183 lines
18 KiB
XML
183 lines
18 KiB
XML
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<mods:title id="A6A7165B2E326B6CCF4F9DA8CAF22856">New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)</mods:title>
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<mods:namePart id="E584E62E830BC7D9E220A1C666C1A95A">Perrichot, Vincent</mods:namePart>
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<mods:namePart id="7AEAF5117399824C74A33A13887C7ECF">Ortega-Blanco, Jaime</mods:namePart>
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<mods:namePart id="7D00276186493C98ADA8D3B0C576FCFA">McKellar, Ryan C.</mods:namePart>
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<mods:namePart id="3176D52F88A76B41BE35C5FC8EF05A6F">Delclos, Xavier</mods:namePart>
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<mods:namePart id="56754542EEA83099879C343D037AE2CD">Azar, Dany</mods:namePart>
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<mods:namePart id="CC927529FAE5CCDBB28A0C4319570E76">Nel, Andre</mods:namePart>
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<mods:namePart id="21BDAE3D2FE12341B718A21CA1D9FF4C">Tafforeau, Paul</mods:namePart>
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<mods:namePart id="7D3967B38A4BD42155E068B7A5CBE269">Engel, Michael S.</mods:namePart>
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<mods:title id="6C86F59E043C6C05020D47919ACDB6D9">ZooKeys</mods:title>
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<mods:date id="5E4BF80FD291F03F6082699AA1E00A2C">2011</mods:date>
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<mods:number id="115B8501E17F27C8BC6D0D5DF1B9F954">130</mods:number>
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<mods:url id="C484D29D78944A9CE82D84A5E4A34081">http://dx.doi.org/10.3897/zookeys.130.1453</mods:url>
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<mods:classification id="AFB16B20AECDF4AC088C12A2A9EFC719">journal article</mods:classification>
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<mods:identifier id="943DE857D083311B8A99ECCDE489B678" type="DOI">http://dx.doi.org/10.3897/zookeys.130.1453</mods:identifier>
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<mods:identifier id="A21B9C058E161B775C9F87AB36227904" type="Pensoft-Pub">1313-2970-130-421</mods:identifier>
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<treatment id="B1F3C15D03A6D4CD8AFE538755050FE3" ID-GBIF-Taxon="152030994" LSID="urn:lsid:plazi:treatment:B1F3C15D03A6D4CD8AFE538755050FE3" httpUri="http://treatment.plazi.org/id/B1F3C15D03A6D4CD8AFE538755050FE3" lastPageId="8" lastPageNumber="429" pageId="5" pageNumber="426" scope_family="Maimetshidae">
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<subSubSection id="87C44F7AF2952D17E16A91C331B299B8" pageId="5" pageNumber="426" type="nomenclature">
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<paragraph id="E7520DD65D83BF3F163D85FFF9F97690" pageId="5" pageNumber="426">
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<taxonomicName id="248D612C2191415B8CAF8597010782F2" LSID="http://species-id.net/wiki/Guyotemaimetsha_enigmatica" authority="Perrichot, Nel & Neraudeau, 2004 a" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">
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<pageBreakToken id="2CEECFCE9264DCCB2320B0ED05B990D0" pageId="5" pageNumber="426" start="start">Guyotemaimetsha</pageBreakToken>
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enigmatica Perrichot, Nel &
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<normalizedToken id="E535A71469DAFE9EFBAF87DF3AE60414" originalValue="Néraudeau">Neraudeau</normalizedToken>
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, 2004a
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</taxonomicName>
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Figs 14
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</paragraph>
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</subSubSection>
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<subSubSection id="5D22EEA9BF4B1270806040A928A747ED" pageId="5" pageNumber="426" type="reference_group">
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<paragraph id="04174335BAF9863C51575E96DF1529C0" pageId="5" pageNumber="426">
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<taxonomicName id="9CBA6292F0FB2438EC4E3274716D7061" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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Perrichot, Nel &
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<normalizedToken id="8E95F73D26C74F86ACD2FFA2849A7CAE" originalValue="Néraudeau">Neraudeau</normalizedToken>
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, 2004a: 159, figs. 1A-B, 2 [♂].
