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<document id="664D5BE99A21A904796A934148AB2565" ID-CLB-Dataset="54407" ID-DOI="10.5733/afin.049.0105" ID-GBIF-Dataset="01968b77-015c-45df-aa06-4a7b1090904e" ID-ISSN="2305-2562" ID-Zenodo-Dep="7667189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639426290" checkinUser="felipe" docAuthor="Morita, Shelah" docDate="2008" docId="B17087D2593AFFF3300BFE6FFEE4FC0F" docLanguage="en" docName="AfrInvertebr.49.1.129-158.pdf" docOrigin="African Invertebrates 49 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Philoliche (Philoliche) umbratipennis Morita 2008, stat. rev." docType="treatment" docVersion="7" lastPageNumber="148" masterDocId="4D49FFAA5928FFE73120FF81FFD8FFD3" masterDocTitle="A revision of the Philoliche aethiopica species complex (Diptera: Tabanidae)" masterLastPageNumber="158" masterPageNumber="129" pageNumber="146" updateTime="1699475318295" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="30EC9001ED6B5CE9287C05BD1C501945">A revision of the Philoliche aethiopica species complex (Diptera: Tabanidae)</mods:title>
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<mods:namePart id="7BA33032B6682FC770C7502AD1C4DB01">Morita, Shelah</mods:namePart>
<mods:affiliation id="B86B06237EF0B2719096FA71E4D20869">Population Biology Graduate Group, University of California at Davis, 1 Shields Ave., Davis, CA 95616, USA; Current address: Department of Botany &amp; Zoology, Stellenbosch University, P. Bag X 1, Matieland, 7602 South Africa</mods:affiliation>
<mods:nameIdentifier id="7DEB4E7A9D44011B349931454BB69E81" type="email">simorita@ncsu.edu</mods:nameIdentifier>
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<mods:title id="5F9B6FDD4D0EE7A21353B3706CBE3BB4">African Invertebrates</mods:title>
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<mods:date id="AB8DCBD0BCDB55A4FC2F9445576003F8">2008</mods:date>
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<treatment id="B17087D2593AFFF3300BFE6FFEE4FC0F" ID-DOI="http://doi.org/10.5281/zenodo.7667608" ID-GBIF-Taxon="209340607" ID-Zenodo-Dep="7667608" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F" httpUri="http://treatment.plazi.org/id/B17087D2593AFFF3300BFE6FFEE4FC0F" lastPageId="20" lastPageNumber="148" pageId="18" pageNumber="146">
<subSubSection id="71C3654F593AFFF5300BFE6FFBD8FDD4" box="[299,1024,494,519]" pageId="18" pageNumber="146" type="nomenclature">
<paragraph id="396636C4593AFFF5300BFE6FFBD8FDD4" blockId="18.[299,1024,494,519]" box="[299,1024,494,519]" pageId="18" pageNumber="146">
<heading id="622E81A8593AFFF5300BFE6FFBD8FDD4" box="[299,1024,494,519]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="146" reason="3">
<taxonomicName id="FED94D47593AFFF5300BFE6FFC57FDD4" ID-CoL="88D3K" authority="(Ricardo, 1900)" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" baseAuthorityYear="1900" box="[299,911,494,519]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="umbratipennis" status="stat. nov." subGenus="Philoliche">
<emphasis id="0BADEAD6593AFFF5300BFE6FFE46FDD5" box="[299,414,494,518]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
(
<emphasis id="0BADEAD6593AFFF5308FFE6FFDFAFDD5" box="[431,546,494,518]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
)
<emphasis id="0BADEAD6593AFFF53312FE6FFD0CFDD5" box="[562,724,494,518]" italics="true" pageId="18" pageNumber="146">umbratipennis</emphasis>
(
<bibRefCitation id="5D484B35593AFFF533C3FE6FFC5FFDD4" author="RICARDO, G." box="[739,903,494,519]" pageId="18" pageNumber="146" pagination="167 - 182" refId="ref22149" refString="RICARDO, G. 1900. Notes on the Pangoniinae of the family Tabanidae in the British Museum collection. Annals &amp; Magazine of Natural History (7) 5: 97 - 121, 167 - 182." type="journal article" year="1900">Ricardo, 1900</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="0BADEAD6593AFFF532BAFE6FFBD8FDD4" bold="true" box="[922,1024,494,519]" pageId="18" pageNumber="146">
<taxonomicNameLabel id="109E57AD593AFFF532BAFE6FFBD8FDD4" box="[922,1024,494,519]" pageId="18" pageNumber="146">stat. rev.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593AFFF53370FD97FD03FDFC" box="[592,731,534,559]" pageId="18" pageNumber="146" type="description">
<paragraph id="396636C4593AFFF53370FD97FD03FDFC" blockId="18.[181,1142,534,1671]" box="[592,731,534,559]" pageId="18" pageNumber="146">
<figureCitation id="A1E22A41593AFFF53370FD97FD7DFDFC" box="[592,677,534,559]" captionStart="Fig" captionStartId="10.[182,215,1548,1568]" captionTargetBox="[212,1113,224,1514]" captionTargetId="figure-85@10.[212,1113,209,1526]" captionTargetPageId="10" captionText="Fig. 5. (a) Morphotypes of Philoliche aethiopica. From left to right: Philoliche aethiopica Natal (female and male), P. aethiopica Albany (female and male). Note that the Natal form pictured here has a white stripe formed by a continuous fringe of white hairs on the posterior margin of the fourth abdominal segment. (b) Females of the rondani species group. From left to right: Philoliche umbratipennis stat. rev., P. rondani, P. rubiginosa, and P. elegans." figureDoi="http://doi.org/10.5281/zenodo.7667203" httpUri="https://zenodo.org/record/7667203/files/figure.png" pageId="18" pageNumber="146">Figs 5b</figureCitation>
,
<figureCitation id="A1E22A41593AFFF53391FD97FD18FDFC" box="[689,704,534,559]" captionStart="Fig" captionStartId="17.[144,177,897,917]" captionTargetBox="[164,1084,203,882]" captionTargetId="figure-317@17.[164,1084,205,876]" captionTargetPageId="17" captionText="Fig. 8. Geographic distribution of the rondani species group over South African provinces (GP Gauteng, MPU Mpumalanga, KZN KwaZulu-Natal) and in neighbouring countries (SW Swaziland). Note that all taxa overlap in distribution in the Eastern Cape, with the exception of Philoliche rubiginosa." figureDoi="http://doi.org/10.5281/zenodo.7667209" httpUri="https://zenodo.org/record/7667209/files/figure.png" pageId="18" pageNumber="146">8</figureCitation>
,
<figureCitation id="A1E22A41593AFFF533EDFD97FD03FDFC" box="[717,731,534,559]" captionStart="Fig" captionStartId="19.[144,177,943,963]" captionTargetBox="[148,1100,203,927]" captionTargetId="figure-248@19.[149,1099,206,921]" captionTargetPageId="19" captionText="Fig. 9. Map showing geographic and taxonomic variation in proboscis length for Philoliche rubiginosa, P. rondani and P. umbratipennis. Circle diameter is proportional to the proboscis length of the individual specimen collected at that site." figureDoi="http://doi.org/10.5281/zenodo.7667211" httpUri="https://zenodo.org/record/7667211/files/figure.png" pageId="18" pageNumber="146">9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593AFFF53195FDBCFD26FD4B" pageId="18" pageNumber="146" type="reference_group">
<treatmentCitationGroup id="19C911EA593AFFF53195FDBCFD18FDBB" pageId="18" pageNumber="146">
<paragraph id="396636C4593AFFF53195FDBCFBABFD83" blockId="18.[181,1142,534,1671]" box="[181,1139,572,593]" pageId="18" pageNumber="146">
<treatmentCitation id="B87810D5593AFFF53195FDBCFD22FD83" author="RICARDO, G." box="[181,762,572,593]" page="111" pageId="18" pageNumber="146" year="1900">
<taxonomicName id="FED94D47593AFFF53195FDBCFD22FD83" authority="Ricardo, 1900: 111" authorityName="Ricardo" authorityPageNumber="111" authorityYear="1900" box="[181,762,572,593]" class="Insecta" family="Tabanidae" genus="Diatomineura" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="umbratipennis" subGenus="Corizoneura">
<emphasis id="0BADEAD6593AFFF53195FDBCFEECFD82" box="[181,308,573,593]" italics="true" pageId="18" pageNumber="146">Diatomineura</emphasis>
(
<emphasis id="0BADEAD6593AFFF53062FDBCFE6DFD82" box="[322,437,573,593]" italics="true" pageId="18" pageNumber="146">Corizoneura</emphasis>
)
<emphasis id="0BADEAD6593AFFF530E2FDBCFD9DFD82" box="[450,581,573,593]" italics="true" pageId="18" pageNumber="146">umbratipennis</emphasis>
<bibRefCitation id="5D484B35593AFFF5336AFDBDFD22FD83" author="RICARDO, G." box="[586,762,572,592]" pageId="18" pageNumber="146" pagination="167 - 182" refId="ref22149" refString="RICARDO, G. 1900. Notes on the Pangoniinae of the family Tabanidae in the British Museum collection. Annals &amp; Magazine of Natural History (7) 5: 97 - 121, 167 - 182." type="journal article" year="1900">Ricardo, 1900: 111</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Annshaw [ECP],
<collectingCountry id="41CE7654593AFFF53281FDBDFBCCFD83" box="[929,1044,572,592]" name="South Africa" pageId="18" pageNumber="146">South Africa</collectingCountry>
(BMNH).
