153 lines
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153 lines
20 KiB
XML
<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="B1B4E121EFDF5693B99A96A381BC23B1" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina brachysoma Ancey 1904" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:titleInfo id="66B6A2F119B04BB5533F04C7BFD76134">
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<mods:title id="D99D78BE3AEF72A4800DF8C4E1732E9B">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name id="D322330A9B87C47E010E031758D6B949" type="personal">
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<mods:namePart id="7F062D61164DA3A0DDD4DB64D4612700">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="C4454CA493236A1332A92A55697AC05B" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="99012EE68B68F63F606246CEC4A0E4C9">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="EB2EAD284D777067E20A1915156854D7" type="personal">
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<mods:roleTerm id="FD3DE208DEDC386C462594519F1A2541">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A4341CC9FC72849A3AD8CF45CC8635CA">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="094B89C192DFD0CCC2F71EA2AD349D53">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="F98AE5B2202D6AAF34417C053DA51629" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:name id="55C6C0B9F16E4AD7CC67EB3ADB5A764E" type="personal">
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<mods:namePart id="FCF1B8E149F65EC32FB8749DD88085FD">Panha, Somsak</mods:namePart>
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<mods:affiliation id="1C6351CC4A53B92E8CAD52046080EE7E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="78A12F8E0DC9AA72D30BABCE484063BE" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="35F3F2EB4D8CA1DCDA41692D174B3562">text</mods:typeOfResource>
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<mods:title id="A57AD9D22AEB7034C011AF9E63C6C186">ZooKeys</mods:title>
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<mods:part id="7F3E202B9C0F408407E23EC62260A477">
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<mods:date id="FCB775FEF2F77D76AF7E0BBF0A6497AA">2022</mods:date>
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<mods:number id="1B66E6B428E20256DF08CA70E7547CB1">2022-08-25</mods:number>
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<mods:detail id="23CA93D7C9CA678EAC456D45BB76ED5B" type="volume">
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<mods:number id="E5B0A9D5848643E9510E7F630A914381">1119</mods:number>
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<mods:start id="CCE42730DD710A4DA188D931C44EADA8">1</mods:start>
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<mods:end id="904FE090050AF7ACC40D30972FD67DE1">115</mods:end>
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<mods:url id="979CE0CEE890936098A0B92644EA21DB">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification id="D07A8C25D465981CDE6FDA68B3ECCF46">journal article</mods:classification>
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<mods:identifier id="E87CF5B09D51A1E978364823F4331063" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier id="7A9EEE1140F0BF972F6FED9B420931E7" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier id="2E09C6676A0A4E3ED03A16A3EFA55C51" type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<treatment id="B1B4E121EFDF5693B99A96A381BC23B1" ID-GBIF-Taxon="199838992" LSID="urn:lsid:plazi:treatment:B1B4E121EFDF5693B99A96A381BC23B1" httpUri="http://treatment.plazi.org/id/B1B4E121EFDF5693B99A96A381BC23B1" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="43B8751877B24929EDFDBA13BFBDC840" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="FB65948DAEFB0932B617FD1FC98589C5" pageId="0" pageNumber="1">
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<taxonomicName id="636B713A6FD8DE5306B4F637DC896499" ID-CoL="4QJQX" LSID="B1B4E121-EFDF-5693-B99A-96A381BC23B1" authority="Ancey, 1904" authorityName="Ancey" authorityYear="1904" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">Pupina brachysoma Ancey, 1904</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="E54D977CD56E722970975D3073C4D0DB" pageId="0" pageNumber="1" type="description">
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<paragraph id="C4E0ACE514AD74566AE6FDE73F7145E9" pageId="0" pageNumber="1">
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<figureCitation id="7974C324837EDDAD3BEFCA6A52811ED9" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">Figs 36M</figureCitation>
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<figureCitation id="7E6A95E151A6F506C60A88F6399EEA83" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. A Pupina dorri dorri, specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 from Haiphong, Vietnam B Pupina tongupensis, syntype NHMUK 1906.4.4.38 from Tongoop Pass, Arakan Hills, east side C Pupina anceyi, syntype MNHN-IM- 2000 - 35833 from Lang-Son and That-Khe D Pupina laffonti, syntype MNHN-IM- 2000 - 9656 from Poulo-Condor E Pupina solidula, lectotype SMF 109915 / 1 from Lang Son, Tonkin F Pupina brachysoma, syntype MNHN-IM- 2000 - 9652 from Haut Tonkin. Photo: J. Ablett, H. Taylor, NHM (B), P. Maestrati, M. Caballer, MNHN (C-F)." figureDoi="10.3897/zookeys.1119.85400.figure41" httpUri="https://binary.pensoft.net/fig/735402" pageId="0" pageNumber="1">, 41F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FDB9E3DFD79DE938DB00A9BEADFF2646" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="006363CC870E22CB6A6E80A4DB4A7D07" pageId="0" pageNumber="1">
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<taxonomicName id="1865F4D2E0568248ACDAB9974CED32A9" authorityName="Ancey" authorityYear="1904" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">Pupina brachysoma</taxonomicName>
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Ancey in Bavay & Dautzenberg, 1904 [1903]: 230, 231, pl. 10, figs 15, 16. Type locality: Haut-Tonkin [northern Vietnam].
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<bibRefCitation id="D437C42014C44F1FD407F12721C1D5BE" author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette 1950</bibRefCitation>
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: 171.
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<bibRefCitation id="C1A4EB38C075E893EF178A5910CECA07" author="Wood, H" journalOrPublisher="The Dorset Press, Dorchester" pageId="0" pageNumber="1" refId="B197" refString="Wood, H, Gallichan, J, 2008. The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3. The Dorset Press, Dorchester" title="The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3." year="2008">Wood and Gallichan 2008</bibRefCitation>
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: 31, pl. 25, figs 5, vi.
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</paragraph>
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</subSubSection>
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<subSubSection id="7FA28CD9D8B75A28AA7E337C685590A1" pageId="0" pageNumber="1" type="type material">
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<paragraph id="42E48340687D805E0450B4D0FC5211FD" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="DD9BB69F0A4B53193E598CF01160786F" pageId="0" pageNumber="1">
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<emphasis id="7CCA5F534402CE3C9978BD230F30BF2F" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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MNHN-IM-2000-9652 (Figs
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<figureCitation id="0DC1D46E3D2452D3A5858866C44FB541" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36M</figureCitation>
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,
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<figureCitation id="70E5F69BF44E5C7E86BBDF65DEE946F5" captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. A Pupina dorri dorri, specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 from Haiphong, Vietnam B Pupina tongupensis, syntype NHMUK 1906.4.4.38 from Tongoop Pass, Arakan Hills, east side C Pupina anceyi, syntype MNHN-IM- 2000 - 35833 from Lang-Son and That-Khe D Pupina laffonti, syntype MNHN-IM- 2000 - 9656 from Poulo-Condor E Pupina solidula, lectotype SMF 109915 / 1 from Lang Son, Tonkin F Pupina brachysoma, syntype MNHN-IM- 2000 - 9652 from Haut Tonkin. Photo: J. Ablett, H. Taylor, NHM (B), P. Maestrati, M. Caballer, MNHN (C-F)." figureDoi="10.3897/zookeys.1119.85400.figure41" httpUri="https://binary.pensoft.net/fig/735402" pageId="0" pageNumber="1">41F</figureCitation>
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) from Haut Tonkin. Paralectotypes NMW.1955.158.24153 figured in
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<bibRefCitation id="FEFABF1295C00F592F15DF220D7CDE66" author="Wood, H" journalOrPublisher="The Dorset Press, Dorchester" pageId="0" pageNumber="1" refId="B197" refString="Wood, H, Gallichan, J, 2008. The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3. The Dorset Press, Dorchester" title="The new molluscan names of Cesar-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organism from the National Museum of Wales. Biotir Report 3." year="2008">Wood and Gallichan (2008</bibRefCitation>
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: pl. 25, figs 5, vi).
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</paragraph>
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</subSubSection>
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<subSubSection id="10DEBE8CCA53027D86F33E17991AEEC2" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="1A5E82AED8342913B8131564EA1F962C" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="F7E5FC068FF1A4A5BA7865A787D73B76" pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened, slightly expanded. Parietal tooth sharp, tooth-like, thickened; columellar tooth fin-shaped, slightly thickened, located next to slit-like anterior canal. Posterior canal widened.</paragraph>
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</subSubSection>
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<subSubSection id="EE1AE904C1D796681C2F0F0C5EE89420" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="E7AC1673036E7C85D9DD266889032D9E" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="0AC6EA94FC2486A43F2D3965F3731764" pageId="0" pageNumber="1">
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<taxonomicName id="490DA90FB99E7557280352F71ADA541E" authorityName="Ancey" authorityYear="1904" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">
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<emphasis id="B4E49BE28760B33ACF6C842A8D83D547" italics="true" pageId="0" pageNumber="1">Pupina brachysoma</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName id="74EE761F0466A77FDBA73F24D0543993" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tchehelensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tchehelensis">
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<emphasis id="8A9CC9B23B5529FCB19CAD448D529ECD" italics="true" pageId="0" pageNumber="1">P. tchehelensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="6F47F0D99D44FCAB5DE46DD936616043" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. lowi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lowi">
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<emphasis id="4930F9EE19A2998525D238BF35DB4C5F" italics="true" pageId="0" pageNumber="1">P. lowi</emphasis>
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</taxonomicName>
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in having a sharp, tooth-like, thickened parietal tooth, a fin-shaped, thickened columellar tooth that is located next to a slit-like anterior canal, and a posterior canal that is gradually widening. However,
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<taxonomicName id="A551A34E7DDAAD19440E482D597B76A8" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">
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<emphasis id="0029A57500BDDF0601356C6C1915F428" italics="true" pageId="0" pageNumber="1">P. brachysoma</emphasis>
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</taxonomicName>
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is different from both
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<taxonomicName id="F02781A85621167BD7E49FDCC5F70912" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tchehelensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tchehelensis">
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<emphasis id="D56E6DFD30498B0F35E635B5DB7FDF09" italics="true" pageId="0" pageNumber="1">P. tchehelensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="3606ABE4F13421F8525F9A192F6F05C9" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. lowi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lowi">
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<emphasis id="E1416386469ED14C3496142C4048727C" italics="true" pageId="0" pageNumber="1">P. lowi</emphasis>
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</taxonomicName>
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by a more ovate-fusiform shell shape, and a less thickened but slightly expanded apertural lip.
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<taxonomicName id="3082F07DC719D3308B996B60152DFA32" authorityName="Ancey" authorityYear="1904" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">
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<emphasis id="8F35EACFCB5781482324B5249280B374" italics="true" pageId="0" pageNumber="1">Pupina brachysoma</emphasis>
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</taxonomicName>
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is also similar to
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<taxonomicName id="D7C490233B8C80409F9C644AE39088EC" authority="dorri" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. dorri subsp. dorri" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dorri" subSpecies="dorri">
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<emphasis id="0429C4C2C6DA88326833C03AEB38E242" italics="true" pageId="0" pageNumber="1">P. dorri dorri</emphasis>
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</taxonomicName>
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in shell shape, but differs in having a gradually widening posterior canal.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2C2C26809C832BE0842CF5A127E98F5" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="5C8D9429DC736FC830C934671596AFE3" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="51470529E7FE3809789F98DDEC64693F" pageId="0" pageNumber="1">
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Known only from the type locality (
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<bibRefCitation id="FBF054E8B006AD3E6CE1B932CA9289B2" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="201 - 236" refId="B7" refString="Bavay, A, Dautzenberg, P, 1904. Description de coquilles nouvelles de ľIndo-Chine (3e suite). Journal de Conchyliologie 51 (1903): 201 - 236" title="Description de coquilles nouvelles de ľIndo-Chine (3 e suite)." volume="51" year="1904">Bavay and Dautzenberg 1904</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="4C787B775D51B1BFAC0F1FD4C0C6BDCE" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="D49529E5E92425F4F9643E5C9F3A47EA" pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph id="2844380A8251845F351FF59E02B55B17" pageId="0" pageNumber="1">
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As the original description did not explicitly state that the description of this species was based on a single specimen (nor could this be inferred), the designation of a holotype by
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<bibRefCitation id="42CD6ADB3C4A47B7CC10B8FC16F384B4" author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette (1950)</bibRefCitation>
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in fact constitutes a lectotype designation (
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<bibRefCitation id="0E26F2C317C0B4DB6208F18C4C925898" author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
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: Art. 74.6).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |