (
<geoCoordinate id="5F8C5CD5FFD0FF9DA73A03FAFE16B7DE" box="[366,461,187,212]" direction="north" orientation="latitude" pageId="4" pageNumber="381" precision="925" value="4.5333333">N04°32′</geoCoordinate>
24″, 
<geoCoordinate id="5F8C5CD5FFD0FF9DA45703FAFDB5B7DE" box="[515,622,187,212]" direction="east" orientation="longitude" pageId="4" pageNumber="381" precision="925" value="114.65">E114°39′</geoCoordinate>
19″), 
<date id="4E061CD2FFD0FF9DA4FF03FDFCEAB7D9" box="[683,817,187,212]" pageId="4" pageNumber="381" value="2008-07-07">7 July 2008</date>
, J.K.H. Koh leg. (BM, JK.08.06.07.0021). 
<emphasis id="08CCE600FFD0FF9DA34F03FAFE4EB7FD" bold="true" pageId="4" pageNumber="381">
<typeStatus id="E50384B0FFD0FF9DA34F03FAFA4CB7DE" box="[1307,1431,187,212]" pageId="4" pageNumber="381" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="42AF7A82FFD0FF9DA6C3039FFEDDB7FD" box="[151,262,222,247]" name="Brunei" pageId="4" pageNumber="381">BRUNEI</collectingCountry>
, 
<emphasis id="08CCE600FFD0FF9DA747039FFE4EB7FD" bold="true" box="[275,405,222,247]" italics="true" pageId="4" pageNumber="381">Temburong</emphasis>
</emphasis>
: Sungai Batu Apoi, river bank along disturbed forest (
<geoCoordinate id="5F8C5CD5FFD0FF9DA253039FFBB3B7FD" box="[1031,1128,222,247]" direction="north" orientation="latitude" pageId="4" pageNumber="381" precision="925" value="4.7">N04°42′</geoCoordinate>
19″, 
<geoCoordinate id="5F8C5CD5FFD0FF9DA2C9039FFAD3B7FD" box="[1181,1288,222,247]" direction="east" orientation="longitude" pageId="4" pageNumber="381" precision="925" value="115.11667">E115°07′</geoCoordinate>
13″), 
<date id="4E061CD2FFD0FF9DA310039EFF09B610" pageId="4" pageNumber="381" value="2007-06-17">17 June 2007</date>
, J.K.H. Koh leg., 1♀ (BM, JK.07.04.17.3008). 
<emphasis id="08CCE600FFD0FF9DA4AE0243FC8DB610" bold="true" box="[762,854,257,282]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA4AE0243FC97B613" bold="true" box="[762,844,258,281]" italics="true" pageId="4" pageNumber="381">Tutong</emphasis>
:
</emphasis>
Tasek Merimbun Heritage Park, Zone C2, Sungai Melunchur, Base Camp I, mixed dipterocarp forest (N4O35’13 E114O41’14), 
<date id="4E061CD2FFD0FF9DA24C0266FB15B64A" box="[1048,1230,295,320]" pageId="4" pageNumber="381" value="2012-03-25">25 March 2012</date>
, J.K.H. Koh leg., Kampung Bukit, foliage in logged forest (N04O35'31&quot; E114O37'10&quot;), 
<date id="4E061CD2FFD0FF9DA5E8020FFB7BB66C" box="[956,1184,333,358]" pageId="4" pageNumber="381" value="2006-09-30">30 September 2006</date>
, J.K.H. Koh leg., 1♀ (LKCNHM, JK.06.09.30.0066). 
<emphasis id="08CCE600FFD0FF9DA45E0231FD43B683" bold="true" box="[522,664,368,393]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA45E0231FD56B683" bold="true" box="[522,653,368,393]" italics="true" pageId="4" pageNumber="381">Temburong</emphasis>
:
</emphasis>
Sungai Esu, vegetation along stream in primary mixed dipterocarp forest (N04O32’17 E115O09’35), 
<date id="4E061CD2FFD0FF9DA45D02D6FD12B6A5" box="[521,713,406,431]" pageId="4" pageNumber="381" value="2007-10-24">24 October 2007</date>
, J.K.H. Koh leg., 
<specimenCount id="2CBEF19BFFD0FF9DA5C002D6FC6CB6A5" box="[916,951,407,431]" pageId="4" pageNumber="381" type="generic">1♂</specimenCount>
(LKCNHM, JK.07.10.24.0016). Kampong Labu, foliage in disturbed forest (N04O45’41 E15O10’51), 
<date id="4E061CD2FFD0FF9DA56302FCFBC3B6DF" box="[823,1048,444,469]" pageId="4" pageNumber="381" value="2008-11-10">10 November 2008</date>
, J.K.H. Koh leg., 1♀ (LKCNHM JK.08.11.10.1001). Perdayan Forest Reserve, foliage in lowland mixed dipterocarp forest (
<geoCoordinate id="5F8C5CD5FFD0FF9DA35F02A2FA94B6F1" box="[1291,1359,483,507]" direction="north" orientation="latitude" pageId="4" pageNumber="381" precision="925" value="0.06666667">N 0 4</geoCoordinate>
O45’41 E115O10’51), 
<date id="4E061CD2FFD0FF9DA7620148FE02B52B" box="[310,473,520,545]" pageId="4" pageNumber="381" value="2012-04-20">20 April 2012</date>
, J.K.H. Koh leg., 1♀ (LKCNHM, JK.12.04.20.0020). 
<emphasis id="08CCE600FFD0FF9DA2100149FA94B52B" bold="true" box="[1092,1359,520,545]" pageId="4" pageNumber="381">
<taxonomicName id="FDB84191FFD0FF9DA2100149FA9DB52A" authority="Seri Begawan" authorityName="Seri Begawan" box="[1092,1350,520,545]" class="Insecta" family="Cerambycidae" genus="Bandar" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="08CCE600FFD0FF9DA2100149FA9DB52A" bold="true" box="[1092,1350,520,545]" italics="true" pageId="4" pageNumber="381">Bandar Seri Begawan</emphasis>
</taxonomicName>
:
</emphasis>
Wasai Sungai Akar, foliage in disturbed secondary forest (N04O56’50 E114O59’13), 
<date id="4E061CD2FFD0FF9DA24C016EFB78B54D" box="[1048,1187,558,583]" pageId="4" pageNumber="381" value="2011-07-09">9 July 2011</date>
, J.K.H. Koh leg., 
<specimenCount id="2CBEF19BFFD0FF9DA32D016EFA47B54D" box="[1401,1436,559,583]" pageId="4" pageNumber="381" type="generic">1♂</specimenCount>
(LKCNHM, JK.11.07.09.0023).
</paragraph>
</subSubSection>
<subSubSection id="72A26999FFD0FF9DA6930134FE4EB31B" pageId="4" pageNumber="381" type="diagnosis">
<paragraph id="3A073A12FFD0FF9DA6930134FE4EB31B" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA6930134FE9AB584" bold="true" box="[199,321,629,654]" pageId="4" pageNumber="381">Diagnosis.</emphasis>
Males of 
<taxonomicName id="FDB84191FFD0FF9DA7E40137FDAFB587" box="[432,628,629,653]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="merimbunenis" status="sp. nov.">
<emphasis id="08CCE600FFD0FF9DA7E40137FDAFB587" box="[432,628,629,653]" italics="true" pageId="4" pageNumber="381">M. merimbunenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13FF5B7BFFD0FF9DA42E0134FD08B584" box="[634,723,629,654]" pageId="4" pageNumber="381" rank="species">
<emphasis id="08CCE600FFD0FF9DA42E0134FD08B584" bold="true" box="[634,723,629,654]" pageId="4" pageNumber="381">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily recognised by the dark, elevated, circular tubercle on the proximal dorsal surface of femora I–III (
<figureCitation id="A2832697FFD0FF9DA4EE01D9FCDAB5BB" box="[698,769,664,689]" captionStart="FIGURES 4 – 11. 4" captionStartId="2.[151,264,1887,1910]" captionTargetBox="[172,1415,1139,1867]" captionTargetId="figure@2.[172,1416,1139,1868]" captionTargetPageId="2" captionText="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="4" pageNumber="381">Fig. 4</figureCitation>
), in combination with posterior metatarsi distinctly longer than femora, as well as their distinctive leg colour pattern (see species description below). These tubercles are stridulatory organs (
<bibRefCitation id="5E2947E3FFD0FF9DA722019FFD4AB5FD" author="Dankittipakul" box="[374,657,734,759]" pageId="4" pageNumber="381" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella, Strand, 1906, with description of new genera and new species from Southeast Asia (Araneae, Zodariidae). Zootaxa, 3369, 1 - 327." type="journal article" year="2012">
Dankittipakul 
<emphasis id="08CCE600FFD0FF9DA44C01A1FD92B5FD" box="[536,585,735,759]" italics="true" pageId="4" pageNumber="381">et al</emphasis>
. 2012
</bibRefCitation>
; 
<bibRefCitation id="5E2947E3FFD0FF9DA4C8019EFCF1B5FD" author="Jocque" box="[668,810,734,759]" pageId="4" pageNumber="381" refString="Jocque, R. (2005) Six stridulating organs on one spider (Araneae: Zodariidae): Is this the limit? The Journal of Arachnology, 33, 597 - 603. http: // dx. doi. org / 10.1636 / 04 - 107.1" type="journal article" year="2005">Jocqué 2005</bibRefCitation>
) of which three pairs are present also in male specimens of 
<taxonomicName id="FDB84191FFD0FF9DA6E20042FDD9B410" authority="Thorell, 1892" authorityName="Thorell" authorityYear="1892" box="[182,514,770,795]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="08CCE600FFD0FF9DA6E20042FE91B410" box="[182,330,770,794]" italics="true" pageId="4" pageNumber="381">M. annulipes</emphasis>
(Thorell, 1892)
</taxonomicName>
, found in 
<collectingCountry id="42AF7A82FFD0FF9DA4230043FD30B411" box="[631,747,770,795]" name="Singapore" pageId="4" pageNumber="381">Singapore</collectingCountry>
, peninsular 
<collectingCountry id="42AF7A82FFD0FF9DA5210043FC05B411" box="[885,990,770,795]" name="Malaysia" pageId="4" pageNumber="381">Malaysia</collectingCountry>
, and in the province of Riau Islands of 
<collectingCountry id="42AF7A82FFD0FF9DA6C30064FED3B434" box="[151,264,805,830]" name="Indonesia" pageId="4" pageNumber="381">Indonesia</collectingCountry>
. However, they are absent in the closely related 
<taxonomicName id="FDB84191FFD0FF9DA54A0067FAF9B434" authority="Bosmans &amp; Hillyard, 1990" authorityName="Bosmans &amp; Hillyard" authorityYear="1990" box="[798,1314,805,830]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="08CCE600FFD0FF9DA54A0067FC06B437" box="[798,989,805,829]" italics="true" pageId="4" pageNumber="381">M. albomaculata</emphasis>
(
<bibRefCitation id="5E2947E3FFD0FF9DA5BF0067FAC1B434" author="Bosmans" box="[1003,1306,805,830]" pageId="4" pageNumber="381" refString="Bosmans, R. &amp; Hillyard, P. (1990) Spiders of the family Zodariidae from Sulawesi, Indonesia (Arachnida: Araneae: Zodariidae). Bulletin of the British Arachnological Society, 8, 147 - 160." type="journal article" year="1990">Bosmans &amp; Hillyard, 1990</bibRefCitation>
)
</taxonomicName>
(also from Borneo, 
<typeStatus id="E50384B0FFD0FF9DA6AD0008FEF2B46B" box="[249,297,841,865]" pageId="4" pageNumber="381">type</typeStatus>
deposited at NHML, examined). Males of this new species are separated from those of the closely related 
<taxonomicName id="FDB84191FFD0FF9DA6BE002DFE72B48E" box="[234,425,876,900]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="08CCE600FFD0FF9DA6BE002DFE72B48E" box="[234,425,876,900]" italics="true" pageId="4" pageNumber="381">M. albomaculata</emphasis>
</taxonomicName>
by the smaller RTA (
<figureCitation id="A2832697FFD0FF9DA4CD002AFD32B48E" box="[665,745,875,900]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="4" pageNumber="381">Fig. 17</figureCitation>
) and the larger distal extremities on both rami of the embolus (
<figureCitation id="A2832697FFD0FF9DA6CB00CFFEF8B4AD" box="[159,291,910,935]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="4" pageNumber="381">Figs 15–16</figureCitation>
). Females resemble those of 
<taxonomicName id="FDB84191FFD0FF9DA42500D1FCE8B4AD" box="[625,819,911,935]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="08CCE600FFD0FF9DA42500D1FCE8B4AD" box="[625,819,911,935]" italics="true" pageId="4" pageNumber="381">M. albomaculata</emphasis>
</taxonomicName>
(
<figureCitation id="A2832697FFD0FF9DA51000CFFC52B4AD" box="[836,905,910,935]" captionStart="FIGURES 4 – 11. 4" captionStartId="2.[151,264,1887,1910]" captionTargetBox="[172,1415,1139,1867]" captionTargetId="figure@2.[172,1416,1139,1868]" captionTargetPageId="2" captionText="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="4" pageNumber="381">Fig. 7</figureCitation>
) in possessing the deeply invaginated anterior margin of the epigyne and the distinctly broad lateral lobes. They can be distinguished by the longer median groove reaching the posterior margin of the epigynal septum (
<figureCitation id="A2832697FFD0FF9DA5440094FC8EB4E4" box="[784,853,981,1006]" captionStart="FIGURES 4 – 11. 4" captionStartId="2.[151,264,1887,1910]" captionTargetBox="[172,1415,1139,1867]" captionTargetId="figure@2.[172,1416,1139,1868]" captionTargetPageId="2" captionText="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="4" pageNumber="381">Fig. 8</figureCitation>
) and by the different configuration of the internal genitalia (
<figureCitation id="A2832697FFD0FF9DA75200B9FE52B31B" box="[262,393,1016,1041]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="4" pageNumber="381">Figs 18–21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72A26999FFD0FF9DA693075AFC0AB35D" pageId="4" pageNumber="381" type="etymology">
<paragraph id="3A073A12FFD0FF9DA693075AFC0AB35D" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA693075AFE95B33E" bold="true" box="[199,334,1051,1076]" pageId="4" pageNumber="381">Etymology.</emphasis>
The specific epithet is derived from the 
<typeStatus id="E50384B0FFD0FF9DA575075DFC8AB33E" box="[801,849,1052,1076]" pageId="4" pageNumber="381">type</typeStatus>
locality, Tasek Merimbun, a wildlife sanctuary and designated ASEAN Heritage Park in Tutong District, 
<collectingCountry id="42AF7A82FFD0FF9DA4A5077FFC16B35D" box="[753,973,1086,1111]" name="Brunei" pageId="4" pageNumber="381">Brunei Darussalam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="72A26999FFD0FF9CA6930723FDBDB58E" lastPageId="5" lastPageNumber="382" pageId="4" pageNumber="381" type="description">
<paragraph id="3A073A12FFD0FF9DA6930723FDBFB221" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA6930723FDCEB371" bold="true" box="[199,533,1122,1147]" pageId="4" pageNumber="381">
Description. Male 
<typeStatus id="E50384B0FFD0FF9DA7F30723FDCBB371" box="[423,528,1122,1147]" pageId="4" pageNumber="381" type="holotype">holotype</typeStatus>
.
</emphasis>
Total length 6.9; prosoma 3.1 long, 2.8 wide; opisthosoma 3.8 long, 2.6 wide. Leg measurements: I 13.0 (3.4, 0.8, 3.2, 3.4, 2.2), II 12.2 (3.2, 0.8, 3.0, 3.2, 2.0), III 12.0 (3.0, 0.6, 2.9, 3.1, 2.4), IV 16.2 (3.8, 0.8, 4.2, 4.4, 3.0). Eyes circular, pale, from above AER straight, PER strongly procurved. MOQ longer than wide, wider in front than behind. Eye sizes and interdistances: AME largest, others subequal; AME separated by their diameter or less, about their diameter to ALE; PME smallest, separated by 1.5 diameter, from PLE by 2.5 diameter; ALE from PLE by their radius.
</paragraph>
<paragraph id="3A073A12FFD0FF9DA6930674FE17B2F4" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA6930674FE14B247" box="[199,463,1333,1357]" italics="true" pageId="4" pageNumber="381">Coloration and pattern</emphasis>
. Prosoma strongly convex, relatively high, widest at coxae II, in profile highest mid distance between PME and fovea, integument smooth. Fovea deep, longitudinal. Carapace dark reddish brown, distinctly darker peripherally. Chilum trapezoidal, orange, lined with few hairs. Chelicerae, labium and sternum reddish brown. Gnathocoxae brown, pale distally. Leg coxae white; femora dark green, proximally with white dorsoprolateral band running obliquely; dark, circular tubercle present on dorsoproximal portion of legs I–III (
<figureCitation id="A2832697FFD0FF9DA33B0683FF73B2F4" captionStart="FIGURES 4 – 11. 4" captionStartId="2.[151,264,1887,1910]" captionTargetBox="[172,1415,1139,1867]" captionTargetId="figure@2.[172,1416,1139,1868]" captionTargetPageId="2" captionText="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="4" pageNumber="381">Fig. 4</figureCitation>
); patellae to tibiae brown.
</paragraph>
<paragraph id="3A073A12FFD0FF9DA6930549FD72B181" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">Opisthosoma sepia, mottled with white spots: Medially with five pairs, first pair represented by two irregularshaped spots, second to fifth pairs by oval spots; peripherally with three pairs of oblong spots; posteriorly with two distinctly large spots situated behind fifth median pair. Dorsal scutum dark brown, calcareous, represented by longitudinal band with mesolateral concavities.</paragraph>
<paragraph id="3A073A12FFD0FF9DA69305D4FC5CB08B" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA69305D4FF27B1A7" box="[199,252,1685,1709]" italics="true" pageId="4" pageNumber="381">Palp</emphasis>
(
<figureCitation id="A2832697FFD0FF9DA75905D4FE49B1A4" box="[269,402,1685,1710]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="4" pageNumber="381">Figs 15–17</figureCitation>
). Palpal tibia relatively short, with broad ventral ridge. RTA digitiform, short and slender, clearly separable from palpal tibia in ventral view, gradually tapering towards blunt apex in retrolateral view. Tegular spine absent. TA rostrate; bifid apical process elongated, obliquely directed mesad, with mesal branch twice as long as lateral branch; apico-retrolateral flange subtriangular, curved inwards, apex pointing prolaterad; posterior tooth triangular, apex sharply pointed, directed anteriad. Embolic base originating at 270°. Embolus broad, with basal groove, divided into two rami near base; mesal ramus subterminally narrow, its extremity bladelike; lateral ramus distally expand, resulting in beak-shaped flange.
</paragraph>
<paragraph id="3A073A12FFD0FF9DA69304CAFE43B0E0" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="381">
<emphasis id="08CCE600FFD0FF9DA69304CAFE45B0AE" bold="true" box="[199,414,1931,1956]" pageId="4" pageNumber="381">
Female 
<typeStatus id="E50384B0FFD0FF9DA77804CAFE41B0AE" box="[300,410,1931,1956]" pageId="4" pageNumber="381" type="paratype">paratype</typeStatus>
.
</emphasis>
Total length 7.6; prosoma 3.6 long, 3.0 wide; opisthosoma 4.0 long, 3.4 wide. Leg measurements: I 13.5 (3.5, 0.8, 3.0, 3.4, 2.8), II 12.3 (3.3, 0.8, 2.6, 3.2, 2.4), III 11.9 (3.0, 0.6, 3.0, 3.1, 2.2), IV 16.2 (3.8, 0.8, 4.0, 4.6, 3.0).
</paragraph>
<paragraph id="3A073A12FFD1FF9CA69303D9FC08B7F2" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="382">
<emphasis id="08CCE600FFD1FF9CA69303D9FE15B7BA" box="[199,462,152,176]" italics="true" pageId="5" pageNumber="382">Coloration and pattern</emphasis>
(
<figureCitation id="A2832697FFD1FF9CA78B03D6FDFEB7BA" box="[479,549,151,176]" captionStart="FIGURES 1 – 3. 1 – 3" captionStartId="2.[151,264,1049,1072]" captionTargetBox="[177,1418,195,1028]" captionTargetId="figure@2.[175,1419,195,1029]" captionTargetPageId="2" captionText="FIGURES 1 – 3. 1 – 3. Habitus. 1. Mallinella myrmecophaga sp. nov., female holotype. 2. M. platyrhyncha sp. nov., female holotype. 3. M. merimbunenis sp. nov., female paratype." httpUri="https://zenodo.org/record/225301/files/figure.png" pageId="5" pageNumber="382">Fig. 3</figureCitation>
). Carapace dark chestnut brown. Chelicerae, mouthparts and sternum orangebrown. Legs with pale yellow coxae; femora bicolored, proximal portion pale, median and distal portion dark green, without semi-circular tubercle on legs I–IV; other segments brown.
</paragraph>
<paragraph id="3A073A12FFD1FF9CA6930245FAABB682" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="382">Opisthosoma almost black, mottled with white spots: Medially with five pairs, first to fourth pairs represented by oblique oval spots, spots of first and second pairs widely separated, spots of third and fourth pairs situated close together, fifth pairs onwards fused together, represented by short median band; peripherally with three pairs of oblong spots; posteriorly with two distinctly large spots situated behind fused sixth pair. Dorsal scutum absent.</paragraph>
<paragraph id="3A073A12FFD1FF9CA69302D5FD60B512" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="382">
<emphasis id="08CCE600FFD1FF9CA69302D5FEEAB6A6" box="[199,305,404,428]" italics="true" pageId="5" pageNumber="382">Genitalia</emphasis>
(
<figureCitation id="A2832697FFD1FF9CA71402D5FE53B6A6" box="[320,392,404,429]" captionStart="FIGURES 4 – 11. 4" captionStartId="2.[151,264,1887,1910]" captionTargetBox="[172,1415,1139,1867]" captionTargetId="figure@2.[172,1416,1139,1868]" captionTargetPageId="2" captionText="FIGURES 4 – 11. 4. Leg I – III, showing tubercle (indicated by arrow). 5 – 9, 11. Epigyne, ventral. 10. Internal female genitalia, anterior. 4. Mallinella merimbunenis sp. nov., male holotype. 5. M. bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new material from Brunei. 6. M. bigemina, female holotype. 7. M. albomaculata (Bosmans &amp; Hillyard, 1990), female paratype. 8. M. merimbunenis sp. nov., female paratype. 9 – 10. M. myrmecophaga sp. nov., female holotype. 11. M. platyrhyncha sp. nov., female holotype." httpUri="https://zenodo.org/record/225302/files/figure.png" pageId="5" pageNumber="382">Figs 8</figureCitation>
, 
<figureCitation id="A2832697FFD1FF9CA7C102D5FE04B6A6" box="[405,479,404,428]" captionStart="FIGURES 12 – 21. 12, 18" captionStartId="3.[151,264,1862,1885]" captionTargetBox="[167,1417,236,1838]" captionTargetId="figure@3.[151,1436,225,1841]" captionTargetPageId="3" captionText="FIGURES 12 – 21. 12, 18. Internal female genitalia, dorsal. 20. Ditto, lateral. 13. Left spermatheca, anterior. 14, 21. Right spermatheca, contralateral. 15. Male palp, prolateral. 16. Ditto, ventral. 17. Ditto, retrolateral. 19. Right spermatheca, anterior. 12 – 14. Mallinella bigemina Dankittipakul, Jocqué &amp; Singtripop, 2012, new female from Brunei. 15 – 17. M. merimbunenis sp. nov., male holotype. 18 – 21. M. merimbunenis sp. nov., female paratype. Scale bar: 0.1 mm (15 – 17); 0.25 mm (12 – 14, 18 – 21)." httpUri="https://zenodo.org/record/225303/files/figure.png" pageId="5" pageNumber="382">18–21</figureCitation>
). Epigynal plate bipartite, deeply invaginated anteriorly, with median incision almost reaching rebordered posterior margin. Lateral borders distinctly enlarged, extending inwards, apex broad and blunt. Insemination ducts relatively short, almost indistinct. Spermathecae heavily sclerotised, pyriform in ventral view, with small circular elevation situated posteriorly.
</paragraph>
<paragraph id="3A073A12FFD1FF9CA6930165FDBDB58E" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="382">
<emphasis id="08CCE600FFD1FF9CA6930165FE5EB537" bold="true" box="[199,389,548,573]" pageId="5" pageNumber="382">Natural history.</emphasis>
<typeStatus id="E50384B0FFD1FF9CA7D90164FE1EB537" box="[397,453,549,573]" pageId="5" pageNumber="382">Type</typeStatus>
specimens were invariably collected by beating foliage along hiking trails at eye level. A more thorough search is required to investigate if they also inhabit the litter layer, as all other 
<taxonomicName id="FDB84191FFD1FF9CA3730109FA47B56A" box="[1319,1436,584,608]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="08CCE600FFD1FF9CA3730109FA47B56A" box="[1319,1436,584,608]" italics="true" pageId="5" pageNumber="382">Mallinella</emphasis>
</taxonomicName>
reportedly do (
<bibRefCitation id="5E2947E3FFD1FF9CA76F012DFD81B58E" author="Dankittipakul" box="[315,602,620,645]" pageId="5" pageNumber="382" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella, Strand, 1906, with description of new genera and new species from Southeast Asia (Araneae, Zodariidae). Zootaxa, 3369, 1 - 327." type="journal article" year="2012">
Dankittipakul 
<emphasis id="08CCE600FFD1FF9CA78A012CFDD4B58E" box="[478,527,620,644]" italics="true" pageId="5" pageNumber="382">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72A26999FFD1FF9CA69301CEFB1BB41F" pageId="5" pageNumber="382" type="distribution">
<paragraph id="3A073A12FFD1FF9CA69301CEFB1BB41F" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="382">
<emphasis id="08CCE600FFD1FF9CA69301CEFEBBB5A2" bold="true" box="[199,352,655,680]" pageId="5" pageNumber="382">Distribution.</emphasis>
The species is common and widespread in primary and secondary forests in all four districts in 
<collectingCountry id="42AF7A82FFD1FF9CA6C301F5FF3DB5C7" box="[151,230,692,717]" name="Brunei" pageId="5" pageNumber="382">Brunei</collectingCountry>
, viz., 
<taxonomicName id="FDB84191FFD1FF9CA76501F5FDEAB5C7" authority="Seri Begawan" authorityName="Seri Begawan" box="[305,561,692,717]" class="Insecta" family="Cerambycidae" genus="Bandar" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="genus">Bandar Seri Begawan</taxonomicName>
, Tutong, Temburong and Belait (juveniles only, not listed under “Material Examined”). Given the fact that it is so common in 
<collectingCountry id="42AF7A82FFD1FF9CA48B0196FCF5B5FA" box="[735,814,727,752]" name="Brunei" pageId="5" pageNumber="382">Brunei</collectingCountry>
, it will be not surprising that it will be found at least in Malaysia’s 
<collectingCountry id="42AF7A82FFD1FF9CA74301BDFEA1B41F" box="[279,378,764,789]" name="Malaysia" pageId="5" pageNumber="382">Sarawak</collectingCountry>
, as part of it straddles between Brunei’s districts of Belait and Temburong.
</paragraph>
</subSubSection>
</treatment>
</document>