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</paragraph>
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</subSubSection>
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<subSubSection id="3F39352330929A378D767EB5E497EEFC" pageId="5" pageNumber="426" type="type material">
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<paragraph id="F9521E5936A9714B57A0D8658FCA1EE7" pageId="5" pageNumber="426">Type material.</paragraph>
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<paragraph id="26C4D9EFD64E6F0F2802A04EDC063179" pageId="5" pageNumber="426">
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Holotype MNHN.F.A30174 (♂); paratype MNHN.F.A30175 (♂); in amber collection of the Museum National
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<normalizedToken id="4A8DFC87D6CFB4DB4A7312EBC5DC6124" originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle, Paris, France.
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</paragraph>
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</subSubSection>
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<subSubSection id="11BA643E74651F4B3C80D5BA093480B2" pageId="5" pageNumber="426" type="other material examined">
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<paragraph id="C02CDA9F0B8442E331634FBDCE0ABBB5" pageId="5" pageNumber="426">Other material examined.</paragraph>
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<paragraph id="132A80E82D10211C1B822A1EE7996C20" pageId="5" pageNumber="426">Specimens IGR.ARC-1 (♀, missing anterior part of the head and with cuticle somewhat altered by taphonomy), IGR.ARC-309.11 (sex unknown, missing apices of right forewing and metasoma), IGR.ARC-370.7 (= scan ESRF-A-018g, reconstructed, ♀, in opaque amber), IGR.ARC-378.1 (♂, missing head, most metasoma and apices of all legs), IGR.ARC-378.2 (scan ERSF A-401-378, not reconstructed; ♂), IGR.ARC-393.4 (= radiograph ESRF-A-053d, not reconstructed; ♂), IGR.CDL-2.5 (♂, with forewing fragment in IGR.CDL-2.16, missing apical flagellomeres of left antenna; ex collection Arnaud), IGR.CDL-2.33 (♂, missing left antenna and apices of right antenna, right forewing, and all legs; ex collection Arnaud). All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.</paragraph>
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</subSubSection>
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<subSubSection id="9D2758F6B481F83BC3E572C9ED8FB126" pageId="5" pageNumber="426" type="indeterminate specimens">
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<paragraph id="D42EF95621E50A7349D364DF00314D94" pageId="5" pageNumber="426">Indeterminate specimens.</paragraph>
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<paragraph id="85128FF2D0900425B05EBE5525086593" pageId="5" pageNumber="426">
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IGR.ARC-171 (♂, missing head, anterodorsal part of metasoma, and forelegs); IGR.ARC-191 (♂?, missing part of the head and mostly obscured, preserved in very dark amber); IGR.ARC-275 (♀, missing parts of mesosoma, left legs, and with metasoma mostly obscured by a bacterial film). These specimens agree in most respects with the definition of
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<taxonomicName id="AC8E3DE17F04B8781C5DFD286741894F" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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but little is preserved and attribution to this species cannot be asserted. All specimens deposited in the amber collection of the Department of Geosciences of the University Rennes I, France.
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</paragraph>
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<caption id="1B5D3943DD097148C7DA912210BFE96C" pageId="5" pageNumber="426">
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<paragraph id="8A3E623114FE4F5DEFF64F3A90E7D24C" pageId="5" pageNumber="426">
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Figure 1. Photomicrographs of male of
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<taxonomicName id="02F537C8855778054242B383E619CB2C" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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Perrichot et al., 2004a (IGR.CDL-2.5) A Left lateral habitus B Right lateral habitus.
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</paragraph>
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</caption>
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<caption id="C33E5A3C1F5A0CF78E88BEAA9EE5F617" pageId="5" pageNumber="426">
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<paragraph id="FEAE070920022B8EC20C7C19C7E88DA8" pageId="5" pageNumber="426">
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Figure 2. Photomicrograph detail of male of
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<taxonomicName id="248F3C3818237059713C7D975B6B9DA8" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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Perrichot et al., 2004a (IGR.CDL-2.5).
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</paragraph>
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</caption>
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<caption id="8EAC6FAF78BBBBFE74DED98F07AD5373" pageId="5" pageNumber="426">
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<paragraph id="BE14079E187336705FA278E360E18340" pageId="5" pageNumber="426">
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Figure 3. Forewing venation of
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<taxonomicName id="8ACE00FE6763CD78D13BC152D3E73F31" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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Perrichot et al., 2004a (paratype MNHN-A30175), annotated with vein and cell nomenclature employed herein.
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</paragraph>
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</caption>
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<caption id="EB82711E45F8AB98EBDCE974EEAF9D3F" pageId="5" pageNumber="426">
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<paragraph id="3A17CF50B41B643A7131834AD35A08F5" pageId="5" pageNumber="426">
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Figure 4. 3D virtual extraction of female of
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<taxonomicName id="1A46339BCD83D23B4BC90A65EB08BE00" genus="Guyotemaimetsha" lsidName="Guyotemaimetsha enigmatica" pageId="5" pageNumber="426" rank="species" species="enigmatica">Guyotemaimetsha enigmatica</taxonomicName>
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Perrichot et al., 2004a (IGR.ARC-370.7, scan ESRF-A-018g), using
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<normalizedToken id="53E8D7E03AA102C35B8561932A967B9F" originalValue="PPC-SRµCT">PPC-SRµCT</normalizedToken>
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(voxel size 5.06
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<normalizedToken id="469E548D1EFC7A68BDAA3C2E9FA90ED6" originalValue="µm">µm</normalizedToken>
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, propagation distance 990 mm, 30 keV) A Dorsal habitus B Left lateral habitus C Facial aspect D Right lateral habitus. The arrows indicate tarsal plantulae.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="671BF78FC845E5C089FA82982D5C4897" pageId="5" pageNumber="426" type="type locality">
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<paragraph id="CC726624EA8FDE6A0F2F744791B628B2" pageId="5" pageNumber="426">Type locality.</paragraph>
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<paragraph id="F16467B0AC98AED580B0D3497F1EF515" pageId="5" pageNumber="426">Font-de-Benon quarry, 1 km east of Archingeay, Charente-Maritime, France.</paragraph>
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</subSubSection>
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<subSubSection id="58A556F17BCB867432E19D584FE39B33" pageId="5" pageNumber="426" type="stratigraphic horizon">
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<paragraph id="5BDF6C2BD420BEB41BEE3D193DF398E4" pageId="5" pageNumber="426">Stratigraphic horizon.</paragraph>
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<paragraph id="E6F66B59A4A7A88310F0FEBC4CE04CEA" pageId="5" pageNumber="426">
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Lithological subunit A1sl-A, Uppermost
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<normalizedToken id="14BBA295712AA3F82C7BE96C055ABF94" originalValue="Albian–Lowermost">Albian-Lowermost</normalizedToken>
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Cenomanian, middle Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="D84882753450F71DD2822325069EBB51" pageId="5" pageNumber="426" type="diagnosis">
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<paragraph id="2ADE84CFFBDBC535ADD05EB91FF4629F" pageId="5" pageNumber="426">Diagnosis.</paragraph>
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<paragraph id="4F90F07C2A087171B711D0AB599CFED9" pageId="5" pageNumber="426">As for the genus (see above).</paragraph>
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</subSubSection>