</paragraph>
<paragraph id="396636C4593AFFF53195FDD4FD18FDBB" blockId="18.[181,1142,534,1671]" box="[181,704,596,617]" pageId="18" pageNumber="146">
<taxonomicName id="FED94D47593AFFF53195FDD4FDC8FDBB" ID-CoL="88D3K" authority="(Ricardo)" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[181,528,596,617]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593AFFF53195FDD4FE69FDBA" box="[181,433,597,617]" italics="true" pageId="18" pageNumber="146">Corizoneura umbratipennis</emphasis>
(Ricardo)
</taxonomicName>
:
<treatmentCitation id="B87810D5593AFFF53338FDD5FD64FDBB" author="AUSTEN, E. E." box="[536,700,596,616]" page="140" pageId="18" pageNumber="146" year="1920">
<bibRefCitation id="5D484B35593AFFF53338FDD5FD64FDBB" author="AUSTEN, E. E." box="[536,700,596,616]" pageId="18" pageNumber="146" pagination="139 - 152" refId="ref19979" refString="AUSTEN, E. E. 1920. Notes on the nomenclature of certain African Tabanidae (sub-family Pangoniinae), with descriptions of a new genus and new species. Bulletin of Entomological Research 11 (43 - 45): 139 - 152." type="journal article" year="1920">Austen 1920: 140</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</treatmentCitationGroup>
<paragraph id="396636C4593AFFF53195FDECFB8AFD53" blockId="18.[181,1142,534,1671]" box="[181,1106,620,641]" pageId="18" pageNumber="146">
<treatmentCitationGroup id="19C911EA593AFFF53195FDECFB8AFD53" box="[181,1106,620,641]" pageId="18" pageNumber="146">
<taxonomicName id="FED94D47593AFFF53195FDECFD3FFD53" ID-CoL="8D9WF" authority="(Thunberg)" baseAuthorityName="Thunberg" box="[181,743,620,641]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="subSpecies" species="aethiopica" subGenus="Philoliche" subSpecies="aethiopica">
<emphasis id="0BADEAD6593AFFF53195FDECFECAFD52" box="[181,274,621,641]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
(
<emphasis id="0BADEAD6593AFFF53000FDECFEA5FD52" box="[288,381,621,641]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
)
<emphasis id="0BADEAD6593AFFF530AAFDECFE32FD52" box="[394,490,621,641]" italics="true" pageId="18" pageNumber="146">aethiopica</emphasis>
ssp.
<emphasis id="0BADEAD6593AFFF53338FDECFDA0FD52" box="[536,632,621,641]" italics="true" pageId="18" pageNumber="146">aethiopica</emphasis>
(Thunberg)
</taxonomicName>
:
<treatmentCitation id="B87810D5593AFFF533D0FDEDFC78FD53" author="OLDROYD, H." box="[752,928,620,640]" page="337" pageId="18" pageNumber="146" year="1957">
<bibRefCitation id="5D484B35593AFFF533D0FDEDFC78FD53" author="OLDROYD, H." box="[752,928,620,640]" pageId="18" pageNumber="146" refId="ref22111" refString="OLDROYD, H. 1957. The horseflies of the Ethiopian region. Vol. III. Subfamiles Chrysopsinae, Scepsidinae and Pangoniinae, and a revised classification. London: British Museum (Natural History)." type="book" year="1957">Oldroyd 1957: 337</bibRefCitation>
</treatmentCitation>
. Syn. rejected here.
</treatmentCitationGroup>
</paragraph>
<paragraph id="396636C4593AFFF53195FD04FD26FD4B" blockId="18.[181,1142,534,1671]" box="[181,766,644,665]" pageId="18" pageNumber="146">
<taxonomicName id="FED94D47593AFFF53195FD04FD82FD4B" ID-CoL="8D9WF" authority="(Thunberg)" baseAuthorityName="Thunberg" baseAuthorityYear="1789" box="[181,602,644,665]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="aethiopica" subGenus="Philoliche">
<emphasis id="0BADEAD6593AFFF53195FD04FECAFD4A" box="[181,274,645,665]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
(
<emphasis id="0BADEAD6593AFFF53000FD04FEA5FD4A" box="[288,381,645,665]" italics="true" pageId="18" pageNumber="146">Philoliche</emphasis>
)
<emphasis id="0BADEAD6593AFFF530AAFD04FE32FD4A" box="[394,490,645,665]" italics="true" pageId="18" pageNumber="146">aethiopica</emphasis>
(Thunberg)
</taxonomicName>
: Dias 1966: 1176.
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593AFFF53195FD27FC50FCEC" pageId="18" pageNumber="146" type="diagnosis">
<paragraph id="396636C4593AFFF53195FD27FC50FCEC" blockId="18.[181,1142,534,1671]" pageId="18" pageNumber="146">
Diagnosis:
<taxonomicName id="FED94D47593AFFF53013FD27FE36FD6D" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[307,494,678,702]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593AFFF53013FD27FE36FD6D" box="[307,494,678,702]" italics="true" pageId="18" pageNumber="146">P. umbratipennis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="FED94D47593AFFF53236FD27FC50FD6D" authorityName="Bertoloni" authorityYear="1861" box="[790,904,678,702]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593AFFF53236FD27FC50FD6D" box="[790,904,678,702]" italics="true" pageId="18" pageNumber="146">P. rondani</emphasis>
</taxonomicName>
by its relatively paler hind coxa and hind femur, by the third abdominal segment which is relatively paler than the other segments, and by having both black and white tufts of hair posterolaterally on segment 4. This species appears to occur only in the Albany
<collectingRegion id="FB1DF826593AFFF53256FC87FC19FCCC" box="[886,961,774,799]" country="France" name="Centre" pageId="18" pageNumber="146">Centre</collectingRegion>
of endemism. It usually has a proboscis 2 times greater in length than
<taxonomicName id="FED94D47593AFFF5322EFCA7FC59FCED" authorityName="Bertoloni" authorityYear="1861" box="[782,897,806,830]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593AFFF5322EFCA7FC59FCED" box="[782,897,806,830]" italics="true" pageId="18" pageNumber="146">P. rondani</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593AFFF43195FCCFFCBAF9F3" lastPageId="19" lastPageNumber="147" pageId="18" pageNumber="146" type="description">
<paragraph id="396636C4593AFFF53195FCCFFE99FCB4" blockId="18.[181,1142,534,1671]" box="[181,321,846,871]" pageId="18" pageNumber="146">Description:</paragraph>
<paragraph id="396636C4593AFFF53195FCF7FEC9FC5C" blockId="18.[181,1142,534,1671]" box="[181,273,886,911]" pageId="18" pageNumber="146">
<emphasis id="0BADEAD6593AFFF53195FCF7FED1FC5D" box="[181,265,886,910]" italics="true" pageId="18" pageNumber="146">Female</emphasis>
.
</paragraph>
<paragraph id="396636C4593AFFF53195FC1FFDE4FB44" blockId="18.[181,1142,534,1671]" pageId="18" pageNumber="146">
<emphasis id="0BADEAD6593AFFF53195FC1FFF2AFC65" box="[181,242,926,950]" italics="true" pageId="18" pageNumber="146">Head</emphasis>
: Eyes dichoptic; facets of similar size throughout. Frons only slightly divergent towards antennae, almost parallel; covered with yellow-grey or golden pruinescence (pollinosity) with shiny, reddish surface visible underneath; convex. Face conically produced forward, distance just greater than sagittal depth of the eye. Antennae with first two segments light brown to brown with black hairs; flagellum orange-brown to bright orange, sometimes with dark apex. Palps slender; distal segment wider with black hairs and sensory groove leading to an aperture; tip ending in a point. Proboscis length usually 1.52× body length.
</paragraph>
<paragraph id="396636C4593AFFF53195FB27FC3BFAEC" blockId="18.[181,1142,534,1671]" pageId="18" pageNumber="146">
<emphasis id="0BADEAD6593AFFF53195FB27FEDEFB6D" box="[181,262,1190,1214]" italics="true" pageId="18" pageNumber="146">Thorax</emphasis>
: Mesonotum dark brown with faint or no sublateral stripes. If sublateral stripes are evident, they are not narrow and grey but instead broad and brown or brownish red. Supra-alar areas of scutum sharply contrasting with white-grey pruinescence beginning with the postpronotal lobes and ending at the post-alar calli. Pleura light grey to pinkish grey with numerous white to yellow hairs, also with very few dark hairs.
</paragraph>
<paragraph id="396636C4593AFFF53195FAC7FE13FA4C" blockId="18.[181,1142,534,1671]" pageId="18" pageNumber="146">Wings: Smoky, becoming clearer posteriorly; darkened at cross-veins. Fifth radial cell usually open but closed at margin or even closed and stalked in some specimens; third medial cell always open.</paragraph>
<paragraph id="396636C4593AFFF53195FA27FBA9F9EC" blockId="18.[181,1142,534,1671]" pageId="18" pageNumber="146">Legs: Fore and mid coxae grey to pinkish grey, paler dorsally and with white and a few black hairs; hind coxa yellow, trochanters orange; fore and mid femora orange; hind femur yellow, same as hind coxa. Tibia of foreleg and midleg paler than femora. Hind tibia darker than hind femur with black hairs. Tarsi concolourous with tibia of same leg; darkened at apex of fifth tarsomere in fore and midtarsi. Tarsomeres with black hairs.</paragraph>
<paragraph id="396636C4593AFFF43195F9CFFB94FB73" blockId="18.[181,1142,534,1671]" lastBlockId="19.[144,1104,1063,1674]" lastPageId="19" lastPageNumber="147" pageId="18" pageNumber="146">
<emphasis id="0BADEAD6593AFFF53195F9CFFEC7F9B5" box="[181,287,1614,1638]" italics="true" pageId="18" pageNumber="146">Abdomen</emphasis>
: Orange-yellow with black medial spot on segments 1 and 2; segments 3 orange-yellow with scattered spots of pigment, but no true spot; segments 47 dark yellow-brown. Central posterior margin of segment 2 sometimes with white or yellow hairs. Posterior margin of segment 4 with black hairs (as rest of segment), except on posterolateral margin. Posterolateral margin of segments 37 with tufts of black or white hairs as follows: 3 black, 4 black and white, 5 black, 6 white, 7 white.
</paragraph>
<caption id="6DA6664C593BFFF431B0FC2EFDB0FC20" ID-DOI="http://doi.org/10.5281/zenodo.7667211" ID-Zenodo-Dep="7667211" httpUri="https://zenodo.org/record/7667211/files/figure.png" pageId="19" pageNumber="147" startId="19.[144,177,943,963]" targetBox="[148,1100,203,927]" targetPageId="19">
<paragraph id="396636C4593BFFF431B0FC2EFDB0FC20" blockId="19.[144,1104,943,1011]" pageId="19" pageNumber="147">
Fig. 9. Map showing geographic and taxonomic variation in proboscis length for
<taxonomicName id="FED94D47593BFFF432A3FC2EFB92FC10" authorityName="Dias, 1966:" authorityYear="1189" box="[899,1098,943,963]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="147" phylum="Arthropoda" rank="species" species="rubiginosa">
<emphasis id="0BADEAD6593BFFF432A3FC2EFB92FC10" box="[899,1098,943,963]" italics="true" pageId="19" pageNumber="147">Philoliche rubiginosa</emphasis>
</taxonomicName>
,
<taxonomicName id="FED94D47593BFFF431D0FC46FE96FC08" authorityName="Bertoloni" authorityYear="1861" box="[240,334,967,987]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="147" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593BFFF431D0FC46FE96FC08" box="[240,334,967,987]" italics="true" pageId="19" pageNumber="147">P. rondani</emphasis>
</taxonomicName>
and
<emphasis id="0BADEAD6593BFFF4305CFC46FDC5FC08" box="[380,541,967,987]" italics="true" pageId="19" pageNumber="147">
<taxonomicName id="FED94D47593BFFF4305CFC46FDC1FC08" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[380,537,967,987]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="147" phylum="Arthropoda" rank="species" species="umbratipennis">P. umbratipennis</taxonomicName>
.
</emphasis>
Circle diameter is proportional to the proboscis length of the individual specimen collected at that site.
</paragraph>
</caption>
<paragraph id="396636C4593BFFF431B0FB31FE4BFB1B" blockId="19.[144,1104,1063,1674]" box="[144,403,1199,1224]" pageId="19" pageNumber="147">
<emphasis id="0BADEAD6593BFFF431B0FB31FF12FB1B" box="[144,202,1200,1224]" italics="true" pageId="19" pageNumber="147">Male</emphasis>
as female except:
</paragraph>
<paragraph id="396636C4593BFFF431B0FB59FDFEFA43" blockId="19.[144,1104,1063,1674]" pageId="19" pageNumber="147">
<emphasis id="0BADEAD6593BFFF431B0FB59FF15FB23" box="[144,205,1240,1264]" italics="true" pageId="19" pageNumber="147">Head</emphasis>
: Eyes holoptic; facets increasing in size from ventral to dorsal very abruptly. Frons extremely small and triangular, pruinescent, covered with long, silky yellow, golden or white hairs; eye margins beginning to diverge a distance from vertex similar to length of antennae. Face conically produced, but not as prominent as in female, clypeus highly reduced. Palps reduced compared to female but with sensory pit. Clypeus reduced to frontoclypeal membrane.
</paragraph>
<paragraph id="396636C4593BFFF431B0FA21FC42FA6B" blockId="19.[144,1104,1063,1674]" box="[144,922,1439,1464]" pageId="19" pageNumber="147">
<emphasis id="0BADEAD6593BFFF431B0FA21FF39FA6B" box="[144,225,1440,1464]" italics="true" pageId="19" pageNumber="147">Thorax</emphasis>
: Mesonotum with longer, more dense, golden or yellow hairs.
</paragraph>
<paragraph id="396636C4593BFFF431B0FA3EFBD4FA2B" blockId="19.[144,1104,1063,1674]" pageId="19" pageNumber="147">Legs: With short, distally projecting dorsal process on tarsomeres 1 and 2 (lappet). Lappet of tarsomere 2 never extending beyond proximal margin of tarsomere 4.</paragraph>
<paragraph id="396636C4593BFFF431B0F989FCBAF9F3" blockId="19.[144,1104,1063,1674]" box="[144,866,1543,1568]" pageId="19" pageNumber="147">
<emphasis id="0BADEAD6593BFFF431B0F989FF22F9F3" box="[144,250,1544,1568]" italics="true" pageId="19" pageNumber="147">Abdomen</emphasis>
: Abdomen narrower and often more brightly coloured.
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593BFFF331B0F9AFFE51FE5B" lastPageId="20" lastPageNumber="148" pageId="19" pageNumber="147" type="materials_examined">
<paragraph id="396636C4593BFFF431B0F9AFFEAAF991" blockId="19.[144,1104,1063,1674]" box="[144,370,1582,1602]" pageId="19" pageNumber="147">Type material examined:</paragraph>
<paragraph id="396636C4593BFFF431B0F9C7FDB5F989" blockId="19.[144,1104,1063,1674]" box="[144,621,1606,1626]" pageId="19" pageNumber="147">
<taxonomicName id="FED94D47593BFFF431B0F9C7FDB5F989" authority="Ricardo" authorityName="Ricardo" authorityYear="1900" box="[144,621,1606,1626]" class="Insecta" family="Tabanidae" genus="Diatomineura" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="147" phylum="Arthropoda" rank="species" species="umbratipennis" subGenus="Corizoneura">
<emphasis id="0BADEAD6593BFFF431B0F9C7FED7F989" box="[144,271,1606,1626]" italics="true" pageId="19" pageNumber="147">Diatomineura</emphasis>
(
<emphasis id="0BADEAD6593BFFF4303CF9C7FE57F989" box="[284,399,1606,1626]" italics="true" pageId="19" pageNumber="147">Corizoneura</emphasis>
)
<emphasis id="0BADEAD6593BFFF430BCF9C7FDC7F989" box="[412,543,1606,1626]" italics="true" pageId="19" pageNumber="147">umbratipennis</emphasis>
Ricardo
</taxonomicName>
</paragraph>
<paragraph id="396636C4593BFFF431B0F9DFFF32F959" blockId="19.[144,1104,1063,1674]" pageId="19" pageNumber="147">
<materialsCitation id="89B13C99593BFFF431B0F9DFFF3DF959" collectionCode="BMNH" country="South Africa" location="Barrett" pageId="19" pageNumber="147" specimenCount="1" stateProvince="Eastern Cape" typeStatus="syntype">
<typeStatus id="E6628866593BFFF431B0F9DFFF3EF9A1" box="[144,230,1630,1650]" pageId="19" pageNumber="147" type="syntype">Syntypes</typeStatus>
:
<emphasis id="0BADEAD6593BFFF431CCF9DFFEFFF9A2" box="[236,295,1629,1650]" italics="true" pageId="19" pageNumber="147">1♀1♂</emphasis>
<collectingCountry id="41CE7654593BFFF4300BF9DFFE17F9A1" box="[299,463,1630,1650]" name="South Africa" pageId="19" pageNumber="147">SOUTH AFRICA</collectingCountry>
:
<emphasis id="0BADEAD6593BFFF430F5F9DFFD96F9A1" box="[469,590,1630,1650]" italics="true" pageId="19" pageNumber="147">
<collectingRegion id="FB1DF826593BFFF430F5F9DFFD96F9A1" box="[469,590,1630,1650]" country="South Africa" name="Eastern Cape" pageId="19" pageNumber="147">Eastern Cape</collectingRegion>
</emphasis>
: “Annshaw” [Mission], [Rev.]
<location id="3C06601F593BFFF43241F9DFFC79F9A1" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593BFFF43241F9DFFC79F9A1" box="[865,929,1630,1650]" country="South Africa" name="Barrett" pageId="19" pageNumber="147" stateProvince="Eastern Cape">Barrett</location>
, #242033, #242036 (
<collectionCode id="5FC8AE01593BFFF431B6F9F7FF07F959" box="[150,223,1654,1674]" pageId="19" pageNumber="147">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="396636C4593CFFF33195FF4DFE51FE5B" blockId="20.[181,1142,203,392]" pageId="20" pageNumber="148">
Additional material examined:
<materialsCitation id="89B13C99593CFFF330E6FF4DFE1FFF2B" collectingDate="2004-02-05" collectionCode="SMPC" collectorName="S. I. Morita" country="South Africa" latitude="-32.666668" location="E of Bedford" longLatPrecision="1209" longitude="26.016666" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<collectingCountry id="41CE7654593CFFF330E6FF4DFDB1FF33" box="[454,617,204,224]" name="South Africa" pageId="20" pageNumber="148">SOUTH AFRICA</collectingCountry>
:
<emphasis id="0BADEAD6593CFFF3334FFF4CFD3FFF32" box="[623,743,205,225]" italics="true" pageId="20" pageNumber="148">
<collectingRegion id="FB1DF826593CFFF3334FFF4CFD3FFF32" box="[623,743,205,225]" country="South Africa" name="Eastern Cape" pageId="20" pageNumber="148">Eastern Cape</collectingRegion>
</emphasis>
:
<emphasis id="0BADEAD6593CFFF333D2FF4DFCF4FF33" box="[754,812,204,224]" italics="true" pageId="20" pageNumber="148">2♀2♂</emphasis>
R63
<location id="3C06601F593CFFF33279FF4DFC12FF33" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF33279FF4DFC12FF33" box="[857,970,204,224]" country="South Africa" latitude="-32.666668" longLatPrecision="1209" longitude="26.016666" name="E of Bedford" pageId="20" pageNumber="148" stateProvince="Eastern Cape">E of Bedford</location>
(
<geoCoordinate id="5CED5003593CFFF332F5FF4DFBC4FF33" box="[981,1052,203,224]" degrees="32" direction="south" minutes="40" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-32.666668">32°40'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF3353FFF4DFBB3FF33" box="[1055,1131,203,224]" degrees="26" direction="east" minutes="01" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.016666">26°01'E</geoCoordinate>
),
<date id="4D671004593CFFF33195FF65FED0FF2B" box="[181,264,228,248]" pageId="20" pageNumber="148" value="2004-02-05">
<collectingDate id="5D23E9EC593CFFF33195FF65FED0FF2B" box="[181,264,228,248]" pageId="20" pageNumber="148" value="2004-02-05">5.ii.2004</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF33031FF65FEACFF2B" box="[273,372,228,248]" pageId="20" pageNumber="148">S.I. Morita</collectorName>
(
<collectionCode id="5FC8AE01593CFFF330A0FF65FE19FF2B" box="[384,449,228,248]" pageId="20" pageNumber="148">SMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF330F0FF65FE92FEC3" collectingDate="2004-02-06" collectionCode="SMPC" collectorName="S. I. Morita" country="South Africa" county="Rd" latitude="32.41" location="Bosberg" longitude="25.65" municipality="Glen Craig Farm" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="0BADEAD6593CFFF330F0FF65FDCEFF2B" box="[464,534,228,248]" italics="true" pageId="20" pageNumber="148">3♀11♂</emphasis>
<collectingCounty id="D0074E48593CFFF3333BFF65FDEDFF2B" box="[539,565,228,248]" pageId="20" pageNumber="148">Rd</collectingCounty>
to
<collectingMunicipality id="D902ACBE593CFFF33372FF65FD34FF2B" box="[594,748,228,248]" pageId="20" pageNumber="148">Glen Craig Farm</collectingMunicipality>
at
<location id="3C06601F593CFFF33227FF65FC8AFF2B" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF33227FF65FC8AFF2B" box="[775,850,228,248]" country="South Africa" county="Rd" longitude="25.65" municipality="Glen Craig Farm" name="Bosberg" pageId="20" pageNumber="148" stateProvince="Eastern Cape">Bosberg</location>
(
<geoCoordinate id="5CED5003593CFFF3327DFF65FC45FF2B" box="[861,925,227,248]" degrees="32" direction="south" minutes="41" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="32.41">32°41':</geoCoordinate>
<geoCoordinate id="5CED5003593CFFF332BDFF65FC32FF2B" box="[925,1002,227,248]" degrees="25" direction="east" minutes="39" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="25.65">25°39'E</geoCoordinate>
),
<date id="4D671004593CFFF332DAFF65FB94FF2B" box="[1018,1100,228,248]" pageId="20" pageNumber="148" value="2004-02-06">
<collectingDate id="5D23E9EC593CFFF332DAFF65FB94FF2B" box="[1018,1100,228,248]" pageId="20" pageNumber="148" value="2004-02-06">6.ii.2004</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF33576FF65FF2BFEC3" pageId="20" pageNumber="148">S.I. Morita</collectorName>
(
<collectionCode id="5FC8AE01593CFFF33023FF7DFE9DFEC3" box="[259,325,252,272]" pageId="20" pageNumber="148">SMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF33077FF7DFC3DFEC3" box="[343,997,251,272]" collectingDate="1922-10-29" collectionCode="NMSA" collectorName="H. K. Munro" country="South Africa" latitude="-33.033333" location="East London" longLatPrecision="1207" longitude="27.916666" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<location id="3C06601F593CFFF33077FF7DFE16FEC3" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF33077FF7DFE16FEC3" box="[343,462,252,272]" country="South Africa" latitude="-33.033333" longLatPrecision="1207" longitude="27.916666" name="East London" pageId="20" pageNumber="148" stateProvince="Eastern Cape">East London</location>
(
<geoCoordinate id="5CED5003593CFFF330FDFF7DFDFEFEC3" box="[477,550,251,272]" degrees="33" direction="south" minutes="02" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.033333">33°02'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF3330AFF7DFDAFFEC3" box="[554,631,251,272]" degrees="27" direction="east" minutes="55" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="27.916666">27°55'E</geoCoordinate>
),
<collectorName id="942C5312593CFFF333ABFF7DFCDAFEC3" box="[651,770,252,272]" pageId="20" pageNumber="148">H.K. Munro</collectorName>
: 1
<emphasis id="0BADEAD6593CFFF33238FF7DFCFEFEC3" box="[792,806,252,272]" italics="true" pageId="20" pageNumber="148"></emphasis>
<date id="4D671004593CFFF3320FFF7DFC52FEC3" box="[815,906,252,272]" pageId="20" pageNumber="148" value="1922-10-29">
<collectingDate id="5D23E9EC593CFFF3320FFF7DFC52FEC3" box="[815,906,252,272]" pageId="20" pageNumber="148" value="1922-10-29">29.x.1922</collectingDate>
</date>
(
<collectionCode id="5FC8AE01593CFFF332B9FF7DFC07FEC3" box="[921,991,252,272]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="89B13C99593CFFF332D2FF7DFED1FEFB" collectingDate="1924-11-15" collectionCode="NMSA" country="South Africa" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<date id="4D671004593CFFF332D2FF7DFBAEFEC3" box="[1010,1142,252,272]" pageId="20" pageNumber="148" value="1924-11-15">
<collectingDate id="5D23E9EC593CFFF332D2FF7DFBAEFEC3" box="[1010,1142,252,272]" pageId="20" pageNumber="148" value="1924-11-15">
<emphasis id="0BADEAD6593CFFF332D2FF7DFBF2FEC3" box="[1010,1066,252,272]" italics="true" pageId="20" pageNumber="148">1♀15</emphasis>
.xi.1924
</collectingDate>
</date>
(
<collectionCode id="5FC8AE01593CFFF3319CFE95FEDBFEFB" box="[188,259,276,296]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF33036FE95FC51FEFB" box="[278,905,275,296]" collectingDate="1928-03-23" collectionCode="NMSA" collectorName="A. Walton" country="South Africa" latitude="-33.166668" location="Resolution" longLatPrecision="1206" longitude="26.716667" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="0BADEAD6593CFFF33036FE95FE44FEFB" box="[278,412,276,296]" italics="true" pageId="20" pageNumber="148">
<location id="3C06601F593CFFF33036FE95FE44FEFB" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF33036FE95FE44FEFB" box="[278,412,276,296]" country="South Africa" latitude="-33.166668" longLatPrecision="1206" longitude="26.716667" name="Resolution" pageId="20" pageNumber="148" stateProvince="Eastern Cape">1♀Resolution</location>
</emphasis>
(
<geoCoordinate id="5CED5003593CFFF3308CFE95FE2DFEFB" box="[428,501,275,296]" degrees="33" direction="south" minutes="10" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.166668">33°10'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF330D8FE95FD9FFEFB" box="[504,583,275,296]" degrees="26" direction="east" minutes="43" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.716667">26°43'E</geoCoordinate>
),
<date id="4D671004593CFFF3337BFE95FD19FEFB" box="[603,705,276,296]" pageId="20" pageNumber="148" value="1928-03-23">
<collectingDate id="5D23E9EC593CFFF3337BFE95FD19FEFB" box="[603,705,276,296]" pageId="20" pageNumber="148" value="1928-03-23">23.iii.1928</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF333ECFE95FCF5FEFB" box="[716,813,276,296]" pageId="20" pageNumber="148">A. Walton</collectorName>
(
<collectionCode id="5FC8AE01593CFFF3321CFE95FC5BFEFB" box="[828,899,276,296]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF332B6FE95FD87FE93" collectingDate="1970-04-05" collectionCode="NMSA" collectorName="J. G. H. Londt" country="South Africa" latitude="-33.3" location="Grahamstown" longLatPrecision="1205" longitude="26.533333" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<location id="3C06601F593CFFF332B6FE95FBCFFEFB" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF332B6FE95FBCFFEFB" box="[918,1047,276,296]" country="South Africa" latitude="-33.3" longLatPrecision="1205" longitude="26.533333" name="Grahamstown" pageId="20" pageNumber="148" stateProvince="Eastern Cape">Grahamstown</location>
(
<geoCoordinate id="5CED5003593CFFF33506FE95FBAAFEFB" box="[1062,1138,275,296]" degrees="33" direction="south" minutes="18" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.3">33°18'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF33195FEADFED9FE93" box="[181,257,299,320]" degrees="26" direction="east" minutes="32" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.533333">26°32'E</geoCoordinate>
):
<date id="4D671004593CFFF33031FEADFE51FE93" box="[273,393,300,320]" pageId="20" pageNumber="148" value="1970-04-05">
<collectingDate id="5D23E9EC593CFFF33031FEADFE51FE93" box="[273,393,300,320]" pageId="20" pageNumber="148" value="1970-04-05">
<emphasis id="0BADEAD6593CFFF33031FEADFEE6FE93" box="[273,318,300,320]" italics="true" pageId="20" pageNumber="148">2♀5</emphasis>
.iv.1970
</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF330B1FEADFDDFFE93" box="[401,519,300,320]" pageId="20" pageNumber="148">J.G.H. Londt</collectorName>
(
<collectionCode id="5FC8AE01593CFFF33332FEADFD81FE93" box="[530,601,300,320]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF33348FEADFBAAFE93" box="[616,1138,300,320]" collectingDate="1949-05-11" collectionCode="NMSA" collectorName="B. Stuckenberg" country="South Africa" location="Dist." municipality="Albany" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
1
<emphasis id="0BADEAD6593CFFF33353FEADFD59FE93" box="[627,641,300,320]" italics="true" pageId="20" pageNumber="148"></emphasis>
<collectingMunicipality id="D902ACBE593CFFF333AFFEADFD0BFE93" box="[655,723,300,320]" pageId="20" pageNumber="148">Albany</collectingMunicipality>
[
<location id="3C06601F593CFFF333FDFEADFCD5FE93" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF333FDFEADFCD5FE93" box="[733,781,300,320]" country="South Africa" municipality="Albany" name="Dist." pageId="20" pageNumber="148" stateProvince="Eastern Cape">Dist.</location>
] ”,
<date id="4D671004593CFFF3320CFEADFC5EFE93" box="[812,902,300,320]" pageId="20" pageNumber="148" value="1949-05-11">
<collectingDate id="5D23E9EC593CFFF3320CFEADFC5EFE93" box="[812,902,300,320]" pageId="20" pageNumber="148" value="1949-05-11">11.v.1949</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF332AFFEADFBC2FE93" box="[911,1050,300,320]" pageId="20" pageNumber="148">B. Stuckenberg</collectorName>
(
<collectionCode id="5FC8AE01593CFFF33505FEADFBB4FE93" box="[1061,1132,300,320]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
;
<emphasis id="0BADEAD6593CFFF33195FEC5FF2AFE8B" box="[181,242,324,344]" italics="true" pageId="20" pageNumber="148">3♀2♂</emphasis>
R67
<materialsCitation id="89B13C99593CFFF3300AFEC5FC61FE8B" box="[298,953,323,344]" collectingDate="2004-02-18" collectionCode="SMPC" collectorName="S. I. Morita" country="South Africa" latitude="-33.383335" location="Port Alfred" longLatPrecision="1205" longitude="26.7" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
N of
<location id="3C06601F593CFFF3307FFEC5FE11FE8B" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF3307FFEC5FE11FE8B" box="[351,457,324,344]" country="South Africa" latitude="-33.383335" longLatPrecision="1205" longitude="26.7" name="Port Alfred" pageId="20" pageNumber="148" stateProvince="Eastern Cape">Port Alfred</location>
(
<geoCoordinate id="5CED5003593CFFF330FAFEC5FDFCFE8B" box="[474,548,323,344]" degrees="33" direction="south" minutes="23" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.383335">33°23'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF33307FEC5FDAFFE8B" box="[551,631,323,344]" degrees="26" direction="east" minutes="42" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.7">26°42'E</geoCoordinate>
),
<date id="4D671004593CFFF333ABFEC5FD33FE8B" box="[651,747,324,344]" pageId="20" pageNumber="148" value="2004-02-18">
<collectingDate id="5D23E9EC593CFFF333ABFEC5FD33FE8B" box="[651,747,324,344]" pageId="20" pageNumber="148" value="2004-02-18">18.ii.2004</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF333D8FEC5FCB8FE8B" box="[760,864,324,344]" pageId="20" pageNumber="148">S.I. Morita</collectorName>
(
<collectionCode id="5FC8AE01593CFFF33250FEC5FC6BFE8B" box="[880,947,324,344]" pageId="20" pageNumber="148">SMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF332E5FEC5FD50FEA3" collectingDate="1924-11" collectionCode="NMSA" collectorName="H. Cronwright" country="South Africa" latitude="-33.45" location="Trappes Valley" longLatPrecision="1205" longitude="26.9" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<location id="3C06601F593CFFF332E5FEC5FBADFE8B" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF332E5FEC5FBADFE8B" box="[965,1141,324,344]" country="South Africa" latitude="-33.45" longLatPrecision="1205" longitude="26.9" name="Trappes Valley" pageId="20" pageNumber="148" stateProvince="Eastern Cape">
<emphasis id="0BADEAD6593CFFF332E5FEC5FBEAFE8B" box="[965,1074,324,344]" italics="true" pageId="20" pageNumber="148">1♀Trappes</emphasis>
Valley
</location>
(
<geoCoordinate id="5CED5003593CFFF3319CFEDDFEDCFEA3" box="[188,260,347,368]" degrees="33" direction="south" minutes="27" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.45">33°27'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF33027FEDDFE8CFEA3" box="[263,340,347,368]" degrees="26" direction="east" minutes="54" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.9">26°54'E</geoCoordinate>
),
<date id="4D671004593CFFF33042FEDDFE70FEA3" box="[354,424,348,368]" pageId="20" pageNumber="148" value="1924-11">
<collectingDate id="5D23E9EC593CFFF33042FEDDFE70FEA3" box="[354,424,348,368]" pageId="20" pageNumber="148" value="1924-11">xi.1924</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF33090FEDDFDEAFEA3" box="[432,562,348,368]" pageId="20" pageNumber="148">H. Cronwright</collectorName>
(
<collectionCode id="5FC8AE01593CFFF3331CFEDDFD5AFEA3" box="[572,642,348,368]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="89B13C99593CFFF333B0FEDDFE5DFE5B" collectingDate="1970-11-21" collectionCode="NMSA" collectorName="J. G. H. Londt" country="South Africa" latitude="-33.683334" location="Kenton On The Sea" longLatPrecision="1204" longitude="26.666666" pageId="20" pageNumber="148" specimenCount="1" stateProvince="Eastern Cape">
<location id="3C06601F593CFFF333B0FEDDFC87FEA3" LSID="urn:lsid:plazi:treatment:B17087D2593AFFF3300BFE6FFEE4FC0F:3C06601F593CFFF333B0FEDDFC87FEA3" box="[656,863,348,368]" country="South Africa" latitude="-33.683334" longLatPrecision="1204" longitude="26.666666" name="Kenton On The Sea" pageId="20" pageNumber="148" stateProvince="Eastern Cape">
<emphasis id="0BADEAD6593CFFF333B0FEDDFD2CFEA3" box="[656,756,348,368]" italics="true" pageId="20" pageNumber="148">3♀Kenton</emphasis>
On The Sea
</location>
(
<geoCoordinate id="5CED5003593CFFF3324AFEDDFC68FEA3" box="[874,944,347,368]" degrees="33" direction="south" minutes="41" orientation="latitude" pageId="20" pageNumber="148" precision="925" value="-33.683334">33°41'S</geoCoordinate>
:
<geoCoordinate id="5CED5003593CFFF33294FEDDFBD9FEA3" box="[948,1025,347,368]" degrees="26" direction="east" minutes="40" orientation="longitude" pageId="20" pageNumber="148" precision="925" value="26.666666">26°40'E</geoCoordinate>
),
<date id="4D671004593CFFF33530FEDDFBAAFEA3" box="[1040,1138,348,368]" pageId="20" pageNumber="148" value="1970-11-21">
<collectingDate id="5D23E9EC593CFFF33530FEDDFBAAFEA3" box="[1040,1138,348,368]" pageId="20" pageNumber="148" value="1970-11-21">21.xi.1970</collectingDate>
</date>
,
<collectorName id="942C5312593CFFF33195FEF5FEF4FE5B" box="[181,300,372,392]" pageId="20" pageNumber="148">J.G.H. Londt</collectorName>
(
<collectionCode id="5FC8AE01593CFFF33018FEF5FEA7FE5B" box="[312,383,372,392]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="148" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593CFFF33195FE18FC51FE61" box="[181,905,409,434]" pageId="20" pageNumber="148" type="distribution">
<paragraph id="396636C4593CFFF33195FE18FC51FE61" blockId="20.[181,905,409,434]" box="[181,905,409,434]" pageId="20" pageNumber="148">
Distribution: Southern Africa; known only in the Albany
<collectingRegion id="FB1DF826593CFFF33216FE18FC5CFE61" box="[822,900,409,434]" country="France" name="Centre" pageId="20" pageNumber="148">Centre</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="71C3654F593CFFF33195FE42FEE4FC0F" pageId="20" pageNumber="148" type="discussion">
<paragraph id="396636C4593CFFF33195FE42FC4BFD8F" blockId="20.[181,1142,451,988]" pageId="20" pageNumber="148">
Remarks:
<bibRefCitation id="5D484B35593CFFF33004FE42FE11FE0F" author="RICARDO, G." box="[292,457,451,476]" pageId="20" pageNumber="148" pagination="167 - 182" refId="ref22149" refString="RICARDO, G. 1900. Notes on the Pangoniinae of the family Tabanidae in the British Museum collection. Annals &amp; Magazine of Natural History (7) 5: 97 - 121, 167 - 182." type="journal article" year="1900">Ricardo (1900)</bibRefCitation>
described
<taxonomicName id="FED94D47593CFFF3331BFE45FCF2FE0F" authorityName="Ricardo" authorityYear="1900" box="[571,810,451,476]" class="Insecta" family="Tabanidae" genus="Diatomineura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis" subGenus="Corizoneura">
<emphasis id="0BADEAD6593CFFF3331BFE45FD8FFE0F" box="[571,599,452,476]" italics="true" pageId="20" pageNumber="148">D.</emphasis>
(
<emphasis id="0BADEAD6593CFFF33344FE45FDA6FE0F" box="[612,638,452,476]" italics="true" pageId="20" pageNumber="148">C.</emphasis>
)
<emphasis id="0BADEAD6593CFFF333ABFE45FCF2FE0F" box="[651,810,452,476]" italics="true" pageId="20" pageNumber="148">umbratipennis</emphasis>
</taxonomicName>
based on a series of specimens collected by Rev. Barrett at the Annshaw Mission in the former Ciskei (currently
<collectingRegion id="FB1DF826593CFFF33502FE62FF2BFDCF" country="South Africa" name="Eastern Cape" pageId="20" pageNumber="148">Eastern Cape</collectingRegion>
). It was later reclassified as
<taxonomicName id="FED94D47593CFFF33302FD85FD77FDCF" authorityName="Rondani" authorityYear="1864" box="[546,687,516,540]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="0BADEAD6593CFFF33302FD85FD77FDCF" box="[546,687,516,540]" italics="true" pageId="20" pageNumber="148">Corizoneura</emphasis>
</taxonomicName>
by
<bibRefCitation id="5D484B35593CFFF333F4FD82FCADFDCF" author="AUSTEN, E. E." box="[724,885,515,540]" pageId="20" pageNumber="148" pagination="139 - 152" refId="ref19979" refString="AUSTEN, E. E. 1920. Notes on the nomenclature of certain African Tabanidae (sub-family Pangoniinae), with descriptions of a new genus and new species. Bulletin of Entomological Research 11 (43 - 45): 139 - 152." type="journal article" year="1920">Austen (1920)</bibRefCitation>
, and then synonymised under
<taxonomicName id="FED94D47593CFFF331DDFDA5FD9AFDEF" box="[253,578,547,572]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="subSpecies" species="aethiopica" subSpecies="aethiopica">
<emphasis id="0BADEAD6593CFFF331DDFDA5FE48FDEF" box="[253,400,548,572]" italics="true" pageId="20" pageNumber="148">P. aethiopica</emphasis>
ssp.
<emphasis id="0BADEAD6593CFFF330ECFDA5FD9AFDEF" box="[460,578,548,572]" italics="true" pageId="20" pageNumber="148">aethiopica</emphasis>
</taxonomicName>
by
<bibRefCitation id="5D484B35593CFFF3334FFDA2FCFAFDEF" author="OLDROYD, H." box="[623,802,547,572]" pageId="20" pageNumber="148" refId="ref22111" refString="OLDROYD, H. 1957. The horseflies of the Ethiopian region. Vol. III. Subfamiles Chrysopsinae, Scepsidinae and Pangoniinae, and a revised classification. London: British Museum (Natural History)." type="book" year="1957">Oldroyd (1957)</bibRefCitation>
. It was left in synonymy with
<taxonomicName id="FED94D47593CFFF33195FDC5FE9FFD8F" baseAuthorityName="Thunberg" baseAuthorityYear="1789" box="[181,327,580,604]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis id="0BADEAD6593CFFF33195FDC5FE9FFD8F" box="[181,327,580,604]" italics="true" pageId="20" pageNumber="148">P. aethiopica</emphasis>
</taxonomicName>
when Dias separated
<taxonomicName id="FED94D47593CFFF3331DFDC5FD68FD8F" authorityName="Bertoloni" authorityYear="1861" box="[573,688,580,604]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593CFFF3331DFDC5FD68FD8F" box="[573,688,580,604]" italics="true" pageId="20" pageNumber="148">P. rondani</emphasis>
</taxonomicName>
as a species (1966).
</paragraph>
<paragraph id="396636C4593CFFF331F0FDE2FEE4FC0F" blockId="20.[181,1142,451,988]" pageId="20" pageNumber="148">
The molecular (
<bibRefCitation id="5D484B35593CFFF3305FFDE2FDD7FDAF" author="MORITA, S. I." box="[383,527,611,636]" pageId="20" pageNumber="148" refId="ref21967" refString="MORITA, S. I. 2006. The evolution of long-proboscis horse flies (Diptera: Tabanidae) in the genus Philoliche (Widemann). Unpublished Dissertation, Univeristy of California, Davis." type="book" year="2006">Morita 2006</bibRefCitation>
, in press
<emphasis id="0BADEAD6593CFFF33354FDE5FD5AFDAF" box="[628,642,612,636]" italics="true" pageId="20" pageNumber="148">a</emphasis>
) and morphological evidence presented here show that
<taxonomicName id="FED94D47593CFFF3300FFD05FD39FD4F" authority="(Ricardo)" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[303,737,643,668]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593CFFF3300FFD05FDB3FD4F" box="[303,619,644,668]" italics="true" pageId="20" pageNumber="148">Corizoneura umbratipennis</emphasis>
(Ricardo)
</taxonomicName>
was incorrectly synonymised with
<taxonomicName id="FED94D47593CFFF33195FD25FE9FFD6F" baseAuthorityName="Thunberg" baseAuthorityYear="1789" box="[181,327,676,700]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis id="0BADEAD6593CFFF33195FD25FE9FFD6F" box="[181,327,676,700]" italics="true" pageId="20" pageNumber="148">P. aethiopica</emphasis>
</taxonomicName>
. Examination of the
<typeStatus id="E6628866593CFFF33310FD22FDB9FD6F" box="[560,609,675,700]" pageId="20" pageNumber="148">type</typeStatus>
material reveals clear dimorphism in eye facets from dorsal to ventral, only short lappets on the fore tarsi of the male, and the index of frons divergence less than 2.1 in the female. These characters place
<taxonomicName id="FED94D47593CFFF33299FD65FBADFD2F" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[953,1141,740,764]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593CFFF33299FD65FBADFD2F" box="[953,1141,740,764]" italics="true" pageId="20" pageNumber="148">P. umbratipennis</emphasis>
</taxonomicName>
in the
<taxonomicName id="FED94D47593CFFF331DAFC85FE8AFCCF" authorityName="Bertoloni" authorityYear="1861" box="[250,338,772,796]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593CFFF331DAFC85FE8AFCCF" box="[250,338,772,796]" italics="true" pageId="20" pageNumber="148">rondani</emphasis>
</taxonomicName>
species group. Additionally, molecular data resolve
<taxonomicName id="FED94D47593CFFF332BAFC85FB80FCCF" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[922,1112,772,796]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593CFFF332BAFC85FB80FCCF" box="[922,1112,772,796]" italics="true" pageId="20" pageNumber="148">P. umbratipennis</emphasis>
</taxonomicName>
as a monophyletic clade within the
<taxonomicName id="FED94D47593CFFF3331AFCA5FD4AFCEF" authorityName="Bertoloni" authorityYear="1861" box="[570,658,804,828]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593CFFF3331AFCA5FD4AFCEF" box="[570,658,804,828]" italics="true" pageId="20" pageNumber="148">rondani</emphasis>
</taxonomicName>
clade (
<figureCitation id="A1E22A41593CFFF333CCFCA2FCEDFCEF" box="[748,821,803,828]" captionStart="Fig" captionStartId="5.[144,177,1479,1499]" captionTargetBox="[144,1091,1008,1460]" captionTargetId="figure-361@5.[144,1091,1008,1460]" captionTargetPageId="5" captionText="Fig. 2. Phylogenetic relationships in the subgenus Philoliche (Philoliche) from southern Africa.Relationships and chronographic data are based on Morita (2006, in press a, b). Numbers under branches represent clade support from maximum parsimony (bootstrap proportions from 1000 replicates, MP), maximum likelihood (bootstrap proportions from 1000 replicates, ML), Bayesian (posterior probability from 20 million generations of MCMC, PP) analyses presented as MP/ML/PP (Morita in press a). Branch lengths are proportional to millions of years (MYA) (Morita in press b). Major clades are marked with letters in circles:A P. aethiopica species complex, B aethiopica species group, C rondani species group, D P. rubiginosa, E P. umbratipennis." figureDoi="http://doi.org/10.5281/zenodo.7667193" httpUri="https://zenodo.org/record/7667193/files/figure.png" pageId="20" pageNumber="148">Fig. 2</figureCitation>
). However, the sister clade of
<taxonomicName id="FED94D47593CFFF331F8FCC5FE43FC8F" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[216,411,836,860]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593CFFF331F8FCC5FE43FC8F" box="[216,411,836,860]" italics="true" pageId="20" pageNumber="148">P. umbratipennis</emphasis>
</taxonomicName>
is still unclear. It forms a strongly supported clade with both
<taxonomicName id="FED94D47593CFFF33195FCE5FEF1FCAF" authorityName="Bertoloni" authorityYear="1861" box="[181,297,868,892]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593CFFF33195FCE5FEF1FCAF" box="[181,297,868,892]" italics="true" pageId="20" pageNumber="148">P. rondani</emphasis>
</taxonomicName>
and
<taxonomicName id="FED94D47593CFFF33040FCE5FE2EFCAF" authorityName="Dias, 1966:" authorityYear="1189" box="[352,502,868,892]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rubiginosa">
<emphasis id="0BADEAD6593CFFF33040FCE5FE2EFCAF" box="[352,502,868,892]" italics="true" pageId="20" pageNumber="148">P. rubiginosa</emphasis>
</taxonomicName>
, which is itself a sister one to
<taxonomicName id="FED94D47593CFFF3326EFCE5FC18FCAF" baseAuthorityName="Bigot" baseAuthorityYear="1892" box="[846,960,868,892]" class="Insecta" family="Tabanidae" genus="Philoliche" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="0BADEAD6593CFFF3326EFCE5FC18FCAF" box="[846,960,868,892]" italics="true" pageId="20" pageNumber="148">P. elegans</emphasis>
</taxonomicName>
. Strong support for monophyly of
<taxonomicName id="FED94D47593CFFF330A5FC05FD9CFC4F" authorityName="Morita" authorityYear="2008" baseAuthorityName="Ricardo" box="[389,580,900,924]" class="Insecta" family="Tabanidae" genus="Corizoneura" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="umbratipennis">
<emphasis id="0BADEAD6593CFFF330A5FC05FD9CFC4F" box="[389,580,900,924]" italics="true" pageId="20" pageNumber="148">P. umbratipennis</emphasis>
</taxonomicName>
and its morphological distinctness, even where it overlaps geographically with
<taxonomicName id="FED94D47593CFFF3332CFC25FD5AFC6F" authorityName="Bertoloni" authorityYear="1861" box="[524,642,932,956]" class="Insecta" family="Tabanidae" genus="Pangonia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="148" phylum="Arthropoda" rank="species" species="rondani">
<emphasis id="0BADEAD6593CFFF3332CFC25FD5AFC6F" box="[524,642,932,956]" italics="true" pageId="20" pageNumber="148">P. rondani</emphasis>
</taxonomicName>
, testifies to this entitys species status (see discussion).
</paragraph>
</subSubSection>
</treatment>
</document>