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<subSubSection id="5A206AD9C6F2CF37E281BFF133215759" lastPageId="8" lastPageNumber="429" pageId="5" pageNumber="426" type="redescription">
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<paragraph id="E748A11FABA168CF80C3AABE44687480" pageId="5" pageNumber="426">Redescription.</paragraph>
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<paragraph id="E0B7BA0DC2D3CFDCE3BF759A52F2DA34" lastPageId="8" lastPageNumber="429" pageId="5" pageNumber="426">
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Male: Integument reddish-brown in color, with dense, minute punctures and fine, decumbent pubescence. Head transverse, with length approximately 0.8x width; ocelli separated from each other by one ocellar diameter; antennae inserted between compound eyes, closer to each other than to compound eye margin or posterior clypeal margin; scape globular in lateral view, laterally compressed in frontal view, with faintly concave apex; pedicel with sides convex; flagellum with dense coat of short, inclined setae, with first article longest, twice length of pedicel; penultimate article as broad as long; apical article with rounded apex; clypeus transverse, about twice as broad as long, with anterior margin rounded and posterior margin straight; mandibles large, with outer margin convex, overlapping apically when closed; right
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<pageBreakToken id="840DC5A947F5392730EFC70777CEA432" pageId="6" pageNumber="427" start="start">mandible</pageBreakToken>
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with basal tooth large, second and third teeth equally smaller, apical tooth largest; left mandible with basal tooth large, median tooth smallest, apical tooth largest. Mesosoma compact, 0.7x as high as long; pronotum in dorsal view reduced to a short neck; mesoscutum broader than long, about 0.4x mesosomal length, abruptely truncate anteriorly to form a flat vertical surface, bordered by a transverse carina at anterolateral corners, with low dorsal convexity; median mesoscutal sulcus crenulate, notauli deeply impressed, slightly diverging anteriorly; mesoscutellum with low dorsal convexity, small triangular axillae separated anteriorly by large foveate groove; propodeum areolate, gradually sloping to pronounced posterior lip; meso- and metapleuro
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<pageBreakToken id="51BDD80CF94FE0BBA625DCBCEB44A13C" pageId="7" pageNumber="428" start="start">n</pageBreakToken>
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with anterior margin foveolate. Forewing hyaline, with costal cell distinctly enlarged around midlength; Rs junction with Sc+R forming an acute angle; cell [1M] with vein M slightly arched, roughly aligned with Rs, vein Cu shortest, M and Rs+M subequal in length; cells [1M] and [1Rs] well distant from each other, separated by long vein 2Rs+M; 2Cu significantly longer than 1Cu and 1m-cu; M and Cu nebulous at apex, reaching wing margin; 2A fading to nebulous vein posterior of [2Cu]. Hind wing with Sc+R, R, M+Cu, Cu, and A tubular, other veins nebulous; Sc+R and R more heavily sclerotized than M+Cu, Cu, and A; Cu fading to nebulous vein apically; free apex of Rs and Cu short. Metacoxa approximately 1.4x length of pro-and mesocoxae, all with broad bases and distinct apical constriction, in lateral view inserted posteroventrally on mesosoma; metafemur distinctly enlarged around midlength; all tibiae approximately subequal to femora in length, with basitarsus length slightly less than combined length
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<pageBreakToken id="C8F0BC4C7269F35BD5C067784B2FCA14" pageId="8" pageNumber="429" start="start">of</pageBreakToken>
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all subsequent tarsomeres. Metasoma compact, as long as mesosoma; parameres trapezoidal, narrowed toward apex, with outer surface slightly convex, with short erect setae around apex; cerci small, spatulate, inserted just anterior to parameres.
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</paragraph>
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<paragraph id="15556C7523A3FA479F5B3A65710EB14A" pageId="8" pageNumber="429">
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Female: Very similar to male but with pedicel cylindrical and each leg with a small plantar lobe (or tarsal plantula) on ventral apex of tarsomeres
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<normalizedToken id="5143DA57949EEF0369484309156D3B20" originalValue="I–IV">I-IV</normalizedToken>
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; ovipositor approximately as long as metasomal length, apex acute. Integument sculpturing and pubescence not visible, altered on specimen IGR.ARC.1 and not reconstructed in microtomographic scan of specimen IGR.ARC-370.7.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |